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<document id="E97D1BEB6C8CFB2104959278D6BA49EB" ID-CLB-Dataset="25154" ID-DOI="10.11646/zootaxa.4688.4.1" ID-GBIF-Dataset="c4072224-34c3-4b12-9415-a59d7c9484b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3517608" ID-ZooBank="BABABF8C-30B3-45D0-89B5-6F4DC1B19B70" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571896092680" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2019" docId="3706C715FFA6FFF4FF5B679BFC97FA9A" docLanguage="en" docName="zootaxa.4688.4.1.pdf" docOrigin="Zootaxa 4688 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium crenulatum Shavrin &amp; Smetana 2019, sp.n." docType="treatment" docVersion="8" lastPageNumber="465" masterDocId="CB3FBF6DFFAAFFFAFFCC611AFF99FFCD" masterDocTitle="A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group" masterLastPageNumber="482" masterPageNumber="451" pageNumber="463" updateTime="1698760874617" updateUser="ExternalLinkService">
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<mods:title id="7EBCFAD384AC6F75F7799ABE5831F385">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group</mods:title>
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<mods:name id="0B2BA95386A1DFC7854D38A8B9240969" type="personal">
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<mods:namePart id="19D23F95717655C8A74B25708AC86B4F">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="595F002056BBD0AE56E9179A8848250C">Smetana, Aleš</mods:namePart>
</mods:name>
<mods:typeOfResource id="11566ADE79FC19C31E76D5C968E20C10">text</mods:typeOfResource>
<mods:relatedItem id="91566825450F2580B6370A8669D2F6F4" type="host">
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<mods:title id="1B8E433B3119DE2D6ADEB922985BB573">Zootaxa</mods:title>
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<mods:part id="B53062BD0EAA26D084D85BA632C7A71B">
<mods:date id="A81BCC9F01DEB39FB0D3A81A5DD1BE8C">2019</mods:date>
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<mods:number id="3908C6F6374B0DCDB217A554164B64D5">2019-10-24</mods:number>
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<mods:number id="F76B42BF4980CD8DA28D7ADDF4DA70E4">4688</mods:number>
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<mods:classification id="E621F5F6291D19F438E86792DB5CB55A">journal article</mods:classification>
<mods:identifier id="41A945B19DE1B4EE05F2EBC41F94F1C3" type="CLB-Dataset">25154</mods:identifier>
<mods:identifier id="86F1758EE4AE4E0CA93D1C8554AF23FE" type="DOI">10.11646/zootaxa.4688.4.1</mods:identifier>
<mods:identifier id="B46DE31B5F6FEAE9B6203D31517BF14E" type="GBIF-Dataset">c4072224-34c3-4b12-9415-a59d7c9484b6</mods:identifier>
<mods:identifier id="2ABAB2B110F81DD0D21F0238F11EB2F0" type="ISSN">1175-5326</mods:identifier>
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<treatment id="3706C715FFA6FFF4FF5B679BFC97FA9A" ID-GBIF-Taxon="159834458" LSID="urn:lsid:plazi:treatment:3706C715FFA6FFF4FF5B679BFC97FA9A" httpUri="http://treatment.plazi.org/id/3706C715FFA6FFF4FF5B679BFC97FA9A" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463">
<subSubSection id="F7B52588FFA6FFF6FF5B679BFD65F956" box="[151,764,1665,1692]" pageId="12" pageNumber="463" type="nomenclature">
<paragraph id="BF107603FFA6FFF6FF5B679BFD65F956" blockId="12.[151,764,1665,1728]" box="[151,764,1665,1692]" pageId="12" pageNumber="463">
<heading id="E458C16FFFA6FFF6FF5B679BFD65F956" bold="true" box="[151,764,1665,1692]" fontSize="11" level="1" pageId="12" pageNumber="463" reason="1">
