treatments-xml/data/37/6A/87/376A87F52748FF8E5BA1FB9C45C4FD4B.xml

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<mods:title id="8C2DE42F285E06F2AFF6D9AE831556A0">Japanese species of the genus Proutia Tutt, 1899 (Lepidoptera: Psychidae)</mods:title>
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<mods:namePart id="7C8E48636C21428A67E8382FB6126B6A">Saigusa, Toyohei</mods:namePart>
<mods:affiliation id="59B57B0D544B6F46211A924757669B98">7 - 1 - 402 Baikoen 2 - chome, Chuo-ku, Fukuoka-shi 810 - 0035, Japan. E-mail: toyohei _ saigusa @ yahoo. co. jp</mods:affiliation>
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<mods:namePart id="7813C0F97980195916671C51041E15D2">Sugimoto, Mika</mods:namePart>
<mods:affiliation id="4DFD25A1CE361A7F2C486B7999020CD9">Kyushu University Museum, 10 - 1 Hakozaki 6 - chome, Higashi-ku, Fukuoka-shi 812 - 8581, Japan. E-mail: mksgmt 03 gi @ yahoo. co. jp Correspondence author</mods:affiliation>
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<heading id="E434818F2748FF895BA1FB9C43FDFB93" bold="true" box="[151,758,1088,1115]" fontSize="11" level="1" pageId="1" pageNumber="144" reason="1">
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<taxonomicName id="78C34D602748FF895BA1FB9C4386FB93" ID-CoL="B72JN" authority="Saigusa et Sugimoto" authorityName="Saigusa et Sugimoto" authorityYear="2014" box="[151,653,1088,1115]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="maculatella" status="sp. nov.">
<emphasis id="8DB7EAF12748FF895BA1FB9C408CFB92" bold="true" box="[151,391,1088,1114]" italics="true" pageId="1" pageNumber="144">Proutia maculatella</emphasis>
Saigusa et Sugimoto
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, 
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<paragraph id="BF7C36E32748FF895BA1FBB84079FBB5" blockId="1.[151,758,1088,1149]" box="[151,370,1124,1149]" pageId="1" pageNumber="144">
(
<figureCitation id="27F82A662748FF895BA9FBB841E7FBB5" box="[159,236,1124,1149]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="1" pageNumber="144">Figs. 1</figureCitation>
, 
<figureCitation id="27F82A662748FF895BCCFBB84001FBB5" box="[250,266,1124,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="1" pageNumber="144">2</figureCitation>
, 
<figureCitation id="27F82A662748FF895A20FBB84030FBB5" box="[278,315,1124,1149]" captionStart="FIGURE 3" captionStartId="4.[151,250,720,742]" captionTargetBox="[343,1242,197,695]" captionTargetId="figure-181@4.[340,1247,193,699]" captionTargetPageId="4" captionText="FIGURE 3. Wing venation of Proutia spp. males. A: Proutia maculatella sp. nov.; B: Proutia nigra sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4947205" httpUri="https://zenodo.org/record/4947205/files/figure.png" pageId="1" pageNumber="144">3A</figureCitation>
, 
<figureCitation id="27F82A662748FF895A70FBB84061FBB5" box="[326,362,1124,1149]" captionStart="FIGURE 4" captionStartId="4.[151,250,1748,1770]" captionTargetBox="[452,1136,801,1718]" captionTargetId="figure-190@4.[446,1141,792,1726]" captionTargetPageId="4" captionText="FIGURE 4. Lateral aspect of male genitalia of Proutia spp. A: Proutia maculatella sp. nov.; B: Proutia nigra sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4947207" httpUri="https://zenodo.org/record/4947207/files/figure.png" pageId="1" pageNumber="144">4A</figureCitation>
)
</paragraph>
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<subSubSection id="F7D965682748FF895BA1FB70426BFB28" pageId="1" pageNumber="144" type="reference_group">
<paragraph id="BF7C36E32748FF895BA1FB704330FB0A" blockId="1.[151,864,1196,1248]" box="[151,571,1196,1218]" pageId="1" pageNumber="144">
<treatmentCitationGroup id="9FD311CD2748FF895BA1FB704330FB0A" box="[151,571,1196,1218]" pageId="1" pageNumber="144">
<taxonomicName id="78C34D602748FF895BA1FB704001FB09" box="[151,266,1196,1217]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8DB7EAF12748FF895BA1FB7041EFFB09" box="[151,228,1196,1217]" italics="true" pageId="1" pageNumber="144">Proutia</emphasis>
sp.
</taxonomicName>
: 
<treatmentCitation id="3E6210F22748FF895A22FB7040FAFB0A" author="Saigusa, T." box="[276,497,1196,1218]" page="3" pageId="1" pageNumber="144" year="1992">
<bibRefCitation id="DB524B122748FF895A22FB7040FAFB0A" author="Saigusa, T." box="[276,497,1196,1218]" pageId="1" pageNumber="144" pagination="168 - 169" refId="ref5880" refString="Saigusa, T. (1992) The ultimate evolution in the Psychidae. Dobutsu-tachi no chikyu, 78, 168 - 169. [in Japanese]" type="journal article" year="1992">Saigusa, 1992: 3–169</bibRefCitation>
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, 2 figs.
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</paragraph>
<paragraph id="BF7C36E32748FF895BA1FB17426BFB28" blockId="1.[151,864,1196,1248]" box="[151,864,1226,1248]" pageId="1" pageNumber="144">
<treatmentCitationGroup id="9FD311CD2748FF895BA1FB17426BFB28" box="[151,864,1226,1248]" pageId="1" pageNumber="144">
<taxonomicName id="78C34D602748FF895BA1FB174001FB28" box="[151,266,1227,1248]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8DB7EAF12748FF895BA1FB1741EFFB28" box="[151,228,1227,1248]" italics="true" pageId="1" pageNumber="144">Proutia</emphasis>
sp.
</taxonomicName>
: 
<treatmentCitation id="3E6210F22748FF895A22FB164395FB28" author="Saigusa, T. &amp; Sugimoto, M." box="[276,670,1226,1248]" page="146" pageId="1" pageNumber="144" year="2013">
<bibRefCitation id="DB524B122748FF895A22FB164395FB28" author="Saigusa, T. &amp; Sugimoto, M." box="[276,670,1226,1248]" pageId="1" pageNumber="144" pagination="3 - 14" refId="ref5908" refString="Saigusa, T. &amp; Sugimoto, M. (2013) Psychidae. In: Hirowatari, T. et al. (Eds.), The Standard of Moths in Japan III, Gakken Education Publishing, Tokyo, pp. 136 - 155. [pls 3 - 13, 3 - 14]" type="journal article" year="2013">Saigusa and Sugimoto, 2013: 146–147</bibRefCitation>
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, pl. 3–14, figs 1, 2.
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<paragraph id="BF7C36E32748FF895BA1FAD1437FFA5A" blockId="1.[151,1437,1293,2038]" pageId="1" pageNumber="144">
<emphasis id="8DB7EAF12748FF895BA1FAD1401AFAEE" bold="true" box="[151,273,1293,1318]" inLineHeading="true" pageId="1" pageNumber="144" reason="1">Diagnosis.</emphasis>
Male: Large for the genus 
<taxonomicName id="78C34D602748FF89590BFAD2439AFAED" authorityName="Tutt" authorityYear="1899" box="[573,657,1294,1317]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="8DB7EAF12748FF89590BFAD2439AFAED" box="[573,657,1294,1317]" italics="true" pageId="1" pageNumber="144">Proutia</emphasis>
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, with more or less semi-transparent apically rounded wings, forewing upperside greyish brown with indistinct greyish yellow speckles scattering on distal 1/2, and semi-transparent pale grey hindwing covered with hair-like upper scales. Female: Antenna long, more than 1.5 times head width; abdominal membrane light reddish brown.
</paragraph>
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<subSubSection id="F7D965682748FF8D5BF1FA41449EFA9B" lastPageId="5" lastPageNumber="148" pageId="1" pageNumber="144" type="description">
<paragraph id="BF7C36E32748FF895BF1FA41404DF83E" blockId="1.[151,1437,1293,2038]" pageId="1" pageNumber="144">
<emphasis id="8DB7EAF12748FF895BF1FA414053FA7D" box="[199,344,1437,1462]" inLineHeading="true" pageId="1" pageNumber="144" reason="1">
<emphasis id="8DB7EAF12748FF895BF1FA41405AFA7E" bold="true" box="[199,337,1437,1462]" pageId="1" pageNumber="144">Description</emphasis>
.
