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<mods:title id="E631C508B89A17120B6A31F4BDDE0258">New species and records of pollen and sap beetles for Iran (Coleoptera: Kateretidae, Nitidulidae)</mods:title>
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<emphasis id="A0D0FD7C7F1DFF87FF343093FCEA61DC" bold="true" box="[151,865,152,178]" pageId="2" pageNumber="371">
<taxonomicName id="55A45AED7F1DFF87FF343093FDEC61DC" authority="Jelinek, 1981" authorityName="Jelinek" authorityYear="1981" box="[151,615,152,178]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="ahriman">
<emphasis id="A0D0FD7C7F1DFF87FF343093FE2C61DC" bold="true" box="[151,423,152,178]" italics="true" pageId="2" pageNumber="371">Thymogethes ahriman</emphasis>
(
<bibRefCitation id="F6355C9F7F1DFF87FE143093FDD561DC" author="Jelinek" box="[439,606,152,178]" pageId="2" pageNumber="371" refString="Jelinek, J. (1981) Results of the Czechoslovak - Iranian entomological expeditions to Iran 1970 and 1973. Coleoptera: Nitidulidae. Acta Entomologica Musei Nationalis Pragae, 40, 105 - 119." type="journal article" year="1981">Jelínek, 1981</bibRefCitation>
)
</taxonomicName>
, species resurrected
</emphasis>
</heading>
</paragraph>
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<collectingMunicipality id="727FBB147F1DFF87FF3430D5FE9B6199" box="[151,272,222,247]" pageId="2" pageNumber="371">Examined</collectingMunicipality>
material.
<collectingCountry id="EAB361FE7F1DFF87FE2930D5FE5A6199" box="[394,465,222,247]" name="Iran" pageId="2" pageNumber="371">IRAN</collectingCountry>
</emphasis>
:
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,
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,
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,
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,
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</date>
/
</materialsCitation>
<materialsCitation id="22CC2B337F1DFF87FB5A30D4FBC26074" ID-GBIF-Occurrence="1424563924" collectingDate="1973-05-03" collectorName="Exp. Nat." country="Iran" location="Mus" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Mus">
<date id="E61A07AE7F1DFF87FB5A30D4FAD76199" box="[1273,1372,223,247]" pageId="2" pageNumber="371" value="1973-05-03">
<collectingDate id="F65EFE467F1DFF87FB5A30D4FAD76199" box="[1273,1372,223,247]" pageId="2" pageNumber="371" value="1973-05-03">3.V.1973</collectingDate>
</date>
, river bed and irrigation channel,
<collectedFrom id="21B862937F1DFF87FE423108FD456075" box="[481,718,258,283]" pageId="2" pageNumber="371">
on
<taxonomicName id="55A45AED7F1DFF87FDAA3109FD456075" box="[521,718,258,283]" class="Magnoliopsida" family="Lamiaceae" genus="Mentha" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="371" phylum="Tracheophyta" rank="species" species="aquatica">
<emphasis id="A0D0FD7C7F1DFF87FDAA3109FD456075" box="[521,718,258,283]" italics="true" pageId="2" pageNumber="371">Mentha aquatica</emphasis>
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, (loc. no. 186),
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<collectingRegion id="5060EF8C7F1DFF87FBB73108FBC26074" box="[1044,1097,259,282]" country="Turkey" name="Mus" pageId="2" pageNumber="371">Mus</collectingRegion>
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.
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<collectingRegion id="5060EF8C7F1DFF87FBFA3108FB226075" box="[1113,1193,259,283]" country="Czech Republic" name="Praha" pageId="2" pageNumber="371">Prague</collectingRegion>
,
<specimenCount id="84A2EAE77F1DFF87FB183108FA926075" box="[1211,1305,258,283]" pageId="2" pageNumber="371" type="male">3 males</specimenCount>
,
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(
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and
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;
<collectionCode id="F4B5B9AB7F1DFF87FE63312CFD996051" box="[448,530,295,319]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="371">NMPC</collectionCode>
,
<collectingCountry id="EAB361FE7F1DFF87FD82312CFDD46051" box="[545,607,295,319]" name="Central African Republic" pageId="2" pageNumber="371">CAR</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="22CC2B337F1DFF87FDD4312CFDA5600C" ID-GBIF-Occurrence="1424563932" collectingDate="1973-06-11" collectorName="Exp. Nat" country="Central African Republic" elevation="1125" latitude="29.783333" location="Tang-e Chogan" longLatPrecision="1225" longitude="51.633335" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Fars Province" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FDD4312CFC916051" box="[631,794,294,319]" country="Iran" name="Fars" pageId="2" pageNumber="371">Fars Province</collectingRegion>
,
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,
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<geoCoordinate id="F79047A97F1DFF87FBF4312DFB3C6051" box="[1111,1207,294,319]" direction="east" orientation="longitude" pageId="2" pageNumber="371" precision="925" value="51.633335">51°38´E</geoCoordinate>
,
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, 10/
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, (loc. no. 234),
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.
<materialsCitation id="22CC2B337F1DFF87FD9B3140FDE6600C" ID-GBIF-Occurrence="1424563930" box="[568,621,331,354]" country="Central African Republic" location="Mus" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Mus" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FD9B3140FDE6600C" box="[568,621,331,354]" country="Turkey" name="Mus" pageId="2" pageNumber="371">Mus</collectingRegion>
</materialsCitation>
.