<emphasis id="8DDBAA11FFA6FFF6FF5B679BFD65F956" bold="true" box="[151,764,1665,1692]" pageId="12" pageNumber="463">
<taxonomicName id="78AF0D80FFA6FFF6FF5B679BFD2CF956" ID-CoL="5VCD7" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[151,693,1665,1692]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="crenulatum" status="sp. nov.">
<emphasis id="8DDBAA11FFA6FFF6FF5B679BFE2AF951" bold="true" box="[151,435,1665,1692]" italics="true" pageId="12" pageNumber="463">Anthobium crenulatum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="96E8176AFFA6FFF6FD0D679BFD65F956" box="[705,764,1665,1691]" pageId="12" pageNumber="463" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFA6FFF6FF5B67BFFE1BF90D" box="[151,386,1700,1728]" pageId="12" pageNumber="463" type="description">
<paragraph id="BF107603FFA6FFF6FF5B67BFFE1BF90D" blockId="12.[151,764,1665,1728]" box="[151,386,1700,1728]" pageId="12" pageNumber="463">
(
<figureCitation id="27946A86FFA6FFF6FF5367BFFF64F972" box="[159,253,1700,1727]" captionStart="FIGURES 56–59" captionStartId="11.[151,264,1782,1807]" captionTargetBox="[158,1424,347,1754]" captionTargetId="figure@11.[151,1436,337,1762]" captionTargetPageId="11" captionText="FIGURES 56–59. Habitus of species of Anthobium crassum-group: 56—A. crassum (holotype), 57—A. crenulatum (holotype), 58—A. explanatum (holotype), 59—A. farkaci (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517618" httpUri="https://zenodo.org/record/3517618/files/figure.png" pageId="12" pageNumber="463">Figs. 57</figureCitation>
,
<figureCitation id="27946A86FFA6FFF6FEC567BFFECBF972" box="[265,338,1701,1727]" captionStart-0="FIGURES 60–63" captionStart-1="FIGURE 64" captionStartId-0="13.[151,264,1973,1998]" captionStartId-1="15.[151,250,1958,1983]" captionTargetBox-0="[293,1300,179,1943]" captionTargetBox-1="[151,1437,1273,1938]" captionTargetId-0="figure@13.[275,1312,166,1954]" captionTargetId-1="figure@15.[151,1437,1273,1938]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionText-0="FIGURES 60–63. Aedeagus of Anthobium crassum (60–61: paratype) and A. crenulatum (62–63: holotype): 60, 62—parameral view, 61, 63—lateral view. Scale bar: 0.1 mm." captionText-1="FIGURE 64. Distribution of Anthobium crassum (black circles), A. crenulatum (black square), A. explanatum (black rhomb), and A. farkaci (black triangle) in the south-western China." figureDoi-0="http://doi.org/10.5281/zenodo.3517620" figureDoi-1="http://doi.org/10.5281/zenodo.3517622" httpUri-0="https://zenodo.org/record/3517620/files/figure.png" httpUri-1="https://zenodo.org/record/3517622/files/figure.png" pageId="12" pageNumber="463">62–64</figureCitation>
,
<figureCitation id="27946A86FFA6FFF6FE9167BFFEE3F90D" box="[349,378,1701,1728]" captionStart="FIGURES 65–68" captionStartId="16.[151,264,1727,1752]" captionTargetBox="[158,1412,486,1697]" captionTargetId="figure@16.[151,1420,481,1708]" captionTargetPageId="16" captionText="FIGURES 65–68. Type locality of Anthobium crassum (65–66), A. crenulatum (67) and A. gaoligongshanense (68) in Yunnan, China (photograph: M. Schülke)." figureDoi="http://doi.org/10.5281/zenodo.3517624" httpUri="https://zenodo.org/record/3517624/files/figure.png" pageId="12" pageNumber="463">67</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFA6FFF6FF5B67F7FD0FF8BE" pageId="12" pageNumber="463" type="materials_examined">