</emphasis>
<emphasis id="8DB7EAF12748FF895A57FA414096FA7E" bold="true" box="[353,413,1437,1462]" pageId="1" pageNumber="144">Male</emphasis>
(
<figureCitation id="27F82A662748FF895A99FA4140E7FA7E" box="[431,492,1437,1462]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="1" pageNumber="144">Fig.1</figureCitation>
). Wing expanse 
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. Coloration of vestiture. Head clothed with greyish brown hair-like scales. Dorsal surface of antennal shaft covered with greyish brown scales, mixed with greyish yellow ones on apical part and posterior surface of each flagellomere. Thoracic nota clothed with brown hair-like scales; abdomen clothed above with greyish brown hair-like scales, somewhat darker on anterior part. Legs covered with pale brown scales, mixed with yellowish grey ones on apical parts of tarsomeres. Forewing upperside. In well-reticulated specimens (
<figureCitation id="27F82A662748FF895AC2F98D4382F9A2" box="[500,649,1617,1642]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="1" pageNumber="144">Figs. 1A, 1C</figureCitation>
), costal area and anterior 1/2 of discoidal cell brown, remaining area pale greyish yellow and reticulated with dark brown vein stripes and narrow transverse stripes; 3–4 transverse stripes present beyond discoidal cell, and width of stripes slightly wider than dark vein stripes; reticulation indistinct on basal 1/2 of wing. In darker specimens (
<figureCitation id="27F82A662748FF8959CFF961425AF91E" box="[761,849,1725,1750]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="1" pageNumber="144">Fig. 1B</figureCitation>
), transverse stripes wider, as wide as or wider than yellowish speckles between stripes. Fringe uniformly brown. Hindwing upperside semitransparent brownish grey, vein lines brown; fringe brown, apical part of fringe on termen posterior to space 2 yellowish white.?Antenna (
<figureCitation id="27F82A662748FF895E59F8D941B2F88A" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="1" pageNumber="144">Fig. 1K</figureCitation>
) slightly shorter than 1/2 length of forewing including fringe (0.40–0.45); flagellum bipectinated and consisting of 20–22 (M. 21.1) flagellomeres; pectination on 9th to 10th flagellomeres longest, 
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(M. 
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) long, 2.1–2.8 (M. 2.42) times as long as flagellomere; pectinations gradually elongated from basal flagellomeres to flagellomere 8, almost keeping maximum length until 10th, then gradually shortened towards antennal tip. Fore tibia with epiphysis, 0.36–0.57 (M. 0.51) times as long as fore tibia, arising from 0.34–0.46 (M. 0.41) length of tibia from base.
</paragraph>
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<paragraph id="BF7C36E3274BFF8A5BA1F90345FDF899" blockId="2.[151,1436,1759,1873]" pageId="2" pageNumber="145">
<emphasis id="8DB7EAF1274BFF8A5BA1F9034012F93C" bold="true" box="[151,281,1759,1781]" pageId="2" pageNumber="145">FIGURE 1.</emphasis>
<taxonomicName id="78C34D60274BFF8A5A13F93C40FCF93D" authority="Saigusa &amp; Sugimoto, 2014" authorityName="Saigusa &amp; Sugimoto" authorityYear="2014" box="[293,503,1759,1781]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="maculatella" status="sp. nov.">
<emphasis id="8DB7EAF1274BFF8A5A13F93C40FCF93D" box="[293,503,1759,1781]" italics="true" pageId="2" pageNumber="145">Proutia maculatella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684578A274BFF8A5935F9034351F93D" box="[515,602,1759,1781]" pageId="2" pageNumber="145" rank="species">
<emphasis id="8DB7EAF1274BFF8A5935F903435FF93C" bold="true" box="[515,596,1759,1780]" pageId="2" pageNumber="145">sp. nov</emphasis>
.
</taxonomicNameLabel>
A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated.
</paragraph>
</caption>
<caption id="EBBC666B274AFF8B5BA1F8A64083F823" ID-DOI="http://doi.org/10.5281/zenodo.4947203" ID-Zenodo-Dep="4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="3" pageNumber="146" startId="3.[151,250,1914,1936]" targetBox="[165,1420,200,1879]" targetPageId="3">
<paragraph id="BF7C36E3274AFF8B5BA1F8A64083F823" blockId="3.[151,1436,1914,2027]" pageId="3" pageNumber="146">
<emphasis id="8DB7EAF1274AFF8B5BA1F8A64018F847" bold="true" box="[151,275,1914,1936]" pageId="3" pageNumber="146">FIGURE 2.</emphasis>
<taxonomicName id="78C34D60274AFF8B5A2CF8A740EFF858" authority="Saigusa &amp; Sugimoto, 2014" authorityName="Saigusa &amp; Sugimoto" authorityYear="2014" box="[282,484,1914,1936]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="maculatella" status="sp. nov.">
<emphasis id="8DB7EAF1274AFF8B5A2CF8A740EFF858" box="[282,484,1914,1936]" italics="true" pageId="3" pageNumber="146">Proutia maculatella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684578A274AFF8B5ADDF8A64330F847" box="[491,571,1914,1935]" pageId="3" pageNumber="146" rank="species">
<emphasis id="8DB7EAF1274AFF8B5ADDF8A6433DF847" bold="true" box="[491,566,1914,1935]" pageId="3" pageNumber="146">sp. nov</emphasis>
.
</taxonomicNameLabel>
A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa.
</paragraph>
</caption>
<caption id="EBBC666B274DFF8C5BA1FD0C45EDFD2E" ID-DOI="http://doi.org/10.5281/zenodo.4947205" ID-Zenodo-Dep="4947205" box="[151,1254,720,742]" httpUri="https://zenodo.org/record/4947205/files/figure.png" pageId="4" pageNumber="147" startId="4.[151,250,720,742]" targetBox="[343,1242,197,695]" targetPageId="4">
<paragraph id="BF7C36E3274DFF8C5BA1FD0C45EDFD2E" blockId="4.[151,1254,720,742]" box="[151,1254,720,742]" pageId="4" pageNumber="147">
<emphasis id="8DB7EAF1274DFF8C5BA1FD0C4019FD2D" bold="true" box="[151,274,720,742]" pageId="4" pageNumber="147">FIGURE 3.</emphasis>
Wing venation of 
<taxonomicName id="78C34D60274DFF8C5AF9FD0D4347FD2E" box="[463,588,721,742]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8DB7EAF1274DFF8C5AF9FD0D4317FD2E" box="[463,540,721,742]" italics="true" pageId="4" pageNumber="147">Proutia</emphasis>
spp.
</taxonomicName>
males. A: 
<taxonomicName id="78C34D60274DFF8C598FFD0D428FFD2E" authorityName="Saigusa &amp; Sugimoto" authorityYear="2014" box="[697,900,720,742]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="maculatella">
<emphasis id="8DB7EAF1274DFF8C598FFD0D428FFD2E" box="[697,900,720,742]" italics="true" pageId="4" pageNumber="147">Proutia maculatella</emphasis>
</taxonomicName>
s 
<emphasis id="8DB7EAF1274DFF8C58A2FD0C42DFFD2D" bold="true" box="[916,980,720,741]" pageId="4" pageNumber="147">p. nov</emphasis>
.; B: 
<taxonomicName id="78C34D60274DFF8C5F30FD0D459BFD2E" authority="Saigusa &amp; Sugimoto, 2014" authorityName="Saigusa &amp; Sugimoto" authorityYear="2014" box="[1030,1168,721,742]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
<emphasis id="8DB7EAF1274DFF8C5F30FD0D459BFD2E" box="[1030,1168,721,742]" italics="true" pageId="4" pageNumber="147">Proutia nigra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684578A274DFF8C5FA0FD0C45EDFD2E" box="[1174,1254,720,742]" pageId="4" pageNumber="147" rank="species">
<emphasis id="8DB7EAF1274DFF8C5FA0FD0C45EBFD2D" bold="true" box="[1174,1248,720,741]" pageId="4" pageNumber="147">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="EBBC666B274DFF8C5BA1F9084443F921" ID-DOI="http://doi.org/10.5281/zenodo.4947207" ID-Zenodo-Dep="4947207" box="[151,1352,1748,1770]" httpUri="https://zenodo.org/record/4947207/files/figure.png" pageId="4" pageNumber="147" startId="4.[151,250,1748,1770]" targetBox="[452,1136,801,1718]" targetPageId="4">
<paragraph id="BF7C36E3274DFF8C5BA1F9084443F921" blockId="4.[151,1352,1748,1770]" box="[151,1352,1748,1770]" pageId="4" pageNumber="147">
<emphasis id="8DB7EAF1274DFF8C5BA1F9084019F921" bold="true" box="[151,274,1748,1770]" pageId="4" pageNumber="147">FIGURE 4.</emphasis>
Lateral aspect of male genitalia of 
<taxonomicName id="78C34D60274DFF8C594EF908421AF922" box="[632,785,1748,1770]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="8DB7EAF1274DFF8C594EF90843CEF921" box="[632,709,1748,1769]" italics="true" pageId="4" pageNumber="147">Proutia</emphasis>
spp. A
</taxonomicName>
: 
<taxonomicName id="78C34D60274DFF8C582DF90842EEF922" authorityName="Saigusa &amp; Sugimoto" authorityYear="2014" box="[795,997,1748,1770]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="maculatella">
<emphasis id="8DB7EAF1274DFF8C582DF90842EEF922" box="[795,997,1748,1770]" italics="true" pageId="4" pageNumber="147">Proutia maculatella</emphasis>
</taxonomicName>
s 
<emphasis id="8DB7EAF1274DFF8C58C3F908453DF921" bold="true" box="[1013,1078,1748,1769]" pageId="4" pageNumber="147">p. nov</emphasis>
.; B: 
<taxonomicName id="78C34D60274DFF8C5F51F90845FAF921" authority="Saigusa &amp; Sugimoto, 2014" authorityName="Saigusa &amp; Sugimoto" authorityYear="2014" box="[1127,1265,1748,1769]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">
<emphasis id="8DB7EAF1274DFF8C5F51F90845FAF921" box="[1127,1265,1748,1769]" italics="true" pageId="4" pageNumber="147">Proutia nigra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684578A274DFF8C5FC1F9084443F921" box="[1271,1352,1748,1769]" pageId="4" pageNumber="147" rank="species">
<emphasis id="8DB7EAF1274DFF8C5FC1F9084449F921" bold="true" box="[1271,1346,1748,1769]" pageId="4" pageNumber="147">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="BF7C36E3274DFF8D5BF1F8CD421FFF1D" blockId="4.[151,1437,1809,2014]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="148" pageId="4" pageNumber="147">
Forewing 
<quantity id="783B9B06274DFF8C5A0FF8CD40B4F8E1" box="[313,447,1809,1834]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="7.6" metricValueMin="6.6" pageId="4" pageNumber="147" unit="mm" value="7.1" valueMax="7.6" valueMin="6.6">6.6–7.6 mm</quantity>
(M. 7.0 mm) in length excluding fringe, 2.30–2.52 (M. 2.41) times as long as wide, with termen distinctly rounded; forewing length including fringe 
<quantity id="783B9B06274DFF8C587EF8E942DAF885" box="[840,977,1845,1870]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="8.7" metricValueMin="7.4" pageId="4" pageNumber="147" unit="mm" value="8.05" valueMax="8.7" valueMin="7.4">7.4–8.7 mm</quantity>
(M. 
