treatments-xml/data/2C/A2/9E/2CA29E2FBF6C5E4EAC4002448FB57B55.xml
2024-06-21 12:32:25 +02:00

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<mods:title id="ABAA3C71E613B54FEFDB468E5570F181">Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species</mods:title>
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<taxonomicName id="72D6CC861552EDD0512528CCE1FD1C0C" LSID="2CA29E2F-BF6C-5E4E-AC40-02448FB57B55" authority="(Stoliczka, 1871)" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Stoliczka" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Acinolaemus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Acinolaemus dayanum" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="dayanum" status="comb. nov.">Acinolaemus dayanum (Stoliczka, 1871)</taxonomicName>
<taxonomicNameLabel id="FB5815DBF6F6647608DBFA218DB1E4D5" pageId="0" pageNumber="157">comb. nov.</taxonomicNameLabel>
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<figureCitation id="A80A159ECD74DC0B1A4819C14E7AABAD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A Acinolaemus dayanum, topotype specimen HA Collection from Mawlamyine Township, Mon State B Acinolaemus cryptidentatus, specimen CUMZ 14413 from Taunggyi District, Shan State. The white arrow indicates constriction, the yellow arrow indicates a cervical crest. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure2" httpUri="https://binary.pensoft.net/fig/1002051" pageId="0" pageNumber="157">Figs 2A</figureCitation>
<figureCitation id="1315BA5980D0C72FE3B5B2C8799F40B7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">, 13A</figureCitation>
</paragraph>
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<paragraph id="FAE8A1D4EC9FCC86E0D77655B5732FD8" pageId="0" pageNumber="157">
<taxonomicName id="3A746F687438B3FA3029EDF233B217E1" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma dayanum" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="dayanum" status="comb. nov.">Hypselostoma dayanum</taxonomicName>
Stoliczka, 1871: 172, 173, pl. 7, fig. 2. Type locality: Damotha, prope Moulmein [Kayon Hill, Dhammasa Village, Mawlamyine District, Mon State].
<bibRefCitation id="431BA1BC7764DD0B3A00CA834A2FBD4A" author="Hanley, SC" journalOrPublisher="Taylor and Francis Group, New York" pageId="0" pageNumber="157" publicationUrl="https://www.biodiversitylibrary.org/page/46204402" refId="B27" refString="Hanley, SC, Theobald, W, 1870 1874 1876. [1870-1876] Conchologia Indica, being Illustrations of the Land and Freshwater Shells of British India. L. Reeve &amp; Co., London,65 pp. 160 pls. [pp. 1-18, pls 1-40 (1870); pp. 35-48, pls 81-120 (1874); pp. 35-48, pls 81-120 (1874); pp. 57-65, pls 141-160 (1876)] https://www.biodiversitylibrary.org/page/46204402" title="[1870 - 1876] Conchologia Indica, being Illustrations of the Land and Freshwater Shells of British India. L. Reeve &amp; Co., London, 65 pp. 160 pls. [pp. 1 - 18, pls 1 - 40 (1870); pp. 35 - 48, pls 81 - 120 (1874); pp. 35 - 48, pls 81 - 120 (1874); pp. 57 - 65, pls 141 - 160 (1876)]" url="https://www.biodiversitylibrary.org/page/46204402" year="1870">Hanley and Theobald 1876</bibRefCitation>
: 59, pl. 147, fig. 10.
<bibRefCitation id="32BC68DAAD725771ECE215C7060C2BE3" author="Pfeiffer, L" journalOrPublisher="Tome 1, Cassel: Th. Fischer" pageId="0" pageNumber="157" publicationUrl="https://www.biodiversitylibrary.org/page/11248775" refId="B68" refString="Pfeiffer, L, 1876. [1875-1876] Monographia Heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum (Vol. 7). Lipsiae: F.A. Brockhaus. 674 pp. [pp. 481-674 (1876)] https://www.biodiversitylibrary.org/page/11248775" title="[1875 - 1876] Monographia Heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum (Vol. 7). Lipsiae: F. A. Brockhaus. 674 pp. [pp. 481 - 674 (1876)]" url="https://www.biodiversitylibrary.org/page/11248775" year="1876">Pfeiffer 1876</bibRefCitation>
: 488.