<paragraph id="BF107603FFA6FFF6FF5B67F7FD0FF8BE" blockId="12.[151,1437,1772,1979]" pageId="12" pageNumber="463">
<emphasis id="8DDBAA11FFA6FFF6FF5B67F7FE26F8CA" bold="true" box="[151,447,1772,1799]" pageId="12" pageNumber="463">Type material examined:</emphasis>
<materialsCitation id="0FC77C5EFFA6FFF6FE0967F7FC03F8CA" ID-GBIF-Occurrence="2429182984" box="[453,922,1772,1799]" country="China" location="Yunnan" pageId="12" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="6014C8A1FFA6FFF6FE0967F7FDB4F8CA" box="[453,557,1773,1799]" pageId="12" pageNumber="463" type="holotype">Holotype</typeStatus>
<specimenCount id="A9A9BD8AFFA6FFF6FDF867F7FDD0F8CA" box="[564,585,1773,1799]" pageId="12" pageNumber="463" type="male">♂</specimenCount>
[dissected]: ‘
<collectingCountry id="C7B83693FFA6FFF6FD1667F7FCA5F8CA" box="[730,828,1773,1799]" name="China" pageId="12" pageNumber="463">CHINA</collectingCountry>
:
<collectingRegion id="7D6BB8E1FFA6FFF6FC8867F7FC03F8CA" box="[836,922,1773,1799]" country="China" name="Yunnan" pageId="12" pageNumber="463">Yunnan</collectingRegion>
</materialsCitation>
<materialsCitation id="0FC77C5EFFA6FFF6FC6C67F6FDB6F882" ID-GBIF-Occurrence="2429182968" collectingDate="2007-05-28" collectorName="Diacang Shan W &amp; M. Schulke'" country="Switzerland" elevation="3160" latitude="25.688889" location="Dali" longLatPrecision="21" longitude="100.10333" pageId="12" pageNumber="463" specimenCount="1" typeStatus="holotype">
[
<collectingCountry id="C7B83693FFA6FFF6FC6B67F6FC48F8CA" box="[935,977,1772,1799]" name="Switzerland" pageId="12" pageNumber="463">CH</collectingCountry>
07-02A], | Dali Bai Auton. Pref.,
<collectorName id="125A13D5FFA6FFF6FAF367F7FF6BF8E6" pageId="12" pageNumber="463">Diacang Shan W</collectorName>
|
<location id="BA7020D8FFA6FFF6FEC6660BFEA4F8E6" LSID="urn:lsid:plazi:treatment:3706C715FFA6FFF4FF5B679BFC97FA9A:BA7020D8FFA6FFF6FEC6660BFEA4F8E6" box="[266,317,1809,1835]" country="Switzerland" latitude="25.688889" longLatPrecision="21" longitude="100.10333" name="Dali" pageId="12" pageNumber="463">Dali</location>
,
<geoCoordinate id="DA9B10C4FFA6FFF6FE86660AFE43F8E6" box="[330,474,1808,1835]" degrees="25" direction="north" minutes="41" orientation="latitude" pageId="12" pageNumber="463" precision="15" seconds="20" value="25.688889">25°41’20’’N</geoCoordinate>
,
<geoCoordinate id="DA9B10C4FFA6FFF6FE24660BFD18F8E6" box="[488,641,1809,1835]" degrees="100" direction="east" minutes="06" orientation="longitude" pageId="12" pageNumber="463" precision="15" seconds="12" value="100.10333">100°06’12’’E</geoCoordinate>
|
<quantity id="7857DBE6FFA6FFF6FD54660BFD6BF8E6" box="[664,754,1809,1835]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="12" pageNumber="463" unit="m" value="3160.0">
<elevation id="34829130FFA6FFF6FD54660BFD6BF8E6" box="[664,754,1809,1835]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="12" pageNumber="463" unit="m" value="3160.0">3160 m</elevation>
</quantity>
, small creek valley, litter and | debris sifted,
<date id="CB1150C3FFA6FFF6FB35660BFAF7F8E6" box="[1273,1390,1809,1835]" pageId="12" pageNumber="463" value="2007-05-28">
<collectingDate id="DB55A92BFFA6FFF6FB35660BFAF7F8E6" box="[1273,1390,1809,1835]" pageId="12" pageNumber="463" value="2007-05-28">28.V.2007</collectingDate>
</date>
,
<collectorName id="125A13D5FFA6FFF6FAB0660BFF61F882" pageId="12" pageNumber="463">M. Schülke’</collectorName>
&lt;rectangular label, printed
</materialsCitation>
&gt;, ‘