<quantity id="783B9B06274DFF8C5F3AF8E9457CF886" box="[1036,1143,1845,1870]" metricMagnitude="-3" metricUnit="m" metricValue="8.31" pageId="4" pageNumber="147" unit="mm" value="8.31">8.31 mm</quantity>
); discoidal cell 0.68–0.74 (M. 0.71) times as long as forewing. Hindwing 
<quantity id="783B9B06274DFF8C59F2F8854245F8B9" box="[708,846,1881,1906]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.9" metricValueMin="5.2" pageId="4" pageNumber="147" unit="mm" value="5.55" valueMax="5.9" valueMin="5.2">5.2–5.9 mm</quantity>
(M 
<quantity id="783B9B06274DFF8C58B2F88542D5F8BA" box="[900,990,1881,1906]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="147" unit="mm" value="5.5">5.5 mm</quantity>
) long excluding fringe, 2.00–2.16 (M. 2.08) times as long as wide, with termen rounded; discoidal cell 0.54–0.59 (M. 0.57) times as long as hindwing. Forewing venation (
<figureCitation id="27F82A66274DFF8C5A4DF87D40DCF872" box="[379,471,1953,1978]" captionStart="FIGURE 3" captionStartId="4.[151,250,720,742]" captionTargetBox="[343,1242,197,695]" captionTargetId="figure-181@4.[340,1247,193,699]" captionTargetPageId="4" captionText="FIGURE 3. Wing venation of Proutia spp. males. A: Proutia maculatella sp. nov.; B: Proutia nigra sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4947205" httpUri="https://zenodo.org/record/4947205/files/figure.png" pageId="4" pageNumber="147">Fig. 3A</figureCitation>
): Accessory cell absent; intercalary cell present; forking point of vein M in discoidal cell at level of origin of vein CuA2 or at middle between origins of veins R1 and R2; bases of all veins from cell separated from each other. Hindwing venation (
<figureCitation id="27F82A66274CFF8D59F7FF4B4214FF78" box="[705,799,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,720,742]" captionTargetBox="[343,1242,197,695]" captionTargetId="figure-181@4.[340,1247,193,699]" captionTargetPageId="4" captionText="FIGURE 3. Wing venation of Proutia spp. males. A: Proutia maculatella sp. nov.; B: Proutia nigra sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4947205" httpUri="https://zenodo.org/record/4947205/files/figure.png" pageId="5" pageNumber="148">Fig. 3A</figureCitation>
): Veins Rs and M1 usually separated, but occasionally connate or short-stalked; vein M in discoidal cell simple.
</paragraph>
<paragraph id="BF7C36E3274CFF8D5BF1FF03428CFD38" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="148">
Wing vestiture. Forewing upperside covered with hair-like upper scales (
<figureCitation id="27F82A66274CFF8D5F32FF03456DFF30" box="[1028,1126,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="5" pageNumber="148">Fig. 1 H</figureCitation>
) (approximately 
<quantity id="783B9B06274CFF8D5E1CFF034497FF30" box="[1322,1436,223,248]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="5" pageNumber="148" unit="m" value="375.0" valueMax="500.0" valueMin="250.0">250–500µ</quantity>
long, 
<quantity id="783B9B06274CFF8D5BE1FED8403EFED4" box="[215,309,260,285]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="148" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">10–20 µ</quantity>
wide) and apically bidentate lower scales (
<figureCitation id="27F82A66274CFF8D582BFED84262FED5" box="[797,873,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="5" pageNumber="148">Fig. 1I</figureCitation>
) (approximately 
<quantity id="783B9B06274CFF8D5F1BFED84580FED4" box="[1069,1163,260,285]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="5" pageNumber="148" unit="m" value="55.0" valueMax="70.0" valueMin="40.0">40–70 µ</quantity>
long, 
<quantity id="783B9B06274CFF8D5FE5FED8443AFED4" box="[1235,1329,260,285]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="148" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 µ</quantity>
wide) on basal 1/3, apically bidentate upper scales (approximately 
<quantity id="783B9B06274CFF8D5803FEFB42B9FE88" box="[821,946,295,320]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="5" pageNumber="148" unit="m" value="140.0" valueMax="150.0" valueMin="130.0">130–150 µ</quantity>
long, 
<quantity id="783B9B06274CFF8D5F37FEFB4569FE88" box="[1025,1122,295,320]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="148" unit="m" value="35.0" valueMax="40.0" valueMin="30.0">30–40 µ</quantity>
wide) and bidentate lower scales (approximately 
<quantity id="783B9B06274CFF8D5AA5FE9040FBFEAC" box="[403,496,332,357]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="5" pageNumber="148" unit="m" value="50.0" valueMax="60.0" valueMin="40.0">40–60 µ</quantity>
long, 
<quantity id="783B9B06274CFF8D590EFE904360FEAC" box="[568,619,332,357]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="148" unit="m" value="20.0">20 µ</quantity>
wide) on apical 1/2 of discoidal cell, and mostly tridentate (occasionally bi- or quadridentate) upper scales (approximately 
<quantity id="783B9B06274CFF8D59FBFEB3424CFE40" box="[717,839,367,392]" metricMagnitude="2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="5" pageNumber="148" unit="m" value="190.0" valueMax="200.0" valueMin="180.0">180–200 µ</quantity>
long, 
<quantity id="783B9B06274CFF8D58A6FEB342E5FE40" box="[912,1006,367,392]" metricMagnitude="1" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="5" pageNumber="148" unit="m" value="47.5" valueMax="50.0" valueMin="45.0">45–50 µ</quantity>
wide, 
<quantity id="783B9B06274CFF8D5F0DFEB34592FE40" box="[1083,1177,367,392]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="5" pageNumber="148" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">15–20 µ</quantity>
long of dentation) and bidentate lower scales (approximately 
<quantity id="783B9B06274CFF8D597DFE4843A3FE64" box="[587,680,404,429]" metricMagnitude="1" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="148" unit="m" value="45.0" valueMax="50.0" valueMin="40.0">40–50 µ</quantity>
long, 
<quantity id="783B9B06274CFF8D59C7FE484244FE64" box="[753,847,404,429]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="5" pageNumber="148" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">15–20 µ</quantity>
wide) on apical 1/3 of wing. Measurement of slidemounted scales taken mainly from distal 2/3 of upperside of forewing as follows (larger upper scales and smaller lower scales were measured). Upper scales 
<quantity id="783B9B06274CFF8D59B2FE0043F4FE3C" box="[644,767,476,501]" metricMagnitude="2" metricUnit="m" metricValue="1.625" metricValueMax="1.8" metricValueMin="1.45" pageId="5" pageNumber="148" unit="m" value="162.5" valueMax="180.0" valueMin="145.0">145–180 µ</quantity>
long (M 
<quantity id="783B9B06274CFF8D585DFE0042A4FE3D" box="[875,943,476,501]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="148" unit="m" value="150.0">150 µ</quantity>
), 
<quantity id="783B9B06274CFF8D58F3FE004528FE3C" box="[965,1059,476,501]" metricMagnitude="1" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="5" pageNumber="148" unit="m" value="51.0" valueMax="54.0" valueMin="48.0">48–54 µ</quantity>
wide (M 
<quantity id="783B9B06274CFF8D5FA2FE0045D4FE3D" box="[1172,1247,476,501]" metricMagnitude="1" metricUnit="m" metricValue="4.45" pageId="5" pageNumber="148" unit="m" value="44.5">44.5 µ</quantity>
), with serrations 
<quantity id="783B9B06274CFF8D5BA1FE2341FFFDD0" box="[151,244,511,536]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="148" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">20–30 µ</quantity>
(M 
<quantity id="783B9B06274CFF8D5A15FE234053FDD0" box="[291,344,511,536]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="148" unit="m" value="24.0">24 µ</quantity>
); lower scales 