<bibRefCitation id="45077ACF222B5B0F937E95CD277ADCE2" author="Pfeiffer, L" journalOrPublisher="Fischer, Cassel" pageId="0" pageNumber="157" refId="B70" refString="Pfeiffer, L, 1880. [1878-1881] Nomenclator heliceorum viventium. Fischer, Cassel" title="[1878 - 1881] Nomenclator heliceorum viventium." year="1880">Pfeiffer 1880</bibRefCitation>
: 344.
</paragraph>
<paragraph id="CF6AA828F2A9BAB6A4E60F57AE94D99D" pageId="0" pageNumber="157">
<taxonomicName id="E3E466D81DD077F624BA882DA4EF3BFB" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus" status="comb. nov.">Boysidia</taxonomicName>
(?)
<taxonomicName id="F51856FB780FB0753B1DDA55F6740310" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia dayana" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="dayana" status="comb. nov.">Boysidia dayana</taxonomicName>
.
<bibRefCitation id="F0E44BD4946BA5C9A122CB1A8D75C84F" author="Pilsbry, HA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="157" refId="B74" refString="Pilsbry, HA, 1917 1918. [1916-1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113-256, pls. 14-38 (1917); pp. 257-380, pls 39-49 (1918)]" title="[1916 - 1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113 - 256, pls. 14 - 38 (1917); pp. 257 - 380, pls 39 - 49 (1918)]" year="1917">Pilsbry 1917</bibRefCitation>
: 205, 206, pl. 34, figs 5, 6.
</paragraph>
<paragraph id="F22B86839CDFB2F2F088B70BBBC10262" pageId="0" pageNumber="157">
<taxonomicName id="36F2564EEB599BDE8FDD1A85267ADA1F" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus" status="comb. nov.">Hypselostoma</taxonomicName>
<emphasis id="08F4DE90FFAA55C92D14350E69AA6135" italics="true" pageId="0" pageNumber="157">Pupa (Hypselostoma) dayana</emphasis>
.
<bibRefCitation id="7F8BA7A85A8C3ADA7EB006E3C8DC6600" DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="157" refId="B44" refString="Nevill, G, 1878. Hand List of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Calcutta, 338 pp. https://doi.org/10.5962/bhl.title.11957" title="Hand List of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Calcutta, 338 pp." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
: 193.
</paragraph>
<paragraph id="5A880F8FD49E5FDC48DAED33F541701E" pageId="0" pageNumber="157">
<taxonomicName id="0458BC1F1611E99C69469391C5AD1788" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus" status="comb. nov.">Hypselostoma</taxonomicName>
<emphasis id="08D1D7CF75CF433FD977E1D7225155A9" italics="true" pageId="0" pageNumber="157">Pupa (Hypselostoma) dayanum</emphasis>
.
<bibRefCitation id="4F6FE4F3F805B887C7C9B16F8AC63097" DOI="https://doi.org/10.5962/bhl.title.12891" author="Gude, GK" journalOrPublisher="Taylor &amp; Francis, London" pageId="0" pageNumber="157" refId="B25" refString="Gude, GK, 1914. Mollusca-II. Trochomorphidae-Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma. Taylor &amp; Francis, London, DOI: https://doi.org/10.5962/bhl.title.12891" title="Mollusca-II. Trochomorphidae-Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma." url="https://doi.org/10.5962/bhl.title.12891" year="1914">Gude 1914</bibRefCitation>
: 300, 301.