<materialsCitation id="22CC2B337F1DFF87FDDB3140FC1D600D" ID-GBIF-Occurrence="1424563919" box="[632,918,330,355]" collectionCode="NMPC" country="Central African Republic" location="Prague" pageId="2" pageNumber="371" specimenCount="4" specimenCount-male="4" stateProvince="Prague" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FDDB3140FD42600D" box="[632,713,331,355]" country="Czech Republic" name="Praha" pageId="2" pageNumber="371">Prague</collectingRegion>
,
<specimenCount id="84A2EAE77F1DFF87FD703140FCA2600D" box="[723,809,330,355]" pageId="2" pageNumber="371" type="male">4 males</specimenCount>
(
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)
</materialsCitation>
;
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<collectingRegion id="5060EF8C7F1DFF87FC023141FB13600D" box="[929,1176,330,355]" country="Iran" name="Mazandaran" pageId="2" pageNumber="371">Mazandaran Province</collectingRegion>
, E
<collectingRegion id="5060EF8C7F1DFF87FB183141FA82600D" box="[1211,1289,330,355]" country="Iran" name="Alborz" pageId="2" pageNumber="371">Alborz</collectingRegion>
Mts,
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,
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,
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,
<date id="E61A07AE7F1DFF87FE1C3164FDCD60E9" box="[447,582,367,391]" pageId="2" pageNumber="371" value="1970-08-02">
<collectingDate id="F65EFE467F1DFF87FE1C3164FDCD60E9" box="[447,582,367,391]" pageId="2" pageNumber="371" value="1970-08-02">2.VIII.1970</collectingDate>
</date>
, (loc. no. 81),
<collectorName id="3F5144B87F1DFF87FD4B3164FCC460E8" box="[744,847,367,390]" pageId="2" pageNumber="371">Exp. Nat</collectorName>
</materialsCitation>
.
<materialsCitation id="22CC2B337F1DFF87FCFA3164FC0560E8" ID-GBIF-Occurrence="1424563920" box="[857,910,367,390]" country="Central African Republic" location="Mus" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Mus" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FCFA3164FC0560E8" box="[857,910,367,390]" country="Turkey" name="Mus" pageId="2" pageNumber="371">Mus</collectingRegion>
</materialsCitation>
.
<materialsCitation id="22CC2B337F1DFF87FC3A3164FB2660E9" ID-GBIF-Occurrence="1424563917" box="[921,1197,366,391]" collectionCode="NMPC" country="Central African Republic" location="Prague" pageId="2" pageNumber="371" specimenCount="1" specimenCount-male="1" stateProvince="Prague" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FC3A3164FC6260E9" box="[921,1001,367,391]" country="Czech Republic" name="Praha" pageId="2" pageNumber="371">Prague</collectingRegion>
,
<specimenCount id="84A2EAE77F1DFF87FC563164FBCB60E9" box="[1013,1088,366,391]" pageId="2" pageNumber="371" type="male">1 male</specimenCount>
(
<collectionCode id="F4B5B9AB7F1DFF87FBEC3164FB2E60E9" box="[1103,1189,367,391]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="371">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22CC2B337F1DFF87FB1B3164FA1260C4" ID-GBIF-Occurrence="1424563925" collectingDate="1973-05-30" collectingDateMax="1973-06-05" collectingDateMin="1973-05-30" collectorName="Exp. Nat" country="Central African Republic" latitude="29.15" location="Mian Jangal" longLatPrecision="1228" longitude="53.7" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Fars Province" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FB1B3164FADC60E9" box="[1208,1367,366,391]" country="Iran" name="Fars" pageId="2" pageNumber="371">Fars Province</collectingRegion>
,
<location id="977B77B57F1DFF87FAC23165FF6960C5" LSID="urn:lsid:plazi:treatment:1A0D90787F1DFF87FF343093FC8E6769:977B77B57F1DFF87FAC23165FF6960C5" country="Central African Republic" latitude="29.15" longLatPrecision="1228" longitude="53.7" name="Mian Jangal" pageId="2" pageNumber="371" stateProvince="Fars Province">Mian Jangal</location>
,
<geoCoordinate id="F79047A97F1DFF87FF4C3198FEDB60C5" box="[239,336,403,427]" direction="north" orientation="latitude" pageId="2" pageNumber="371" precision="925" value="29.15">29°09´N</geoCoordinate>
<geoCoordinate id="F79047A97F1DFF87FEFA3199FE3360C5" box="[345,440,402,427]" direction="east" orientation="longitude" pageId="2" pageNumber="371" precision="925" value="53.7">53 42´E</geoCoordinate>
,
<date id="E61A07AE7F1DFF87FE653198FD0660C5" box="[454,653,402,427]" pageId="2" pageNumber="371" value="1973-05-30" valueMax="1973-06-05" valueMin="1973-05-30">
<collectingDate id="F65EFE467F1DFF87FE653198FD0660C5" box="[454,653,402,427]" pageId="2" pageNumber="371" value="1973-05-30" valueMax="1973-06-05" valueMin="1973-05-30">30.V. 5.VI.1973</collectingDate>
</date>
,
<collectedFrom id="21B862937F1DFF87FD383198FC0A60C5" box="[667,897,402,427]" pageId="2" pageNumber="371">
on
<taxonomicName id="55A45AED7F1DFF87FD633199FC0A60C5" box="[704,897,402,427]" class="Magnoliopsida" family="Lamiaceae" genus="Mentha" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="371" phylum="Tracheophyta" rank="species" species="aquatica">
<emphasis id="A0D0FD7C7F1DFF87FD633199FC0A60C5" box="[704,897,402,427]" italics="true" pageId="2" pageNumber="371">Mentha aquatica</emphasis>
</taxonomicName>
</collectedFrom>
and/or
<taxonomicName id="55A45AED7F1DFF87FC7F319FFBFA60C5" box="[988,1137,402,427]" class="Magnoliopsida" family="Lamiaceae" genus="Mentha" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="371" phylum="Tracheophyta" rank="species" species="longifolia">
<emphasis id="A0D0FD7C7F1DFF87FC7F319FFBFA60C5" box="[988,1137,402,427]" italics="true" pageId="2" pageNumber="371">M. longifolia</emphasis>
</taxonomicName>
, (loc. no. 223),
<collectorName id="3F5144B87F1DFF87FA8C3198FA1260C4" box="[1327,1433,403,426]" pageId="2" pageNumber="371">Exp. Nat</collectorName>
</materialsCitation>
.
