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<document id="D57ED3758951F1663EC1B4BEA7305EE2" ID-DOI="10.26107/RBZ-2020-0092" ID-ISSN="2345-7600" ID-Zenodo-Dep="5350725" ID-ZooBank="6963D2F4-0E08-4455-AEFC-735A912DA735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630205214526" checkinUser="diego" docAuthor="Zettel, Herbert, Laciny, Alice &amp; Freitag, Hendrik" docDate="2020" docId="1A4987922772780AFCD3FD813813670C" docLanguage="en" docName="RBZ-2020-0092.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Rhagovelia Mayr 1865" docType="key" docVersion="4" lastPageNumber="813" masterDocId="E670FFEA27707809FFFDFFD43B166441" masterDocTitle="Review of the genus Rhagovelia Mayr, 1865 (Insecta: Heteroptera: Veliidae) in the Palawan biogeographic region, the Philippines" masterLastPageNumber="837" masterPageNumber="810" pageNumber="812" updateTime="1699184686890" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C9398E6F006BF55E5E553A5796B52F86">Review of the genus Rhagovelia Mayr, 1865 (Insecta: Heteroptera: Veliidae) in the Palawan biogeographic region, the Philippines</mods:title>
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<mods:namePart id="6AC9770552E998D79FD64B28AA9E93EF">Zettel, Herbert</mods:namePart>
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<treatment id="1A4987922772780AFCD3FD813813670C" ID-DOI="http://doi.org/10.5281/zenodo.5449432" ID-Zenodo-Dep="5449432" LSID="urn:lsid:plazi:treatment:1A4987922772780AFCD3FD813813670C" httpUri="http://treatment.plazi.org/id/1A4987922772780AFCD3FD813813670C" lastPageId="3" lastPageNumber="813" pageId="2" pageNumber="812">
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<emphasis id="A094EA962772780BFCD3FD813EBC662C" bold="true" box="[814,1450,597,621]" pageId="2" pageNumber="812">
Key to the species of
<taxonomicName id="55E04D072772780BFBDBFD813FB4662C" ID-CoL="78BP" authorityName="Mayr" authorityYear="1865" box="[1062,1186,597,621]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="genus">
<emphasis id="A094EA962772780BFBDBFD813FB4662C" bold="true" box="[1062,1186,597,621]" italics="true" pageId="2" pageNumber="812">Rhagovelia</emphasis>
</taxonomicName>
in the
<collectingRegion id="5024F8662772780BFB0BFD813EBC662C" box="[1270,1450,597,621]" country="Philippines" name="Palawan" pageId="2" pageNumber="812">Palawan region</collectingRegion>
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<subSubSection id="DAFA650F2772780AFCD0FD403813670C" lastPageId="3" lastPageNumber="813" pageId="2" pageNumber="812" type="key">
<keyStep id="29142E212772780BFCD0FD403EBC6043" pageId="2" pageNumber="812">
<paragraph id="925F36842772780BFCD0FD403EBC6778" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
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1. Anterior margin of pronotum entirely yellowish; connexival margin yellowish (
<figureCitation id="0ADB2A012772780BFBF7FD643F686687" box="[1034,1150,688,710]" captionStart="Fig" captionStartId="14.[113,148,1621,1643]" captionTargetBox="[113,1451,189,1606]" captionTargetId="figure-12@14.[113,1451,189,1606]" captionTargetPageId="14" captionText="Fig. 5. Habitus of Rhagovelia kawakamii hoberlandti. A, apterous male; B, apterous female; C, macropterous male; D, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350735" httpUri="https://zenodo.org/record/5350735/files/figure.png" pageId="2" pageNumber="812">Figs. 5AD</figureCitation>
,
<figureCitation id="0ADB2A012772780BFB77FD643FD16687" box="[1162,1223,688,710]" captionStart="Fig" captionStartId="16.[113,148,1727,1749]" captionTargetBox="[113,1451,189,1712]" captionTargetId="figure-12@16.[113,1451,189,1712]" captionTargetPageId="16" captionText="Fig. 6. Habitus of Rhagovelia gapudi, new species. A, apterous male; B, apterous female; C, macropterous male." figureDoi="http://doi.org/10.5281/zenodo.5350737" httpUri="https://zenodo.org/record/5350737/files/figure.png" pageId="2" pageNumber="812">6AC</figureCitation>
). Pronotum of apterous morph much longer than eye length, with strongly convex hind margin, covering most of mesonotum (e.g.,
<figureCitation id="0ADB2A012772780BFB0EFD3E3E286741" box="[1267,1342,746,768]" captionStart="Fig" captionStartId="14.[113,148,1621,1643]" captionTargetBox="[113,1451,189,1606]" captionTargetId="figure-12@14.[113,1451,189,1606]" captionTargetPageId="14" captionText="Fig. 5. Habitus of Rhagovelia kawakamii hoberlandti. A, apterous male; B, apterous female; C, macropterous male; D, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350735" httpUri="https://zenodo.org/record/5350735/files/figure.png" pageId="2" pageNumber="812">Fig. 5A</figureCitation>
). Forewing blackish with light streak at base (e.g.,
<figureCitation id="0ADB2A012772780BFB29FCD33E36675C" box="[1236,1312,775,797]" captionStart="Fig" captionStartId="14.[113,148,1621,1643]" captionTargetBox="[113,1451,189,1606]" captionTargetId="figure-12@14.[113,1451,189,1606]" captionTargetPageId="14" captionText="Fig. 5. Habitus of Rhagovelia kawakamii hoberlandti. A, apterous male; B, apterous female; C, macropterous male; D, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350735" httpUri="https://zenodo.org/record/5350735/files/figure.png" pageId="2" pageNumber="812">Fig. 5C</figureCitation>
). Body length
<quantity id="55189B612772780BFCA9FCF038DA6778" box="[852,972,804,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.9" metricValueMin="2.8" pageId="2" pageNumber="812" unit="mm" value="3.3499999999999996" valueMax="3.9" valueMin="2.8">2.83.9 mm</quantity>
. (III.