</paragraph>
</subSubSection>
<subSubSection id="9EAD2D44CC3D6B8712662185F626D8DA" pageId="0" pageNumber="157" type="materials_examined">
<paragraph id="24254FB9A42D6769CEAB6B16CFEA4DCF" pageId="0" pageNumber="157">Material examined.</paragraph>
<paragraph id="3339F2D750121695AAAB3F551643AA72" pageId="0" pageNumber="157">
<materialsCitation id="504908E344F07542747D58D71ED183D1" collectingDate="2018-10-11" collectorName="Otani, J. U., Okubo, K., Hunyadi, A., HA Collection" country="Myanmar" elevation="8" latitude="16.506716" location="Dhammasa Cave" longLatPrecision="1" longitude="97.81076" municipality="Mawlamyine" specimenCount="1" stateProvince="Mon State">
<location id="C4443A19F8FADCC84B6D94DFEF91EDD0" LSID="urn:lsid:plazi:treatment:2CA29E2FBF6C5E4EAC4002448FB57B55:C4443A19F8FADCC84B6D94DFEF91EDD0" country="Myanmar" latitude="16.506716" longLatPrecision="1" longitude="97.81076" municipality="Mawlamyine" name="Dhammasa Cave" stateProvince="Mon State">Dhammasa Cave</location>
(
<quantity id="74F755E42FCF6DAE8B7F738FE005ABB0" metricMagnitude="0" metricUnit="m" metricValue="8.0" unit="m" value="8.0">
<elevation id="ECB0AA6758D502633B3C6F3F26391787" metricMagnitude="0" metricUnit="m" metricValue="8.0" unit="m" value="8.0">8 m</elevation>
</quantity>
),
<collectingMunicipality id="395ABF753015CE08CA053359EB724E8F">Mawlamyine</collectingMunicipality>
centre NEE ca
<quantity id="35D90315A2876E84C2CD243514521011" metricMagnitude="4" metricUnit="m" metricValue="2.6" unit="km" value="26.0">26 km</quantity>
,
<collectingRegion id="EC858B3D477CBF170F7C8F9080BBD673" country="Myanmar" name="Mon">Mon State</collectingRegion>
,
<collectingCountry id="F9CA8BF13C9555884E4679F13F0F0C6D" name="Myanmar">Myanmar</collectingCountry>
(
<geoCoordinate id="434F0D3F770FBC6BBD50C1A866E6222A" degrees="16.506715" direction="north" orientation="latitude" precision="1" value="16.506716">16.506715°N</geoCoordinate>
,
<geoCoordinate id="7BC5A7826818FC65B8B47300BC401BB0" degrees="97.810763" direction="east" orientation="longitude" precision="1" value="97.81076">97.810763°E</geoCoordinate>
), leg.
<collectorName id="E76824256DCA5CC16BCF5984F55E7593">Otani, J.U.</collectorName>
,
<collectorName id="174254BB871D1F9C2F068E266E405746">Okubo, K.</collectorName>
and
<collectorName id="7D2B79F6C28AF4B53C766EF0028633B1">Hunyadi, A.</collectorName>
<collectingDate id="F2E5ACC0E376A23FC5043AF13FF3979B" value="2018-10-11">11 October 2018</collectingDate>
:
<collectorName id="E514D707107B319982438723271EC4C8">HA Collection</collectorName>
(11 shells; Figs
<figureCitation id="91CA879584A8C1EB52FE65CF5CE1D0CA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A Acinolaemus dayanum, topotype specimen HA Collection from Mawlamyine Township, Mon State B Acinolaemus cryptidentatus, specimen CUMZ 14413 from Taunggyi District, Shan State. The white arrow indicates constriction, the yellow arrow indicates a cervical crest. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure2" httpUri="https://binary.pensoft.net/fig/1002051" pageId="0" pageNumber="157">2A</figureCitation>
,
<figureCitation id="EB31386B9DEE344996C73F8A4BB3CA54" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">13A</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A400E369C65A7454E3FCB9AFDFF98C8A" pageId="0" pageNumber="157" type="description">
<paragraph id="8E1AACE620F9D2CC2CFE9B64672C9785" pageId="0" pageNumber="157">Description.</paragraph>
<paragraph id="0CE0AB90DFCD85AE9FFE3ACE4C18B80E" pageId="0" pageNumber="157">Shell turban shaped, low spire, ~ 3-4 whorls and whitish in colour. Shell height 1.2-1.3 mm and shell width 1.3-1.4 mm. Apex blunt; protoconch with fine spiral striae. Teleoconch sculptured with regularly spaced radial ribs, paralleled with strongly wrinkled and malleated pits; suture well impressed and deep. Last whorl shouldered and flattened below periphery. Peristome thickened and weakly expanded; constriction and cervical crest very weak; lip whitish. Aperture subquadrate with eight dentitions marking peristome. Parietal lamella large and long, deeper inside aperture with tall ridge; infraparietal lamella small with long and low rise. Angular lamella smaller and weaker than parietal lamella, long, low rise with slope somewhat bent. Palatal tubercle strongly developed with triangular shape. Upper-, inter-, and lower- palatal plicae small, connecting peristome, and continuous with thin and straight ridges deeper inside aperture. Basal plica small, tubercle-like. Columellar lamella strong, continuing deep inside aperture with undulated ridge. Umbilicus widely perforate, ~ 1/3 of shell width, rounded and deep.</paragraph>