<materialsCitation id="22CC2B337F1DFF87FF3431BCFF4760A0" ID-GBIF-Occurrence="1424563918" box="[151,204,439,462]" country="Central African Republic" location="Mus" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Mus" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FF3431BCFF4760A0" box="[151,204,439,462]" country="Turkey" name="Mus" pageId="2" pageNumber="371">Mus</collectingRegion>
</materialsCitation>
.
<materialsCitation id="22CC2B337F1DFF87FF7B31BCFDE060A1" ID-GBIF-Occurrence="1424563927" box="[216,619,438,463]" collectionCode="NMPC" country="Central African Republic" location="Prague" pageId="2" pageNumber="371" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Prague" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FF7B31BCFEA260A1" box="[216,297,439,463]" country="Czech Republic" name="Praha" pageId="2" pageNumber="371">Prague</collectingRegion>
,
<specimenCount id="84A2EAE77F1DFF87FE9731BCFE0460A1" box="[308,399,438,463]" pageId="2" pageNumber="371" type="male">3 males</specimenCount>
,
<specimenCount id="84A2EAE77F1DFF87FE3831BCFE7660A1" box="[411,509,438,463]" pageId="2" pageNumber="371" type="female">1 female</specimenCount>
(
<collectionCode id="F4B5B9AB7F1DFF87FDAE31BCFDE860A1" box="[525,611,439,463]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="371">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22CC2B337F1DFF87FDD431BDFCFB609D" ID-GBIF-Occurrence="1424563914" collectingDate="1970-07-01" collectionCode="NMPC" country="Central African Republic" elevation="2000" latitude="32.816666" location="Singerd" longLatPrecision="1208" longitude="50.433334" municipality="Eskandari" pageId="2" pageNumber="371" specimenCount="30" specimenCount-female="24" specimenCount-male="6" stateProvince="Esfahan Province" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FDD431BDFCB460A1" box="[631,831,438,463]" country="Iran" name="Esfahan" pageId="2" pageNumber="371">Esfahan Province</collectingRegion>
,
<collectingMunicipality id="727FBB147F1DFF87FCEF31BDFC3460A1" box="[844,959,438,463]" pageId="2" pageNumber="371">Eskandari</collectingMunicipality>
, near
<location id="977B77B57F1DFF87FBA031BDFBD660A1" LSID="urn:lsid:plazi:treatment:1A0D90787F1DFF87FF343093FC8E6769:977B77B57F1DFF87FBA031BDFBD660A1" box="[1027,1117,438,463]" country="Central African Republic" latitude="32.816666" longLatPrecision="1208" longitude="50.433334" municipality="Eskandari" name="Singerd" pageId="2" pageNumber="371" stateProvince="Esfahan Province">Singerd</location>
,
<geoCoordinate id="F79047A97F1DFF87FBC931BCFB4060A1" box="[1130,1227,439,463]" direction="north" orientation="latitude" pageId="2" pageNumber="371" precision="925" value="32.816666">32°49´N</geoCoordinate>
<geoCoordinate id="F79047A97F1DFF87FB7131BDFAB860A1" box="[1234,1331,438,463]" direction="east" orientation="longitude" pageId="2" pageNumber="371" precision="925" value="50.433334">50°26´E</geoCoordinate>
,
<quantity id="555C8C8B7F1DFF87FA9C31BCFA1C60A0" box="[1343,1431,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="371" unit="m" value="2000.0">
<elevation id="1989C65D7F1DFF87FA9C31BCFA1C60A0" box="[1343,1431,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="371" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="E61A07AE7F1DFF87FF3431D0FE9F609D" box="[151,276,475,499]" pageId="2" pageNumber="371" value="1970-07-01">
<collectingDate id="F65EFE467F1DFF87FF3431D0FE9F609D" box="[151,276,475,499]" pageId="2" pageNumber="371" value="1970-07-01">1.VII.1970</collectingDate>
</date>
, (loc. no. 36)
<specimenCount id="84A2EAE77F1DFF87FE6731D1FDA9609D" box="[452,546,474,499]" pageId="2" pageNumber="371" type="male">6 males</specimenCount>
,
<specimenCount id="84A2EAE77F1DFF87FD9131D0FD3A609D" box="[562,689,474,499]" pageId="2" pageNumber="371" type="female">24 females</specimenCount>
(
<collectionCode id="F4B5B9AB7F1DFF87FD6631D0FC92609D" box="[709,793,475,499]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="371">NMPC</collectionCode>
,
<collectingCountry id="EAB361FE7F1DFF87FC8A31D0FCEC609D" box="[809,871,475,499]" name="Central African Republic" pageId="2" pageNumber="371">CAR</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="22CC2B337F1DFF87FC2331D0FBD66355" ID-GBIF-Occurrence="1424563928" collectingDate="2014-06-14" collectionCode="HMIM, NMPC" collectorName="Zagros Mts, SE &amp; P. Audisio &amp; Huds." country="Central African Republic" county="Dorud" elevation="1695" latitude="33.418056" location="Chamnar" longLatPrecision="20" longitude="49.162224" municipality="Oshtoran Kuh" pageId="2" pageNumber="371" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Lorestan Province" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FC2331D0FBDE609D" box="[896,1109,474,499]" country="Iran" name="Lorestan" pageId="2" pageNumber="371">Lorestan Province</collectingRegion>
,
<collectorName id="3F5144B87F1DFF87FBC631D0FA95609D" box="[1125,1310,475,499]" pageId="2" pageNumber="371">Zagros Mts, SE</collectorName>
of
<collectingCounty id="7B7A59E27F1DFF87FAEE31D1FA1C609D" box="[1357,1431,474,499]" pageId="2" pageNumber="371">Dorud</collectingCounty>
,
<collectingMunicipality id="727FBB147F1DFF87FF3431F5FEB76379" box="[151,316,510,535]" pageId="2" pageNumber="371">Oshtoran Kuh</collectingMunicipality>
