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<document id="ED104A4280E582B83B2BA5DEDB22242D" ID-CLB-Dataset="25154" ID-DOI="10.11646/zootaxa.4688.4.1" ID-GBIF-Dataset="c4072224-34c3-4b12-9415-a59d7c9484b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3517608" ID-ZooBank="BABABF8C-30B3-45D0-89B5-6F4DC1B19B70" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571896092680" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2019" docId="3706C715FFB1FFE7FF5B63E8FDBDFC92" docLanguage="en" docName="zootaxa.4688.4.1.pdf" docOrigin="Zootaxa 4688 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium transversale Shavrin &amp; Smetana 2019, sp.n." docType="treatment" docVersion="10" lastPageNumber="480" masterDocId="CB3FBF6DFFAAFFFAFFCC611AFF99FFCD" masterDocTitle="A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group" masterLastPageNumber="482" masterPageNumber="451" pageNumber="478" updateTime="1698760874617" updateUser="ExternalLinkService">
<mods:mods id="72EE1C9867752EF686D3F372B9D2E599" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="899BC82F22AE1F0A7CD7DC7F93863A81">
<mods:title id="99C326F542601191BD3F1923F2336E78">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group</mods:title>
</mods:titleInfo>
<mods:name id="1B7227A8C1975B9455B1CC05D1273D58" type="personal">
<mods:role id="4EDD89644A7E03F84CF250AD9772B424">
<mods:roleTerm id="D2E6D66A69EA07F5320D29987EE259D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE69462E6E406BF74BC2E53FF3E4D781">Shavrin, Alexey V.</mods:namePart>
</mods:name>
<mods:name id="AAAF5F05B891068D2246C56EF9CF05FF" type="personal">
<mods:role id="CEAB28067A29453DA28730BAC66D7C5E">
<mods:roleTerm id="E3C77BE233EF8575077379EF96BD7A23">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E6D95DBD040FDDE0AF655B536454D5B">Smetana, Aleš</mods:namePart>
</mods:name>
<mods:typeOfResource id="6D9D4216DB379ED39DB36ADED173237C">text</mods:typeOfResource>
<mods:relatedItem id="462E32E83082C2C0899DC60772E689F7" type="host">
<mods:titleInfo id="F792DCF07133019EF10E9D85FA96C1D0">
<mods:title id="0934E404B73FFCEB90E9BED275356FBD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A8D39354477E83B709EA32E1794B2084">
<mods:date id="E36A1342767400B93A44231996DA8F9F">2019</mods:date>
<mods:detail id="BB850FD48F33BE75DE10CCDBF58F9C6B" type="pubDate">
<mods:number id="011A4AC64F112FF142485DD2F4D26065">2019-10-24</mods:number>
</mods:detail>
<mods:detail id="7842248E74131689D348E7790039E72A" type="volume">
<mods:number id="6D6E7FEBEF390FC1B5073482D22C9F50">4688</mods:number>
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<mods:number id="44D4A7A61A701398775EB35870034FD7">4</mods:number>
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<mods:start id="78296D2CA0C21DF31A6B9BD20FA96B8D">451</mods:start>
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<mods:classification id="FDC423E4A9DDF7FAA609FEDD14029324">journal article</mods:classification>
<mods:identifier id="2CEEA934D082F77EE54AE8EDFAE6D181" type="CLB-Dataset">25154</mods:identifier>
<mods:identifier id="8D484D90391F41104767EB85771A51A2" type="DOI">10.11646/zootaxa.4688.4.1</mods:identifier>
<mods:identifier id="AEE82A4CC6B1AD37CE227ED8AB581FDE" type="GBIF-Dataset">c4072224-34c3-4b12-9415-a59d7c9484b6</mods:identifier>
<mods:identifier id="43584DCA0F93974410B55B1F08E6C2A7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B739CDFAF0D61D34EFB35C6235D12E43" type="Zenodo-Dep">3517608</mods:identifier>
<mods:identifier id="CA0F23B750ECF9A036DDC3FB2C47ACE9" type="ZooBank">BABABF8C-30B3-45D0-89B5-6F4DC1B19B70</mods:identifier>
</mods:mods>