<quantity id="783B9B06274CFF8D5AC8FE234351FDD0" box="[510,602,511,536]" metricMagnitude="1" metricUnit="m" metricValue="7.25" metricValueMax="8.5" metricValueMin="6.0" pageId="5" pageNumber="148" unit="m" value="72.5" valueMax="85.0" valueMin="60.0">60–85 µ</quantity>
(M 
<quantity id="783B9B06274CFF8D59BCFE2343B6FDD0" box="[650,701,511,536]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="148" unit="m" value="66.0">66 µ</quantity>
) long, 
<quantity id="783B9B06274CFF8D583AFE234263FDD0" box="[780,872,511,536]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="5" pageNumber="148" unit="m" value="20.0" valueMax="25.0" valueMin="15.0">15–25 µ</quantity>
wide (M 
<quantity id="783B9B06274CFF8D58E3FE234514FDD0" box="[981,1055,511,536]" metricMagnitude="1" metricUnit="m" metricValue="1.97" pageId="5" pageNumber="148" unit="m" value="19.7">19.7 µ</quantity>
), with serrations 
<quantity id="783B9B06274CFF8D5FD6FE234437FDD0" box="[1248,1340,511,536]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="148" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 µ</quantity>
(M 
<quantity id="783B9B06274CFF8D5E5AFE2341A2FDF5" metricMagnitude="1" metricUnit="m" metricValue="1.18" pageId="5" pageNumber="148" unit="m" value="11.8">11.8 µ</quantity>
). Fringe consisting of long slender scales, scales very narrow on basal 2/3, widened subapically, and ending in pointed tip, with 2–3 small dentations on both lateral margins towards apex. Hindwing rather sparsely clothed with hair-like upper scales (approximately 
<quantity id="783B9B06274CFF8D5978FDB043C7FD4C" box="[590,716,620,645]" metricMagnitude="2" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="5" pageNumber="148" unit="m" value="255.0" valueMax="290.0" valueMin="220.0">220–290 µ</quantity>
long, 
<quantity id="783B9B06274CFF8D582CFDB04266FD4C" box="[794,877,620,645]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="5" pageNumber="148" unit="m" value="8.5" valueMax="10.0" valueMin="7.0">7–10 µ</quantity>
wide) and usually short bidentate (occasionally simple distally) lower scales (approximately 
<quantity id="783B9B06274CFF8D5990FD53420CFD60" box="[678,775,655,680]" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="5" pageNumber="148" unit="m" value="55.0" valueMax="60.0" valueMin="50.0">50–60 µ</quantity>
long, 
<quantity id="783B9B06274CFF8D586EFD5342A7FD60" box="[856,940,655,680]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="5" pageNumber="148" unit="m" value="8.5" valueMax="10.0" valueMin="7.0">7–10 µ</quantity>
wide) on almost whole surface; fringe of hindwing similar to forewing, but inner margin to basal part of outer margin almost hair-like. Hindwing with semitransparent appearance owing to covering of hair-like upper scales.
</paragraph>
<paragraph id="BF7C36E3274CFF8D5BF1FD20442EFC25" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="148">
Male genitalia (
<figureCitation id="27F82A66274CFF8D5A4FFD2040F9FCDD" box="[377,498,764,789]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="5" pageNumber="148">Fig. 1D–G</figureCitation>
, 
<figureCitation id="27F82A66274CFF8D5AC8FD20432EFCDD" box="[510,549,764,789]" captionStart="FIGURE 4" captionStartId="4.[151,250,1748,1770]" captionTargetBox="[452,1136,801,1718]" captionTargetId="figure-190@4.[446,1141,792,1726]" captionTargetPageId="4" captionText="FIGURE 4. Lateral aspect of male genitalia of Proutia spp. A: Proutia maculatella sp. nov.; B: Proutia nigra sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4947207" httpUri="https://zenodo.org/record/4947207/files/figure.png" pageId="5" pageNumber="148">4A</figureCitation>
): Similar to those of 
<taxonomicName id="78C34D60274CFF8D5822FD214287FCDC" baseAuthorityName="Zeller" baseAuthorityYear="1839" box="[788,908,764,788]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="species" species="betulina">
<emphasis id="8DB7EAF1274CFF8D5822FD214287FCDC" box="[788,908,764,788]" italics="true" pageId="5" pageNumber="148">P. betulina</emphasis>
</taxonomicName>
except as follows: ampulla (dorsal process) of valva (
<figureCitation id="27F82A66274CFF8D5BD3FCC34036FCF0" box="[229,317,799,824]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="5" pageNumber="148">Fig. 1D</figureCitation>
) longer and stouter, harpe (ventral process of valva) larger, saccus shorter, in lateral aspect dorsal part of ring (element of tegumen) (
<figureCitation id="27F82A66274CFF8D5928FC98437DFC95" box="[542,630,836,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1915,1937]" captionTargetBox="[156,1429,633,1875]" captionTargetId="figure-267@7.[151,1435,625,1894]" captionTargetPageId="7" captionText="FIGURE 5. Proutia nigra sp. nov. A: male; B: slide-mounted male wings; C: dorsal aspect of male genital dorsum; D: dorsal aspect of male genital anellus; E: male right genital valva; F: male phallus; G: upper (larger) and lower (smaller) scales of upperside of forewing; H: upper (larger) and lower (smaller) scales of upperside of hind wing; I: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947209" httpUri="https://zenodo.org/record/4947209/files/figure.png" pageId="5" pageNumber="148">Fig. 5A</figureCitation>
) wider; (compare illustrations of 
<taxonomicName id="78C34D60274CFF8D58C6FC984547FC94" baseAuthorityName="Zeller" baseAuthorityYear="1839" box="[1008,1100,836,860]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="species" species="betulina">
<emphasis id="8DB7EAF1274CFF8D58C6FC984547FC94" box="[1008,1100,836,860]" italics="true" pageId="5" pageNumber="148">betulina</emphasis>
</taxonomicName>
in 
<bibRefCitation id="DB524B12274CFF8D5F46FC984467FC95" author="Kozhantshikov, I. V." box="[1136,1388,836,861]" pageId="5" pageNumber="148" refId="ref5770" refString="Kozhantshikov, I. V. (1956) Fauna SSSR, Hansekomye cheshuekrylye 3 (2) Chekhlonosy-meshechnitsy (sem. Psychidae). Izdatelstvo Akademii Nauk SSSR, Moskva-Leningrad, 517 pp." type="book" year="1956">Kozhantshikov (1956)</bibRefCitation>
and dissected genitalia of one 
<taxonomicName id="78C34D60274CFF8D5AF2FCB4432BFC48" baseAuthorityName="Zeller" baseAuthorityYear="1839" box="[452,544,872,896]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="species" species="betulina">
<emphasis id="8DB7EAF1274CFF8D5AF2FCB4432BFC48" box="[452,544,872,896]" italics="true" pageId="5" pageNumber="148">betulina</emphasis>
</taxonomicName>
specimen from Berlin). Relative length of ampulla to dorsal margin of valva (including ampulla and excluding transtilla) 0.48–0.52; ampulla 2.4–2.6 times as long as narrowest width; distal margin of dorsum (
<figureCitation id="27F82A66274CFF8D5AB5FC7340D0FC00" box="[387,475,943,968]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="5" pageNumber="148">Fig. 1F</figureCitation>
) weakly produced to pair of short, apically round projections; phallus (Fig. E) moderately curved without denticles on vesica; anellus (
<figureCitation id="27F82A66274CFF8D583BFC084269FC25" box="[781,866,980,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1759,1781]" captionTargetBox="[159,1429,200,1727]" captionTargetId="figure-101@2.[150,1436,193,1738]" captionTargetPageId="2" captionText="FIGURE 1. Proutia maculatella sp. nov. A: male with well-reticulated forewings; B: male with obscurely maculated forewings; C: slide-mounted male wings; D: male genital right valva; E: male phallus; F: dorsal aspect of male genital dorsum; G: dorsal aspect of male genital anellus; H: upper scale of male forewing upperside; I: lower scale of male forewing upperside; J: upper scales (long ones) and two lower scales of male hindwing upperside; K: male left antenna, macerated." figureDoi="http://doi.org/10.5281/zenodo.4947199" httpUri="https://zenodo.org/record/4947199/files/figure.png" pageId="5" pageNumber="148">Fig. 1G</figureCitation>
, valvae penis) bearing only fine setulae.