<taxonomicName id="55E04D072772780BFBF8FCF03F976778" ID-CoL="4RZG3" authorityName="Lundblad" authorityYear="1936" box="[1029,1153,804,825]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="A094EA962772780BFBF8FCF03F976778" box="[1029,1153,804,825]" italics="true" pageId="2" pageNumber="812">R. papuensis</emphasis>
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group) ...................................2
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</paragraph>
<paragraph id="925F36842772780BFCD0FC953EBC6043" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
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Pronotum black, with short orange or yellowish mark anteriorly; connexival margin black (
<figureCitation id="0ADB2A012772780BFBAEFC893FDE6733" box="[1107,1224,861,883]" captionStart="Fig" captionStartId="4.[113,148,1573,1595]" captionTargetBox="[113,1451,190,1558]" captionTargetId="figure-13@4.[113,1451,190,1558]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Rhagovelia species. A, R. minutissima, apterous male; B, R. minutissima, apterous female; C, R. palawanensis, apterous male; D, R. palawanensis, apterous female." figureDoi="http://doi.org/10.5281/zenodo.5350727" httpUri="https://zenodo.org/record/5350727/files/figure.png" pageId="2" pageNumber="812">Figs. 1AD</figureCitation>
,
<figureCitation id="0ADB2A012772780BFB29FC893E046733" box="[1236,1298,861,882]" captionStart="Fig" captionStartId="7.[136,171,1498,1520]" captionTargetBox="[136,1474,189,1482]" captionTargetId="figure-7@7.[136,1474,189,1482]" captionTargetPageId="7" captionText="Fig. 2. Habitus ofRhagovelia matillanoi, new species. A, apterous male; B, apterous female; C, macropterous dealate, male; D, macropterous dealate, female." figureDoi="http://doi.org/10.5281/zenodo.5350729" httpUri="https://zenodo.org/record/5350729/files/figure.png" pageId="2" pageNumber="812">2AD</figureCitation>
,
<figureCitation id="0ADB2A012772780BFAE3FC893E406733" box="[1310,1366,861,882]" captionStart="Fig" captionStartId="9.[136,171,2064,2086]" captionTargetBox="[166,1445,189,2049]" captionTargetId="figure-8@9.[166,1445,189,2049]" captionTargetPageId="9" captionText="Fig. 3. Habitus of Rhagovelia species. R. estrella: A, apterous male; B, apterous female. R. abbreviata, new species: C, apterous male; D, macropterous dealate, male; E, apterous female; F, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350731" httpUri="https://zenodo.org/record/5350731/files/figure.png" pageId="2" pageNumber="812">3AF</figureCitation>
,
<figureCitation id="0ADB2A012772780BFA9EFC893EB66732" box="[1379,1440,861,883]" captionStart="Fig" captionStartId="12.[112,147,1308,1330]" captionTargetBox="[113,1451,189,1293]" captionTargetId="figure-303@12.[113,1451,189,1293]" captionTargetPageId="12" captionText="Fig. 4. Habitus of Rhagovelia lansburyi. A, apterous male; B, apterous female; C, macropterous male." figureDoi="http://doi.org/10.5281/zenodo.5350733" httpUri="https://zenodo.org/record/5350733/files/figure.png" pageId="2" pageNumber="812">4AC</figureCitation>
). Pronotum of apterous morph shorter than eye length, with straight hind margin; mesonotum largely exposed (e.g.,
<figureCitation id="0ADB2A012772780BFA78FC433864678B" captionStart="Fig" captionStartId="4.[113,148,1573,1595]" captionTargetBox="[113,1451,190,1558]" captionTargetId="figure-13@4.[113,1451,190,1558]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Rhagovelia species. A, R. minutissima, apterous male; B, R. minutissima, apterous female; C, R. palawanensis, apterous male; D, R. palawanensis, apterous female." figureDoi="http://doi.org/10.5281/zenodo.5350727" httpUri="https://zenodo.org/record/5350727/files/figure.png" pageId="2" pageNumber="812">Fig. 1A</figureCitation>
). Forewing without light streak at base (e.g.,