, small village near
<location id="977B77B57F1DFF87FD8A31F5FD1A6379" LSID="urn:lsid:plazi:treatment:1A0D90787F1DFF87FF343093FC8E6769:977B77B57F1DFF87FD8A31F5FD1A6379" box="[553,657,510,535]" country="Central African Republic" county="Dorud" latitude="33.418056" longLatPrecision="20" longitude="49.162224" municipality="Oshtoran Kuh" name="Chamnar" pageId="2" pageNumber="371" stateProvince="Lorestan Province">Chamnar</location>
,
<geoCoordinate id="F79047A97F1DFF87FD0231F5FCA86379" box="[673,803,510,535]" direction="north" orientation="latitude" pageId="2" pageNumber="371" precision="15" value="33.418056">33°25'05''N</geoCoordinate>
<geoCoordinate id="F79047A97F1DFF87FC9331F4FC396379" box="[816,946,511,535]" direction="east" orientation="longitude" pageId="2" pageNumber="371" precision="15" value="49.162224">49°09'44''E</geoCoordinate>
,
<quantity id="555C8C8B7F1DFF87FC6131F5FB966378" box="[962,1053,510,535]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="2" pageNumber="371" unit="m" value="1695.0">
<elevation id="1989C65D7F1DFF87FC6131F5FB966378" box="[962,1053,510,535]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="2" pageNumber="371" unit="m" value="1695.0">1695 m</elevation>
</quantity>
,
<date id="E61A07AE7F1DFF87FB8D31F4FB3B6379" box="[1070,1200,511,535]" pageId="2" pageNumber="371" value="2014-06-14">
<collectingDate id="F65EFE467F1DFF87FB8D31F4FB3B6379" box="[1070,1200,511,535]" pageId="2" pageNumber="371" value="2014-06-14">14.VI.2014</collectingDate>
</date>
, leg.
<collectorName id="3F5144B87F1DFF87FB5631F4FAFA6379" box="[1269,1393,510,535]" pageId="2" pageNumber="371">P. Audisio</collectorName>
,
<collectedFrom id="21B862937F1DFF87FA2331F4FEEB6355" pageId="2" pageNumber="371">
on
<taxonomicName id="55A45AED7F1DFF87FF343229FEEB6355" box="[151,352,546,571]" class="Magnoliopsida" family="Lamiaceae" genus="Mentha" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="371" phylum="Tracheophyta" rank="species" species="longifolia">
<emphasis id="A0D0FD7C7F1DFF87FF343229FEEB6355" box="[151,352,546,571]" italics="true" pageId="2" pageNumber="371">Mentha longifolia</emphasis>
</taxonomicName>
</collectedFrom>
(L.)
<collectorName id="3F5144B87F1DFF87FE3A3229FE506355" box="[409,475,546,571]" pageId="2" pageNumber="371">Huds.</collectorName>
(
<taxonomicName id="55A45AED7F1DFF87FE4A3229FDE36355" box="[489,616,546,571]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="371" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
),
<specimenCount id="84A2EAE77F1DFF87FDDF3228FD5F6355" box="[636,724,546,571]" pageId="2" pageNumber="371" type="male">4 males</specimenCount>
,
<specimenCount id="84A2EAE77F1DFF87FD433229FCC76355" box="[736,844,546,571]" pageId="2" pageNumber="371" type="female">5 females</specimenCount>
(
<collectingCountry id="EAB361FE7F1DFF87FCF83228FC126355" box="[859,921,547,571]" name="Central African Republic" pageId="2" pageNumber="371">CAR</collectingCountry>
,
<collectionCode id="F4B5B9AB7F1DFF87FC003228FC7E6354" box="[931,1013,547,570]" country="Spain" httpUri="http://grbio.org/cool/gj99-xj2c" name="Jard<72> Bot<6F>nic Marimurtra" pageId="2" pageNumber="371">HMIM</collectionCode>
,
<collectionCode id="F4B5B9AB7F1DFF87FBA33228FBDE6355" box="[1024,1109,547,571]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="371">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="22CC2B337F1DFF87FBCB3229FE8063C9" ID-GBIF-Occurrence="1424563921" collectingDate="2008-05-23" collectionCode="NKME" collectorName="Muhle &amp; Refer &amp; Jelinek &amp; Audisio &amp; Lason &amp; Ghahari" country="Central African Republic" elevation="1650" location="Shir-Kuh" pageId="2" pageNumber="371" specimenCount="1" stateProvince="Yazd Province" typeStatus="holotype">
<collectingRegion id="5060EF8C7F1DFF87FBCB3229FA856355" box="[1128,1294,546,571]" country="Iran" name="Yazd" pageId="2" pageNumber="371">Yazd Province</collectingRegion>
,
<location id="977B77B57F1DFF87FABB3229FA086355" LSID="urn:lsid:plazi:treatment:1A0D90787F1DFF87FF343093FC8E6769:977B77B57F1DFF87FABB3229FA086355" box="[1304,1411,546,571]" country="Central African Republic" name="Shir-Kuh" pageId="2" pageNumber="371" stateProvince="Yazd Province">Shir-Kuh</location>
,
<locationDeviation id="83A9384C7F1DFF87FA2D3229FE8C6331" pageId="2" pageNumber="371">
<quantity id="555C8C8B7F1DFF87FA2D3229FF306331" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="371" unit="km" value="5.0">5 km</quantity>
S Taft
</locationDeviation>
,
<quantity id="555C8C8B7F1DFF87FEB2324DFE256330" box="[273,430,582,607]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="2" pageNumber="371" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<elevation id="1989C65D7F1DFF87FEB2324DFE256330" box="[273,430,582,607]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="2" pageNumber="371" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">16001700 m</elevation>
</quantity>
,
<date id="E61A07AE7F1DFF87FE1A324CFDA56331" box="[441,558,583,607]" pageId="2" pageNumber="371" value="2008-05-23">
<collectingDate id="F65EFE467F1DFF87FE1A324CFDA56331" box="[441,558,583,607]" pageId="2" pageNumber="371" value="2008-05-23">23.V.2008</collectingDate>
</date>
, leg.
