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<mods:title id="53A9AC32BD8B7C5C8BA7A4C1142E3259">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group</mods:title>
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, 
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; 
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<bibRefCitation id="DB3E0BF2FFB9FFE9FD2E65C0FB81FB3E" author="Schulke, M. &amp; Smetana, A." box="[738,1048,1242,1267]" pageId="19" pageNumber="470" pagination="304 - 1134" refId="ref19533" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke &amp; Smetana 2015: 307</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFB9FFEEFF5B643BFE29FE8E" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470" type="materials_examined">
<paragraph id="BF107603FFB9FFE9FF5B643BFBC1FA6A" blockId="19.[151,1437,1312,2023]" pageId="19" pageNumber="470">
<emphasis id="8DDBAA11FFB9FFE9FF5B643BFE59FAF6" bold="true" box="[151,448,1312,1339]" pageId="19" pageNumber="470">Type material examined:</emphasis>
<typeStatus id="6014C8A1FFB9FFE9FE0B643BFDAEFAF6" box="[455,567,1313,1339]" pageId="19" pageNumber="470" type="lectotype">Lectotype</typeStatus>
(here designated) of 
<taxonomicName id="78AF0D80FFB9FFE9FCEE643BFB2EFAF6" authority="Reitter, 1891" authorityName="Reitter" authorityYear="1891" box="[802,1207,1313,1339]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFB9FFE9FCEE643BFBB8FAF6" box="[802,1057,1313,1339]" italics="true" pageId="19" pageNumber="470">Lathrimaeum reflexum</emphasis>
<bibRefCitation id="DB3E0BF2FFB9FFE9FBEB643BFB2EFAF6" author="Reitter, E." box="[1063,1207,1313,1339]" pageId="19" pageNumber="470" pagination="195 - 199" refId="ref19463" refString="Reitter, E. (1891) Zweiter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische Zeitung, 10, 195 - 199." type="journal article" year="1891">Reitter, 1891</bibRefCitation>
</taxonomicName>
<specimenCount id="A9A9BD8AFFB9FFE9FB72643BFB4AFAF6" box="[1214,1235,1313,1339]" pageId="19" pageNumber="470" type="male">♂</specimenCount>
(very small silver round label without data is pinned under the card with the beetle): ‘ 
<taxonomicName id="78AF0D80FFB9FFE9FC58645FFA81FA92" authority="Tashkend." authorityName="Tashkend." box="[916,1304,1349,1375]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">Lathrimaeum reflexum Tashkend.</taxonomicName>
Typ’ &lt;rectangular label, printed&gt;, ‘ 
<typeStatus id="6014C8A1FFB9FFE9FE786473FDB7FA4E" box="[436,558,1385,1411]" pageId="19" pageNumber="470" type="holotype">Holotypus</typeStatus>
<taxonomicName id="78AF0D80FFB9FFE9FDF56473FCA6FA4E" box="[569,831,1385,1411]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">Lathrimaeum reflexum</taxonomicName>
Reitter’ &lt;rectangular label, printed&gt;, ‘ 
<taxonomicName id="78AF0D80FFB9FFE9FACC6473FEB4FA6A" authority="Rtt." authorityName="Reitter" authorityYear="1891" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">Lathrimaeum reflexum Rtt.</taxonomicName>
[handwritten] V.I. Gusarov det. 1996’ &lt;rectagular label, printed&gt; (
<collectionCode id="D9BEEEC6FFB9FFE9FBDB6497FBD2FA6A" box="[1047,1099,1421,1447]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
).
</paragraph>
<paragraph id="BF107603FFB9FFE9FF0B64ABFB88F9FA" blockId="19.[151,1437,1312,2023]" pageId="19" pageNumber="470">
<typeStatus id="6014C8A1FFB9FFE9FF0B64ABFEF6FA06" box="[199,367,1457,1483]" pageId="19" pageNumber="470" type="paralectotype">Paralectotypes</typeStatus>
: 
<specimenCount id="A9A9BD8AFFB9FFE9FEB664ABFE31FA06" box="[378,424,1457,1483]" pageId="19" pageNumber="470" type="male">1 ♂</specimenCount>
: ‘460’ &lt;small rectangular label, printed&gt;, ‘ 
<taxonomicName id="78AF0D80FFB9FFE9FC4B64ABFB09FA06" authorityName="Reitter" authorityYear="1891" box="[903,1168,1457,1483]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">Lathrimaeum reflexum</taxonomicName>
Rtt’ &lt;rectangular label, handwritten&gt;, ‘ 
<taxonomicName id="78AF0D80FFB9FFE9FEF164CFFDEAFA22" authority="Rtt." authorityName="Reitter" authorityYear="1891" box="[317,627,1493,1519]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">Lathrimaeum reflexum Rtt.</taxonomicName>
[handwritten] V.I. Gusarov det. 1996’ &lt;rectangular label, printed&gt; (
<collectionCode id="D9BEEEC6FFB9FFE9FA9764CFFA16FA22" box="[1371,1423,1493,1519]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
); 
<specimenCount id="A9A9BD8AFFB9FFE9FF5B64E3FF75F9DE" box="[151,236,1529,1555]" pageId="19" pageNumber="470" type="generic">1 spec.</specimenCount>
: ‘Ср[едНЯЯ]. АЗиЯ [Middle Asia] | Tashkent [sic]’ &lt;rectangular label, handwritten&gt;, ‘ 
<typeStatus id="6014C8A1FFB9FFE9FB7164E3FAA8F9DE" box="[1213,1329,1529,1555]" pageId="19" pageNumber="470" type="paratype">Paratypus</typeStatus>
[printed] Lathrima | eum reflexum | Reitter [handwritten]’ &lt;red rectangular label&gt; (
<collectionCode id="D9BEEEC6FFB9FFE9FC1C6707FB9DF9FA" box="[976,1028,1565,1591]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
).
</paragraph>
<paragraph id="BF107603FFB9FFEEFF0B675AFE29FE8E" blockId="19.[151,1437,1312,2023]" lastBlockId="20.[151,1436,152,432]" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470">
<materialsCitation id="0FC77C5EFFB9FFE9FF0B675AFC6BF9B2" ID-GBIF-Occurrence="2429182972" collectingDate="1985-05-17" collectorName="V. Kastcheev" country="Kazakhstan" location="KaZak" pageId="19" pageNumber="470" specimenCount="2" specimenCount-male="2">
<emphasis id="8DDBAA11FFB9FFE9FF0B675AFD0DF996" bold="true" box="[199,660,1600,1627]" pageId="19" pageNumber="470">
Additional material: 
<collectingCountry id="C7B83693FFB9FFE9FE0A675AFD09F996" box="[454,656,1600,1627]" name="Kazakhstan" pageId="19" pageNumber="470">KAZAKHSTAN</collectingCountry>
:
</emphasis>
<specimenCount id="A9A9BD8AFFB9FFE9FD50675BFD78F996" box="[668,737,1601,1627]" pageId="19" pageNumber="470" type="male">2 ♂♂</specimenCount>
: ‘ 
<location id="BA7020D8FFB9FFE9FD3D675AFCA6F996" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FD3D675AFCA6F996" box="[753,831,1600,1627]" country="Kazakhstan" name="KaZak" pageId="19" pageNumber="470">КаЗах</location>
[ста]-Н; Аксу- | ДЖабаглы | В. КаЩеев [Kazakhstan; Aks-Dzhabagly (=Aksu-Zhabagly Nature Reserve) 
<collectorName id="125A13D5FFB9FFE9FD1B677FFCF4F9B2" box="[727,877,1636,1663]" pageId="19" pageNumber="470">V. Kastcheev</collectorName>
] 
<date id="CB1150C3FFB9FFE9FCB5677EFC6BF9B2" box="[889,1010,1636,1663]" pageId="19" pageNumber="470" value="1985-05-17">
<collectingDate id="DB55A92BFFB9FFE9FCB5677EFC6BF9B2" box="[889,1010,1636,1663]" pageId="19" pageNumber="470" value="1985-05-17">17.V.1985</collectingDate>
</date>
</materialsCitation>
’, ‘ 
<materialsCitation id="0FC77C5EFFB9FFE9FBCA677FFDEAF96E" ID-GBIF-Occurrence="2429182946" collectionCode="ZIN" collectorName="Kastcheev'" country="Kazakhstan" pageId="19" pageNumber="470" specimenCount="1" typeStatus="holotype">
<typeStatus id="6014C8A1FFB9FFE9FBCA677FFB19F9B2" box="[1030,1152,1637,1663]" pageId="19" pageNumber="470" type="holotype">Holotypus</typeStatus>
Lathrima- | eum (Priono- thorax) | turcestanicum | 
<collectorName id="125A13D5FFB9FFE9FE606793FDB1F96E" box="[428,552,1673,1699]" pageId="19" pageNumber="470">Kastcheev’</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FDFB6793FDF2F96E" box="[567,619,1673,1699]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FDB36793FAB5F90A" ID-GBIF-Occurrence="2429182969" collectingDate="1985-05" collectingDateMax="1985-05-20" collectingDateMin="1985-05" collectionCode="ZIN" collectorName="Dzhabagly &amp; Dzhabagly R. &amp; Kastcheev &amp; Kastcheev'" country="Kazakhstan" pageId="19" pageNumber="470" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FDB36793FD59F96E" box="[639,704,1673,1699]" pageId="19" pageNumber="470" type="female">3 ♀♀</specimenCount>
: ‘Аксу-ДЖабаглы, р. ДЖабаглы, 20.5.[19] 
<date id="CB1150C3FFB9FFE9FB646792FB7FF96E" box="[1192,1254,1672,1699]" pageId="19" pageNumber="470" value="1985-05">
<collectingDate id="DB55A92BFFB9FFE9FB646792FB7FF96E" box="[1192,1254,1672,1699]" pageId="19" pageNumber="470" value="1985-05">85 В.</collectingDate>
</date>
КаЩеев [Aksu- 
<collectorName id="125A13D5FFB9FFE9FF5B67B7FE88F90A" box="[151,273,1709,1735]" pageId="19" pageNumber="470">Dzhabagly</collectorName>
, 
<collectorName id="125A13D5FFB9FFE9FEEC67B7FE26F90A" box="[288,447,1709,1735]" pageId="19" pageNumber="470">Dzhabagly R.</collectorName>
, 
<date id="CB1150C3FFB9FFE9FE0267B6FDD5F90A" box="[462,588,1708,1735]" pageId="19" pageNumber="470" value="1985-05-20">
<collectingDate id="DB55A92BFFB9FFE9FE0267B6FDD5F90A" box="[462,588,1708,1735]" pageId="19" pageNumber="470" value="1985-05-20">20.05.1985</collectingDate>
</date>
V. 