</subSubSection>
<subSubSection id="E154D1F5C59A7DDCB45BD2A2826D7514" pageId="0" pageNumber="157" type="distribution">
<paragraph id="FBBD844ACD342F9AD5FDF291D86519FD" pageId="0" pageNumber="157">Distribution.</paragraph>
<paragraph id="671860B9D33D998B6EA96404B00A81A3" pageId="0" pageNumber="157">Currently, this species is known only from the type locality, a limestone outcrop in Mon State, Myanmar.</paragraph>
</subSubSection>
<subSubSection id="D40178D61283E80D5295D8D688DABBF7" pageId="0" pageNumber="157" type="differential diagnosis">
<paragraph id="A1EDBAF95E562D0F42A5FE0251EA2687" pageId="0" pageNumber="157">Differential diagnosis.</paragraph>
<paragraph id="86BA92AB499AC6CB4C151870B3B49673" pageId="0" pageNumber="157">
<taxonomicName id="BB05F8FECD6476BFB6A69C1CFF1222B7" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Stoliczka" baseAuthorityYear="1871" class="Gastropoda" family="Hypselostomatidae" genus="Acinolaemus" kingdom="Animalia" lsidName="Acinolaemus dayanum" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="dayanum">
<emphasis id="1E0ED75BD4653A0A4A8526626033C091" italics="true" pageId="0" pageNumber="157">Acinolaemus dayanum</emphasis>
</taxonomicName>
differs from all known
<taxonomicName id="955B5B6B213A9912DE45ABAB428D389F" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="C613F561F5DE168761F1F790DB201D0C" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
</taxonomicName>
species in Myanmar and Thailand by its unique, turban shaped shell and teleoconch surface sculptured with malleated wrinkles and pits. In contrast, the four species from Myanmar and three species from Thailand, namely
<taxonomicName id="C386A02F9B9D77212D5F4D51FC7D84CB" lsidName="B. tamphathai" pageId="0" pageNumber="157" rank="species" species="tamphathai">
<emphasis id="D7377DCDA9A637B9234DACA077EE04B4" italics="true" pageId="0" pageNumber="157">B. tamphathai</emphasis>
</taxonomicName>
(Panha &amp; Burch, 2000),
<taxonomicName id="6CA27B899F46FEB1CC8CC2CDC3F6AB82" lsidName="B. nabhitabhatai" pageId="0" pageNumber="157" rank="species" species="nabhitabhatai">
<emphasis id="A8F8B3FBFFA95958C346B6D0711C65B8" italics="true" pageId="0" pageNumber="157">B. nabhitabhatai</emphasis>
</taxonomicName>
(Panha &amp; Burch, 2002) and
<taxonomicName id="FF1FD827105738B47BC01E45AE0054E5" lsidName="B. pangmapaensis" pageId="0" pageNumber="157" rank="species" species="pangmapaensis">
<emphasis id="1A2C66280CCCE4ECBFFDD7A443C3B2B7" italics="true" pageId="0" pageNumber="157">B. pangmapaensis</emphasis>
</taxonomicName>
(Panha &amp; Burch, 2002) possess a conical shell and a teleoconch surface generally with conspicuous irregular growth lines and fine spiral striae (
<bibRefCitation id="E5E5437B74EE9EBF9DBE5234CE3772D0" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" pagination="107 - 112" refId="B59" refString="Panha, S, Burch, JB, 2000. A new species of Paraboysidia from Thailand (Pulmonata: Pupillidae). Walkerana 11 (25): 107 - 112" title="A new species of Paraboysidia from Thailand (Pulmonata: Pupillidae)." volume="11" year="2000">Panha and Burch 2000</bibRefCitation>
,
<bibRefCitation id="6AC853445154C0252E0017C039A5C7A7" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B61" refString="Panha, S, Burch, JB, 2002b. The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12[2001](28): 77-94. [Date of publication follow Jirapatrasilp et al. (2023)]" title="The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12 [2001] (28): 77 - 94. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2002 b">2002b</bibRefCitation>
).