<figureCitation id="0ADB2A012772780BFADBFC603E616788" box="[1318,1399,948,970]" captionStart="Fig" captionStartId="9.[136,171,2064,2086]" captionTargetBox="[166,1445,189,2049]" captionTargetId="figure-8@9.[166,1445,189,2049]" captionTargetPageId="9" captionText="Fig. 3. Habitus of Rhagovelia species. R. estrella: A, apterous male; B, apterous female. R. abbreviata, new species: C, apterous male; D, macropterous dealate, male; E, apterous female; F, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350731" httpUri="https://zenodo.org/record/5350731/files/figure.png" pageId="2" pageNumber="812">Figs. 3F</figureCitation>
,
<figureCitation id="0ADB2A012772780BFA7CFC603EB7678B" box="[1409,1441,948,970]" captionStart="Fig" captionStartId="12.[112,147,1308,1330]" captionTargetBox="[113,1451,189,1293]" captionTargetId="figure-303@12.[113,1451,189,1293]" captionTargetPageId="12" captionText="Fig. 4. Habitus of Rhagovelia lansburyi. A, apterous male; B, apterous female; C, macropterous male." figureDoi="http://doi.org/10.5281/zenodo.5350733" httpUri="https://zenodo.org/record/5350733/files/figure.png" pageId="2" pageNumber="812">4C</figureCitation>
). Body length
<quantity id="55189B612772780BFC29FC053F5B67A7" box="[980,1101,977,998]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.9" metricValueMin="1.9" pageId="2" pageNumber="812" unit="mm" value="2.4" valueMax="2.9" valueMin="1.9">1.92.9 mm</quantity>
, except
<quantity id="55189B612772780BFB5DFC043E0067A7" box="[1184,1302,976,998]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="2" pageNumber="812" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.33.5 mm</quantity>
in the females of
<taxonomicName id="55E04D072772780BFC93FC3A38F06043" ID-CoL="4RZDM" authorityName="Zettel" authorityYear="1995" box="[878,998,1005,1027]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="lansburyi">
<emphasis id="A094EA962772780BFC93FC3A38F06043" box="[878,998,1005,1027]" italics="true" pageId="2" pageNumber="812">R. lansburyi</emphasis>
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.........................................................................3
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</keyStep>
<keyStep id="29142E212772780BFCD0FBDE3EBC61D7" pageId="2" pageNumber="812">
<paragraph id="925F36842772780BFCD0FBDE3EBC608D" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
<keyLead id="291195B12772780BFCD0FBDE3EBC608D" pageId="2" pageNumber="812">
2. Anterior part of head black (not in freshly moulted specimens). Tergites 7 and 8 without shiny areas. Juga (pair of plates at sides of labrum and rostrum base) with small black spinulae. Paramere of male distally broad, truncate (
<figureCitation id="0ADB2A012772780BFB0EFBB43E576036" box="[1267,1345,1120,1143]" captionStart="Fig" captionStartId="17.[136,171,808,830]" captionTargetBox="[136,1474,191,791]" captionTargetId="figure-8@17.[134,1477,188,792]" captionTargetPageId="17" captionText="Fig. 7. Left parameres of Rhagovelia species, lateral view. A, R. minutissima; B, R. palawanensis; C, R. matillanoi, new species; D, R. estrella; E, R. abbreviata, new species; F, R. lansburyi; G, R. kawakamii hoberlandti; H, R. gapudi, new species." figureDoi="http://doi.org/10.5281/zenodo.5350739" httpUri="https://zenodo.org/record/5350739/files/figure.png" pageId="2" pageNumber="812">Fig. 7G</figureCitation>
). Apterous female with connexival margins widely separated all over length (
<figureCitation id="0ADB2A012772780BFC60FB4E38FC60F1" box="[925,1002,1178,1200]" captionStart="Fig" captionStartId="14.[113,148,1621,1643]" captionTargetBox="[113,1451,189,1606]" captionTargetId="figure-12@14.[113,1451,189,1606]" captionTargetPageId="14" captionText="Fig. 5. Habitus of Rhagovelia kawakamii hoberlandti. A, apterous male; B, apterous female; C, macropterous male; D, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350735" httpUri="https://zenodo.org/record/5350735/files/figure.png" pageId="2" pageNumber="812">Fig. 5B</figureCitation>
). Common and widespread in the region, also on Borneo ........................................