<collectorName id="125A13D5FFB9FFE9FDBB67B6FD6FF90A" box="[631,758,1708,1735]" pageId="19" pageNumber="470">Kastcheev</collectorName>
]’, ‘ 
<taxonomicName id="78AF0D80FFB9FFE9FCD567B7FBC1F90A" box="[793,1112,1709,1735]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="turcestanicum">
<emphasis id="8DDBAA11FFB9FFE9FCD567B7FBC1F90A" box="[793,1112,1709,1735]" italics="true" pageId="19" pageNumber="470">Lathrimaeum turcestanicum</emphasis>
</taxonomicName>
<collectorName id="125A13D5FFB9FFE9FBAE67B7FB47F90A" box="[1122,1246,1709,1735]" pageId="19" pageNumber="470">Kastcheev’</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FB3C67B7FABDF90A" box="[1264,1316,1709,1735]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FAF767B7FC45F926" ID-GBIF-Occurrence="2429182950" collectingDate="1985-06-22" collectionCode="CS, ZIN" collectorName="Kastcheev" country="Kazakhstan" location="Nature Reserve" pageId="19" pageNumber="470" specimenCount="15" specimenCount-female="6" specimenCount-male="9" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FAF767B7FAE6F90A" box="[1339,1407,1709,1735]" pageId="19" pageNumber="470" type="male">9 ♂♂</specimenCount>
, 
<specimenCount id="A9A9BD8AFFB9FFE9FA4267B7FF5AF926" pageId="19" pageNumber="470" type="female">6 ♀♀</specimenCount>
: same 
<location id="BA7020D8FFB9FFE9FEDF67CBFE58F926" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FEDF67CBFE58F926" box="[275,449,1745,1771]" country="Kazakhstan" name="Nature Reserve" pageId="19" pageNumber="470">Nature Reserve</location>
and river. 
<date id="CB1150C3FFB9FFE9FDF067CAFD27F926" box="[572,702,1744,1771]" pageId="19" pageNumber="470" value="1985-06-22">
<collectingDate id="DB55A92BFFB9FFE9FDF067CAFD27F926" box="[572,702,1744,1771]" pageId="19" pageNumber="470" value="1985-06-22">22.06.1985</collectingDate>
</date>
. V. 
<collectorName id="125A13D5FFB9FFE9FD2567CBFCC7F926" box="[745,862,1745,1771]" pageId="19" pageNumber="470">Kastcheev</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FCA267CBFC0AF926" box="[878,915,1745,1771]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="19" pageNumber="470">CS</collectionCode>
, 
<collectionCode id="D9BEEEC6FFB9FFE9FC6C67CBFC4AF926" box="[928,979,1745,1771]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FC2467CBFD06F8C2" ID-GBIF-Occurrence="2429182963" collectingDate="1979-04-25" collectionCode="ZIN" collectorName="V. Kastcheev" country="Kazakhstan" location="Kshi-Kaindy." municipality="Nature Reserve" pageId="19" pageNumber="470" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FC2467CBFBB5F926" box="[1000,1068,1745,1771]" pageId="19" pageNumber="470" type="male">2 ♂♂</specimenCount>
: same 
<collectingMunicipality id="5F74EC79FFB9FFE9FBB667CBFAB1F926" box="[1146,1320,1745,1771]" pageId="19" pageNumber="470">Nature Reserve</collectingMunicipality>
and river, 
<location id="BA7020D8FFB9FFE9FF5B67EFFEB7F8C2" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FF5B67EFFEB7F8C2" box="[151,302,1781,1807]" country="Kazakhstan" municipality="Nature Reserve" name="Kshi-Kaindy." pageId="19" pageNumber="470">Kshi-Kaindy.</location>
<date id="CB1150C3FFB9FFE9FEF867EEFE2FF8C2" box="[308,438,1780,1807]" pageId="19" pageNumber="470" value="1979-04-25">
<collectingDate id="DB55A92BFFB9FFE9FEF867EEFE2FF8C2" box="[308,438,1780,1807]" pageId="19" pageNumber="470" value="1979-04-25">25.04.1979</collectingDate>
</date>
. 
<collectorName id="125A13D5FFB9FFE9FE0C67EFFDCDF8C2" box="[448,596,1781,1807]" pageId="19" pageNumber="470">V. Kastcheev</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FDAF67EFFD0EF8C2" box="[611,663,1781,1807]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FD6767EFFEC0F8FE" ID-GBIF-Occurrence="2429182996" collectingDate="2001-08-15" collectionCode="ZIN" collectorName="V. Kastcheev" country="Kazakhstan" location="Ulken-Kaindy." municipality="Nature Reserve" pageId="19" pageNumber="470" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FD6767EFFD4EF8C2" box="[683,727,1781,1807]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
: same 
<collectingMunicipality id="5F74EC79FFB9FFE9FCE967EFFC4AF8C2" box="[805,979,1781,1807]" pageId="19" pageNumber="470">Nature Reserve</collectingMunicipality>
and river, 
<location id="BA7020D8FFB9FFE9FB8067EFFB6BF8C2" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FB8067EFFB6BF8C2" box="[1100,1266,1781,1807]" country="Kazakhstan" municipality="Nature Reserve" name="Ulken-Kaindy." pageId="19" pageNumber="470">Ulken-Kaindy.</location>
<date id="CB1150C3FFB9FFE9FB3567EEFAE3F8C2" box="[1273,1402,1780,1807]" pageId="19" pageNumber="470" value="2001-08-15">
<collectingDate id="DB55A92BFFB9FFE9FB3567EEFAE3F8C2" box="[1273,1402,1780,1807]" pageId="19" pageNumber="470" value="2001-08-15">15.08.2001</collectingDate>
</date>
. 
<collectorName id="125A13D5FFB9FFE9FA4867EFFE95F8FE" pageId="19" pageNumber="470">V. Kastcheev</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FED16603FEC8F8FE" box="[285,337,1817,1843]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FEAB6603FADCF8FE" ID-GBIF-Occurrence="2429182970" box="[359,1349,1816,1843]" collectingDate="1979-05-20" collectionCode="ZIN" collectorName="B. Iskakov" country="Kazakhstan" location="Taldybulak River" municipality="Nature Reserve" pageId="19" pageNumber="470" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FEAB6603FE33F8FE" box="[359,426,1817,1843]" pageId="19" pageNumber="470" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="A9A9BD8AFFB9FFE9FE746603FE7FF8FE" box="[440,486,1817,1843]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
: same 
<collectingMunicipality id="5F74EC79FFB9FFE9FDFB6603FD70F8FE" box="[567,745,1817,1843]" pageId="19" pageNumber="470">Nature Reserve</collectingMunicipality>
, 
<location id="BA7020D8FFB9FFE9FD3A6603FC25F8FE" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FD3A6603FC25F8FE" box="[758,956,1817,1843]" country="Kazakhstan" municipality="Nature Reserve" name="Taldybulak River" pageId="19" pageNumber="470">Taldybulak River</location>
. 10- 
<date id="CB1150C3FFB9FFE9FC266602FBF6F8FE" box="[1002,1135,1816,1843]" pageId="19" pageNumber="470" value="1979-05-20">
<collectingDate id="DB55A92BFFB9FFE9FC266602FBF6F8FE" box="[1002,1135,1816,1843]" pageId="19" pageNumber="470" value="1979-05-20">20.05.1979</collectingDate>
</date>
. 