<collectorName id="3F5144B87F1DFF87FDCA324DFD3E6331" box="[617,693,582,607]" pageId="2" pageNumber="371">Mühle</collectorName>
, 1 ex. (
<collectionCode id="F4B5B9AB7F1DFF87FCA7324CFCD66331" box="[772,861,583,607]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13752" name="Naturkundemuseum Erfurt" pageId="2" pageNumber="371">NKME</collectionCode>
).
<collectorName id="3F5144B87F1DFF87FCCC324DFC266331" box="[879,941,582,607]" pageId="2" pageNumber="371">Refer</collectorName>
to
<bibRefCitation id="F6355C9F7F1DFF87FC73324DFBF86331" author="Jelinek" box="[976,1139,582,607]" pageId="2" pageNumber="371" refString="Jelinek, J. (1981) Results of the Czechoslovak - Iranian entomological expeditions to Iran 1970 and 1973. Coleoptera: Nitidulidae. Acta Entomologica Musei Nationalis Pragae, 40, 105 - 119." type="journal article" year="1981">
<collectorName id="3F5144B87F1DFF87FC73324DFB946331" box="[976,1055,582,607]" pageId="2" pageNumber="371">Jelínek</collectorName>
(1981)
</bibRefCitation>
,
<bibRefCitation id="F6355C9F7F1DFF87FBDD324DFAB36331" author="Audisio" box="[1150,1336,582,607]" pageId="2" pageNumber="371" refString="Audisio, P. (1993 b) Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. 32. Calderini ed., Bologna, XVI + 971 pp." type="book" year="1993" yearSuffix="b">
<collectorName id="3F5144B87F1DFF87FBDD324DFB5C6331" box="[1150,1239,582,607]" pageId="2" pageNumber="371">Audisio</collectorName>
(1993b)
</bibRefCitation>
,
<bibRefCitation id="F6355C9F7F1DFF87FAE0324DFEAE63ED" pageId="2" pageNumber="371" refString="Audisio, P., Jelinek, J., Mariotti, A. &amp; De Biase, A. (2000) The Coleoptera Nitidulidae and Kateretidae from Anatolian, Caucasian and Middle East regions. Biogeographia, 21, 241 - 354." type="journal article">
<collectorName id="3F5144B87F1DFF87FAE0324DFA176331" box="[1347,1436,582,607]" pageId="2" pageNumber="371">Audisio</collectorName>
<emphasis id="A0D0FD7C7F1DFF87FF343267FF5963ED" box="[151,210,618,643]" italics="true" pageId="2" pageNumber="371">et al.</emphasis>
(2000)
</bibRefCitation>
and
<bibRefCitation id="F6355C9F7F1DFF87FEC33260FDF063ED" author="Lason" box="[352,635,618,643]" pageId="2" pageNumber="371" refString="Lason, A. &amp; Ghahari, H. (2013) A checklist of the Kateretidae and Nitidulidae of Iran (Coleoptera: Cucujoidea). Zootaxa, 3746, 101 - 122. http: // dx. doi. org / 10.11646 / zootaxa. 3746.1.4" type="journal article" year="2013">
<collectorName id="3F5144B87F1DFF87FEC33260FE2F63ED" box="[352,420,619,643]" pageId="2" pageNumber="371">Lasoń</collectorName>
&amp;
<collectorName id="3F5144B87F1DFF87FE6E3261FDAC63ED" box="[461,551,618,643]" pageId="2" pageNumber="371">Ghahari</collectorName>
(2013)
</bibRefCitation>
for references where part of the above listed material was previously published.
</materialsCitation>
</paragraph>
<paragraph id="921B216E7F1DFF87FF6432B9FE696259" blockId="2.[151,1437,222,1543]" pageId="2" pageNumber="371">
This taxon, described as
<taxonomicName id="55A45AED7F1DFF87FE4C32B9FD5F63A5" box="[495,724,690,715]" class="Insecta" family="Nitidulidae" genus="Meligethes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="ahriman">
<emphasis id="A0D0FD7C7F1DFF87FE4C32B9FD5F63A5" box="[495,724,690,715]" italics="true" pageId="2" pageNumber="371">Meligethes ahriman</emphasis>
</taxonomicName>
by
<bibRefCitation id="F6355C9F7F1DFF87FCA432B9FC2663A5" author="Jelinek" box="[775,941,690,715]" pageId="2" pageNumber="371" refString="Jelinek, J. (1981) Results of the Czechoslovak - Iranian entomological expeditions to Iran 1970 and 1973. Coleoptera: Nitidulidae. Acta Entomologica Musei Nationalis Pragae, 40, 105 - 119." type="journal article" year="1981">Jelínek (1981)</bibRefCitation>
from
<collectingCountry id="EAB361FE7F1DFF87FC5832B8FBA063A4" box="[1019,1067,691,714]" name="Iran" pageId="2" pageNumber="371">Iran</collectingCountry>
, was tentatively and doubtfully considered a possible synonym of
<taxonomicName id="55A45AED7F1DFF87FDBE32DDFC6E6381" authority="Easton, 1957" authorityName="Easton" authorityYear="1957" box="[541,997,726,751]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="A0D0FD7C7F1DFF87FDBE32DDFCB36381" box="[541,824,726,751]" italics="true" pageId="2" pageNumber="371">Thymogethes klapperichi</emphasis>
(
<bibRefCitation id="F6355C9F7F1DFF87FCE432DCFC566381" author="Easton" box="[839,989,726,751]" pageId="2" pageNumber="371" refString="Easton, A. M. (1957) The Meligethes (Col., Nitidulidae) of Afghanistan. Entomologist's Monthly Magazine, 92, 385 - 401." type="journal article" year="1957">Easton, 1957</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="EAB361FE7F1DFF87FB8F32DDFB3E6381" box="[1068,1205,726,751]" name="Afghanistan" pageId="2" pageNumber="371">Afghanistan</collectingCountry>
and Middle Asia by
<bibRefCitation id="F6355C9F7F1DFF87FF3432F1FED8627D" author="Audisio" box="[151,339,762,787]" pageId="2" pageNumber="371" refString="Audisio, P. (1993 b) Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. 32. Calderini ed., Bologna, XVI + 971 pp." type="book" year="1993" yearSuffix="b">Audisio (1993b)</bibRefCitation>
, and later reported as