<taxonomicName id="55E04D072772780BFB53FB6C3EBC608D" ID-CoL="7L327" authorityName="Hungerford &amp; Matsuda" authorityYear="1961" box="[1198,1450,1207,1229]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="subSpecies" species="kawakamii" subSpecies="hoberlandti">
<emphasis id="A094EA962772780BFB53FB6C3EBC608D" box="[1198,1450,1207,1229]" italics="true" pageId="2" pageNumber="812">R. kawakamii hoberlandti</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="925F36842772780BFCD0FB003EBC61D7" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
<keyLead id="291195B12772780BFCD0FB003EBC61D7" pageId="2" pageNumber="812">
Anterior part of head (in frontal view) usually pale orange to brown, rarely blackish. Juga without spinulae. Apterous specimens with shiny areas on tergites 7 and 8 (more distinct in female). Paramere of male distally slender, with small ventrally directed lobe (
<figureCitation id="0ADB2A012772780BFC26FA933F3E611D" box="[987,1064,1351,1373]" captionStart="Fig" captionStartId="17.[136,171,808,830]" captionTargetBox="[136,1474,191,791]" captionTargetId="figure-8@17.[134,1477,188,792]" captionTargetPageId="17" captionText="Fig. 7. Left parameres of Rhagovelia species, lateral view. A, R. minutissima; B, R. palawanensis; C, R. matillanoi, new species; D, R. estrella; E, R. abbreviata, new species; F, R. lansburyi; G, R. kawakamii hoberlandti; H, R. gapudi, new species." figureDoi="http://doi.org/10.5281/zenodo.5350739" httpUri="https://zenodo.org/record/5350739/files/figure.png" pageId="2" pageNumber="812">Fig. 7H</figureCitation>
). Connexival margin of apterous female strongly convergent near apex (
<figureCitation id="0ADB2A012772780BFB7EFAB03FC1613B" box="[1155,1239,1380,1402]" captionStart="Fig" captionStartId="16.[113,148,1727,1749]" captionTargetBox="[113,1451,189,1712]" captionTargetId="figure-12@16.[113,1451,189,1712]" captionTargetPageId="16" captionText="Fig. 6. Habitus of Rhagovelia gapudi, new species. A, apterous male; B, apterous female; C, macropterous male." figureDoi="http://doi.org/10.5281/zenodo.5350737" httpUri="https://zenodo.org/record/5350737/files/figure.png" pageId="2" pageNumber="812">Figs. 6B</figureCitation>
,
<figureCitation id="0ADB2A012772780BFB1FFAB03E19613B" box="[1250,1295,1380,1402]" captionStart="Fig" captionStartId="19.[136,171,1404,1426]" captionTargetBox="[136,1475,188,1387]" captionTargetId="figure-332@19.[132,1480,183,1392]" captionTargetPageId="19" captionText="Fig. 10. Abdominal terminalia of females of Rhagovelia species, pilosity partly omitted. AC, R. matillanoi, new species, arrow in A pointing to characteristic setae; DF, R. abbreviata, new species; GI, R. gapudi, new species, arrows in H, I pointing to concave ventral outline of gonocoxa. A, D, G, apterous specimen, dorsal aspect; B, E, H, apterous specimen, lateral aspect; C, F, I, macropterous specimen in 48 dealate, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5350745" httpUri="https://zenodo.org/record/5350745/files/figure.png" pageId="2" pageNumber="812">10G</figureCitation>
). Widespread in the region.............................................
<taxonomicName id="55E04D072772780BFB35FA553E3161D4" authority="Zettel &amp; Laciny &amp; Freitag, 2020" authorityName="Zettel &amp; Laciny &amp; Freitag" authorityYear="2020" box="[1224,1319,1408,1430]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="gapudi" status="sp. nov.">
<emphasis id="A094EA962772780BFB35FA553E3161D4" box="[1224,1319,1408,1430]" italics="true" pageId="2" pageNumber="812">R. gapudi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BBA757ED2772780BFAC8FA553EBC61D7" box="[1333,1450,1408,1430]" pageId="2" pageNumber="812" rank="species">new species</taxonomicNameLabel>
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</paragraph>
</keyStep>
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<paragraph id="925F36842772780BFCD0FA493EBC6203" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
<keyLead id="291195B12772780BFCD0FA493EBC6203" pageId="2" pageNumber="812">
3. Stout species, strongly dimorphic; female much larger than male, in apterous morph
<quantity id="55189B612772780BFBC7FA6E3FD56191" box="[1082,1219,1466,1488]" metricMagnitude="-3" metricUnit="m" metricValue="3.1849999999999996" metricValueMax="3.34" metricValueMin="3.03" pageId="2" pageNumber="812" unit="mm" value="3.1849999999999996" valueMax="3.34" valueMin="3.03">3.033.34 mm</quantity>
vs.