<treatment id="3706C715FFB1FFE7FF5B63E8FDBDFC92" ID-DOI="http://doi.org/10.5281/zenodo.5921727" ID-GBIF-Taxon="159834454" ID-Zenodo-Dep="5921727" LSID="urn:lsid:plazi:treatment:3706C715FFB1FFE7FF5B63E8FDBDFC92" httpUri="http://treatment.plazi.org/id/3706C715FFB1FFE7FF5B63E8FDBDFC92" lastPageId="29" lastPageNumber="480" pageId="27" pageNumber="478">
<subSubSection id="F7B52588FFB1FFE1FF5B63E8FC9AFCC0" box="[151,771,754,781]" pageId="27" pageNumber="478" type="nomenclature">
<paragraph id="BF107603FFB1FFE1FF5B63E8FC9AFCC0" blockId="27.[151,771,754,817]" box="[151,771,754,781]" pageId="27" pageNumber="478">
<heading id="E458C16FFFB1FFE1FF5B63E8FC9AFCC0" bold="true" box="[151,771,754,781]" fontSize="11" level="1" pageId="27" pageNumber="478" reason="1">
<emphasis id="8DDBAA11FFB1FFE1FF5B63E8FC9AFCC0" bold="true" box="[151,771,754,781]" pageId="27" pageNumber="478">
<taxonomicName id="78AF0D80FFB1FFE1FF5B63E8FD22FCC1" ID-CoL="8XTTS" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[151,699,754,781]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="transversale" status="sp. nov.">
<emphasis id="8DDBAA11FFB1FFE1FF5B63E8FE23FCC0" bold="true" box="[151,442,754,781]" italics="true" pageId="27" pageNumber="478">Anthobium transversale</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="96E8176AFFB1FFE1FD0463E9FC9AFCC0" box="[712,771,755,781]" pageId="27" pageNumber="478" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFB1FFE1FF5B620CFEC1FCFD" box="[151,344,790,817]" pageId="27" pageNumber="478" type="description">
<paragraph id="BF107603FFB1FFE1FF5B620CFEC1FCFD" blockId="27.[151,771,754,817]" box="[151,344,790,817]" pageId="27" pageNumber="478">
(
<figureCitation id="27946A86FFB1FFE1FF53620CFF64FCFC" box="[159,253,790,817]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1834,1859]" captionTargetBox="[165,1424,451,1805]" captionTargetId="figure@20.[157,1431,445,1815]" captionTargetPageId="20" captionText="FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="27" pageNumber="478">Figs. 77</figureCitation>
,
<figureCitation id="27946A86FFB1FFE1FEC5620CFEC9FCFD" box="[265,336,790,816]" captionStart-0="FIGURE 87" captionStart-1="FIGURES 88–89" captionStartId-0="29.[151,250,1614,1639]" captionStartId-1="30.[151,264,1425,1450]" captionTargetBox-0="[151,1435,913,1541]" captionTargetBox-1="[163,1424,179,1399]" captionTargetId-0="figure@29.[151,1435,913,1541]" captionTargetId-1="figure@30.[151,1436,174,1406]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURE 87. Distribution of Anthobium deplanatum (black circles), A. gaoligongshanense (black square) and A. transversale (black triangle) in the south-western China." captionText-1="FIGURES 88–89. Aedeagus of Anthobium transversale (paratype): 88—parameral view, 89—lateral view. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3517636" figureDoi-1="http://doi.org/10.5281/zenodo.3517638" httpUri-0="https://zenodo.org/record/3517636/files/figure.png" httpUri-1="https://zenodo.org/record/3517638/files/figure.png" pageId="27" pageNumber="478">87–89</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFB1FFE1FF5B6244FE23FB9D" pageId="27" pageNumber="478" type="materials_examined">
<paragraph id="BF107603FFB1FFE1FF5B6244FDB6FC29" blockId="27.[151,1437,861,2040]" pageId="27" pageNumber="478">
<emphasis id="8DDBAA11FFB1FFE1FF5B6244FE51FCB5" bold="true" box="[151,456,861,888]" pageId="27" pageNumber="478">Type material examined:</emphasis>