<collectorName id="125A13D5FFB9FFE9FBB06603FB61F8FE" box="[1148,1272,1817,1843]" pageId="19" pageNumber="470">B. Iskakov</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FAC46603FAA5F8FE" box="[1288,1340,1817,1843]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="470" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FA9E6603FC36F89A" ID-GBIF-Occurrence="2429182960" collectingDate="1988-05-17" collectionCode="CS" collectorName="Aksu R. &amp; V. Kastcheev" country="Kazakhstan" location="Aksu-Dzhabagly" pageId="19" pageNumber="470" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FA9E6603FA19F8FE" box="[1362,1408,1817,1843]" pageId="19" pageNumber="470" type="male">1 ♂</specimenCount>
, 
<specimenCount id="A9A9BD8AFFB9FFE9FA426603FF5AF89A" pageId="19" pageNumber="470" type="female">2 ♀♀</specimenCount>
: 
<location id="BA7020D8FFB9FFE9FF026627FE14F89A" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FF026627FE14F89A" box="[206,397,1853,1879]" country="Kazakhstan" name="Aksu-Dzhabagly" pageId="19" pageNumber="470">Aksu-Dzhabagly</location>
, kanyon 
<collectorName id="125A13D5FFB9FFE9FE236627FDD2F89A" box="[495,587,1853,1879]" pageId="19" pageNumber="470">Aksu R.</collectorName>
<date id="CB1150C3FFB9FFE9FD9E6626FD4AF89A" box="[594,723,1852,1879]" pageId="19" pageNumber="470" value="1988-05-17">
<collectingDate id="DB55A92BFFB9FFE9FD9E6626FD4AF89A" box="[594,723,1852,1879]" pageId="19" pageNumber="470" value="1988-05-17">17.05.1988</collectingDate>
</date>
. 
<collectorName id="125A13D5FFB9FFE9FD116627FCE9F89A" box="[733,880,1853,1879]" pageId="19" pageNumber="470">V. Kastcheev</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FCB36627FC3CF89A" box="[895,933,1853,1879]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="19" pageNumber="470">CS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FC766626FC8DF8B6" ID-GBIF-Occurrence="2429182976" collectingDate="1996-01-05" collectionCode="CK" collectorName="Grebennikov" country="Uzbekistan" location="Akbulak River" municipality="Chatkal River" pageId="19" pageNumber="470" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="8DDBAA11FFB9FFE9FC766626FBE0F89A" bold="true" box="[954,1145,1852,1879]" pageId="19" pageNumber="470">
<collectingCountry id="C7B83693FFB9FFE9FC766626FBEDF89A" box="[954,1140,1852,1879]" name="Uzbekistan" pageId="19" pageNumber="470">UZBEKISTAN</collectingCountry>
:
</emphasis>
<specimenCount id="A9A9BD8AFFB9FFE9FBB36627FB35F89A" box="[1151,1196,1853,1879]" pageId="19" pageNumber="470" type="male">1 ♂</specimenCount>
: right side of 
<collectingMunicipality id="5F74EC79FFB9FFE9FA896627FF4EF8B6" pageId="19" pageNumber="470">Chatkal River</collectingMunicipality>
, 
<location id="BA7020D8FFB9FFE9FF2C667BFE1EF8B6" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FF2C667BFE1EF8B6" box="[224,391,1889,1915]" country="Uzbekistan" municipality="Chatkal River" name="Akbulak River" pageId="19" pageNumber="470">Akbulak River</location>
. 
<date id="CB1150C3FFB9FFE9FE5D667AFD8AF8B6" box="[401,531,1888,1915]" pageId="19" pageNumber="470" value="1996-01-05">
<collectingDate id="DB55A92BFFB9FFE9FE5D667AFD8AF8B6" box="[401,531,1888,1915]" pageId="19" pageNumber="470" value="1996-01-05">01.05.1996</collectingDate>
</date>
. K. 
<collectorName id="125A13D5FFB9FFE9FDF3667BFD48F8B6" box="[575,721,1889,1915]" pageId="19" pageNumber="470">Grebennikov</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FD2C667BFC95F8B6" box="[736,780,1889,1915]" pageId="19" pageNumber="470">CK</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FCD3667BFC20F852" ID-GBIF-Occurrence="2429182944" collectingDate="1974-07-04" collectionCode="ZMM" collectorName="S. Yu. Gryuntal" country="Uzbekistan" location="Bottom" municipality="Chimgan" pageId="19" pageNumber="470" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FCD3667BFCD2F8B6" box="[799,843,1889,1915]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
: 
<collectingMunicipality id="5F74EC79FFB9FFE9FC9B667BFC27F8B6" box="[855,958,1889,1915]" pageId="19" pageNumber="470">Chimgan</collectingMunicipality>
[=Chimgon]. 
<location id="BA7020D8FFB9FFE9FB91667BFB28F8B6" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FB91667BFB28F8B6" box="[1117,1201,1889,1915]" country="Uzbekistan" municipality="Chimgan" name="Bottom" pageId="19" pageNumber="470">Bottom</location>
of the ravine, stream overgrown with horsetail, in litter. 
<date id="CB1150C3FFB9FFE9FDEE669FFD3DF852" box="[546,676,1925,1951]" pageId="19" pageNumber="470" value="1974-07-04">
<collectingDate id="DB55A92BFFB9FFE9FDEE669FFD3DF852" box="[546,676,1925,1951]" pageId="19" pageNumber="470" value="1974-07-04">07.04.1974</collectingDate>
</date>
. 
<collectorName id="125A13D5FFB9FFE9FD63669FFCC1F852" box="[687,856,1925,1951]" pageId="19" pageNumber="470">S.Yu. Gryuntal</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FCA4669FFC28F852" box="[872,945,1925,1951]" pageId="19" pageNumber="470">ZMM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FC0A669EFCEDF80E" ID-GBIF-Occurrence="2429182975" collectingDate="1983-09-24" collectorName="A. B. Ryvkin" country="Kyrgyzstan" location="Sary-Chelek Nature Reserve Sary-Chelek Lake" pageId="19" pageNumber="470" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="8DDBAA11FFB9FFE9FC0A669EFB0AF852" bold="true" box="[966,1171,1924,1951]" pageId="19" pageNumber="470">
<collectingCountry id="C7B83693FFB9FFE9FC0A669EFB17F852" box="[966,1166,1924,1951]" name="Kyrgyzstan" pageId="19" pageNumber="470">KYRGYZSTAN</collectingCountry>
:
</emphasis>
<specimenCount id="A9A9BD8AFFB9FFE9FB57669FFB51F852" box="[1179,1224,1925,1951]" pageId="19" pageNumber="470" type="male">1 ♂</specimenCount>
, 
<specimenCount id="A9A9BD8AFFB9FFE9FB19669FFA9BF852" box="[1237,1282,1925,1951]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
: 
<location id="BA7020D8FFB9FFE9FADC669FFD8FF80E" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FADC669FFD8FF80E" country="Kyrgyzstan" name="Sary-Chelek Nature Reserve Sary-Chelek Lake" pageId="19" pageNumber="470">Sary-Chelek Nature Reserve Sary-Chelek Lake</location>
. 
<date id="CB1150C3FFB9FFE9FDD366B3FD39F80E" box="[543,672,1961,1987]" pageId="19" pageNumber="470" value="1983-09-24">
<collectingDate id="DB55A92BFFB9FFE9FDD366B3FD39F80E" box="[543,672,1961,1987]" pageId="19" pageNumber="470" value="1983-09-24">24.09.1983</collectingDate>
</date>
. 
<collectorName id="125A13D5FFB9FFE9FD6466B3FCADF80E" box="[680,820,1961,1987]" pageId="19" pageNumber="470">A.B. Ryvkin</collectorName>
(CR)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFE9FCB366B3FE6DF82A" ID-GBIF-Occurrence="2429182994" collectingDate="1996-04-06" collectionCode="CS" collectorName="Klimenko" country="Kyrgyzstan" elevation="2800" location="Nature Reserve" pageId="19" pageNumber="470" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFB9FFE9FCB366B3FC33F80E" box="[895,938,1961,1987]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
: same 
<location id="BA7020D8FFB9FFE9FC3866B3FB39F80E" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFB9FFE9FC3866B3FB39F80E" box="[1012,1184,1961,1987]" country="Kyrgyzstan" name="Nature Reserve" pageId="19" pageNumber="470">Nature Reserve</location>
and lake. 
<elevation id="34829130FFB9FFE9FAC366B3FA04F80E" box="[1295,1437,1961,1987]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="470" unit="m" value="2800.0">
<quantity id="7857DBE6FFB9FFE9FAC366B3FAFAF80E" box="[1295,1379,1961,1987]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="470" unit="m" value="2800.0">2800 m</quantity>
a.s.l.
</elevation>
<date id="CB1150C3FFB9FFE9FF5B66D7FE80F82A" box="[151,281,1997,2023]" pageId="19" pageNumber="470" value="1996-04-06">
<collectingDate id="DB55A92BFFB9FFE9FF5B66D7FE80F82A" box="[151,281,1997,2023]" pageId="19" pageNumber="470" value="1996-04-06">04.06.1996</collectingDate>
</date>
. A. 