</paragraph>
<paragraph id="D27B43D2DB79CC95890D05B853FF0F28" pageId="0" pageNumber="157">
Compared to the
<taxonomicName id="6E05F075E2DBF12D6C89CC0BCC3DAE48" authorityName="Thompson &amp; Upatham" authorityYear="1997" class="Gastropoda" family="Hypselostomatidae" genus="Acinolaemus" kingdom="Animalia" lsidName="Acinolaemus" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="5CA717537DBE9279924EEF1D229D3F92" italics="true" pageId="0" pageNumber="157">Acinolaemus</emphasis>
</taxonomicName>
species from Thailand,
<taxonomicName id="704CA096EE51E9D66B2B8B62F06E11E6" lsidName="A. dayanum" pageId="0" pageNumber="157" rank="species" species="dayanum">
<emphasis id="EEBBBC314947E43B737CFBAD6628AE32" italics="true" pageId="0" pageNumber="157">A. dayanum</emphasis>
</taxonomicName>
differs by having pitted sculpture on the shell surface, weaker spiral and radial striae, a larger parietal than angular lamella, and a shouldered last whorl.
<taxonomicName id="32C38E04C85B6B3415B13683CC412FAC" authorityName="F. G. Thompson &amp; Upatham" authorityYear="1997" class="Gastropoda" family="Hypselostomatidae" genus="Acinolaemus" kingdom="Animalia" lsidName="Acinolaemus ptychochilus" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="ptychochilus">
<emphasis id="F84C408A15C94286088A0E9DA2801B72" italics="true" pageId="0" pageNumber="157">Acinolaemus ptychochilus</emphasis>
</taxonomicName>
Thompson &amp; Upatham, 1997 differs from
<taxonomicName id="D0C84FB72C4715FDC84348F0BC6AFEB9" lsidName="A. dayanum" pageId="0" pageNumber="157" rank="species" species="dayanum">
<emphasis id="5FAF27B5F2A8A5B78AF207DE9538B558" italics="true" pageId="0" pageNumber="157">A. dayanum</emphasis>
</taxonomicName>
by its rounded last whorl, strong tubercles on the peristomal lip, and the presence of supra- and subcolumellar lamellae.