<materialsCitation id="0FC77C5EFFB1FFE1FE1E6244FE58FC0D" ID-GBIF-Occurrence="2429182971" collectingDate="1996-07-06" collectorName="A. Smetana &amp; J. Farkac &amp; P. Kabatek'" country="China" county="Gongga" elevation="3000" latitude="29.583334" location="Hailuogou" longLatPrecision="1226" longitude="102.0" municipality="Shan" pageId="27" pageNumber="478" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="6014C8A1FFB1FFE1FE1E6244FDA3FCB5" box="[466,570,862,888]" pageId="27" pageNumber="478" type="holotype">Holotype</typeStatus>
<specimenCount id="A9A9BD8AFFB1FFE1FD886245FDC5FCB4" box="[580,604,863,889]" pageId="27" pageNumber="478" type="male">♂</specimenCount>
: ‘
<collectingCountry id="C7B83693FFB1FFE1FDBC6244FD55FCB5" box="[624,716,862,888]" name="China" pageId="27" pageNumber="478">CHINA</collectingCountry>
<collectingRegion id="7D6BB8E1FFB1FFE1FD1A6244FCAAFCB5" box="[726,819,862,888]" country="China" name="Sichuan" pageId="27" pageNumber="478">Sichuan</collectingRegion>
,
<collectingCounty id="56710E8FFFB1FFE1FC8E6244FC02FCB5" box="[834,923,862,888]" pageId="27" pageNumber="478">Gongga</collectingCounty>
|
<collectingMunicipality id="5F74EC79FFB1FFE1FC786244FC76FCB5" box="[948,1007,862,888]" pageId="27" pageNumber="478">Shan</collectingMunicipality>
,
<location id="BA7020D8FFB1FFE1FC326244FBEEFCB5" LSID="urn:lsid:plazi:treatment:3706C715FFB1FFE7FF5B63E8FDBDFC92:BA7020D8FFB1FFE1FC326244FBEEFCB5" box="[1022,1143,862,888]" country="China" county="Gongga" latitude="29.583334" longLatPrecision="1226" longitude="102.0" municipality="Shan" name="Hailuogou" pageId="27" pageNumber="478" stateProvince="Sichuan">Hailuogou</location>
, above |
<location id="BA7020D8FFB1FFE1FB2E6244FABCFCB5" LSID="urn:lsid:plazi:treatment:3706C715FFB1FFE7FF5B63E8FDBDFC92:BA7020D8FFB1FFE1FB2E6244FABCFCB5" box="[1250,1317,862,888]" country="China" county="Gongga" latitude="29.583334" longLatPrecision="1226" longitude="102.0" municipality="Shan" name="Camp" pageId="27" pageNumber="478" stateProvince="Sichuan">Camp</location>
3,
<quantity id="7857DBE6FFB1FFE1FA826244FA05FCB5" box="[1358,1436,862,888]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="478" unit="m" value="3000.0">
<elevation id="34829130FFB1FFE1FA826244FA05FCB5" box="[1358,1436,862,888]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="478" unit="m" value="3000.0">3000m</elevation>
</quantity>
<date id="CB1150C3FFB1FFE1FF5B6298FF6FFC51" box="[151,246,898,924]" pageId="27" pageNumber="478" value="1996-07-06">
<collectingDate id="DB55A92BFFB1FFE1FF5B6298FF6FFC51" box="[151,246,898,924]" pageId="27" pageNumber="478" value="1996-07-06">6.VII.96</collectingDate>
</date>
|
<geoCoordinate id="DA9B10C4FFB1FFE1FECB629BFEC6FC51" box="[263,351,897,924]" degrees="29" direction="north" minutes="35" orientation="latitude" pageId="27" pageNumber="478" precision="925" value="29.583334">29°35N</geoCoordinate>
<geoCoordinate id="DA9B10C4FFB1FFE1FEA96298FE5EFC51" box="[357,455,898,924]" degrees="102" direction="east" minutes="00" orientation="longitude" pageId="27" pageNumber="478" precision="925" value="102.0">102°00E</geoCoordinate>
C53’ &lt;rectangular label, printed&gt;, ‘collected by |
<collectorName id="125A13D5FFB1FFE1FC206298FBE9FC51" box="[1004,1136,898,924]" pageId="27" pageNumber="478">A. Smetana</collectorName>
,
<collectorName id="125A13D5FFB1FFE1FBB66299FB47FC51" box="[1146,1246,898,925]" pageId="27" pageNumber="478">J. Farkač</collectorName>
| and
<collectorName id="125A13D5FFB1FFE1FAD26299FA04FC51" box="[1310,1437,898,925]" pageId="27" pageNumber="478">P. Kabátek’</collectorName>
&lt;rectangular label, printed
</materialsCitation>
&gt;, ‘