<collectorName id="125A13D5FFB9FFE9FE8866D7FE2DF82A" box="[324,436,1997,2023]" pageId="19" pageNumber="470">Klimenko</collectorName>
(
<collectionCode id="D9BEEEC6FFB9FFE9FE0866D7FE73F82A" box="[452,490,1997,2023]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="19" pageNumber="470">CS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFB9FFEEFDCC66D6FA01FF79" ID-GBIF-Occurrence="2429182993" collectingDate="1965-05-09" collectingDateMax="1965-09-05" collectingDateMin="1965-05-09" collectionCode="ZMM" collectorName="Tajikistan &amp; Tichomirova. &amp; A. L. Tichomirova" country="Tajikistan" lastPageId="20" lastPageNumber="471" location="Kondara" pageId="19" pageNumber="470" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="8DDBAA11FFB9FFE9FDCC66D6FD29F82A" bold="true" box="[512,688,1996,2023]" pageId="19" pageNumber="470">
<collectingCountry id="C7B83693FFB9FFE9FDCC66D6FD35F82A" box="[512,684,1996,2023]" name="Tajikistan" pageId="19" pageNumber="470">TAJIKISTAN</collectingCountry>
:
</emphasis>
<specimenCount id="A9A9BD8AFFB9FFE9FD7B66D7FD7DF82A" box="[695,740,1997,2023]" pageId="19" pageNumber="470" type="male">1 ♂</specimenCount>
, 
<specimenCount id="A9A9BD8AFFB9FFE9FD3C66D7FC85F82A" box="[752,796,1997,2023]" pageId="19" pageNumber="470" type="female">1 ♀</specimenCount>
: ‘ТадЖикистаН | КоНдара 
<date id="CB1150C3FFB9FFE9FB9C66D6FB2DF82A" box="[1104,1204,1996,2023]" pageId="19" pageNumber="470" value="1965-05-09">
<collectingDate id="DB55A92BFFB9FFE9FB9C66D6FB2DF82A" box="[1104,1204,1996,2023]" pageId="19" pageNumber="470" value="1965-05-09">9.V.1965</collectingDate>
</date>
| А. Тихомирова | в подстилке [
<collectorName id="125A13D5FFBEFFEEFED16183FE08FF7E" box="[285,401,153,179]" pageId="20" pageNumber="471">Tajikistan</collectorName>
, 
<location id="BA7020D8FFBEFFEEFE516183FD99FF7E" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFBEFFEEFE516183FD99FF7E" box="[413,512,153,179]" country="Tajikistan" name="Kondara" pageId="20" pageNumber="471">Kondara</location>
, 
<date id="CB1150C3FFBEFFEEFDC76182FD14FF7E" box="[523,653,152,179]" pageId="20" pageNumber="471" value="1965-09-05">
<collectingDate id="DB55A92BFFBEFFEEFDC76182FD14FF7E" box="[523,653,152,179]" pageId="20" pageNumber="471" value="1965-09-05">09.05.1965</collectingDate>
</date>
. A.L. 
<collectorName id="125A13D5FFBEFFEEFD036183FCFFFF7E" box="[719,870,153,179]" pageId="20" pageNumber="471">Tichomirova.</collectorName>
in litter]’ (
<collectionCode id="D9BEEEC6FFBEFFEEFC116183FBBCFF7E" box="[989,1061,153,179]" pageId="20" pageNumber="471">ZMM</collectionCode>
; collection of 
<collectorName id="125A13D5FFBEFFEEFB0E6183FA09FF79" box="[1218,1424,153,180]" pageId="20" pageNumber="471">A.L. Tichomirova</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFBEFFEEFF5B61A7FA1DFF31" ID-GBIF-Occurrence="2429182958" collectingDate="1965-11-05" collectionCode="ZMM" collectorName="A. L. Tichomirova" country="Tajikistan" county="Tajik" location="Kondara" municipality="Gissar Mt." pageId="20" pageNumber="471" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFBEFFEEFF5B61A7FF43FF1A" box="[151,218,189,215]" pageId="20" pageNumber="471" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="A9A9BD8AFFBEFFEEFF2461A7FEB2FF1A" box="[232,299,189,215]" pageId="20" pageNumber="471" type="female">3 ♀♀</specimenCount>
: ‘ТадЖикс[каЯ] ССР. Гис- | сарск. хр. КоНдара | в подстилке | 11/5-[19] 
<quantity id="7857DBE6FFBEFFEEFBB061A6FB29FF1A" box="[1148,1200,188,215]" metricMagnitude="-2" metricUnit="kg" metricValue="6.5" pageId="20" pageNumber="471" unit="g" value="65.0">65 г</quantity>
. [А.Л.] Тихомирова [
<collectingCounty id="56710E8FFFBEFFEEFF5161FBFF43FF36" box="[157,218,225,251]" pageId="20" pageNumber="471">Tajik</collectingCounty>
SSR. 
<collectingMunicipality id="5F74EC79FFBEFFEEFED361FBFE0FFF36" box="[287,406,225,251]" pageId="20" pageNumber="471">Gissar Mt.</collectingMunicipality>
, 
<location id="BA7020D8FFBEFFEEFE6D61FBFD9DFF36" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFBEFFEEFE6D61FBFD9DFF36" box="[417,516,225,251]" country="Tajikistan" county="Tajik" municipality="Gissar Mt." name="Kondara" pageId="20" pageNumber="471">Kondara</location>
, in litter, 
<date id="CB1150C3FFBEFFEEFDA761FAFD44FF36" box="[619,733,224,251]" pageId="20" pageNumber="471" value="1965-11-05">
<collectingDate id="DB55A92BFFBEFFEEFDA761FAFD44FF36" box="[619,733,224,251]" pageId="20" pageNumber="471" value="1965-11-05">11.5.1965</collectingDate>
</date>
. 
<collectorName id="125A13D5FFBEFFEEFD2961FBFC2DFF36" box="[741,948,224,251]" pageId="20" pageNumber="471">A.L. Tichomirova</collectorName>
]’ (
<collectionCode id="D9BEEEC6FFBEFFEEFC0061FBFB8DFF36" box="[972,1044,225,251]" pageId="20" pageNumber="471">ZMM</collectionCode>
; collection of 
<collectorName id="125A13D5FFBEFFEEFB6361FBFAE5FF31" box="[1199,1404,225,252]" pageId="20" pageNumber="471">A.L. Tichomirova</collectorName>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFBEFFEEFA4261FBFB10FED2" ID-GBIF-Occurrence="2429182997" collectingDate="1985-05-20" collectionCode="CS" collectorName="Shilenkov" country="Tajikistan" elevation="1000" location="Kvak Gorge." municipality="Kandara" pageId="20" pageNumber="471" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFBEFFEEFA4261FBFF37FED2" pageId="20" pageNumber="471" type="female">1 ♀</specimenCount>
: 
<collectingMunicipality id="5F74EC79FFBEFFEEFF77601FFE84FED2" box="[187,285,261,287]" pageId="20" pageNumber="471">Kandara</collectingMunicipality>
, 
<location id="BA7020D8FFBEFFEEFEEB601FFE21FED2" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFEDFF5B62EDFCCAF8C0:BA7020D8FFBEFFEEFEEB601FFE21FED2" box="[295,440,261,287]" country="Tajikistan" municipality="Kandara" name="Kvak Gorge." pageId="20" pageNumber="471">Kvak Gorge.</location>
<elevation id="34829130FFBEFFEEFE72601FFDD7FED2" box="[446,590,261,287]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="471" unit="m" value="1000.0">
<quantity id="7857DBE6FFBEFFEEFE72601FFD8BFED2" box="[446,530,261,287]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="471" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
, near snow field. 
<date id="CB1150C3FFBEFFEEFCD8601EFC0FFED2" box="[788,918,260,287]" pageId="20" pageNumber="471" value="1985-05-20">
<collectingDate id="DB55A92BFFBEFFEEFCD8601EFC0FFED2" box="[788,918,260,287]" pageId="20" pageNumber="471" value="1985-05-20">20.05.1985</collectingDate>
</date>
. V.G. 
<collectorName id="125A13D5FFBEFFEEFC15601FFBD2FED2" box="[985,1099,261,287]" pageId="20" pageNumber="471">Shilenkov</collectorName>
(
<collectionCode id="D9BEEEC6FFBEFFEEFB96601FFB19FED2" box="[1114,1152,261,287]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="20" pageNumber="471">CS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC77C5EFFBEFFEEFB59601FFE35FE8E" ID-GBIF-Occurrence="2429182986" collectionCode="ZMM" collectorName="Tadzhikistan" country="Tajikistan" location="Dushanbe" pageId="20" pageNumber="471" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Dushanbe" typeStatus="holotype">
<specimenCount id="A9A9BD8AFFBEFFEEFB59601FFB59FED2" box="[1173,1216,261,287]" pageId="20" pageNumber="471" type="male">1 ♂</specimenCount>
, 
<specimenCount id="A9A9BD8AFFBEFFEEFB00601FFB61FED2" box="[1228,1272,261,287]" pageId="20" pageNumber="471" type="female">1 ♀</specimenCount>
: ‘ 
<collectorName id="125A13D5FFBEFFEEFAC4601FFA05FED2" box="[1288,1436,261,287]" pageId="20" pageNumber="471">Tadzhikistan</collectorName>
| 
<collectingRegion id="7D6BB8E1FFBEFFEEFF686033FE8DFE8E" box="[164,276,297,323]" country="Tajikistan" name="Dushanbe" pageId="20" pageNumber="471">Dushanbe</collectingRegion>
| IV’ (
<collectionCode id="D9BEEEC6FFBEFFEEFE976033FE3DFE8E" box="[347,420,297,323]" pageId="20" pageNumber="471">ZMM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFBEFFEDFF0B6056FAABFA93" lastPageId="23" lastPageNumber="474" pageId="20" pageNumber="471" type="description">
<paragraph id="BF107603FFBEFFEEFF0B6056FA9DFE62" blockId="20.[151,1436,152,432]" pageId="20" pageNumber="471">
<emphasis id="8DDBAA11FFBEFFEEFF0B6056FEF4FEAA" bold="true" box="[199,365,332,359]" pageId="20" pageNumber="471">Redescription</emphasis>
. Measurements (n=40): HW: 0.67–0.77, HL: 0.42–0.52; AL (averaged): 1.33; OL: 0.13–0.17; PL: 0.60–0.67; PW: 1.22–1.35; ESL: 1.17–1.47; EW: 1.37–1.60; AW: 1.31–1.42; MTbL (averaged): 0.80, MTrL (averaged): 0.35 (MTrL 1–4: 0.20; MTrL 5: 0.15); AedL: 0.63–0.66; TL: 2.49–3.90 (
<typeStatus id="6014C8A1FFBEFFEEFB87608FFB21FE62" box="[1099,1208,405,431]" pageId="20" pageNumber="471" type="lectotype">lectotype</typeStatus>
: 2.55).