<taxonomicName id="4CC4A6BCF7A4357417EE476398BD70CB" authorityName="F. G. Thompson &amp; Upatham" authorityYear="1997" class="Gastropoda" family="Hypselostomatidae" genus="Acinolaemus" kingdom="Animalia" lsidName="Acinolaemus colpodon" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="colpodon">
<emphasis id="87D230E3A90A92BD536D572D4EB98E86" italics="true" pageId="0" pageNumber="157">Acinolaemus colpodon</emphasis>
</taxonomicName>
Thompson &amp; Upatham, 1997 also has a rounded last whorl, possesses a hooked columellar lamella, and lacks an interpalatal plica, a basal plica and an infraparietal lamella. In addition,
<taxonomicName id="0CE2822B11CAC11A0E1D3F1237D9C80A" lsidName="A. sphinctinion" pageId="0" pageNumber="157" rank="species" species="sphinctinion">
<emphasis id="52E6587262FFD352112F22A099A96C0F" italics="true" pageId="0" pageNumber="157">A. sphinctinion</emphasis>
</taxonomicName>
Thompson &amp; Upatham, 1997 has a shouldered last whorl with blunt periphery, a short, weakly ascending tuba, and lacks the angular lamella, the parietal fold, the basal plica and the interpalatal plica (
<bibRefCitation id="243A7DBAF1EBAD1FCF613793FDFA0CBC" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida Museum of Natural History" pageId="0" pageNumber="157" pagination="221 - 245" publicationUrl="http://ufdc.ufl.edu/UF00095785/00001" refId="B87" refString="Thompson, FG, Upatham, S, 1997. Vertiginid land snails from Thailand (Gastropoda, Pulmonata, Pupilloidea). Bulletin of the Florida Museum of Natural History 39 (7): 221 - 245, http://ufdc.ufl.edu/UF00095785/00001" title="Vertiginid land snails from Thailand (Gastropoda, Pulmonata, Pupilloidea)." url="http://ufdc.ufl.edu/UF00095785/00001" volume="39" year="1997">Thompson and Upatham 1997</bibRefCitation>
;
<bibRefCitation id="7E29CD325587960EA35522E20972C48C" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B63" refString="Panha, S, Burch, JB, 2005. An introduction to the microsnails of Thailand. Malacological Review 37/38(2004-2005): 1-155. [Date of publication follow Jirapatrasilp et al. (2023)]" title="An introduction to the microsnails of Thailand. Malacological Review 37 / 38 (2004 - 2005): 1 - 155. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2005">Panha and Burch 2005</bibRefCitation>
).
</paragraph>
<paragraph id="DD3E642698E3B000665F17651F32FCF8" pageId="0" pageNumber="157">
Additionally,
<taxonomicName id="D71859E591EC51DD3F985C9F10F0394D" lsidName="A. dayanum" pageId="0" pageNumber="157" rank="species" species="dayanum">
<emphasis id="4E5E38A0902CE80CECEC1D03629023F0" italics="true" pageId="0" pageNumber="157">A. dayanum</emphasis>
</taxonomicName>
has a similar shell form and sculpture to the two
<taxonomicName id="2A9A47BBAA480878C951848B922F1DBB" authorityName="Thompson &amp; Upatham" authorityYear="1997" class="Gastropoda" family="Hypselostomatidae" genus="Acinolaemus" kingdom="Animalia" lsidName="Acinolaemus" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="D7EFEC62650762E2D6067748BBD45394" italics="true" pageId="0" pageNumber="157">Acinolaemus</emphasis>
</taxonomicName>
species recently described from northern Thailand. It differs from
<taxonomicName id="E69B51203736BE98DFCD27DD6021B73E" lsidName="A. cryptidentatus" pageId="0" pageNumber="157" rank="species" species="cryptidentatus">
<emphasis id="C11C1FBA65E82C9A4123A4634F1768C3" italics="true" pageId="0" pageNumber="157">A. cryptidentatus</emphasis>
</taxonomicName>
<bibRefCitation id="744764C04D1C64C1A255704A450A301F" author="Changlom, B" journalOrPublisher="The Thailand Natural History Museum Journal" pageId="0" pageNumber="157" pagination="155 - 161" refId="B15" refString="Changlom, B, Chan-ard, T, Dumrongrojwattana, P, 2019. Two new species of the genus Acinolaemus (Pulmonata: Pupillidae) from Thailand. The Thailand Natural History Museum Journal 13: 155 - 161" title="Two new species of the genus Acinolaemus (Pulmonata: Pupillidae) from Thailand." volume="13" year="2019">Changlom et al., 2019</bibRefCitation>
, which has 4-5 whorls, strong spiral striae, and discontinuous infraparietal and angular lamellae. It is also distinct from
<taxonomicName id="DD2F51A2266A36C014A3DDAAFB26C4B6" lsidName="A. mueangonensis" pageId="0" pageNumber="157" rank="species" species="mueangonensis">
<emphasis id="720DE59F576C87CD8F4C64E0D2AA2512" italics="true" pageId="0" pageNumber="157">A. mueangonensis</emphasis>
</taxonomicName>
<bibRefCitation id="C2A1F271A42F785B4ACEA1DDF2E0794B" author="Changlom, B" journalOrPublisher="The Thailand Natural History Museum Journal" pageId="0" pageNumber="157" pagination="155 - 161" refId="B15" refString="Changlom, B, Chan-ard, T, Dumrongrojwattana, P, 2019. Two new species of the genus Acinolaemus (Pulmonata: Pupillidae) from Thailand. The Thailand Natural History Museum Journal 13: 155 - 161" title="Two new species of the genus Acinolaemus (Pulmonata: Pupillidae) from Thailand." volume="13" year="2019">Changlom et al., 2019</bibRefCitation>
[corrected original spelling], which has 4-5 whorls, brownish shell colour, and a more prominent subcolumellar than columellar lamella.