<materialsCitation id="0FC77C5EFFB1FFE1FE1262BCFDB2FC29" ID-GBIF-Occurrence="2429182987" collectionCode="NSMT" collectorName="Shavrin A. V. &amp; Smetana A." country="China" pageId="27" pageNumber="478" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="6014C8A1FFB1FFE1FE1262BCFDEEFC0D" box="[478,631,934,960]" pageId="27" pageNumber="478" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="78AF0D80FFB1FFE1FD4B62BCFC9EFC0D" authorityName="Leach" authorityYear="1819" box="[647,775,934,960]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="genus">Anthobium</taxonomicName>
| transversale
<emphasis id="8DDBAA11FFB1FFE1FC6862BCFC45FC0D" bold="true" box="[932,988,934,960]" pageId="27" pageNumber="478">
<taxonomicNameLabel id="96E8176AFFB1FFE1FC6862BCFC45FC0D" box="[932,988,934,960]" pageId="27" pageNumber="478" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="125A13D5FFB1FFE1FC2162BCFBE5FC0D" box="[1005,1148,934,960]" pageId="27" pageNumber="478">Shavrin A.V.</collectorName>
&amp;
<collectorName id="125A13D5FFB1FFE1FB5162BCFA87FC0D" box="[1181,1310,934,960]" pageId="27" pageNumber="478">Smetana A.</collectorName>
2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="D9BEEEC6FFB1FFE1FE0362D0FDBAFC29" box="[463,547,970,996]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="27" pageNumber="478">NSMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BF107603FFB1FFE1FF0B62F5FE23FB9D" blockId="27.[151,1437,861,2040]" pageId="27" pageNumber="478">
<typeStatus id="6014C8A1FFB1FFE1FF0B62F5FEADFBC4" box="[199,308,1007,1033]" pageId="27" pageNumber="478" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="A9A9BD8AFFB1FFE1FE8F62F4FE4DFBC4" box="[323,468,1006,1033]" pageId="27" pageNumber="478" type="generic">4 specimens</specimenCount>
):
<specimenCount id="A9A9BD8AFFB1FFE1FE2962F4FDBDFBC4" box="[485,548,1006,1033]" pageId="27" pageNumber="478" type="male">2 ♂♂</specimenCount>
[one specimen dissected],
<specimenCount id="A9A9BD8AFFB1FFE1FC9D62F4FC08FBC4" box="[849,913,1006,1033]" pageId="27" pageNumber="478" type="female">2 ♀♀</specimenCount>
:
<materialsCitation id="0FC77C5EFFB1FFE1FC5262F5FE37FBE1" ID-GBIF-Occurrence="2429182957" collectionCode="NSMT" collectorName="Shavrin A. V. &amp; Smetana A." country="China" pageId="27" pageNumber="478" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">same data as the holotype, with additional red rectangular printed label</materialsCitation>
: ‘
<materialsCitation id="0FC77C5EFFB1FFE1FE706508FE2FFB9D" ID-GBIF-Occurrence="2429182995" collectionCode="NSMT, CS" collectorName="Shavrin A. V. &amp; Smetana A." country="China" pageId="27" pageNumber="478" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="6014C8A1FFB1FFE1FE706508FDD5FBE1" box="[444,588,1042,1068]" pageId="27" pageNumber="478" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="78AF0D80FFB1FFE1FD936508FD46FBE1" authorityName="Leach" authorityYear="1819" box="[607,735,1042,1068]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="genus">Anthobium</taxonomicName>
| transversale
<emphasis id="8DDBAA11FFB1FFE1FCB36508FC2FFBE1" bold="true" box="[895,950,1042,1068]" pageId="27" pageNumber="478">
<taxonomicNameLabel id="96E8176AFFB1FFE1FCB36508FC2FFBE1" box="[895,950,1042,1068]" pageId="27" pageNumber="478" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="125A13D5FFB1FFE1FC076508FBC2FBE1" box="[971,1115,1042,1068]" pageId="27" pageNumber="478">Shavrin A.V.</collectorName>
&amp;
<collectorName id="125A13D5FFB1FFE1FBB26508FB66FBE1" box="[1150,1279,1042,1068]" pageId="27" pageNumber="478">Smetana A.</collectorName>
2018’ (
<specimenCount id="A9A9BD8AFFB1FFE1FA946508FA1BFBE0" box="[1368,1410,1042,1069]" pageId="27" pageNumber="478" type="male">1 ♂</specimenCount>
,
<specimenCount id="A9A9BD8AFFB1FFE1FA426508FF37FB9C" pageId="27" pageNumber="478" type="female">1 ♀</specimenCount>
:
<collectionCode id="D9BEEEC6FFB1FFE1FF77652CFE92FB9D" box="[187,267,1078,1104]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="27" pageNumber="478">NSMT</collectionCode>
;
<specimenCount id="A9A9BD8AFFB1FFE1FEDA652CFEDBFB9C" box="[278,322,1078,1105]" pageId="27" pageNumber="478" type="male">1 ♂</specimenCount>
,
<specimenCount id="A9A9BD8AFFB1FFE1FE82652CFEE3FB9C" box="[334,378,1078,1105]" pageId="27" pageNumber="478" type="female">1 ♀</specimenCount>
:
<collectionCode id="D9BEEEC6FFB1FFE1FE4B652CFE34FB9D" box="[391,429,1078,1104]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="27" pageNumber="478">CS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFB1FFE7FF0B6543FD2BFD4A" lastPageId="29" lastPageNumber="480" pageId="27" pageNumber="478" type="description">
<paragraph id="BF107603FFB1FFE1FF0B6543FBAAFB71" blockId="27.[151,1437,861,2040]" pageId="27" pageNumber="478">
<emphasis id="8DDBAA11FFB1FFE1FF0B6543FEC8FBB9" bold="true" box="[199,337,1113,1140]" pageId="27" pageNumber="478">Description</emphasis>
. Measurements (n=5): HW: 0.75–0.80; HL: 0.42–0.45; AL(
<typeStatus id="6014C8A1FFB1FFE1FBCC6540FBFEFBB8" box="[1024,1127,1114,1141]" pageId="27" pageNumber="478" type="holotype">holotype</typeStatus>
): 1.41; OL: 0.20–0.22; PL: 0.57–0.65;PW:1.17–1.27; ESL:1.20–1.66;EW:1.67–1.71;AW:1.43–1.53; MTbL(
<typeStatus id="6014C8A1FFB1FFE1FBE16564FB0AFB54" box="[1069,1171,1150,1177]" pageId="27" pageNumber="478" type="holotype">holotype</typeStatus>
):0.82;MTrL(
<typeStatus id="6014C8A1FFB1FFE1FAE06564FA08FB54" box="[1324,1425,1150,1177]" pageId="27" pageNumber="478" type="holotype">holotype</typeStatus>
): 0.34 (MTrL 1–4: 0.19; MTrL 5: 0.15); AedL: 0.65; TL: 3.12–3.47(
<typeStatus id="6014C8A1FFB1FFE1FC4D65B8FC7EFB71" box="[897,999,1186,1212]" pageId="27" pageNumber="478" type="holotype">holotype</typeStatus>
: 3.45).
</paragraph>
<paragraph id="BF107603FFB1FFE1FF0B65DCFCFAF9E8" blockId="27.[151,1437,861,2040]" pageId="27" pageNumber="478">
Head, pronotum, abdomen and antennomeres 4–11 reddish-brown to brown; lateral and basal portions of pronotum and elytra yellow-brown (one
<typeStatus id="6014C8A1FFB1FFE1FDF065F1FD05FAC8" box="[572,668,1259,1285]" pageId="27" pageNumber="478" type="paratype">paratype</typeStatus>
with brown elytra); mouthparts, antennomeres 1–3 and legs yellow. Forebody without microsculpture except for posterior portion of infraorbital ridges and lateral portions of neck with distinct transverse meshes; abdomen with distinct isodiametric microsculpture. Head with fine and deep punctation, markedly denser and deeper on infraorbital ridges; neck with fine and sparse punctation; pronotum with fine, moderately deep punctation, finer and sparser in middle and larger and deeper on lateral portions (some
<typeStatus id="6014C8A1FFB1FFE1FB346461FAFAFA58" box="[1272,1379,1403,1429]" pageId="27" pageNumber="478" type="paratype">paratypes</typeStatus>
with denser and deeper punctation on medioapical portion), usually with small impunctated area on mediobasal third; scutellum with several fine punctures; punctation of elytra as that in pronotum, finer and denser in prescutellar area, each elytron bearing indistinct six to seven vague and tangled longitudinal rows of punctures; abdominal tergites with indistinct, fine and dense punctation, Habitus as in
<figureCitation id="27946A86FFB1FFE1FCC06710FCC6F9E8" box="[780,863,1546,1573]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1834,1859]" captionTargetBox="[165,1424,451,1805]" captionTargetId="figure@20.[157,1431,445,1815]" captionTargetPageId="20" captionText="FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="27" pageNumber="478">Fig. 77</figureCitation>
.