</paragraph>
<paragraph id="BF7C36E3274CFF8D5BF1FC2B4365FB68" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="148">
<emphasis id="8DB7EAF1274CFF8D5BF1FC2B40CFFBD8" box="[199,452,1015,1040]" inLineHeading="true" pageId="5" pageNumber="148" reason="1">
<emphasis id="8DB7EAF1274CFF8D5BF1FC2B4015FBD8" bold="true" box="[199,286,1015,1040]" pageId="5" pageNumber="148">Female</emphasis>
(
<figureCitation id="27F82A66274CFF8D5A18FC2B40B3FBD8" box="[302,440,1015,1040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="5" pageNumber="148">Figs. 2D–H</figureCitation>
).
</emphasis>
Coloration. Sclerites including head, thorax, legs and abdominal terga and sterna dark brown. Head (
<figureCitation id="27F82A66274CFF8D5A0AFBC04098FBFD" box="[316,403,1052,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="5" pageNumber="148">Fig. 2E</figureCitation>
) with pair of pale areas on vertex. Meso- and metanotum with dark dorsal and sublateral markings succeeding to markings of mature larvae. Membranous areas of abdomen light reddish brown owing to coloration of epidermal cells, but cuticula colorless. Abdomen sparsely clothed with light greyish brown fine hairlike scales; corethrogyne yellowish white.
</paragraph>
<paragraph id="BF7C36E3274CFF8D5BF1FB704018FAF8" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="148">
Structure. Antenna (Fig. E) rather long for 
<taxonomicName id="78C34D60274CFF8D598CFB714205FB0C" authorityName="Tutt" authorityYear="1899" box="[698,782,1197,1220]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="8DB7EAF1274CFF8D598CFB714205FB0C" box="[698,782,1197,1220]" italics="true" pageId="5" pageNumber="148">Proutia</emphasis>
</taxonomicName>
, 
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long, 1.6–2.1 times as long as head width, flagellum usually well segmented into 12–14 flagellomeres, segmentation partly more or less incomplete in some specimens. Legs (
<figureCitation id="27F82A66274CFF8D5A57FB2840E8FAC5" box="[353,483,1268,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="5" pageNumber="148">Figs. 2F–H</figureCitation>
) with tarsi divided into 3 tarsomeres in most cases, but into 
<quantity id="783B9B06274CFF8D5FA5FB2845B5FAC5" box="[1171,1214,1268,1293]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="148" unit="in" value="4.0">4 in</quantity>
some legs in a few specimens.
</paragraph>
<paragraph id="BF7C36E3274CFF8D5BF1FAE0449EFA9B" blockId="5.[151,1437,151,2013]" box="[199,1429,1340,1365]" pageId="5" pageNumber="148">
Distance from head to apical part of 7th abdominal segment in natural posture curving ventrally: 
<quantity id="783B9B06274CFF8D5E31FAE0449AFA9B" box="[1287,1425,1340,1365]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="5" pageNumber="148" unit="mm" value="3.9" valueMax="4.3" valueMin="3.5">3.5–4.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="F7D96568274CFF8D5BF1FA8342B0F8EF" pageId="5" pageNumber="148" type="materials_examined">
<paragraph id="BF7C36E3274CFF8D5BF1FA8340A8F998" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="148">
<emphasis id="8DB7EAF1274CFF8D5BF1FA83407CFABF" box="[199,375,1375,1400]" inLineHeading="true" pageId="5" pageNumber="148" reason="1">
<emphasis id="8DB7EAF1274CFF8D5BF1FA83407BFAB0" bold="true" box="[199,368,1375,1400]" pageId="5" pageNumber="148">Type material</emphasis>
.
</emphasis>
<materialsCitation id="0FAB3CBE274CFF8D5AB7FA8343E6FA53" ID-GBIF-Occurrence="3131383301" collectingDate="1970-04-16" collectingDateMax="1970-04-18" collectingDateMin="1970-04-16" collectionCode="T" country="Japan" county="Kyushu" location="Kashii" municipality="Fukoka-shi" pageId="5" pageNumber="148" specimenCount="1" specimenCount-male="1" stateProvince="Fukuoka" typeStatus="holotype">
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<typeStatus id="60788841274CFF8D5AB7FA834314FAB0" box="[385,543,1375,1400]" pageId="5" pageNumber="148" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="A9C5FD6A274CFF8D591FFABC434BFABF" box="[553,576,1376,1399]" pageId="5" pageNumber="148" type="male">♂</specimenCount>
, 
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, 
<collectingMunicipality id="5F18AC99274CFF8D599FFA834221FAB0" box="[681,810,1375,1400]" pageId="5" pageNumber="148">Fukoka-shi</collectingMunicipality>
, 
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, 
<collectingCounty id="561D4E6F274CFF8D58DFFA834548FAB0" box="[1001,1091,1375,1400]" pageId="5" pageNumber="148">Kyushu</collectingCounty>
, 
<collectingCountry id="C7D47673274CFF8D5F67FABC459FFABF" box="[1105,1172,1376,1399]" name="Japan" pageId="5" pageNumber="148">Japan</collectingCountry>
, 
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</date>
, 
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. Saigusa. Donated to the Kyushu University Museum
</materialsCitation>
. 
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<typeStatus id="60788841274CFF8D59C1FA584290FA55" box="[759,923,1412,1437]" pageId="5" pageNumber="148" type="paratype">PARATYPES</typeStatus>
</emphasis>
: 
<specimenCount id="A9C5FD6A274CFF8D589CFA5842D5FA55" box="[938,990,1412,1437]" pageId="5" pageNumber="148" type="male">17♂</specimenCount>
(
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macerated, right wings venation on a slide), same locality as holotype, 
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<collectingDate id="DB39E9CB274CFF8D5913FA7B4259FA08" box="[549,850,1447,1472]" pageId="5" pageNumber="148" value="1958-03-31" valueMax="1958-04-05" valueMin="1958-03-31">March 31 to April 5, 1958</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D5869FA74427AFA77" box="[863,881,1448,1471]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D584AFA7B42DCFA08" box="[892,983,1447,1472]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
; 
<materialsCitation id="0FAB3CBE274CFF8D58D5FA7B45C4FA2D" ID-GBIF-Occurrence="3131383310" collectingDate="1958-04-14" collectionCode="T" collectorName="Saigusa" country="Japan" county="Najima" location="Kyushu" municipality="Fukuoka-shi" pageId="5" pageNumber="148" specimenCount="6" specimenCount-male="6" stateProvince="Fukuoka" typeStatus="paratype">
<specimenCount id="A9C5FD6A274CFF8D58D5FA7B450FFA08" box="[995,1028,1447,1472]" pageId="5" pageNumber="148" type="male">4♂</specimenCount>