</paragraph>
</subSubSection>
<subSubSection id="5E40BFCCCA3636748407E9CC7153F0A3" pageId="0" pageNumber="157" type="remarks">
<paragraph id="5F37039C5FEE4F867C051F1692409C05" pageId="0" pageNumber="157">Remarks.</paragraph>
<paragraph id="D70E5EC27C319E129A3497AB340D097C" pageId="0" pageNumber="157">
The holotype (single shell mentioned in the original description) could not be located in the NHM collection, but we were able to examine recently collected topotypic shells. Originally, this species was classified in
<taxonomicName id="CEA22A7BD242B958EA7A3B9447CA58AF" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="37FF758402FECA34B01662819E44F53E" italics="true" pageId="0" pageNumber="157">Hypselostoma</emphasis>
</taxonomicName>
and later reclassified into the genus
<taxonomicName id="64B63EB3B683A3488F9B4E4CFF294758" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="EBB43FF77E7644DB2CC25101099DDDED" italics="true" pageId="0" pageNumber="157">Boysidia</emphasis>
</taxonomicName>
Ancey, 1881 (i.e.,
<bibRefCitation id="8451C8189AB52329D411189761623033" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="157" pagination="143 - 177" publicationUrl="www.biodiversitylibrary.org/page/35630965" refId="B81" refString="Stoliczka, F, 1871. Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 177, www.biodiversitylibrary.org/page/35630965" title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species." url="www.biodiversitylibrary.org/page/35630965" volume="40" year="1871">Stoliczka 1871</bibRefCitation>
;
<bibRefCitation id="EEF3D818DE8B8AE240FDA57581F3DB07" author="Pilsbry, HA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="157" refId="B74" refString="Pilsbry, HA, 1917 1918. [1916-1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113-256, pls. 14-38 (1917); pp. 257-380, pls 39-49 (1918)]" title="[1916 - 1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113 - 256, pls. 14 - 38 (1917); pp. 257 - 380, pls 39 - 49 (1918)]" year="1917">Pilsbry 1917</bibRefCitation>
). Based on the small size, the whitish shell and the strong teleoconch sculpture, we transfer
<taxonomicName id="8D4397E4EEAF154B08F9BE45F7679CB8" lsidName="H. dayanum" pageId="0" pageNumber="157" rank="species" species="dayanum">
<emphasis id="FED3BD5C11B41A630CBF05430881BC23" italics="true" pageId="0" pageNumber="157">H. dayanum</emphasis>
</taxonomicName>
into the genus
<taxonomicName id="8C23109032DD9B477F1FDF5A2183DC2B" authorityName="Thompson &amp; Upatham" authorityYear="1997" class="Gastropoda" family="Hypselostomatidae" genus="Acinolaemus" kingdom="Animalia" lsidName="Acinolaemus" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="2580FEEE08035512631035A29EAB184A" italics="true" pageId="0" pageNumber="157">Acinolaemus</emphasis>
</taxonomicName>
. In contrast, the genus
<taxonomicName id="7B995E7C1A873E86D079728DED369B94" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="87E6AB83456CDAC465918CFD0C229346" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
</taxonomicName>
is characterised by a brownish shell with nearly smooth or finely, spirally striated shell (
<bibRefCitation id="EAFF3326B81747A867A459FA3C09A40E" DOI="https://doi.org/10.1080/00222933.2022.2152750" author="Pall-Gergely, B" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="157" pagination="2011 - 2029" refId="B49" refString="Pall-Gergely, B, White, TS, 2023. Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45-48): 2011 - 2029, DOI: https://doi.org/10.1080/00222933.2022.2152750" title="Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)." url="https://doi.org/10.1080/00222933.2022.2152750" volume="56" year="2023">