</paragraph>
<paragraph id="BF107603FFB1FFE1FF0B6734FDCEF8ED" blockId="27.[151,1437,861,2040]" pageId="27" pageNumber="478">
Head 1.7 times as wide as long; middle portion slightly elevated, with wide triangular or semicircular impression at level of posterior third of eyes between narrow and deep grooves in front of ocelli, almost reaching level of midlength of eyes; surface between punctures on infraorbital ridges slightly elevated; postocular ridges acute; anterior portion between antennal insertion and anterior nargin of eye with distinct, wide, semicircular notch. Distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Length × width of antennomeres (
<typeStatus id="6014C8A1FFB1FFE1FE8E67F8FE32F930" box="[322,427,1762,1789]" pageId="27" pageNumber="478" type="holotype">holotype</typeStatus>
): 1: 0.16 × 0.08; 2: 0.12 × 0.06; 3: 0.13 × 0.05; 4–5: 0.12 × 0.05; 6–8: 0.12 × 0.06; 9: 0.12 × 0.07; 10: 0.11 × 0.07; 11: 0.17 × 0.08.
</paragraph>
<paragraph id="BF107603FFB1FFE1FF0B6631FD08F835" blockId="27.[151,1437,861,2040]" pageId="27" pageNumber="478">
Pronotum about or twice as wide as long, 1.5 times as wide as head; apical margin rounded, slightly protruded anteriad; posterior angles obtuse, with indistinctly emarginated posteriolateral margins; edges of lateral margins of pronotum with irregular, small crenulation; middle portion with wide and moderately deep, transverse, irregular, mediobasal impressions (one
<typeStatus id="6014C8A1FFB1FFE1FE2E668DFDDBF87C" box="[482,578,1943,1969]" pageId="27" pageNumber="478" type="paratype">paratype</typeStatus>
with additional longitudinal impression, connecting medioapical and mediobasal impressions); surface between punctures in medioapical portion with transverse irregular elevations; lateral portions moderately wide, slightly explanate.
</paragraph>
<caption id="EBD0268BFFB6FFE6FF5B66AFFC95F83F" ID-DOI="http://doi.org/10.5281/zenodo.3517634" ID-Zenodo-Dep="3517634" httpUri="https://zenodo.org/record/3517634/files/figure.png" pageId="28" pageNumber="479" startId="28.[151,264,1973,1998]" targetBox="[288,1302,176,1938]" targetPageId="28">
<paragraph id="BF107603FFB6FFE6FF5B66AFFC95F83F" blockId="28.[151,1436,1973,2034]" pageId="28" pageNumber="479">
<emphasis id="8DDBAA11FFB6FFE6FF5B66AFFECFF803" bold="true" box="[151,342,1973,1998]" pageId="28" pageNumber="479">FIGURES 83–86.</emphasis>
Aedeagus of
<taxonomicName id="78AF0D80FFB6FFE6FE2B66ACFD4FF803" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[487,726,1974,1998]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="deplanatum">
<emphasis id="8DDBAA11FFB6FFE6FE2B66ACFD4FF803" box="[487,726,1974,1998]" italics="true" pageId="28" pageNumber="479">Anthobium deplanatum</emphasis>
</taxonomicName>
(83–84: paratype) and
<taxonomicName id="78AF0D80FFB6FFE6FC0766ACFB34F803" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[971,1197,1974,1998]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="gaoligongshanense">
<emphasis id="8DDBAA11FFB6FFE6FC0766ACFB34F803" box="[971,1197,1974,1998]" italics="true" pageId="28" pageNumber="479">A. gaoligongshanense</emphasis>
</taxonomicName>
(85–86: holotype): 83, 85—parameral view, 84, 86—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="BF107603FFB7FFE7FF0B6183FE90FED2" blockId="29.[151,1437,153,863]" pageId="29" pageNumber="480">Elytra wider than long, more than twice as long as pronotum, distinctly widened posteriad, reaching middle or apical margin of abdominal tergite V, with slightly rounded posterior margins truncated at suture; lateral portions moderately wide, slightly explanate; surface of each elytron with very indistinct four to five longitudinal elevations in middle.</paragraph>