</paragraph>
<caption id="EBD0268BFFBEFFEEFF5B6630FBCEF8AB" ID-DOI="http://doi.org/10.5281/zenodo.3517628" ID-Zenodo-Dep="3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="20" pageNumber="471" startId="20.[151,264,1834,1859]" targetBox="[165,1424,451,1805]" targetPageId="20">
<paragraph id="BF107603FFBEFFEEFF5B6630FBCEF8AB" blockId="20.[151,1436,1834,1895]" pageId="20" pageNumber="471">
<emphasis id="8DDBAA11FFBEFFEEFF5B6630FECFF88E" bold="true" box="[151,342,1834,1859]" pageId="20" pageNumber="471">FIGURES 73–77.</emphasis>
Habitus of 
<taxonomicName id="78AF0D80FFBEFFEEFE1F6631FDDBF88E" authorityName="Leach" authorityYear="1819" box="[467,578,1835,1859]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="8DDBAA11FFBEFFEEFE1F6631FDDBF88E" box="[467,578,1835,1859]" italics="true" pageId="20" pageNumber="471">Anthobium</emphasis>
</taxonomicName>
: 73— 
<taxonomicName id="78AF0D80FFBEFFEEFD496631FC13F88E" authority="(Kazakhstan)" authorityName="Reitter" authorityYear="1891" baseAuthorityName="Kazakhstan" box="[645,906,1834,1859]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFBEFFEEFD496631FD62F88E" box="[645,763,1835,1859]" italics="true" pageId="20" pageNumber="471">A. reflexum</emphasis>
(Kazakhstan)
</taxonomicName>
74— 
<taxonomicName id="78AF0D80FFBEFFEEFC096631FBC5F88E" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[965,1116,1835,1859]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="deplanatum">
<emphasis id="8DDBAA11FFBEFFEEFC096631FC4DF88F" box="[965,980,1835,1858]" italics="true" pageId="20" pageNumber="471">A</emphasis>
. 
<emphasis id="8DDBAA11FFBEFFEEFC286631FBC5F88E" box="[996,1116,1835,1859]" italics="true" pageId="20" pageNumber="471">deplanatum</emphasis>
</taxonomicName>
(holotype), 75— 
<taxonomicName id="78AF0D80FFBEFFEEFAC36631FF6DF8AA" authorityName="Shavrin &amp; Smetana" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="gaoligongshanense">
<emphasis id="8DDBAA11FFBEFFEEFAC36631FA87F88F" box="[1295,1310,1835,1858]" italics="true" pageId="20" pageNumber="471">A</emphasis>
. 
<emphasis id="8DDBAA11FFBEFFEEFAE16631FF6DF8AA" italics="true" pageId="20" pageNumber="471">gaoligongshanense</emphasis>
</taxonomicName>
(holotype), 76— 
<taxonomicName id="78AF0D80FFBEFFEEFE6F6655FDDBF8AA" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[419,578,1871,1895]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="kashmiricum">
<emphasis id="8DDBAA11FFBEFFEEFE6F6655FE2BF8AB" box="[419,434,1871,1894]" italics="true" pageId="20" pageNumber="471">A</emphasis>
. 
<emphasis id="8DDBAA11FFBEFFEEFE736655FDDBF8AA" box="[447,578,1871,1895]" italics="true" pageId="20" pageNumber="471">kashmiricum</emphasis>
</taxonomicName>
, 77— 
<taxonomicName id="78AF0D80FFBEFFEEFD4E6655FC82F8AA" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[642,795,1871,1895]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="transversale">
<emphasis id="8DDBAA11FFBEFFEEFD4E6655FD08F8AB" box="[642,657,1871,1894]" italics="true" pageId="20" pageNumber="471">A</emphasis>
. 
<emphasis id="8DDBAA11FFBEFFEEFD526655FC82F8AA" box="[670,795,1871,1895]" italics="true" pageId="20" pageNumber="471">transversale</emphasis>
</taxonomicName>
(holotype). Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="BF107603FFBEFFEFFF0B668AFC3FFD4A" blockId="20.[151,1436,1936,2035]" lastBlockId="21.[151,1437,153,2015]" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471">
Forebody flattened, moderately wide. Body yellow to reddish-brown, usually with somewhat darker apical part of head (some specimens with fully brown head), antennomeres 5–11 (or 6–11) and abdomen (sometimes only apical portions of abdominal tergites darker); mouthparts, antennomeres 1–4 (or 1–5) and legs yellow. Body shiny; head with distinct, transverse microsculpture between punctures, denser in middle portion between eyes and even more on infraorbital ridges; neck with dense isodiametric meshes; middle portion of pronotum with moderately dense transverse meshes, sometimes indistinct or lacking in mediobasal third; middle portion of scutellum with distinct isodiametric or transverse meshes, or without them; abdomen with distinct transverse microsculpture. Head with very dense, fine, coriaceous punctures, sometimes merging in middle portion between eyes, denser, larger and deeper on infraorbital ridges; apical margin of neck with very dense fine punctation; pronotum with dense, irregular punctation, finer and denser in middle and sparser, larger and deeper on lateral portions, sometimes with narrow, transverse impunctated area on mediobasal third; scutellum with moderately dense punctation, as that in middle portion of pronotum; punctation of elytra very dense, large and deep, somewhat sparser and finer in basal portion and along suture, with interspaces between punctures in middle as long as diameter of two-three nearest punctures, each elytron bearing five to six very vague and tangled longitudinal rows of punctures, replaced by confluent punctures in some specimens; abdominal tergites with dense, regular, fine and moderately deep punctation. Body glabrous; clypeus with several semierect moderately long tactile setae; abdominal tergites with regular, dense, very short setation. Habitus as in 
<figureCitation id="27946A86FFBFFFEFFEA66377FE20FD4A" box="[362,441,621,647]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1834,1859]" captionTargetBox="[165,1424,451,1805]" captionTargetId="figure@20.[157,1431,445,1815]" captionTargetPageId="20" captionText="FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="21" pageNumber="472">Fig. 73</figureCitation>
and 
<figureCitation id="27946A86FFBFFFEFFE3C6377FDA5FD4A" box="[496,572,621,647]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1969,1994]" captionTargetBox="[159,1423,995,1946]" captionTargetId="figure@3.[151,1436,985,1950]" captionTargetPageId="3" captionText="FIGURES 1–2. Habitus of species of Anthobium consanguineum-group: 1—A. consanguineum (holotype), 2—A. jaegeri (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517610" httpUri="https://zenodo.org/record/3517610/files/figure.png" pageId="21" pageNumber="472">Figs. 2</figureCitation>
(1a, 1b) in 
<bibRefCitation id="DB3E0BF2FFBFFFEFFD716377FC3BFD4A" author="Tichomirova, A. L." box="[701,930,621,647]" pageId="21" pageNumber="472" refId="ref19870" refString="Tichomirova, A. L. (1973) Morfoekologicheskiye osobennosti i filogenez stafilinid (s katalogom fauny SSSR) [Morpho-ecological features and phylogeny of Staphylinidae (with a catalogue of the fauna of the USSR)]. Nauka, Moscow, 191 pp. [in Russian]" type="book" year="1973">Tichomirova (1973)</bibRefCitation>
.