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. Male with modified foreleg (
<figureCitation id="0ADB2A012772780BFBC6FA033F9A61AC" box="[1083,1164,1495,1517]" captionStart="Fig" captionStartId="12.[112,147,1308,1330]" captionTargetBox="[113,1451,189,1293]" captionTargetId="figure-303@12.[113,1451,189,1293]" captionTargetPageId="12" captionText="Fig. 4. Habitus of Rhagovelia lansburyi. A, apterous male; B, apterous female; C, macropterous male." figureDoi="http://doi.org/10.5281/zenodo.5350733" httpUri="https://zenodo.org/record/5350733/files/figure.png" pageId="2" pageNumber="812">Fig. 4A</figureCitation>
), protibia with distinct bend at distal fourth. In male, dentition of metafemur reduced; in female, metafemur without dentition (
<figureCitation id="0ADB2A012772780BFB47F9C43E136266" box="[1210,1285,1552,1575]" captionStart="Fig" captionStartId="12.[112,147,1308,1330]" captionTargetBox="[113,1451,189,1293]" captionTargetId="figure-303@12.[113,1451,189,1293]" captionTargetPageId="12" captionText="Fig. 4. Habitus of Rhagovelia lansburyi. A, apterous male; B, apterous female; C, macropterous male." figureDoi="http://doi.org/10.5281/zenodo.5350733" httpUri="https://zenodo.org/record/5350733/files/figure.png" pageId="2" pageNumber="812">Fig. 4B</figureCitation>
). (II.
<taxonomicName id="55E04D072772780BFACBF9C53EBD6264" ID-CoL="4RZDM" authorityName="Zettel" authorityYear="1995" box="[1334,1451,1552,1574]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="lansburyi">
<emphasis id="A094EA962772780BFACBF9C53EBD6264" box="[1334,1451,1552,1574]" italics="true" pageId="2" pageNumber="812">R. lansburyi</emphasis>
</taxonomicName>
group).....................................................................
<taxonomicName id="55E04D072772780BFACEF9FA3EBC6203" ID-CoL="4RZDM" authorityName="Zettel" authorityYear="1995" box="[1331,1450,1581,1603]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="lansburyi">
<emphasis id="A094EA962772780BFACEF9FA3EBC6203" box="[1331,1450,1581,1603]" italics="true" pageId="2" pageNumber="812">R. lansburyi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="925F36842772780BFCD0F99F3EBC6293" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
<keyLead id="291195B12772780BFCD0F99F3EBC6293" pageId="2" pageNumber="812">
Usually slender; if stout, then much smaller. Body length
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. Foreleg of males not modified, protibia more or less straight. In males, dentition of metafemur prominent (e.g.,
<figureCitation id="0ADB2A012772780BFCAFF97438F762F7" box="[850,993,1696,1718]" captionStart="Fig" captionStartId="18.[113,148,1468,1490]" captionTargetBox="[112,1451,189,1452]" captionTargetId="figure-239@18.[106,1456,185,1456]" captionTargetPageId="18" captionText="Fig. 9. Hindlegs of Rhagovelia species, showing characteristic armatures pilosity omitted. A, R. matillanoi, new species, apterous male; B, R. matillanoi, new species, apterous female; C, R. abbreviata, new species, apterous male; D, R. abbreviata, new species, macropterous male; E, R. abbreviata, new species, apterous female; F, G, R. gapudi, new species, apterous males, two morphs; H, R. gapudi, new species, apterous female." figureDoi="http://doi.org/10.5281/zenodo.5350743" httpUri="https://zenodo.org/record/5350743/files/figure.png" pageId="2" pageNumber="812">Fig. 9A, C, D</figureCitation>
); in female reduced, but always present (e.g.,
<figureCitation id="0ADB2A012772780BFCAFF96938AA6292" box="[850,956,1725,1747]" captionStart="Fig" captionStartId="18.[113,148,1468,1490]" captionTargetBox="[112,1451,189,1452]" captionTargetId="figure-239@18.[106,1456,185,1456]" captionTargetPageId="18" captionText="Fig. 9. Hindlegs of Rhagovelia species, showing characteristic armatures pilosity omitted. A, R. matillanoi, new species, apterous male; B, R. matillanoi, new species, apterous female; C, R. abbreviata, new species, apterous male; D, R. abbreviata, new species, macropterous male; E, R. abbreviata, new species, apterous female; F, G, R. gapudi, new species, apterous males, two morphs; H, R. gapudi, new species, apterous female." figureDoi="http://doi.org/10.5281/zenodo.5350743" httpUri="https://zenodo.org/record/5350743/files/figure.png" pageId="2" pageNumber="812">Fig. 9B, E</figureCitation>
). (II.