<taxonomicName id="55A45AED7F1DFF87FDF732F7FD7B627D" box="[596,752,762,787]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="A0D0FD7C7F1DFF87FDF732F7FD7B627D" box="[596,752,762,787]" italics="true" pageId="2" pageNumber="371">T. klapperichi</emphasis>
</taxonomicName>
by
<bibRefCitation id="F6355C9F7F1DFF87FCBD32F1FB8C627D" author="Audisio" box="[798,1031,762,787]" pageId="2" pageNumber="371" refString="Audisio, P., Jelinek, J., Mariotti, A. &amp; De Biase, A. (2000) The Coleoptera Nitidulidae and Kateretidae from Anatolian, Caucasian and Middle East regions. Biogeographia, 21, 241 - 354." type="journal article" year="2000">
Audisio
<emphasis id="A0D0FD7C7F1DFF87FC2332F7FC31627D" box="[896,954,762,787]" italics="true" pageId="2" pageNumber="371">et al.</emphasis>
(2000
</bibRefCitation>
,
<bibRefCitation id="F6355C9F7F1DFF87FBB732F0FBD7627D" author="Audisio" box="[1044,1116,763,787]" pageId="2" pageNumber="371" refString="Audisio, P., Cline, A. R., De Biase, A., Antonini, G., Mancini, E., Trizzino, M., Costantini, L., Strika, S., Lamanna, F. &amp; Cerretti, P. (2009 c) Preliminary re-examination of genus-level taxonomy of the pollen beetle subfamily Meligethinae (Coleoptera: Nitidulidae). Acta Entomologica Musei Nationalis Pragae, 49 (2), 341 - 504." type="journal article" year="2009" yearSuffix="c">2009c</bibRefCitation>
),
<bibRefCitation id="F6355C9F7F1DFF87FBD132F1FA1C627D" author="Jelinek" box="[1138,1431,762,787]" pageId="2" pageNumber="371" refString="Jelinek, J. &amp; Audisio, P. (2007) Family Nitidulidae. In: Lobl, I. &amp; Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Fol. 4: Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, pp. 459 - 491." type="journal article" year="2007">Jelínek &amp; Audisio (2007)</bibRefCitation>
, and
<bibRefCitation id="F6355C9F7F1DFF87FF653314FE566259" author="Lason" box="[198,477,798,823]" pageId="2" pageNumber="371" refString="Lason, A. &amp; Ghahari, H. (2013) A checklist of the Kateretidae and Nitidulidae of Iran (Coleoptera: Cucujoidea). Zootaxa, 3746, 101 - 122. http: // dx. doi. org / 10.11646 / zootaxa. 3746.1.4" type="journal article" year="2013">Lasoń &amp; Ghahari (2013)</bibRefCitation>
.
</paragraph>
<paragraph id="921B216E7F1DFF87FF643349FE516419" blockId="2.[151,1437,222,1543]" pageId="2" pageNumber="371">
Comparing the recently collected specimens from W
<collectingCountry id="EAB361FE7F1DFF87FCE33348FCFB6234" box="[832,880,835,858]" name="Iran" pageId="2" pageNumber="371">Iran</collectingCountry>
, with available material in
<collectionCode id="F4B5B9AB7F1DFF87FB193348FA826235" box="[1210,1289,835,859]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="371">NMPC</collectionCode>
allowed the authors to conclude that Iranian populations of
<taxonomicName id="55A45AED7F1DFF87FD12336CFCA06211" box="[689,811,870,895]" class="Insecta" family="Nitidulidae" genus="Meligethes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="ahriman">
<emphasis id="A0D0FD7C7F1DFF87FD12336CFCA06211" box="[689,811,870,895]" italics="true" pageId="2" pageNumber="371">T. ahriman</emphasis>
</taxonomicName>
do not exhibit any significant N/S or E/W variation in the shape of male genitalia, linking them with the Afghan populations of
<taxonomicName id="55A45AED7F1DFF87FC483387FB0362CD" box="[1003,1160,906,931]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="A0D0FD7C7F1DFF87FC483387FB0362CD" box="[1003,1160,906,931]" italics="true" pageId="2" pageNumber="371">T. klapperichi</emphasis>
</taxonomicName>
. Iranian populations, in fact, with the possible exception of a single specimen from southern Zagros Mts drawn in
<bibRefCitation id="F6355C9F7F1DFF87FB0933A5FEDF6285" author="Audisio" pageId="2" pageNumber="371" refString="Audisio, P. (1993 b) Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. 32. Calderini ed., Bologna, XVI + 971 pp." type="book" year="1993" yearSuffix="b">
Audisio (1993b; Figs 129
<emphasis id="A0D0FD7C7F1DFF87FF6F33DFFF656285" box="[204,238,980,1003]" italics="true" pageId="2" pageNumber="371">q-r</emphasis>
therein)
</bibRefCitation>
, probably erroneously attributed to
<taxonomicName id="55A45AED7F1DFF87FD5433DFFC1D6285" box="[759,918,978,1003]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="A0D0FD7C7F1DFF87FD5433DFFC1D6285" box="[759,918,978,1003]" italics="true" pageId="2" pageNumber="371">T. klapperichi</emphasis>
</taxonomicName>
(an immature specimen showing apparently more elongate but poorly sclerotized tegmen), appear on the contrary to be rather uniform, and also the specimens recently collected along the central-southern portion of the Zagros seem to differ distinctly from central and NE Afghan populations (
<bibRefCitation id="F6355C9F7F1DFF87FE203434FD9A6539" author="Easton" box="[387,529,1086,1111]" pageId="2" pageNumber="371" refString="Easton, A. M. (1957) The Meligethes (Col., Nitidulidae) of Afghanistan. Entomologist's Monthly Magazine, 92, 385 - 401." type="journal article" year="1957">Easton 1957</bibRefCitation>
) of the true
<emphasis id="A0D0FD7C7F1DFF87FD393434FCB66539" box="[666,829,1086,1111]" italics="true" pageId="2" pageNumber="371">
<taxonomicName id="55A45AED7F1DFF87FD393434FCB26539" box="[666,825,1086,1111]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="klapperichi">T. klapperichi</taxonomicName>
.