</paragraph>
<paragraph id="BF107603FFBFFFEFFF0B638BFE4EFAC2" blockId="21.[151,1437,153,2015]" pageId="21" pageNumber="472">
Head moderately large (
<figureCitation id="27946A86FFBFFFEFFE16638BFDB6FD66" box="[474,559,657,683]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 17</figureCitation>
), 1.4–1.5 times as wide as long; middle portion slightly elevated, sometimes with transverse paired impressions about at level of midlength of eyes, with wide longitudinal impressions between middle elevated portion and eyes; infraorbital ridges narrowly elevated along dorsal margin of eyes; grooves in front of ocelli indistinct and short; some specimens with semicircular impression in mediobasal portion between ocelli; surface between punctures in middle portion between eyes and on infraorbital ridges irregularly elevated; postocular ridge distinct, obtuse; surface between postocular ridge and posterior margin of eye as long as two-three nearest combined ommatidia; anterior portion between antennal insertion and anterior margin of eye with indistinct to distinct, wide, semicircular notch. Eyes moderately small, convex. Ocelli large, situated about at level of postocular ridges; distance between ocelli about twice as long as distance between ocellus and posterior margin of eye. Labrum wide, transverse, with rounded membranous latero-apical portions, moderately deep apical emargination and short basolateral projections (
<figureCitation id="27946A86FFBFFFEFFDD462E3FDC7FBDE" box="[536,606,1016,1043]" captionStart="FIGURES 3–14" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[159,1421,638,1913]" captionTargetId="figure@4.[151,1436,625,1928]" captionText="FIGURES 3–14. Mouthparts (dorsal view) of Anthobium (3, 6, 9, 12—A. consanguineum, 4, 7, 10, 13—A. crassum, 5, 8, 11, 14—A. reflexum): 3–5—labrum, 6–8—mandibles (left and right), 9–11—maxilla (right), 12–14—labium and submentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523118" httpUri="https://zenodo.org/record/3523118/files/figure.png" pageId="21" pageNumber="472">Fig. 5</figureCitation>
). Mandibles short and moderately narrow, with somewhat obtuse apices, with distinct small, subtriangular tooth in about middle of right mandible (
<figureCitation id="27946A86FFBFFFEFFC166507FB86FBFA" box="[986,1055,1053,1079]" captionStart="FIGURES 3–14" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[159,1421,638,1913]" captionTargetId="figure@4.[151,1436,625,1928]" captionText="FIGURES 3–14. Mouthparts (dorsal view) of Anthobium (3, 6, 9, 12—A. consanguineum, 4, 7, 10, 13—A. crassum, 5, 8, 11, 14—A. reflexum): 3–5—labrum, 6–8—mandibles (left and right), 9–11—maxilla (right), 12–14—labium and submentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523118" httpUri="https://zenodo.org/record/3523118/files/figure.png" pageId="21" pageNumber="472">Fig. 8</figureCitation>
). Submentum and labium narrow, with very short, transverse preapical labial palpomere (
<figureCitation id="27946A86FFBFFFEFFCC6655BFCC6FB96" box="[778,863,1089,1115]" captionStart="FIGURES 3–14" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[159,1421,638,1913]" captionTargetId="figure@4.[151,1436,625,1928]" captionText="FIGURES 3–14. Mouthparts (dorsal view) of Anthobium (3, 6, 9, 12—A. consanguineum, 4, 7, 10, 13—A. crassum, 5, 8, 11, 14—A. reflexum): 3–5—labrum, 6–8—mandibles (left and right), 9–11—maxilla (right), 12–14—labium and submentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523118" httpUri="https://zenodo.org/record/3523118/files/figure.png" pageId="21" pageNumber="472">Fig. 14</figureCitation>
). Apical segment of maxillary palp about twice as long as preceding segment; galea slightly swollen apically; lacinia moderately wide (
<figureCitation id="27946A86FFBFFFEFFBAC657FFB2AFBB2" box="[1120,1203,1125,1151]" captionStart="FIGURES 3–14" captionStartId="4.[151,264,1948,1973]" captionTargetBox="[159,1421,638,1913]" captionTargetId="figure@4.[151,1436,625,1928]" captionText="FIGURES 3–14. Mouthparts (dorsal view) of Anthobium (3, 6, 9, 12—A. consanguineum, 4, 7, 10, 13—A. crassum, 5, 8, 11, 14—A. reflexum): 3–5—labrum, 6–8—mandibles (left and right), 9–11—maxilla (right), 12–14—labium and submentum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523118" httpUri="https://zenodo.org/record/3523118/files/figure.png" pageId="21" pageNumber="472">Fig. 11</figureCitation>
). Gular sutures with shortest distance between posterior margins of eyes (
<figureCitation id="27946A86FFBFFFEFFD256593FCA4FB6E" box="[745,829,1161,1187]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 17</figureCitation>
). Antenna moderately short, slightly exceeding shoulders of elytra when reclined, with antennomeres progressively widened apicad from middle; length × width of antennomeres (averaged): 1: 0.18 × 0.07; 2–3: 0.13 × 0.06; 4: 0.11 × 0.06; 5: 0.12 × 0.06; 6–8: 0.10 × 0.07; 9–10: 0.10 × 0.08; 11: 0.16 × 0.08.
</paragraph>
<paragraph id="BF107603FFBFFFEFFF0B6403FEB0F972" blockId="21.[151,1437,153,2015]" pageId="21" pageNumber="472">
Pronotum large and transverse, about twice as wide as long, 1.7–1.8 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad; apical margin slightly rounded, markedly narrower than posterior margin; anterior angles rounded, very strongly protruded anteriad; posterior angles acute, with slightly emarginated posterolateral margins; edges of lateral margins with small irregular crenulation, somewhat flattened laterobasally; middle portion wide, slightly elevated, sometimes with moderately wide longitudinal impression in middle, usually with very wide, transverse impression in mediobasal third (longitudinal impression of some specimens reaching mediobasal impression, some specimens without distinct impressions); lateral portions wide, markedly reflexed, each with very deep oval pit in middle, sometimes middle elevation with additional very deep, transverse impression connecting with medio-lateral pits. Prothorax (
<figureCitation id="27946A86FFBFFFEFFD606723FD66F99E" box="[684,767,1593,1619]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 20</figureCitation>
) with short acute intercoxal process, and moderately short protruded mesosternal processes, rounded apically. Mesothorax (
<figureCitation id="27946A86FFBFFFEFFCBE6747FC5DF9BA" box="[882,964,1629,1655]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 29</figureCitation>
) with somewhat acute intercoxal process, almost reaching posterior margin of mesocoxae. Scutellum as in 
<figureCitation id="27946A86FFBFFFEFFCA6679BFC25F956" box="[874,956,1665,1691]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 23</figureCitation>
. Metathorax as in 
<figureCitation id="27946A86FFBFFFEFFB46679BFB45F956" box="[1162,1244,1665,1691]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 32</figureCitation>
. Metendosternite as in 
<figureCitation id="27946A86FFBFFFEFFF1E67BFFEBCF972" box="[210,293,1700,1727]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 35</figureCitation>
.
</paragraph>
<paragraph id="BF107603FFBFFFEFFF0B67D3FED7F85A" blockId="21.[151,1437,153,2015]" pageId="21" pageNumber="472">
Elytra flattened, distinctly wider than long, twice as long as pronotum, slightly or distinctly widened posteriad from middle, reaching apical margin of abdominal tergite IV or V, with widely rounded apical angles and somewhat straight or slightly rounded apical margin truncated at suture; shoulders of elytra widely rounded, not protruded anteriad (
<figureCitation id="27946A86FFBFFFEFFECC662FFECAF89D" box="[256,339,1845,1872]" captionStart="FIGURES 15–35" captionStartId="5.[151,264,1907,1932]" captionTargetBox="[164,1422,180,1882]" captionTargetId="figure@5.[151,1436,174,1888]" captionText="FIGURES 15–35. Body parts of Anthobium (15, 18, 21, 24, 27, 30, 33—A. consanguineum, 16, 19, 22, 25, 28, 31, 34—A. crassum, 17, 20, 23, 26, 29, 32, 35—A. reflexum): 15–17—ventral aspect of head, 18–20—ventral aspect of prothorax, 21–23— scutellum (dorsal view), 24–26—ventral aspect of elytron, 27–29—ventral aspect of mesothorax, 30–32—ventral aspect of metathorax (without metanepisternum), 33–35—metendosternite (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523122" httpUri="https://zenodo.org/record/3523122/files/figure.png" pageId="21" pageNumber="472">Fig. 26</figureCitation>
); lateral portions of elytra wide, markedly explanate, latero-apical margins with small, acute crenulation; sometimes surface of each elytron with very indistinct one to three longitudinal elevation in middle. Wings fully developed.
</paragraph>
<paragraph id="BF107603FFBFFFEFFF0B66BBFCE1F812" blockId="21.[151,1437,153,2015]" pageId="21" pageNumber="472">Legs moderately long and slender, tibiae covered by sparse elongate setae, with a few strong setae on apical margin near apex; metatarsus less than twice as long as metatibia.</paragraph>
<caption id="EBD0268BFFBCFFECFF5B66ADFC95F83E" ID-DOI="http://doi.org/10.5281/zenodo.3517630" ID-Zenodo-Dep="3517630" httpUri="https://zenodo.org/record/3517630/files/figure.png" pageId="22" pageNumber="473" startId="22.[151,264,1975,2000]" targetBox="[266,1322,176,1949]" targetPageId="22">
<paragraph id="BF107603FFBCFFECFF5B66ADFC95F83E" blockId="22.[151,1436,1975,2036]" pageId="22" pageNumber="473">
<emphasis id="8DDBAA11FFBCFFECFF5B66ADFECFF81D" bold="true" box="[151,342,1975,2000]" pageId="22" pageNumber="473">FIGURES 78–81.</emphasis>
Aedeagus of 
<taxonomicName id="78AF0D80FFBCFFECFE2B66A2FD2FF81D" authorityName=": Smetana" authorityYear="2004" box="[487,694,1976,2000]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFBCFFECFE2B66A2FD2FF81D" box="[487,694,1976,2000]" italics="true" pageId="22" pageNumber="473">Anthobium reflexum</emphasis>
</taxonomicName>
(78–79: Kazakhstan) and 
<taxonomicName id="78AF0D80FFBCFFECFC0666A2FBF2F81D" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[970,1131,1976,2000]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="kashmiricum">
<emphasis id="8DDBAA11FFBCFFECFC0666A2FBF2F81D" box="[970,1131,1976,2000]" italics="true" pageId="22" pageNumber="473">A. kashmiricum</emphasis>
</taxonomicName>
(80–81: India, Kashmir): 78, 80—parameral view, 79, 81—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="EBD0268BFFBDFFEDFF5B628EFAA5FC60" ID-DOI="http://doi.org/10.5281/zenodo.3517632" ID-Zenodo-Dep="3517632" box="[151,1340,916,941]" httpUri="https://zenodo.org/record/3517632/files/figure.png" pageId="23" pageNumber="474" startId="23.[151,250,916,941]" targetBox="[151,1435,174,897]" targetPageId="23">
<paragraph id="BF107603FFBDFFEDFF5B628EFAA5FC60" blockId="23.[151,1340,916,941]" box="[151,1340,916,941]" pageId="23" pageNumber="474">
<emphasis id="8DDBAA11FFBDFFEDFF5B628EFEB9FC60" bold="true" box="[151,288,916,941]" pageId="23" pageNumber="474">FIGURE 82.</emphasis>
Distribution of 
<taxonomicName id="78AF0D80FFBDFFEDFE0E628FFD09FC60" authorityName=": Smetana" authorityYear="2004" box="[450,656,917,941]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFBDFFEDFE0E628FFD09FC60" box="[450,656,917,941]" italics="true" pageId="23" pageNumber="474">Anthobium reflexum</emphasis>
</taxonomicName>
(black circles) and 
<taxonomicName id="78AF0D80FFBDFFEDFC9B628FFC6FFC60" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[855,1014,917,941]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="kashmiricum">
<emphasis id="8DDBAA11FFBDFFEDFC9B628FFC6FFC60" box="[855,1014,917,941]" italics="true" pageId="23" pageNumber="474">A. kashmiricum</emphasis>
</taxonomicName>
(black squares) in Central Asia.