<paragraph id="BF107603FFB7FFE7FF0B6033FCC6FE62" blockId="29.[151,1437,153,863]" pageId="29" pageNumber="480">
Male. Protarsomeres 1–4 slightly slightly widened. Apical margins of abdominal tergite VIII slightly and sternite VIII widely emarginated. Aedeagus (
<figureCitation id="27946A86FFB7FFE7FDA16057FD59FEAA" box="[621,704,333,359]" captionStart="FIGURES 88–89" captionStartId="30.[151,264,1425,1450]" captionTargetBox="[163,1424,179,1399]" captionTargetId="figure@30.[151,1436,174,1406]" captionTargetPageId="30" captionText="FIGURES 88–89. Aedeagus of Anthobium transversale (paratype): 88—parameral view, 89—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517638" httpUri="https://zenodo.org/record/3517638/files/figure.png" pageId="29" pageNumber="480">Fig. 88</figureCitation>
) with wide median lobe, truncated apically; parameres long and narrow, slightly exceeding apex of median lobe, with two apical and two preapical setae; internal sac long and wide, spirally rolled in basal portion. Aedeagus laterally as in
<figureCitation id="27946A86FFB7FFE7FCC4608FFCC2FE62" box="[776,859,405,431]" captionStart="FIGURES 88–89" captionStartId="30.[151,264,1425,1450]" captionTargetBox="[163,1424,179,1399]" captionTargetId="figure@30.[151,1436,174,1406]" captionTargetPageId="30" captionText="FIGURES 88–89. Aedeagus of Anthobium transversale (paratype): 88—parameral view, 89—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517638" httpUri="https://zenodo.org/record/3517638/files/figure.png" pageId="29" pageNumber="480">Fig. 89</figureCitation>
.
</paragraph>
<paragraph id="BF107603FFB7FFE7FF0B60A3FCB8FE3A" blockId="29.[151,1437,153,863]" pageId="29" pageNumber="480">Female. Protarsomeres 1–4 moderately narrow. Apical margin of abdominal tergite VIII somewhat straight. Apical margin of abdominal sternite VIII widely rounded.</paragraph>
<paragraph id="BF107603FFB7FFE7FF0B631AFD2BFD4A" blockId="29.[151,1437,153,863]" pageId="29" pageNumber="480">
<emphasis id="8DDBAA11FFB7FFE7FF0B631AFE28FDD6" bold="true" box="[199,433,512,539]" pageId="29" pageNumber="480">Comparative notes.</emphasis>
Based on the coloration, body length, and shape of the forebody,
<taxonomicName id="78AF0D80FFB7FFE7FB54631BFADBFDD6" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[1176,1346,513,539]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="transversale" status="sp. nov.">
<emphasis id="8DDBAA11FFB7FFE7FB54631BFADBFDD6" box="[1176,1346,513,539]" italics="true" pageId="29" pageNumber="480">A. transversale</emphasis>
</taxonomicName>
<emphasis id="8DDBAA11FFB7FFE7FA86631BFA1BFDD6" bold="true" box="[1354,1410,513,539]" pageId="29" pageNumber="480">
<taxonomicNameLabel id="96E8176AFFB7FFE7FA86631BFA1BFDD6" box="[1354,1410,513,539]" pageId="29" pageNumber="480" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to the Himalayan
<taxonomicName id="78AF0D80FFB7FFE7FE7F633FFC96FDF2" authority="(Cameron, 1941)" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[435,783,549,575]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="8DDBAA11FFB7FFE7FE7F633FFE5DFDF2" box="[435,452,549,575]" italics="true" pageId="29" pageNumber="480">A</emphasis>
.