<materialsCitation id="0FC77C5EFFA6FFF6FD82662FFD0BF8BE" ID-GBIF-Occurrence="2429182951" collectionCode="CSC" collectorName="Shavrin A. V. &amp; Smetana A." country="Switzerland" pageId="12" pageNumber="463" specimenCount="1" typeStatus="holotype">
<typeStatus id="6014C8A1FFA6FFF6FD82662FFD7EF882" box="[590,743,1845,1871]" pageId="12" pageNumber="463" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="78AF0D80FFA6FFF6FD37662FFCE2F882" authorityName="Leach" authorityYear="1819" box="[763,891,1845,1871]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">Anthobium</taxonomicName>
| crenulatum
<emphasis id="8DDBAA11FFA6FFF6FBDA662FFBD7F882" bold="true" box="[1046,1102,1845,1871]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="96E8176AFFA6FFF6FBDA662FFBD7F882" box="[1046,1102,1845,1871]" pageId="12" pageNumber="463" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="125A13D5FFA6FFF6FBA8662FFB6CF882" box="[1124,1269,1845,1871]" pageId="12" pageNumber="463">Shavrin A.V.</collectorName>
&amp;
<collectorName id="125A13D5FFA6FFF6FAD6662FFA05F882" box="[1306,1436,1845,1871]" pageId="12" pageNumber="463">Smetana A.</collectorName>
2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="D9BEEEC6FFA6FFF6FD9C6643FD13F8BE" box="[592,650,1881,1907]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="12" pageNumber="463" type="Herbarium">CSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFA6FFF4FF0B6666FB85FBB2" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463" type="description">
<paragraph id="BF107603FFA6FFF6FF0B6666FA02F876" blockId="12.[151,1437,1772,1979]" pageId="12" pageNumber="463">
<emphasis id="8DDBAA11FFA6FFF6FF0B6666FEC8F85A" bold="true" box="[199,337,1916,1943]" pageId="12" pageNumber="463">Description</emphasis>
. Measurements of the
<typeStatus id="6014C8A1FFA6FFF6FD966667FD59F85A" box="[602,704,1917,1943]" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
: HW: 0.77; HL: 0.45; AL: 1.49; OL: 0.18; PL: 0.57; PW: 1.27; ESL: 1.52; EW: 1.55; AW: 1.50; MTbL: 0.80; MTrL: 0.35 (MTrL 1–4: 0.20; MTrL 5: 0.15); AedL: 0.72; TL: 3.40.
</paragraph>
<caption id="EBD0268BFFA7FFF7FF5B66AFFDF7F83F" ID-DOI="http://doi.org/10.5281/zenodo.3517620" ID-Zenodo-Dep="3517620" httpUri="https://zenodo.org/record/3517620/files/figure.png" pageId="13" pageNumber="464" startId="13.[151,264,1973,1998]" targetBox="[293,1300,179,1943]" targetPageId="13">
<paragraph id="BF107603FFA7FFF7FF5B66AFFDF7F83F" blockId="13.[151,1436,1973,2034]" pageId="13" pageNumber="464">
<emphasis id="8DDBAA11FFA7FFF7FF5B66AFFECAF803" bold="true" box="[151,339,1973,1998]" pageId="13" pageNumber="464">FIGURES 60–63.</emphasis>
Aedeagus of
<taxonomicName id="78AF0D80FFA7FFF7FE1666ACFD3AF803" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[474,675,1974,1998]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="crassum">
<emphasis id="8DDBAA11FFA7FFF7FE1666ACFD3AF803" box="[474,675,1974,1998]" italics="true" pageId="13" pageNumber="464">Anthobium crassum</emphasis>
</taxonomicName>
(60–61: paratype) and
<taxonomicName id="78AF0D80FFA7FFF7FC4766ACFB81F803" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[907,1048,1974,1998]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="crenulatum">
<emphasis id="8DDBAA11FFA7FFF7FC4766ACFB81F803" box="[907,1048,1974,1998]" italics="true" pageId="13" pageNumber="464">A. crenulatum</emphasis>
</taxonomicName>