<taxonomicName id="55E04D072772780BFC0BF96A3F826293" ID-CoL="4RZJ2" authorityName="J. Polhemus &amp; D. Polhemus" authorityYear="1988" box="[1014,1172,1725,1747]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="sarawakensis">
<emphasis id="A094EA962772780BFC0BF96A3F826293" box="[1014,1172,1725,1747]" italics="true" pageId="2" pageNumber="812">R. sarawakensis</emphasis>
</taxonomicName>
group) ................................4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="29142E212772780BFCD0F90E3EBC63FB" pageId="2" pageNumber="812">
<paragraph id="925F36842772780BFCD0F90E3EBC636B" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
<keyLead id="291195B12772780BFCD0F90E3EBC636B" pageId="2" pageNumber="812">4. Abdomen without spike-like setae. Sternites 36 (7) of male with sharp median carina. Mesofemur of female with short patch of long setae at depression on distal third....................5</keyLead>
</paragraph>
<paragraph id="925F36842772780BFCD0F8E53EBC63FB" blockId="2.[813,1454,660,2093]" pageId="2" pageNumber="812">
<keyLead id="291195B12772780BFCD0F8E53EBC63FB" pageId="2" pageNumber="812">
Connexiva (and in apterous specimens also anterior tergites) with numerous short and stout, spike-like setae (
<figureCitation id="0ADB2A012772780BFB10F8993E506322" box="[1261,1350,1869,1891]" captionStart="Fig" captionStartId="19.[136,171,1404,1426]" captionTargetBox="[136,1475,188,1387]" captionTargetId="figure-332@19.[132,1480,183,1392]" captionTargetPageId="19" captionText="Fig. 10. Abdominal terminalia of females of Rhagovelia species, pilosity partly omitted. AC, R. matillanoi, new species, arrow in A pointing to characteristic setae; DF, R. abbreviata, new species; GI, R. gapudi, new species, arrows in H, I pointing to concave ventral outline of gonocoxa. A, D, G, apterous specimen, dorsal aspect; B, E, H, apterous specimen, lateral aspect; C, F, I, macropterous specimen in 48 dealate, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5350745" httpUri="https://zenodo.org/record/5350745/files/figure.png" pageId="2" pageNumber="812">Fig. 10A</figureCitation>
). Sternites 35 of male without sharp median carina (a very obtuse carina present in
<taxonomicName id="55E04D072772780BFC44F85C3F4163DD" ID-CoL="4RZFV" authorityName="Zettel" authorityYear="1994" box="[953,1111,1927,1949]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="A094EA962772780BFC44F85C3F4163DD" box="[953,1111,1927,1949]" italics="true" pageId="2" pageNumber="812">R. palawanensis</emphasis>
</taxonomicName>
). Mesofemur of female with brush of long setae in distal half ......................................................6
</keyLead>
</paragraph>
</keyStep>
<paragraph id="925F36842772780AFCD0F81438106568" blockId="2.[813,1454,660,2093]" lastBlockId="3.[136,776,190,845]" lastPageId="3" lastPageNumber="813" pageId="2" pageNumber="812">
<keyLead id="291195B12772780AFCD0F81438106568" lastPageId="3" lastPageNumber="813" pageId="2" pageNumber="812">
<keyStep id="29142E212772780AFCD0F81438106568" lastPageId="3" lastPageNumber="813" pageId="2" pageNumber="812">
5. Very stout species (
<figureCitation id="0ADB2A012772780BFBE8F8143F966397" box="[1045,1152,1984,2006]" captionStart="Fig" captionStartId="9.[136,171,2064,2086]" captionTargetBox="[166,1445,189,2049]" captionTargetId="figure-8@9.[166,1445,189,2049]" captionTargetPageId="9" captionText="Fig. 3. Habitus of Rhagovelia species. R. estrella: A, apterous male; B, apterous female. R. abbreviata, new species: C, apterous male; D, macropterous dealate, male; E, apterous female; F, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350731" httpUri="https://zenodo.org/record/5350731/files/figure.png" pageId="2" pageNumber="812">Fig. 3C, E</figureCitation>
); in female, abdomen strongly shortened, upcurved in apterous morph. In male, median carina on sternite 7 reduced, weaker than on sternites 36. Species from North
<collectingRegion id="5024F8662772780BFC34F7C33F0C6C6C" box="[969,1050,2071,2093]" country="Philippines" name="Palawan" pageId="2" pageNumber="812">Palawan</collectingRegion>
......................