</paragraph>
</caption>
<paragraph id="BF107603FFBDFFEDFF0B62C1FF7BFBF0" blockId="23.[151,1437,987,1806]" pageId="23" pageNumber="474">Abdomen distinctly narrower than elytra, with a pair of small rounded tomentose wing-folding spots in middle of tergite V (some specimens without visible spots); apical margin of abdominal tegite VII with very fine palisade fringe.</paragraph>
<paragraph id="BF107603FFBDFFEDFF0B655DFC0AFB00" blockId="23.[151,1437,987,1806]" pageId="23" pageNumber="474">
Male. Protarsomeres 1–4 slightly widened. Apical margin of abdominal tergite VIII straight (
<figureCitation id="27946A86FFBDFFEDFB25655DFAA7FBAC" box="[1257,1342,1095,1121]" captionStart="FIGURES 36–47" captionStartId="6.[151,264,1086,1111]" captionTargetBox="[161,1425,179,1057]" captionTargetId="figure@6.[151,1436,174,1067]" captionText="FIGURES 36–47. Abdominal segments of Anthobium (36, 39, 42, 45, 48—A. consanguineum, 37, 40, 43, 46, 49—A. crassum, 38, 41, 44, 47, 50—A. reflexum): 36–38—male tergite VIII (dorsal view), 39–41—male sternite VIII (ventral view), 42–44—female tergite VIII (dorsal view), 45–47—female sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523124" httpUri="https://zenodo.org/record/3523124/files/figure.png" pageId="23" pageNumber="474">Fig. 38</figureCitation>
). Apical margin of abdominal sternite VIII emarginated (
<figureCitation id="27946A86FFBDFFEDFD746571FC95FB48" box="[696,780,1131,1157]" captionStart="FIGURES 36–47" captionStartId="6.[151,264,1086,1111]" captionTargetBox="[161,1425,179,1057]" captionTargetId="figure@6.[151,1436,174,1067]" captionText="FIGURES 36–47. Abdominal segments of Anthobium (36, 39, 42, 45, 48—A. consanguineum, 37, 40, 43, 46, 49—A. crassum, 38, 41, 44, 47, 50—A. reflexum): 36–38—male tergite VIII (dorsal view), 39–41—male sternite VIII (ventral view), 42–44—female tergite VIII (dorsal view), 45–47—female sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523124" httpUri="https://zenodo.org/record/3523124/files/figure.png" pageId="23" pageNumber="474">Fig. 41</figureCitation>
). Aedeagus (
<figureCitation id="27946A86FFBDFFEDFC526571FC68FB48" box="[926,1009,1131,1157]" captionStart="FIGURES 78–81" captionStartId="22.[151,264,1975,2000]" captionTargetBox="[266,1322,176,1949]" captionTargetId="figure@22.[255,1333,166,1956]" captionTargetPageId="22" captionText="FIGURES 78–81. Aedeagus of Anthobium reflexum (78–79: Kazakhstan) and A. kashmiricum (80–81: India, Kashmir): 78, 80—parameral view, 79, 81—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517630" httpUri="https://zenodo.org/record/3517630/files/figure.png" pageId="23" pageNumber="474">Fig. 78</figureCitation>
) narrow, long, with truncated apex of median lobe; parameres long and wide, slightly exceeding apex of median lobe, with two apical and two preapical setae; internal sac narrow and long. Aedeagus laterally as in 
<figureCitation id="27946A86FFBDFFEDFCF065A9FC16FB00" box="[828,911,1203,1229]" captionStart="FIGURES 78–81" captionStartId="22.[151,264,1975,2000]" captionTargetBox="[266,1322,176,1949]" captionTargetId="figure@22.[255,1333,166,1956]" captionTargetPageId="22" captionText="FIGURES 78–81. Aedeagus of Anthobium reflexum (78–79: Kazakhstan) and A. kashmiricum (80–81: India, Kashmir): 78, 80—parameral view, 79, 81—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517630" httpUri="https://zenodo.org/record/3517630/files/figure.png" pageId="23" pageNumber="474">Fig. 79</figureCitation>
.
</paragraph>
<paragraph id="BF107603FFBDFFEDFF0B65CDFBFAFAD8" blockId="23.[151,1437,987,1806]" pageId="23" pageNumber="474">
Female. Protarsomeres 1–4 narrow. Apical margin of abdominal tergite VIII emarginated (
<figureCitation id="27946A86FFBDFFEDFB7E65CDFA9FFB3C" box="[1202,1286,1239,1265]" captionStart="FIGURES 36–47" captionStartId="6.[151,264,1086,1111]" captionTargetBox="[161,1425,179,1057]" captionTargetId="figure@6.[151,1436,174,1067]" captionText="FIGURES 36–47. Abdominal segments of Anthobium (36, 39, 42, 45, 48—A. consanguineum, 37, 40, 43, 46, 49—A. crassum, 38, 41, 44, 47, 50—A. reflexum): 36–38—male tergite VIII (dorsal view), 39–41—male sternite VIII (ventral view), 42–44—female tergite VIII (dorsal view), 45–47—female sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523124" httpUri="https://zenodo.org/record/3523124/files/figure.png" pageId="23" pageNumber="474">Fig. 44</figureCitation>
). Apical margin of abdominal sternite VIII widely rounded (
<figureCitation id="27946A86FFBDFFEDFD6065E1FC99FAD8" box="[684,768,1275,1301]" captionStart="FIGURES 36–47" captionStartId="6.[151,264,1086,1111]" captionTargetBox="[161,1425,179,1057]" captionTargetId="figure@6.[151,1436,174,1067]" captionText="FIGURES 36–47. Abdominal segments of Anthobium (36, 39, 42, 45, 48—A. consanguineum, 37, 40, 43, 46, 49—A. crassum, 38, 41, 44, 47, 50—A. reflexum): 36–38—male tergite VIII (dorsal view), 39–41—male sternite VIII (ventral view), 42–44—female tergite VIII (dorsal view), 45–47—female sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3523124" httpUri="https://zenodo.org/record/3523124/files/figure.png" pageId="23" pageNumber="474">Fig. 47</figureCitation>
). Genital segment as in 
<figureCitation id="27946A86FFBDFFEDFBC065E1FBC6FAD8" box="[1036,1119,1274,1301]" captionStart="FIGURES 48–50" captionStartId="6.[151,264,1850,1875]" captionTargetBox="[157,1429,1202,1822]" captionTargetId="figure@6.[151,1436,1197,1830]" captionTargetPageId="6" captionText="FIGURES 48–50. Female genital segment (dorsal view) of Anthobium: 48–A. consanguineum, 49—A. crassum, 50—A. reflexum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517612" httpUri="https://zenodo.org/record/3517612/files/figure.png" pageId="23" pageNumber="474">Fig. 50</figureCitation>
.