(62–63: holotype): 60, 62—parameral view, 61, 63—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="BF107603FFA4FFF4FF0B6183FB86FE46" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">
Body and antennomeres 6–11 reddish-brown, with mouthparts, antennomeres 1–5, lateral portions of pronotum, legs and apical tergites of abdomen yellow-brown. Head with irregular fine punctation, denser between grooves and infraorbital ridges, with narrow transverse impunctated portion in middle; neck with fine irregular punctures; pronotum with very dense punctation, markedly larger and deeper than that on head, with merging punctures on medioapical third, slightly sparser on lateral portions, with wide impunctated area after the middle of disc; punctation of elytra as that on pronotum, but slightly larger, somewhat finer on parascutellar portion, each elytron forming six very vague and tangled longitudinal rows of punctures. Habitus as in
<figureCitation id="27946A86FFA4FFF4FC05606BFB82FE46" box="[969,1051,368,395]" captionStart="FIGURES 56–59" captionStartId="11.[151,264,1782,1807]" captionTargetBox="[158,1424,347,1754]" captionTargetId="figure@11.[151,1436,337,1762]" captionTargetPageId="11" captionText="FIGURES 56–59. Habitus of species of Anthobium crassum-group: 56—A. crassum (holotype), 57—A. crenulatum (holotype), 58—A. explanatum (holotype), 59—A. farkaci (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517618" httpUri="https://zenodo.org/record/3517618/files/figure.png" pageId="14" pageNumber="465">Fig. 57</figureCitation>
.
</paragraph>
<paragraph id="BF107603FFA4FFF4FF0B608FFB86FD4A" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">Middle portion of head and infraorbital ridges slightly elevated, with indistinct semicircular impression between eyes at posterior third, with indistinct short grooves in front of ocelli, reaching posterior margin of eye; surface between punctures in middle transversely and in infraorbital ridges irregularly, diagonally elevated; postocular ridge smooth; anterior portion between antennal insertion and anterior margin of eye with distinct, wide semicircular notch. Ocelli situated at level slightly behind of postocular ridges; distance between ocelli about as distance between ocellus and posterior margin of eye. Length × width of antennomeres: 1: 0.21 × 0.07; 2: 0.12 × 0.06; 3: 0.13 × 0.07; 4: 0.12 × 0.07; 5–8: 0.13 × 0.07; 9: 0.12 × 0.07; 10: 0.11 × 0.07; 11: 0.16 × 0.07.</paragraph>
<paragraph id="BF107603FFA4FFF4FF0B638BFA0CFD3E" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">
Pronotum distinctly more than twice as wide as long, 1.6 times as wide as head, widest in middle; lateral margins with four distinct, irregular projections similar to that in
<taxonomicName id="78AF0D80FFA4FFF4FCF363AFFC22FD02" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[831,955,693,719]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="crassum" status="sp. nov.">
<emphasis id="8DDBAA11FFA4FFF4FCF363AFFC22FD02" box="[831,955,693,719]" italics="true" pageId="14" pageNumber="465">A. crassum</emphasis>
</taxonomicName>
<emphasis id="8DDBAA11FFA4FFF4FC0D63AFFC60FD02" bold="true" box="[961,1017,693,719]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="96E8176AFFA4FFF4FC0D63AFFC60FD02" box="[961,1017,693,719]" pageId="14" pageNumber="465" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; middle portion with indistinct longitudinal and mediobasal portion with deep semicircular impressions; lateral portions wide and markedly explanate.