<taxonomicName id="55E04D072772780BFB5EF7CC3E316C6D" authority="Zettel &amp; Laciny &amp; Freitag, 2020" authorityName="Zettel &amp; Laciny &amp; Freitag" authorityYear="2020" box="[1187,1319,2071,2093]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="812" phylum="Arthropoda" rank="species" species="abbreviata" status="sp. nov.">
<emphasis id="A094EA962772780BFB5EF7CC3E316C6D" box="[1187,1319,2071,2093]" italics="true" pageId="2" pageNumber="812">R. abbreviata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BBA757ED2772780BFAC8F7C33EBC6C6C" box="[1333,1450,2071,2093]" pageId="2" pageNumber="812" rank="species">new species</taxonomicNameLabel>
Species of normal shape (
<figureCitation id="0ADB2A012773780AFE4AFF6A393E6492" box="[439,552,190,212]" captionStart="Fig" captionStartId="9.[136,171,2064,2086]" captionTargetBox="[166,1445,189,2049]" captionTargetId="figure-8@9.[166,1445,189,2049]" captionTargetPageId="9" captionText="Fig. 3. Habitus of Rhagovelia species. R. estrella: A, apterous male; B, apterous female. R. abbreviata, new species: C, apterous male; D, macropterous dealate, male; E, apterous female; F, macropterous female." figureDoi="http://doi.org/10.5281/zenodo.5350731" httpUri="https://zenodo.org/record/5350731/files/figure.png" pageId="3" pageNumber="813">Fig. 3A, B</figureCitation>
). Abdomen of female not shortened. In male, median carina on sternite 7 prominent, accentuated by paired impressions, stronger than on sternites 26. Species from Central
<collectingRegion id="5024F8662773780AFE4BFEC03911656B" box="[438,519,276,298]" country="Philippines" name="Palawan" pageId="3" pageNumber="813">Palawan</collectingRegion>
.........................
<taxonomicName id="55E04D072773780AFD5CFEC138106568" ID-CoL="4RZBJ" authorityName="Zettel" authorityYear="1994" box="[673,774,276,298]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="813" phylum="Arthropoda" rank="species" species="estrella">
<emphasis id="A094EA962773780AFD5CFEC138106568" box="[673,774,276,298]" italics="true" pageId="3" pageNumber="813">R. estrella</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<keyStep id="29142E212773780AFF75FEE5381065FB" pageId="3" pageNumber="813">
<paragraph id="925F36842773780AFF75FEE5381065C1" blockId="3.[136,776,190,845]" pageId="3" pageNumber="813">
<keyLead id="291195B12773780AFF75FEE5381065C1" pageId="3" pageNumber="813">
6. Mesocoxa, mesotrochanter, and metafemur entirely black (
<figureCitation id="0ADB2A012773780AFD1DFEE53BFD6522" captionStart="Fig" captionStartId="4.[113,148,1573,1595]" captionTargetBox="[113,1451,190,1558]" captionTargetId="figure-13@4.[113,1451,190,1558]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Rhagovelia species. A, R. minutissima, apterous male; B, R. minutissima, apterous female; C, R. palawanensis, apterous male; D, R. palawanensis, apterous female." figureDoi="http://doi.org/10.5281/zenodo.5350727" httpUri="https://zenodo.org/record/5350727/files/figure.png" pageId="3" pageNumber="813">Fig. 1C, D</figureCitation>
). Species from South and Central
<collectingRegion id="5024F8662773780AFDC2FE9A39866525" box="[575,656,334,356]" country="Philippines" name="Palawan" pageId="3" pageNumber="813">Palawan</collectingRegion>
................... ........................................................................