</paragraph>
<paragraph id="BF107603FFBDFFEDFF0B6404FAABFA93" blockId="23.[151,1437,987,1806]" pageId="23" pageNumber="474">
<emphasis id="8DDBAA11FFBDFFEDFF0B6404FE28FAF4" bold="true" box="[199,433,1310,1337]" pageId="23" pageNumber="474">Comparative notes.</emphasis>
<taxonomicName id="78AF0D80FFBDFFEDFE756405FD05FAF4" authorityName=": Smetana" authorityYear="2004" box="[441,668,1311,1337]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFBDFFEDFE756405FD05FAF4" box="[441,668,1311,1337]" italics="true" pageId="23" pageNumber="474">Anthobium reflexum</emphasis>
</taxonomicName>
differs from the remaining species of the 
<taxonomicName id="78AF0D80FFBDFFEDFBBA6405FB4EFAF4" authorityName=": Smetana" authorityYear="2004" box="[1142,1239,1311,1337]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFBDFFEDFBBA6405FB4EFAF4" box="[1142,1239,1311,1337]" italics="true" pageId="23" pageNumber="474">reflexum</emphasis>
</taxonomicName>
-group by the lateral edges of the pronotum without projections, and by the shape of the aedeagus and its interal structure.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFBDFFEDFF0B647CFF44F9DC" pageId="23" pageNumber="474" type="distribution">
<paragraph id="BF107603FFBDFFEDFF0B647CFC06FA68" blockId="23.[151,1437,987,1806]" pageId="23" pageNumber="474">
<emphasis id="8DDBAA11FFBDFFEDFF0B647CFEF8FA4C" bold="true" box="[199,353,1382,1409]" pageId="23" pageNumber="474">Distribution.</emphasis>
<taxonomicName id="78AF0D80FFBDFFEDFEAB647DFDD3FA4C" authorityName=": Smetana" authorityYear="2004" box="[359,586,1383,1409]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFBDFFEDFEAB647DFDD3FA4C" box="[359,586,1383,1409]" italics="true" pageId="23" pageNumber="474">Anthobium reflexum</emphasis>
</taxonomicName>
is distributed in mountains of the central Asia in southern 
<collectingCountry id="C7B83693FFBDFFEDFB1B647DFAC4FA4C" box="[1239,1373,1383,1409]" name="Kazakhstan" pageId="23" pageNumber="474">Kazakhstan</collectingCountry>
, eastern 
<collectingCountry id="C7B83693FFBDFFEDFF0E6491FEDAFA68" box="[194,323,1419,1445]" name="Uzbekistan" pageId="23" pageNumber="474">Uzbekistan</collectingCountry>
, western 
<collectingCountry id="C7B83693FFBDFFEDFE606496FDB4FA6B" box="[428,557,1420,1446]" name="Kyrgyzstan" pageId="23" pageNumber="474">Kyrgyzstan</collectingCountry>
and western 
<collectingCountry id="C7B83693FFBDFFEDFD0D6491FCB6FA68" box="[705,815,1419,1445]" name="Tajikistan" pageId="23" pageNumber="474">Tajikistan</collectingCountry>
(
<figureCitation id="27946A86FFBDFFEDFCF26491FC0BFA68" box="[830,914,1419,1445]" captionStart="FIGURE 82" captionStartId="23.[151,250,916,941]" captionTargetBox="[151,1435,174,897]" captionTargetId="figure@23.[151,1435,174,897]" captionTargetPageId="23" captionText="FIGURE 82. Distribution of Anthobium reflexum (black circles) and A. kashmiricum (black squares) in Central Asia." figureDoi="http://doi.org/10.5281/zenodo.3517632" httpUri="https://zenodo.org/record/3517632/files/figure.png" pageId="23" pageNumber="474">Fig. 82</figureCitation>
).
</paragraph>
<paragraph id="BF107603FFBDFFEDFF0B64B4FF44F9DC" blockId="23.[151,1437,987,1806]" pageId="23" pageNumber="474">
<emphasis id="8DDBAA11FFBDFFEDFF0B64B4FED2FA04" bold="true" box="[199,331,1454,1481]" pageId="23" pageNumber="474">Bionomics.</emphasis>
The species is known mainly from high elevations and inhabits riparian, forest and swampy (
<bibRefCitation id="DB3E0BF2FFBDFFEDFA8264B5FF49FA20" author="Reitter, E." pageId="23" pageNumber="474" pagination="195 - 199" refId="ref19463" refString="Reitter, E. (1891) Zweiter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische Zeitung, 10, 195 - 199." type="journal article" year="1891">Reitter 1891</bibRefCitation>
) habitats, where it can be found in mosses (
<bibRefCitation id="DB3E0BF2FFBDFFEDFD0C64C9FCC5FA20" author="Coiffait, H." box="[704,860,1491,1517]" pageId="23" pageNumber="474" pagination="142 - 148" refId="ref19255" refString="Coiffait, H. (1970) Staphylinidae d'Ouzbekistan. L'Entomologiste, 26, 142 - 148." type="journal article" year="1970">Coiffait 1970</bibRefCitation>
), near streams, in litter and occasionally near snow fields.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFBDFFEDFF0B6701FCCAF8C0" pageId="23" pageNumber="474" type="discussion">
<paragraph id="BF107603FFBDFFEDFF0B6701FBE5F924" blockId="23.[151,1437,987,1806]" pageId="23" pageNumber="474">
<emphasis id="8DDBAA11FFBDFFEDFF0B6701FEA2F9F8" bold="true" box="[199,315,1563,1589]" pageId="23" pageNumber="474">Remarks.</emphasis>
<taxonomicName id="78AF0D80FFBDFFEDFE8C6701FDA7F9F8" authorityName="Reitter" authorityYear="1891" box="[320,574,1563,1589]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFBDFFEDFE8C6701FDA7F9F8" box="[320,574,1563,1589]" italics="true" pageId="23" pageNumber="474">Lathrimaeum reflexum</emphasis>
</taxonomicName>
was originally described from unspecified number of specimens from “Taschkent” (see above), Turkestan (now 
<collectingCountry id="C7B83693FFBDFFEDFDD56725FD05F997" box="[537,668,1599,1626]" name="Uzbekistan" pageId="23" pageNumber="474">Uzbekistan</collectingCountry>
). 
<bibRefCitation id="DB3E0BF2FFBDFFEDFD61675AFCACF994" author="Luze, G." box="[685,821,1598,1626]" pageId="23" pageNumber="474" pagination="53 - 69" refId="ref19432" refString="Luze, G. (1905) Revision der palaarktischen Arten der Staphyliniden-Gattung Lathrimaeum Er. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 55, 53 - 69." type="journal article" year="1905">Luze (1905)</bibRefCitation>
placed this species to the subgenus 
<taxonomicName id="78AF0D80FFBDFFEDFB0D6725FF56F9B0" authority="Luze, 1905" authorityName="Luze" authorityYear="1905" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="subGenus" subGenus="Prionothorax">
<emphasis id="8DDBAA11FFBDFFEDFB0D6725FAC1F994" box="[1217,1368,1599,1625]" italics="true" pageId="23" pageNumber="474">Prionothorax</emphasis>
<bibRefCitation id="DB3E0BF2FFBDFFEDFA91675AFF56F9B0" author="Luze, G." pageId="23" pageNumber="474" pagination="53 - 69" refId="ref19432" refString="Luze, G. (1905) Revision der palaarktischen Arten der Staphyliniden-Gattung Lathrimaeum Er. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 55, 53 - 69." type="journal article" year="1905">Luze, 1905</bibRefCitation>
</taxonomicName>
and redescribed it. 
<bibRefCitation id="DB3E0BF2FFBDFFEDFE626779FDCEF9B0" author="Coiffait, H." box="[430,599,1635,1661]" pageId="23" pageNumber="474" pagination="142 - 148" refId="ref19255" refString="Coiffait, H. (1970) Staphylinidae d'Ouzbekistan. L'Entomologiste, 26, 142 - 148." type="journal article" year="1970">Coiffait (1970)</bibRefCitation>
recorded the species from “Monts Tschingan” (
<collectingCountry id="C7B83693FFBDFFEDFBBF6779FB61F9B3" box="[1139,1272,1635,1662]" name="Uzbekistan" pageId="23" pageNumber="474">Uzbekistan</collectingCountry>
). Kastcheev &amp; Iskov (2001) recorded it from Aksu-Dzhabagly Natural Reserve, 
<collectingCountry id="C7B83693FFBDFFEDFCA1679DFC69F96C" box="[877,1008,1671,1697]" name="Kazakhstan" pageId="23" pageNumber="474">Kazakhstan</collectingCountry>
(see revised specimens in the material section above). 
<bibRefCitation id="DB3E0BF2FFBDFFEDFE8067B1FDB6F908" author="Tichomirova, A. L." box="[332,559,1707,1733]" pageId="23" pageNumber="474" refId="ref19870" refString="Tichomirova, A. L. (1973) Morfoekologicheskiye osobennosti i filogenez stafilinid (s katalogom fauny SSSR) [Morpho-ecological features and phylogeny of Staphylinidae (with a catalogue of the fauna of the USSR)]. Nauka, Moscow, 191 pp. [in Russian]" type="book" year="1973">Tichomirova (1973)</bibRefCitation>
and Smetana &amp; 
<bibRefCitation id="DB3E0BF2FFBDFFEDFD3E67B1FC38F908" author="Schulke, M. &amp; Smetana, A." box="[754,929,1706,1733]" pageId="23" pageNumber="474" pagination="304 - 1134" refId="ref19533" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke (2015)</bibRefCitation>
noted distribution of the species for Middle Asia (“Ср[едНЯЯ]. АЗ[иЯ]”) and 
<collectingCountry id="C7B83693FFBDFFEDFDC967D5FDEFF924" box="[517,630,1743,1769]" name="Tajikistan" pageId="23" pageNumber="474">Tajikistan</collectingCountry>
, 
<collectingCountry id="C7B83693FFBDFFEDFD4D67D5FD66F924" box="[641,767,1743,1769]" name="Uzbekistan" pageId="23" pageNumber="474">Uzbekistan</collectingCountry>
(“TD TD [sic] UZ”) respectively.
</paragraph>
<paragraph id="BF107603FFBDFFEDFF0B67E9FCCAF8C0" blockId="23.[151,1437,987,1806]" box="[199,851,1779,1806]" pageId="23" pageNumber="474">
The species is reported from 
<collectingCountry id="C7B83693FFBDFFEDFDC167EEFD17F8C3" box="[525,654,1780,1806]" name="Kyrgyzstan" pageId="23" pageNumber="474">Kyrgyzstan</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>