</paragraph>
<paragraph id="BF107603FFA4FFF4FF0B63E7FC87FC92" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">Elytra subparallel-sided, long, longer than combined wide, reaching basal margin of abdominal tergite V; surface of each elytron with diagonal distinct elevations between punctures on parascutellar portion and with indistinct longitudinal impression on mediobasal third along suture.</paragraph>
<paragraph id="BF107603FFA4FFF4FF0B6273FE15FC22" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">
Male. Apical margins of abdominal tergite VIII and sternite VIII slightly emarginated. Aedeagus (
<figureCitation id="27946A86FFA4FFF4FAC66273FAC2FC49" box="[1290,1371,873,900]" captionStart="FIGURES 60–63" captionStartId="13.[151,264,1973,1998]" captionTargetBox="[293,1300,179,1943]" captionTargetId="figure@13.[275,1312,166,1954]" captionTargetPageId="13" captionText="FIGURES 60–63. Aedeagus of Anthobium crassum (60–61: paratype) and A. crenulatum (62–63: holotype): 60, 62—parameral view, 61, 63—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517620" httpUri="https://zenodo.org/record/3517620/files/figure.png" pageId="14" pageNumber="465">Fig. 62</figureCitation>
) with wide median lobe, gradually narrowed toward truncated apex; pararameres short, widened apically, with two small apical and preapical setae; internal sac very long, narrow in apical and markedly wide in basal portion. Aedeagus laterally as in
<figureCitation id="27946A86FFA4FFF4FEF962CFFE1EFC22" box="[309,391,981,1007]" captionStart="FIGURES 60–63" captionStartId="13.[151,264,1973,1998]" captionTargetBox="[293,1300,179,1943]" captionTargetId="figure@13.[275,1312,166,1954]" captionTargetPageId="13" captionText="FIGURES 60–63. Aedeagus of Anthobium crassum (60–61: paratype) and A. crenulatum (62–63: holotype): 60, 62—parameral view, 61, 63—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517620" httpUri="https://zenodo.org/record/3517620/files/figure.png" pageId="14" pageNumber="465">Fig. 63</figureCitation>
.
</paragraph>
<paragraph id="BF107603FFA4FFF4FF0B62E3FE08FBDE" blockId="14.[151,1437,152,1367]" box="[199,401,1017,1043]" pageId="14" pageNumber="465">Female unknown.</paragraph>
<paragraph id="BF107603FFA4FFF4FF0B6506FB85FBB2" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">
<emphasis id="8DDBAA11FFA4FFF4FF0B6506FE2BFBFA" bold="true" box="[199,434,1052,1079]" pageId="14" pageNumber="465">Comparative notes.</emphasis>
Based on the coloration, punctation, shape of the pronotum and general structure of the aedeagus,
<taxonomicName id="78AF0D80FFA4FFF4FEC2655BFE37FB96" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[270,430,1089,1115]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="crenulatum" status="sp. nov.">
<emphasis id="8DDBAA11FFA4FFF4FEC2655BFE37FB96" box="[270,430,1089,1115]" italics="true" pageId="14" pageNumber="465">A. crenulatum</emphasis>
</taxonomicName>
<emphasis id="8DDBAA11FFA4FFF4FE7B655BFE77FB96" bold="true" box="[439,494,1089,1115]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="96E8176AFFA4FFF4FE7B655BFE77FB96" box="[439,494,1089,1115]" pageId="14" pageNumber="465" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="78AF0D80FFA4FFF4FD45655BFC9EFB96" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[649,775,1089,1115]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="crassum" status="sp. nov.">
<emphasis id="8DDBAA11FFA4FFF4FD45655BFC9EFB96" box="[649,775,1089,1115]" italics="true" pageId="14" pageNumber="465">A. crassum</emphasis>
</taxonomicName>
<emphasis id="8DDBAA11FFA4FFF4FCDD655BFCD1FB96" bold="true" box="[785,840,1089,1115]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="96E8176AFFA4FFF4FCDD655BFCD1FB96" box="[785,840,1089,1115]" pageId="14" pageNumber="465" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by larger body, shorter elytra, and details of the external and internal characters of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFA4FFF4FF0B6592FE56FADD" pageId="14" pageNumber="465" type="distribution">