(
<specimenCount id="A9C5FD6A274CFF8D5F3AFA7B4527FA08" box="[1036,1068,1447,1472]" pageId="5" pageNumber="148" type="male">2♂</specimenCount>
macerated, right wings venation on slides), 
<collectingCounty id="561D4E6F274CFF8D5A26FA10406EFA2D" box="[272,357,1484,1509]" pageId="5" pageNumber="148">Najima</collectingCounty>
, 
<collectingMunicipality id="5F18AC99274CFF8D5A59FA1040F5FA2D" box="[367,510,1484,1509]" pageId="5" pageNumber="148">Fukuoka-shi</collectingMunicipality>
, 
<collectingRegion id="7D07F801274CFF8D593FFA1043A1FA2D" box="[521,682,1484,1509]" country="Japan" name="Fukuoka" pageId="5" pageNumber="148">Fukuoka Pref.</collectingRegion>
, 
<location id="BA1C6038274CFF8D5982FA104205FA2D" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D5982FA104205FA2D" box="[692,782,1484,1509]" country="Japan" county="Najima" municipality="Fukuoka-shi" name="Kyushu" pageId="5" pageNumber="148" stateProvince="Fukuoka">Kyushu</location>
, 
<location id="BA1C6038274CFF8D582FFA104269FA2D" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D582FFA104269FA2D" box="[793,866,1484,1509]" country="Japan" county="Najima" municipality="Fukuoka-shi" name="March" pageId="5" pageNumber="148" stateProvince="Fukuoka">March</location>
31 &amp; 
<date id="CB7D1023274CFF8D5891FA104546FA2D" box="[935,1101,1484,1509]" pageId="5" pageNumber="148" value="1958-04-14">
<collectingDate id="DB39E9CB274CFF8D5891FA104546FA2D" box="[935,1101,1484,1509]" pageId="5" pageNumber="148" value="1958-04-14">April 14, 1958</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D5F61FA104561FA2B" box="[1111,1130,1484,1507]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5F42FA1045C4FA2D" box="[1140,1231,1484,1509]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
; 
<materialsCitation id="0FAB3CBE274CFF8D5FEFFA1042F1F9C0" ID-GBIF-Occurrence="3131383317" collectingDate="1970-04-10" collectingDateMax="1970-04-16" collectingDateMin="1970-04-10" collectionCode="T" collectorName="Saigusa" country="Japan" county="Kyushu" location="Kashii" municipality="Fukoka-shi" pageId="5" pageNumber="148" specimenCount="22" specimenCount-female="5" specimenCount-male="17" stateProvince="Fukuoka" typeStatus="paratype">
<specimenCount id="A9C5FD6A274CFF8D5FEFFA104402FA2D" box="[1241,1289,1484,1509]" pageId="5" pageNumber="148" type="male">15♂</specimenCount>
<specimenCount id="A9C5FD6A274CFF8D5E3FFA10442DFA2D" box="[1289,1318,1484,1509]" pageId="5" pageNumber="148" type="female">4♀</specimenCount>
(
<specimenCount id="A9C5FD6A274CFF8D5E19FA104459FA2D" box="[1327,1362,1484,1509]" pageId="5" pageNumber="148" type="male">2♂</specimenCount>
wings only; 
<specimenCount id="A9C5FD6A274CFF8D5BEEFA3341E3F9C0" box="[216,232,1519,1544]" pageId="5" pageNumber="148" type="female">♀</specimenCount>
in ethanol), same locality as holotype, 
<date id="CB7D1023274CFF8D5995FA33427EF9C0" box="[675,885,1519,1544]" pageId="5" pageNumber="148" value="1970-04-10" valueMax="1970-04-16" valueMin="1970-04-10">
<collectingDate id="DB39E9CB274CFF8D5995FA33427EF9C0" box="[675,885,1519,1544]" pageId="5" pageNumber="148" value="1970-04-10" valueMax="1970-04-16" valueMin="1970-04-10">April 10–16, 1970</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D58B4FA2C429EF9CF" box="[898,917,1520,1543]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D58A9FA3342F1F9C0" box="[927,1018,1519,1544]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
; 
<specimenCount id="A9C5FD6A274CFF8D5F30FA33452CF9C0" box="[1030,1063,1519,1544]" pageId="5" pageNumber="148" type="male">3♂</specimenCount>
(macerated, right wings venation on slides) 
<materialsCitation id="0FAB3CBE274CFF8D5A3AF9C84094F998" ID-GBIF-Occurrence="3131383312" collectingDate="2002-04-16" collectionCode="T" collectorName="M. Sugimoto &amp; Saigusa" country="Japan" county="Sakato" location="Kyushu" municipality="Kasuya-machi" pageId="5" pageNumber="148" specimenCount="52" specimenCount-female="52" stateProvince="Fukuoka" typeStatus="paratype">
<specimenCount id="A9C5FD6A274CFF8D5A3AF9C84037F9E5" box="[268,316,1556,1581]" pageId="5" pageNumber="148" type="female">26♀</specimenCount>
(
<specimenCount id="A9C5FD6A274CFF8D5A73F9C84078F9E5" box="[325,371,1556,1581]" pageId="5" pageNumber="148" type="female">21♀</specimenCount>
in ethanol, 
<specimenCount id="A9C5FD6A274CFF8D5AC1F9C8431AF9E5" box="[503,529,1556,1581]" pageId="5" pageNumber="148" type="female">5♀</specimenCount>
macerated), 
<collectingCounty id="561D4E6F274CFF8D5990F9C843FEF9E5" box="[678,757,1556,1581]" pageId="5" pageNumber="148">Sakato</collectingCounty>
, 
<collectingMunicipality id="5F18AC99274CFF8D5837F9C842ADF9E5" box="[769,934,1556,1581]" pageId="5" pageNumber="148">Kasuya-machi</collectingMunicipality>
, 
<collectingRegion id="7D07F801274CFF8D5884F9C8455EF9E5" box="[946,1109,1556,1581]" country="Japan" name="Fukuoka" pageId="5" pageNumber="148">Fukuoka Pref.</collectingRegion>
, 
<location id="BA1C6038274CFF8D5F57F9C845B0F9E5" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D5F57F9C845B0F9E5" box="[1121,1211,1556,1581]" country="Japan" county="Sakato" municipality="Kasuya-machi" name="Kyushu" pageId="5" pageNumber="148" stateProvince="Fukuoka">Kyushu</location>
, 
<date id="CB7D1023274CFF8D5FF1F9C8447BF9E5" box="[1223,1392,1556,1581]" pageId="5" pageNumber="148" value="2002-04-16">
<collectingDate id="DB39E9CB274CFF8D5FF1F9C8447BF9E5" box="[1223,1392,1556,1581]" pageId="5" pageNumber="148" value="2002-04-16">April 16, 2002</collectingDate>
</date>
, 
<collectorName id="12365335274CFF8D5E4AF9C8400FF998" pageId="5" pageNumber="148">M. Sugimoto</collectorName>
&amp; 
<collectionCode id="D9D2AE26274CFF8D5A11F9E44032F987" box="[295,313,1592,1615]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5A72F9EB4094F998" box="[324,415,1591,1616]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="BF7C36E3274CFF8D5BF1F9804088F8CD" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="148">
<emphasis id="8DB7EAF1274CFF8D5BF1F980408DF9BB" box="[199,390,1628,1653]" inLineHeading="true" pageId="5" pageNumber="148" reason="1">
<emphasis id="8DB7EAF1274CFF8D5BF1F9804074F9BD" bold="true" box="[199,383,1628,1653]" pageId="5" pageNumber="148">Other material</emphasis>
.