</emphasis>
Therefore, we are now convinced that the above cited Iranian specimen previously tentatively attributed to
<taxonomicName id="55A45AED7F1DFF87FD58346FFC126515" box="[763,921,1122,1147]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="A0D0FD7C7F1DFF87FD58346FFC126515" box="[763,921,1122,1147]" italics="true" pageId="2" pageNumber="371">T. klapperichi</emphasis>
</taxonomicName>
belongs to
<taxonomicName id="55A45AED7F1DFF87FB86346FFB286515" box="[1061,1187,1122,1147]" class="Insecta" family="Nitidulidae" genus="Meligethes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="ahriman">
<emphasis id="A0D0FD7C7F1DFF87FB86346FFB286515" box="[1061,1187,1122,1147]" italics="true" pageId="2" pageNumber="371">T. ahriman</emphasis>
</taxonomicName>
, and this latter taxon should be resurrected to a specific rank. This species appears to exhibit a rather wide geographic range from the Zagros Chain and the surrounding mountain systems to the E Elborz Mts (Fig. 25). We summarized in
<figureCitation id="0A9F3DEB7F1DFF87FAAE34A1FAE665AD" box="[1293,1389,1194,1219]" captionStart="FIGURES 3 10" captionStartId="4.[151,264,1624,1647]" captionTargetBox="[226,1362,205,1593]" captionTargetId="figure@4.[212,1374,193,1604]" captionTargetPageId="4" captionText="FIGURES 3 10. Male genitalia (median lobe of aedeagus and tegmen) of: 3 4. Thymogethes ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 5 6. T. klapperichi (Easton, 1957) (male from Pakistan, Madaglasht); 7 8. T. kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 9 10. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3 6); = 0.20 mm (Figs 7 10)." httpUri="https://zenodo.org/record/242295/files/figure.png" pageId="2" pageNumber="371">Figs 35</figureCitation>
and in the key to identification the diagnostic differences between these two taxa, as well as those between
<taxonomicName id="55A45AED7F1DFF87FA9234C4FECA6465" authority="Erichson, 1845" authorityName="Erichson" authorityYear="1845" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="egenus">
<emphasis id="A0D0FD7C7F1DFF87FA9234C4FA176588" box="[1329,1436,1231,1254]" italics="true" pageId="2" pageNumber="371">T. egenus</emphasis>
<bibRefCitation id="F6355C9F7F1DFF87FF3434F9FECA6465" author="Erichson" box="[151,321,1266,1291]" pageId="2" pageNumber="371" refString="Erichson, W. F. (1845) [I., II. Lieferungen]. In: Erichson, W. F. (Ed.), Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Dritter Band. Nikolaische Buchhandlung, Berlin, pp. 1 - 320. [issued in parts: pp. 1 - 320: 1845; pp. 321 - 480: 1846; pp. 481 - 800: 1847; pp. 801 - 968: 1848]." type="journal article" year="1845">Erichson, 1845</bibRefCitation>
</taxonomicName>
(s.l.: see comments below),
<taxonomicName id="55A45AED7F1DFF87FD2734FFFCFA6465" authority="Easton, 1954" authorityName="Easton" authorityYear="1954" box="[644,881,1266,1291]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="otini">
<emphasis id="A0D0FD7C7F1DFF87FD2734FFFD586465" box="[644,723,1268,1291]" italics="true" pageId="2" pageNumber="371">T. otini</emphasis>
<bibRefCitation id="F6355C9F7F1DFF87FD7834F8FCFA6465" author="Easton" box="[731,881,1266,1291]" pageId="2" pageNumber="371" refString="Easton, A. M. (1954) A revision of the lugubris complex in the genus Meligethes Stephens (Col. Nitidulidae). Transactions of the Royal Entomological Society, 105 (16), 373 - 392. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1954. tb 00769. x" type="journal article" year="1954">Easton, 1954</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="55A45AED7F1DFF87FC0F34FFFBD66465" box="[940,1117,1266,1291]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="khorasanicus" status="sp. nov.">
<emphasis id="A0D0FD7C7F1DFF87FC0F34FFFBD66465" box="[940,1117,1266,1291]" italics="true" pageId="2" pageNumber="371">T. khorasanicus</emphasis>
</taxonomicName>
<emphasis id="A0D0FD7C7F1DFF87FBC534F9FB4B6465" bold="true" box="[1126,1216,1266,1291]" pageId="2" pageNumber="371">
<taxonomicNameLabel id="BBE340077F1DFF87FBC534F9FB4B6465" box="[1126,1216,1266,1291]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
See
<figureCitation id="0A9F3DEB7F1DFF87FB5434F9FAEC6465" box="[1271,1383,1266,1291]" captionStart-0="FIGURES 3 10" captionStart-1="FIGURES 11 18" captionStart-2="FIGURES 19 22" captionStart-3="FIGURES 23 24" captionStartId-0="4.[151,264,1624,1647]" captionStartId-1="6.[151,264,1565,1588]" captionStartId-2="7.[151,264,1202,1225]" captionStartId-3="7.[151,264,1914,1937]" captionTargetBox-0="[226,1362,205,1593]" captionTargetBox-1="[262,1351,179,1540]" captionTargetBox-2="[158,1430,193,1168]" captionTargetBox-3="[157,1429,1373,1884]" captionTargetId-0="figure@4.[212,1374,193,1604]" captionTargetId-1="figure@6.[236,1352,169,1544]" captionTargetId-2="figure@7.[151,1436,193,1182]" captionTargetId-3="figure@7.