<normalizedToken id="F58B9B70884C9BB42112BF849AA6835B" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
and White 2023
</bibRefCitation>
), whereas
<taxonomicName id="974865512224577124824BC369B1ACEE" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="26808FB6DA598FBAB2D0A17DE8725D0B" italics="true" pageId="0" pageNumber="157">Hypselostoma</emphasis>
</taxonomicName>
(at least the type species and the morphologically similar and geographically close species; see
<bibRefCitation id="D7A15FE18411B02AAC76BD86F46593C2" DOI="https://doi.org/10.1127/arch.moll/151/131-139" author="Gojsina, V" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="157" pagination="131 - 139" refId="B24" refString="Gojsina, V, Chen, Z-U, Pall-Gergely, B, 2022. Notes on Hypselostomatidae (Gastropoda: Eupulmonata: Pupilloidea) from Myanmar. Archiv fuer Molluskenkunde 151 (2): 131 - 139, DOI: https://doi.org/10.1127/arch.moll/151/131-139" title="Notes on Hypselostomatidae (Gastropoda: Eupulmonata: Pupilloidea) from Myanmar." url="https://doi.org/10.1127/arch.moll/151/131-139" volume="151" year="2022">
<normalizedToken id="6A6075C4D29B6CC91DE1EB567DD4A86B" originalValue="Gojšina">Gojsina</normalizedToken>
et al. 2022
</bibRefCitation>
and
<bibRefCitation id="7732DE48D42EAE69947EABDA408C613B" author="Preece, RC" journalOrPublisher="Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="157" refId="B77" refString="Preece, RC, White, TS, Raheem, DC, Ketchum, H, Ablett, J, Taylor, H, Naggs, F, 2022. William Benson and the golden age of malacology in British India: Biography, illustrated catalogue and evaluation of his molluscan types. Tropical Natural History (Supplement 6): 1-434." title="William Benson and the golden age of malacology in British India: Biography, illustrated catalogue and evaluation of his molluscan types. Tropical Natural History (Supplement 6): 1 - 434." year="2022">Preece et al. 2022</bibRefCitation>
) and
<taxonomicName id="59229527D4D6B9482CE20C0F5614AF01" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="46A15C7F76B61EED7489264F036FAF68" italics="true" pageId="0" pageNumber="157">Boysidia</emphasis>
</taxonomicName>
are much larger, and have brownish, finely sculptured shells. Furthermore,
<taxonomicName id="29380F73F33C945D74FA563D00598038" authorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Tongkerd &amp; Lwin &amp; Páll-Gergely &amp; Chanabun &amp; Pholyotha &amp; Prasankok &amp; Seesamut &amp; Siriwut &amp; Srisonchai &amp; Sutcharit &amp; Panha" baseAuthorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Hypselostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypselostoma" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="D0952ED65C965DC4EB5097187B7AC9A6" italics="true" pageId="0" pageNumber="157">Hypselostoma</emphasis>
</taxonomicName>
is defined on the basis of a detached body whorl and a concrescent angular and parietal lamellae, characters which do not appear in
<taxonomicName id="B6BDBA6AAD88DBDAA73F48295EF6FCAA" lsidName="A. dayanum" pageId="0" pageNumber="157" rank="species" species="dayanum">
<emphasis id="18ECEDCE5537E11CF0796EA21E2E27FA" italics="true" pageId="0" pageNumber="157">A. dayanum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>