<paragraph id="BF107603FFA4FFF4FF0B6592FF5DFB05" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">
<emphasis id="8DDBAA11FFA4FFF4FF0B6592FEF8FB6E" bold="true" box="[199,353,1160,1187]" pageId="14" pageNumber="465">Distribution.</emphasis>
The species is known only from the
<typeStatus id="6014C8A1FFA4FFF4FCCD6593FCA8FB6E" box="[769,817,1161,1187]" pageId="14" pageNumber="465">type</typeStatus>
locality in Diacang Shan range in
<collectingRegion id="7D6BB8E1FFA4FFF4FB746593FA88FB6E" box="[1208,1297,1161,1187]" country="China" name="Yunnan" pageId="14" pageNumber="465">Yunnan</collectingRegion>
,
<collectingCountry id="C7B83693FFA4FFF4FAD06593FAC6FB6E" box="[1308,1375,1161,1187]" name="China" pageId="14" pageNumber="465">China</collectingCountry>
(
<figureCitation id="27946A86FFA4FFF4FAA36593FF2EFB05" captionStart="FIGURE 64" captionStartId="15.[151,250,1958,1983]" captionTargetBox="[151,1437,1273,1938]" captionTargetId="figure@15.[151,1437,1273,1938]" captionTargetPageId="15" captionText="FIGURE 64. Distribution of Anthobium crassum (black circles), A. crenulatum (black square), A. explanatum (black rhomb), and A. farkaci (black triangle) in the south-western China." figureDoi="http://doi.org/10.5281/zenodo.3517622" httpUri="https://zenodo.org/record/3517622/files/figure.png" pageId="14" pageNumber="465">Fig. 64</figureCitation>
).
</paragraph>
<paragraph id="BF107603FFA4FFF4FF0B65CAFE56FADD" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">
<emphasis id="8DDBAA11FFA4FFF4FF0B65CAFED2FB26" bold="true" box="[199,331,1232,1259]" pageId="14" pageNumber="465">Bionomics.</emphasis>
The
<typeStatus id="6014C8A1FFA4FFF4FE4A65CBFE71FB26" box="[390,488,1233,1259]" pageId="14" pageNumber="465" type="holotype">holotype</typeStatus>
was collected at elevation
<quantity id="7857DBE6FFA4FFF4FCD165CBFCEAFB26" box="[797,883,1233,1259]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="14" pageNumber="465" unit="m" value="3160.0">3160 m</quantity>
a.s.l. by sifting of litter and debris along a small creek (
<collectingCountry id="C7B83693FFA4FFF4FF2F65EFFE94FAC2" box="[227,269,1269,1295]" name="Switzerland" pageId="14" pageNumber="465">CH</collectingCountry>
07-02A:
<figureCitation id="27946A86FFA4FFF4FEA365EFFE5AFADD" box="[367,451,1269,1296]" captionStart="FIGURES 65–68" captionStartId="16.[151,264,1727,1752]" captionTargetBox="[158,1412,486,1697]" captionTargetId="figure@16.[151,1420,481,1708]" captionTargetPageId="16" captionText="FIGURES 65–68. Type locality of Anthobium crassum (65–66), A. crenulatum (67) and A. gaoligongshanense (68) in Yunnan, China (photograph: M. Schülke)." figureDoi="http://doi.org/10.5281/zenodo.3517624" httpUri="https://zenodo.org/record/3517624/files/figure.png" pageId="14" pageNumber="465">Fig. 67</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7B52588FFA4FFF4FF0B6403FC97FA9A" pageId="14" pageNumber="465" type="etymology">
<paragraph id="BF107603FFA4FFF4FF0B6403FC97FA9A" blockId="14.[151,1437,152,1367]" pageId="14" pageNumber="465">
<emphasis id="8DDBAA11FFA4FFF4FF0B6403FED7FAFE" bold="true" box="[199,334,1305,1331]" pageId="14" pageNumber="465">Etymology.</emphasis>
The specific epithet is the Latin adjective
<emphasis id="8DDBAA11FFA4FFF4FCE26403FC3AFAFE" box="[814,931,1305,1331]" italics="true" pageId="14" pageNumber="465">crenulatus</emphasis>
,-
<emphasis id="8DDBAA11FFA4FFF4FC786400FC62FAFE" box="[948,1019,1305,1331]" italics="true" pageId="14" pageNumber="465">a. -um</emphasis>
(crenulate). It refers to the markedly crenulate lateral margins of the pronotum of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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