</emphasis>
<materialsCitation id="0FAB3CBE274CFF8D5AA4F9804416F9BD" ID-GBIF-Occurrence="3131383321" box="[402,1309,1628,1653]" collectingDate="1957-04-20" collectionCode="T" collectorName="Saigusa" county="Zenkoji" location="Honshu" municipality="Kofu-shi" pageId="5" pageNumber="148" specimenCount="2" specimenCount-male="2" stateProvince="Yamanashi">
<specimenCount id="A9C5FD6A274CFF8D5AA4F98040BCF9BD" box="[402,439,1628,1653]" pageId="5" pageNumber="148" type="male">2♂</specimenCount>
, 
<collectingCounty id="561D4E6F274CFF8D5AF1F980432AF9BD" box="[455,545,1628,1653]" pageId="5" pageNumber="148">Zenkôji</collectingCounty>
, 
<collectingMunicipality id="5F18AC99274CFF8D5907F980439CF9BD" box="[561,663,1628,1653]" pageId="5" pageNumber="148">Kôfu-shi</collectingMunicipality>
, 
<collectingRegion id="7D07F801274CFF8D5991F980426CF9BD" box="[679,871,1628,1653]" country="Japan" name="Yamanashi" pageId="5" pageNumber="148">Yamanashi Pref.</collectingRegion>
, 
<location id="BA1C6038274CFF8D5840F98042DBF9BD" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D5840F98042DBF9BD" box="[886,976,1628,1653]" county="Zenkoji" municipality="Kofu-shi" name="Honshu" pageId="5" pageNumber="148" stateProvince="Yamanashi">Honshu</location>
, 
<date id="CB7D1023274CFF8D58D6F980459AF9BD" box="[992,1169,1628,1653]" pageId="5" pageNumber="148" value="1957-04-20">
<collectingDate id="DB39E9CB274CFF8D58D6F980459AF9BD" box="[992,1169,1628,1653]" pageId="5" pageNumber="148" value="1957-04-20">April 20, 1957</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D5F96F98045B8F9BB" box="[1184,1203,1628,1651]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5FF4F9804416F9BD" box="[1218,1309,1628,1653]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
; 
<materialsCitation id="0FAB3CBE274CFF8D5E1BF980433DF950" ID-GBIF-Occurrence="3131383313" collectingDate="1958-04-14" collectionCode="T" collectorName="Saigusa" county="Zenkoji" location="Honshu" municipality="Kofu-shi" pageId="5" pageNumber="148" specimenCount="6" specimenCount-male="6" stateProvince="Yamanashi">
<specimenCount id="A9C5FD6A274CFF8D5E1BF9804459F9BD" box="[1325,1362,1628,1653]" pageId="5" pageNumber="148" type="male">6♂</specimenCount>
, same locality, 
<date id="CB7D1023274CFF8D5BCAF9A340A0F950" box="[252,427,1663,1688]" pageId="5" pageNumber="148" value="1958-04-14">
<collectingDate id="DB39E9CB274CFF8D5BCAF9A340A0F950" box="[252,427,1663,1688]" pageId="5" pageNumber="148" value="1958-04-14">April 14, 1958</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D5A8DF95C40C5F95F" box="[443,462,1664,1687]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5AEAF9A3433DF950" box="[476,566,1663,1688]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
; 
<materialsCitation id="0FAB3CBE274CFF8D5971F9A3447AF950" ID-GBIF-Occurrence="3131383303" box="[583,1393,1663,1688]" collectingDate="1958-04-10" collectionCode="T" collectorName="Saigusa" county="Kofu-shi" location="Honshu" municipality="Yamanasshi Pref." pageId="5" pageNumber="148" specimenCount="4" specimenCount-male="4" stateProvince="Yamanashi">
<specimenCount id="A9C5FD6A274CFF8D5971F9A34360F950" box="[583,619,1663,1688]" pageId="5" pageNumber="148" type="male">4♂</specimenCount>
, 
<collectingCounty id="561D4E6F274CFF8D594AF9A343E9F950" box="[636,738,1663,1688]" pageId="5" pageNumber="148">Kôfu-shi</collectingCounty>
, 
<collectingMunicipality id="5F18AC99274CFF8D59C4F9A342B7F950" box="[754,956,1663,1688]" pageId="5" pageNumber="148">Yamanasshi Pref.</collectingMunicipality>
, 
<location id="BA1C6038274CFF8D58FDF9A3452EF950" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D58FDF9A3452EF950" box="[971,1061,1663,1688]" county="Kofu-shi" municipality="Yamanasshi Pref." name="Honshu" pageId="5" pageNumber="148" stateProvince="Yamanashi">Honshu</location>
, 
<date id="CB7D1023274CFF8D5F03F9A345EEF950" box="[1077,1253,1663,1688]" pageId="5" pageNumber="148" value="1958-04-10">
<collectingDate id="DB39E9CB274CFF8D5F03F9A345EEF950" box="[1077,1253,1663,1688]" pageId="5" pageNumber="148" value="1958-04-10">April 10, 1958</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D5FC2F95C440CF95F" box="[1268,1287,1664,1687]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5E20F9A3447AF950" box="[1302,1393,1663,1688]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
; 
<materialsCitation id="0FAB3CBE274CFF8D5EB6F95C456EF975" ID-GBIF-Occurrence="3131383316" collectingDate="1958-04-10" collectionCode="T" collectorName="Saigusa" county="Hanabusa" location="Honshu" municipality="Higashiyatsushiro" pageId="5" pageNumber="148" specimenCount="1" specimenCount-male="1" stateProvince="Yamanashi">
<specimenCount id="A9C5FD6A274CFF8D5EB6F95C449CF95F" box="[1408,1431,1664,1687]" pageId="5" pageNumber="148" type="male">♂</specimenCount>
, 
<collectingCounty id="561D4E6F274CFF8D5BA1F9784003F975" box="[151,264,1700,1725]" pageId="5" pageNumber="148">Hanabusa</collectingCounty>
, 
<collectingMunicipality id="5F18AC99274CFF8D5A21F97840EFF975" box="[279,484,1700,1725]" pageId="5" pageNumber="148">Higashiyatsushiro</collectingMunicipality>
, 
<collectingRegion id="7D07F801274CFF8D5AC2F97843B8F975" box="[500,691,1700,1725]" country="Japan" name="Yamanashi" pageId="5" pageNumber="148">Yamanashi Pref.</collectingRegion>
, 
<location id="BA1C6038274CFF8D59F7F9784210F975" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D59F7F9784210F975" box="[705,795,1700,1725]" county="Hanabusa" municipality="Higashiyatsushiro" name="Honshu" pageId="5" pageNumber="148" stateProvince="Yamanashi">Honshu</location>
, 
<date id="CB7D1023274CFF8D581CF97842D1F975" box="[810,986,1700,1725]" pageId="5" pageNumber="148" value="1958-04-10">
<collectingDate id="DB39E9CB274CFF8D581CF97842D1F975" box="[810,986,1700,1725]" pageId="5" pageNumber="148" value="1958-04-10">April 10, 1958</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D58DEF97842F0F973" box="[1000,1019,1700,1723]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5F3CF978456EF975" box="[1034,1125,1700,1725]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
; 
<materialsCitation id="0FAB3CBE274CFF8D5F42F9784228F928" ID-GBIF-Occurrence="3131383323" collectingDate="1998-04-27" collectionCode="T" collectorName="Saigusa &amp; M. Sugimoto" location="Honshu" municipality="Chiya-Ogi" pageId="5" pageNumber="148" specimenCount="1" specimenCount-male="1" stateProvince="Okayama">
<specimenCount id="A9C5FD6A274CFF8D5F42F9784593F975" box="[1140,1176,1700,1725]" pageId="5" pageNumber="148" type="male">1♂</specimenCount>
, 
<collectingMunicipality id="5F18AC99274CFF8D5F9EF978442AF975" box="[1192,1313,1700,1725]" pageId="5" pageNumber="148">Chiya-Ôgi</collectingMunicipality>
, 
<collectingRegion id="7D07F801274CFF8D5E07F97841DBF928" country="Japan" name="Okayama" pageId="5" pageNumber="148">Okayama Pref.</collectingRegion>
, 
<location id="BA1C6038274CFF8D5BEDF91B403EF928" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D5BEDF91B403EF928" box="[219,309,1735,1760]" municipality="Chiya-Ogi" name="Honshu" pageId="5" pageNumber="148" stateProvince="Okayama">Honshu</location>
, 
<date id="CB7D1023274CFF8D5A76F91B40E3F928" box="[320,488,1735,1760]" pageId="5" pageNumber="148" value="1998-04-27">
<collectingDate id="DB39E9CB274CFF8D5A76F91B40E3F928" box="[320,488,1735,1760]" pageId="5" pageNumber="148" value="1998-04-27">April 27, 1998</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D5AC5F914430DF917" box="[499,518,1736,1759]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5926F91B436CF928" box="[528,615,1735,1760]" pageId="5" pageNumber="148">Saigusa</collectorName>
&amp; 
<collectorName id="12365335274CFF8D59BAF9144228F928" box="[652,803,1735,1760]" pageId="5" pageNumber="148">M. Sugimoto</collectorName>
</materialsCitation>
; 
<materialsCitation id="0FAB3CBE274CFF8D5819F91B4074F8CD" ID-GBIF-Occurrence="3131383306" collectingDate="2005-05-24" collectionCode="T" collectorName="Saigusa" county="Sanshudai" location="Kyushu" municipality="Sobosan" pageId="5" pageNumber="148" specimenCount="3" specimenCount-male="3" stateProvince="Miyazaki">
<specimenCount id="A9C5FD6A274CFF8D5819F91B425FF928" box="[815,852,1735,1760]" pageId="5" pageNumber="148" type="male">3♂</specimenCount>
, 
<collectingCounty id="561D4E6F274CFF8D5856F91B42DDF928" box="[864,982,1735,1760]" pageId="5" pageNumber="148">Sanshûdai</collectingCounty>
, 
<collectingMunicipality id="5F18AC99274CFF8D58D7F91B4548F928" box="[993,1091,1735,1760]" pageId="5" pageNumber="148">Sobosan</collectingMunicipality>
, 
<collectingRegion id="7D07F801274CFF8D5F78F91B45F3F928" box="[1102,1272,1735,1760]" country="Japan" name="Miyazaki" pageId="5" pageNumber="148">Miyazaki Pref.</collectingRegion>
, 
<location id="BA1C6038274CFF8D5E35F91B4456F928" LSID="urn:lsid:plazi:treatment:376A87F52748FF8E5BA1FB9C45C4FD4B:BA1C6038274CFF8D5E35F91B4456F928" box="[1283,1373,1735,1760]" county="Sanshudai" municipality="Sobosan" name="Kyushu" pageId="5" pageNumber="148" stateProvince="Miyazaki">Kyushu</location>
, 
<date id="CB7D1023274CFF8D5E5FF91441F7F8CD" pageId="5" pageNumber="148" value="2005-05-24">
<collectingDate id="DB39E9CB274CFF8D5E5FF91441F7F8CD" pageId="5" pageNumber="148" value="2005-05-24">May 24, 2005</collectingDate>
</date>
, 
<collectionCode id="D9D2AE26274CFF8D5A31F9304011F8CB" box="[263,282,1772,1795]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="148">T</collectionCode>
. 