<emphasis id="8DDBAA11FFB7FFE7FE1E633FFDDEFDF2" box="[466,583,549,575]" italics="true" pageId="29" pageNumber="480">altivagans</emphasis>
(
<bibRefCitation id="DB3E0BF2FFB7FFE7FD99633FFC9EFDF2" author="Cameron, M." box="[597,775,549,575]" pageId="29" pageNumber="480" pagination="56 - 60" refId="ref19201" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). The Proceedings of the Royal Entomological Society of London (B), 10, 56 - 60. https: // doi. org / 10.1111 / j. 1365 - 3113.1941. tb 00693. x" type="journal article" year="1941">Cameron, 1941</bibRefCitation>
)
</taxonomicName>
, from which it differs by longer antennomeres 5–10, more protruded apical angles of the pronotum, more irregular crenulation of the lateral edges of the pronotum, and different shape and internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFB7FFE7FF0B638AFC42FCDA" pageId="29" pageNumber="480" type="distribution">
<paragraph id="BF107603FFB7FFE7FF0B638AFAB5FD66" blockId="29.[151,1437,153,863]" box="[199,1324,656,683]" pageId="29" pageNumber="480">
<emphasis id="8DDBAA11FFB7FFE7FF0B638AFEF8FD66" bold="true" box="[199,353,656,683]" pageId="29" pageNumber="480">Distribution.</emphasis>
The species is known from two locations in Gongga Shan in
<collectingRegion id="7D6BB8E1FFB7FFE7FBDD638BFBF7FD66" box="[1041,1134,657,683]" country="China" name="Sichuan" pageId="29" pageNumber="480">Sichuan</collectingRegion>
,
<collectingCountry id="C7B83693FFB7FFE7FBB5638BFB25FD66" box="[1145,1212,657,683]" name="China" pageId="29" pageNumber="480">China</collectingCountry>
(
<figureCitation id="27946A86FFB7FFE7FB07638BFA86FD66" box="[1227,1311,657,683]" captionStart="FIGURE 87" captionStartId="29.[151,250,1614,1639]" captionTargetBox="[151,1435,913,1541]" captionTargetId="figure@29.[151,1435,913,1541]" captionTargetPageId="29" captionText="FIGURE 87. Distribution of Anthobium deplanatum (black circles), A. gaoligongshanense (black square) and A. transversale (black triangle) in the south-western China." figureDoi="http://doi.org/10.5281/zenodo.3517636" httpUri="https://zenodo.org/record/3517636/files/figure.png" pageId="29" pageNumber="480">Fig. 87</figureCitation>
).
</paragraph>
<paragraph id="BF107603FFB7FFE7FF0B63AEFC42FCDA" blockId="29.[151,1437,153,863]" pageId="29" pageNumber="480">
<emphasis id="8DDBAA11FFB7FFE7FF0B63AEFED2FD02" bold="true" box="[199,331,692,719]" pageId="29" pageNumber="480">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="7857DBE6FFB7FFE7FC9363AFFB8EFD02" box="[863,1047,693,719]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="29" pageNumber="480" unit="m" value="2950.0" valueMax="3000.0" valueMin="2900.0">2900 to 3000 m</quantity>
a.s.l. The specimens from Gonga Shan range (locality: C53) were taken by sifting various debris, leaf litter and moss accumulated on sandy flats of a creek in the primary
<taxonomicName id="78AF0D80FFB7FFE7FE4C63E7FE27FCDA" box="[384,446,765,791]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="29" pageNumber="480" phylum="Tracheophyta" rank="genus">
<emphasis id="8DDBAA11FFB7FFE7FE4C63E7FE27FCDA" box="[384,446,765,791]" italics="true" pageId="29" pageNumber="480">Abies</emphasis>
</taxonomicName>
forest with rich undergrowth of rhododendrons.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFB7FFE7FF0B623BFDBDFC92" pageId="29" pageNumber="480" type="etymology">
<paragraph id="BF107603FFB7FFE7FF0B623BFDBDFC92" blockId="29.[151,1437,153,863]" pageId="29" pageNumber="480">
<emphasis id="8DDBAA11FFB7FFE7FF0B623BFED7FCF6" bold="true" box="[199,334,801,827]" pageId="29" pageNumber="480">Etymology.</emphasis>
The specific epithet is the Latin adjective
<emphasis id="8DDBAA11FFB7FFE7FCEC623BFC28FCF6" box="[800,945,801,827]" italics="true" pageId="29" pageNumber="480">transversalis</emphasis>
, -
<emphasis id="8DDBAA11FFB7FFE7FC0A6238FC40FCF6" box="[966,985,802,827]" italics="true" pageId="29" pageNumber="480">is</emphasis>
, -
<emphasis id="8DDBAA11FFB7FFE7FC236238FC62FCF6" box="[1007,1019,802,827]" italics="true" pageId="29" pageNumber="480">e</emphasis>
(transverse). It refers to the markedly transverse pronotum of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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