<taxonomicName id="55E04D072773780AFD9BFEBF381065C1" ID-CoL="4RZFV" authorityName="Zettel" authorityYear="1994" box="[614,774,363,384]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="813" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="A094EA962773780AFD9BFEBF381065C1" box="[614,774,363,384]" italics="true" pageId="3" pageNumber="813">R. palawanensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="925F36842773780AFF75FE5C381065FB" blockId="3.[136,776,190,845]" pageId="3" pageNumber="813">
<keyLead id="291195B12773780AFF75FE5C381065FB" pageId="3" pageNumber="813">
Mesocoxa, base of mesotrochanter, and base of metafemur yellow (
<figureCitation id="0ADB2A012773780AFF03FE703A6565FB" box="[254,371,420,442]" captionStart="Fig" captionStartId="4.[113,148,1573,1595]" captionTargetBox="[113,1451,190,1558]" captionTargetId="figure-13@4.[113,1451,190,1558]" captionTargetPageId="4" captionText="Fig. 1. Habitus of Rhagovelia species. A, R. minutissima, apterous male; B, R. minutissima, apterous female; C, R. palawanensis, apterous male; D, R. palawanensis, apterous female." figureDoi="http://doi.org/10.5281/zenodo.5350727" httpUri="https://zenodo.org/record/5350727/files/figure.png" pageId="3" pageNumber="813">Figs. 1A, B</figureCitation>
,
<figureCitation id="0ADB2A012773780AFE82FE703AAD65FB" box="[383,443,420,442]" captionStart="Fig" captionStartId="7.[136,171,1498,1520]" captionTargetBox="[136,1474,189,1482]" captionTargetId="figure-7@7.[136,1474,189,1482]" captionTargetPageId="7" captionText="Fig. 2. Habitus ofRhagovelia matillanoi, new species. A, apterous male; B, apterous female; C, macropterous dealate, male; D, macropterous dealate, female." figureDoi="http://doi.org/10.5281/zenodo.5350729" httpUri="https://zenodo.org/record/5350729/files/figure.png" pageId="3" pageNumber="813">2AD</figureCitation>
) ...................................................7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="29142E212773780AFF75FE163813670C" pageId="3" pageNumber="813">
<paragraph id="925F36842773780AFF75FE16381066C5" blockId="3.[136,776,190,845]" pageId="3" pageNumber="813">
<keyLead id="291195B12773780AFF75FE16381066C5" pageId="3" pageNumber="813">
7. Spike-like setae sparse, in apterous male scattered from mesonotum to tergite 5, in apterous female until tergite 2, rarely forming a transverse row on tergite 1. Segment 8 of male ventrally convex. Apterous female without distinct constriction at base of sternite 7. Female without distinct hair tuft on connexival corner. Species from Busuanga Island...... ...........................................................................
<taxonomicName id="55E04D072773780AFD88FDBB381066C5" ID-CoL="7WQ6N" authorityName="Hungerford &amp; Matsuda" authorityYear="1961" box="[629,774,623,644]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="813" phylum="Arthropoda" rank="species" species="minutissima">
<emphasis id="A094EA962773780AFD88FDBB381066C5" box="[629,774,623,644]" italics="true" pageId="3" pageNumber="813">R. minutissima</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="925F36842773780AFF75FD5F3813670C" blockId="3.[136,776,190,845]" pageId="3" pageNumber="813">
<keyLead id="291195B12773780AFF75FD5F3813670C" pageId="3" pageNumber="813">
Spike-like setae numerous, in apterous male from mesonotum to tergite 7, in apterous female until tergite 4, with distinct transverse row on tergite 1. Segment 8 of male ventrally flat. Apterous female with strong constriction at base of sternite 7. Female with distinct tuft of black hairs at connexival corner (
<figureCitation id="0ADB2A012773780AFF48FCCF3A1D6771" box="[181,267,795,817]" captionStart="Fig" captionStartId="19.[136,171,1404,1426]" captionTargetBox="[136,1475,188,1387]" captionTargetId="figure-332@19.[132,1480,183,1392]" captionTargetPageId="19" captionText="Fig. 10. Abdominal terminalia of females of Rhagovelia species, pilosity partly omitted. AC, R. matillanoi, new species, arrow in A pointing to characteristic setae; DF, R. abbreviata, new species; GI, R. gapudi, new species, arrows in H, I pointing to concave ventral outline of gonocoxa. A, D, G, apterous specimen, dorsal aspect; B, E, H, apterous specimen, lateral aspect; C, F, I, macropterous specimen in 48 dealate, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5350745" httpUri="https://zenodo.org/record/5350745/files/figure.png" pageId="3" pageNumber="813">Fig. 10A</figureCitation>
; indistinct in some macropterous individuals). Species from North
<collectingRegion id="5024F8662773780AFEDEFCE33A62670C" box="[291,372,823,845]" country="Philippines" name="Palawan" pageId="3" pageNumber="813">Palawan</collectingRegion>
.......................
<taxonomicName id="55E04D072773780AFDF9FCEC3995670D" authority="Zettel &amp; Laciny &amp; Freitag, 2020" authorityName="Zettel &amp; Laciny &amp; Freitag" authorityYear="2020" box="[516,643,823,845]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="813" phylum="Arthropoda" rank="species" species="matillanoi" status="sp. nov.">
<emphasis id="A094EA962773780AFDF9FCEC3995670D" box="[516,643,823,845]" italics="true" pageId="3" pageNumber="813">R. matillanoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BBA757ED2773780AFD6DFCEC3813670C" box="[656,773,823,845]" pageId="3" pageNumber="813" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>