<collectorName id="12365335274CFF8D5A12F9304074F8CD" box="[292,383,1772,1797]" pageId="5" pageNumber="148">Saigusa</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="BF7C36E3274CFF8D5BF1F8D342B0F8EF" blockId="5.[151,1437,151,2013]" box="[199,955,1807,1832]" pageId="5" pageNumber="148">
<emphasis id="8DB7EAF1274CFF8D5BF1F8D34069F8E0" bold="true" box="[199,354,1807,1832]" pageId="5" pageNumber="148">Type-locality</emphasis>
: Kashii, Fukuoka-shi, 
<collectingRegion id="7D07F801274CFF8D5957F8D34208F8E0" box="[609,771,1807,1832]" country="Japan" name="Fukuoka" pageId="5" pageNumber="148">Fukuoka Pref.</collectingRegion>
, Kyushu, 
<collectingCountry id="C7D47673274CFF8D5845F8CC42BDF8EF" box="[883,950,1808,1831]" name="Japan" pageId="5" pageNumber="148">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F7D96568274CFF8D5BF1F8E84320F885" box="[199,555,1844,1869]" pageId="5" pageNumber="148" type="distribution">
<paragraph id="BF7C36E3274CFF8D5BF1F8E84320F885" blockId="5.[151,1437,151,2013]" box="[199,555,1844,1869]" pageId="5" pageNumber="148">
<emphasis id="8DB7EAF1274CFF8D5BF1F8E8406BF883" box="[199,352,1844,1869]" inLineHeading="true" pageId="5" pageNumber="148" reason="1">
<emphasis id="8DB7EAF1274CFF8D5BF1F8E84052F885" bold="true" box="[199,345,1844,1869]" pageId="5" pageNumber="148">Distribution</emphasis>
.
</emphasis>
Honshu, Kyushu.
</paragraph>
</subSubSection>
<subSubSection id="F7D96568274CFF8E5BF1F88B45C4FD4B" lastPageId="6" lastPageNumber="149" pageId="5" pageNumber="148" type="discussion">
<paragraph id="BF7C36E3274CFF8E5BF1F88B4323FF1D" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,645]" lastPageId="6" lastPageNumber="149" pageId="5" pageNumber="148">
<emphasis id="8DB7EAF1274CFF8D5BF1F88B4030F8A7" box="[199,315,1879,1904]" inLineHeading="true" pageId="5" pageNumber="148" reason="1">
<emphasis id="8DB7EAF1274CFF8D5BF1F88B403FF8B8" bold="true" box="[199,308,1879,1904]" pageId="5" pageNumber="148">Remarks</emphasis>
.
</emphasis>
The male of 
<taxonomicName id="78C34D60274CFF8D5AE2F88443BEF8B8" authority="Saigusa &amp; Sugimoto, 2014" authorityName="Saigusa &amp; Sugimoto" authorityYear="2014" box="[468,693,1880,1904]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="species" species="maculatella" status="sp. nov.">
<emphasis id="8DB7EAF1274CFF8D5AE2F88443BEF8B8" box="[468,693,1880,1904]" italics="true" pageId="5" pageNumber="148">Proutia maculatella</emphasis>
</taxonomicName>
<emphasis id="8DB7EAF1274CFF8D598AF88B421CF8B8" bold="true" box="[700,791,1879,1904]" pageId="5" pageNumber="148">
<taxonomicNameLabel id="9684578A274CFF8D598AF88B421CF8B8" box="[700,791,1879,1904]" pageId="5" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is easily distinguished from the known congeners by the ample maculated or reticulated forewings and semi-transparent hindwings covered with hair-like upper scales. The female of this species is distinctive in having light reddish brown abdominal membrane, and long, well segmented antennae consisting of more than 10 flagellomeres and yellowish white corethrogyne. The female of 
<taxonomicName id="78C34D60274CFF8D5FC9F8194489F814" authorityName="Hattenschwiler et Chao" authorityYear="1990" box="[1279,1410,1988,2012]" class="Insecta" family="Psychidae" genus="Proutia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="8DB7EAF1274CFF8D5FC9F8194489F814" box="[1279,1410,1988,2012]" italics="true" pageId="5" pageNumber="148">P. chinensis</emphasis>
</taxonomicName>
is reddish, but it has very short 6–7 segmented antennae (
<bibRefCitation id="DB524B12274FFF8E5832FF4B4559FF78" author="Hattenschwiler, P. &amp; Chao, C. - L." box="[772,1106,151,176]" pageId="6" pageNumber="149" pagination="262 - 268" refId="ref5734" refString="Hattenschwiler, P. &amp; Chao, C. - L. (1990) A new Proutia species from China (Lepidoptera, Psychidae). Nota Lepidopterologica, 12, 262 - 268." type="journal article" year="1990">Hättenschwiler et Chao 1990</bibRefCitation>
). The pupal antenna (
<figureCitation id="27F82A66274FFF8E5E70FF4B449FFF78" box="[1350,1428,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="6" pageNumber="149">Fig. 2I</figureCitation>
) is much longer than the head width.
</paragraph>
<paragraph id="BF7C36E3274FFF8E5BF1FF0340D4FE88" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="149">
The habitat of this species is grasslands along farm roads, railroads, forests, etc., in lowlands to mountain areas up to 
<quantity id="783B9B06274FFF8E5BEEFED8403FFED3" box="[216,308,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="149" unit="m" value="1500.0">1,500 m</quantity>
altitude (Kanayama, Masutomi, 
<collectingRegion id="7D07F801274FFF8E5991FED8426EFED5" box="[679,869,260,285]" country="Japan" name="Yamanashi" pageId="6" pageNumber="149">Yamanashi Pref.</collectingRegion>
). Adults appear in early spring from late March to April in lowlands of Kyushu.
</paragraph>
<paragraph id="BF7C36E3274FFF8E5BF1FE9045C4FD4B" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="149">
The larvae certainly live near the ground and seem to hibernate as the final instar. The mature larvae fix their larval cases on stones, stakes, fences and walls of sheds, tree trunks, etc. in early spring. This species is univoltine. The larval case (
<figureCitation id="27F82A66274FFF8E5A6EFE4840BBFE65" box="[344,432,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="6" pageNumber="149">Fig. 2C</figureCitation>
) of mature larvae is covered with slender, longish pieces of herbaceous plants that are arranged longitudinally and some of them slightly exceeding posteriorly beyond the posterior tip of the case. In captivity larvae feed on pieces of wilted or moist dead leaves of several kinds of herbaceous plants such as 
<taxonomicName id="78C34D60274FFF8E5BA1FDDC404CFDD0" box="[151,327,512,536]" class="Magnoliopsida" family="Asteraceae" genus="Erigeron" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="149" phylum="Tracheophyta" rank="species" species="anuus">
<emphasis id="8DB7EAF1274FFF8E5BA1FDDC404CFDD0" box="[151,327,512,536]" italics="true" pageId="6" pageNumber="149">Erigeron anuus</emphasis>
</taxonomicName>
. The female adult (
<figureCitation id="27F82A66274FFF8E591FFE234388FDD0" box="[553,643,511,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="6" pageNumber="149">Fig. 2A</figureCitation>
) protrudes most of its body from the pupal case except for the apical part of the abdomen. Head of pupal exuviae (
<figureCitation id="27F82A66274FFF8E59A2FDF843EAFDF5" box="[660,737,548,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="6" pageNumber="149">Fig. 2I</figureCitation>
) and sometimes that of forelegs of the female are not shed, so that the female wears the pupal head cuticle as a mask. During copulation (
<figureCitation id="27F82A66274FFF8E5F3EFD9B456AFDA8" box="[1032,1121,583,608]" captionStart="FIGURE 2" captionStartId="3.[151,250,1914,1936]" captionTargetBox="[165,1420,200,1879]" captionTargetId="figure-86@3.[151,1435,193,1892]" captionTargetPageId="3" captionText="FIGURE 2. Proutia maculatella sp. nov. A: female grasping lower end of its case in alive; B: copulation, female is concealed under male wings; C: male larval case with pupal exuviae. D: female preserved in 80% ethanol, somewhat faded in colour; E: female head, macerated; F: female forelegs, macerated; G: female midlegs, macerated; H: female hindlegs, macerated; I: head capsule of female pupa." figureDoi="http://doi.org/10.5281/zenodo.4947203" httpUri="https://zenodo.org/record/4947203/files/figure.png" pageId="6" pageNumber="149">Fig. 2B</figureCitation>
) the male tightly holds the wings ventrally covering its mate. The eggs were laid inside the pupal exuviae in the pupal case.
</paragraph>
</subSubSection>
</treatment>
</document>