[151,1435,1367,1893]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionText-0="FIGURES 3 10. Male genitalia (median lobe of aedeagus and tegmen) of: 3 4. Thymogethes ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 5 6. T. klapperichi (Easton, 1957) (male from Pakistan, Madaglasht); 7 8. T. kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 9 10. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3 6); = 0.20 mm (Figs 7 10)." captionText-1="FIGURES 11 18. Male genitalia (median lobe of aedeagus and tegmen) of: 11 12, 15 16, Thymogethes egenus (Erichson, 1845) s. l. (11 12: male from Italy, Lazio, Tivoli; 1 5 16: male from N Iran, Kadkan); 13 14. T. khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 17 18. T. otini (Easton, 1954) (male from Morocco, near Taza). Scale bar = 0.24 mm." captionText-2="FIGURES 19 22. SEM images of tubercles on last abdominal ventrite in males of: 19. Thymogethes khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 20. T. ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 21. T. kassites sp. nov. (male holotype from Iran, Zagros Mts, Mt. Oshtoran); 22. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.20 mm." captionText-3="FIGURES 23 24. SEM images of prosternal process of: 23. Thymogethes kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 24. T. otini (Easton, 1954) (male from Morocco, near Taza). Scale bar = 0.24 mm." httpUri-0="https://zenodo.org/record/242295/files/figure.png" httpUri-1="https://zenodo.org/record/242296/files/figure.png" httpUri-2="https://zenodo.org/record/242297/files/figure.png" httpUri-3="https://zenodo.org/record/242298/files/figure.png" pageId="2" pageNumber="371">Figs 324</figureCitation>
also for a summary of the diagnostic characters separating
<taxonomicName id="55A45AED7F1DFF87FC98351CFC356441" box="[827,958,1302,1327]" class="Insecta" family="Nitidulidae" genus="Meligethes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="ahriman">
<emphasis id="A0D0FD7C7F1DFF87FC98351CFC356441" box="[827,958,1302,1327]" italics="true" pageId="2" pageNumber="371">T. ahriman</emphasis>
</taxonomicName>
from
<taxonomicName id="55A45AED7F1DFF87FBB7351CFAC76441" authority="Easton, 1957" authorityName="Easton" authorityYear="1957" box="[1044,1356,1302,1327]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="rebmanni">
<emphasis id="A0D0FD7C7F1DFF87FBB7351CFB296441" box="[1044,1186,1302,1327]" italics="true" pageId="2" pageNumber="371">T. rebmanni</emphasis>
<bibRefCitation id="F6355C9F7F1DFF87FB11351CFAC76441" author="Easton" box="[1202,1356,1302,1327]" pageId="2" pageNumber="371" refString="Easton, A. M. (1957) The Meligethes (Col., Nitidulidae) of Afghanistan. Entomologist's Monthly Magazine, 92, 385 - 401." type="journal article" year="1957">Easton, 1957</bibRefCitation>
</taxonomicName>
(from
<collectingCountry id="EAB361FE7F1DFF87FF343531FEAB643D" box="[151,288,1338,1363]" name="Afghanistan" pageId="2" pageNumber="371">Afghanistan</collectingCountry>
and NW
<collectingCountry id="EAB361FE7F1DFF87FE3A3531FE77643D" box="[409,508,1338,1363]" name="Pakistan" pageId="2" pageNumber="371">Pakistan</collectingCountry>
:
<bibRefCitation id="F6355C9F7F1DFF87FDAF3530FD17643D" author="Easton" box="[524,668,1338,1363]" pageId="2" pageNumber="371" refString="Easton, A. M. (1957) The Meligethes (Col., Nitidulidae) of Afghanistan. Entomologist's Monthly Magazine, 92, 385 - 401." type="journal article" year="1957">Easton 1957</bibRefCitation>
) and
<taxonomicName id="55A45AED7F1DFF87FD473537FCD0643D" box="[740,859,1338,1363]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="kassites" status="sp. nov.">
<emphasis id="A0D0FD7C7F1DFF87FD473537FCD0643D" box="[740,859,1338,1363]" italics="true" pageId="2" pageNumber="371">T. kassites</emphasis>
</taxonomicName>
<emphasis id="A0D0FD7C7F1DFF87FCC53531FC4F643D" bold="true" box="[870,964,1338,1363]" pageId="2" pageNumber="371">
<taxonomicNameLabel id="BBE340077F1DFF87FCC53531FC4F643D" box="[870,964,1338,1363]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(the latter taxon being syntopic with
<taxonomicName id="55A45AED7F1DFF87FA243537FF7D6419" class="Insecta" family="Nitidulidae" genus="Meligethes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="ahriman">
<emphasis id="A0D0FD7C7F1DFF87FA243537FF7D6419" italics="true" pageId="2" pageNumber="371">T. ahriman</emphasis>
</taxonomicName>
on the Zagros Mts).
</paragraph>
<paragraph id="921B216E7F1DFF87FF64358FFC8E6769" blockId="2.[151,1437,222,1543]" pageId="2" pageNumber="371">
<emphasis id="A0D0FD7C7F1DFF87FF64358FFF7764F5" box="[199,252,1412,1435]" italics="true" pageId="2" pageNumber="371">Note</emphasis>
. According to the thus far known distributions (
<bibRefCitation id="F6355C9F7F1DFF87FCE73588FC5164F5" author="Easton" box="[836,986,1410,1435]" pageId="2" pageNumber="371" refString="Easton, A. M. (1954) A revision of the lugubris complex in the genus Meligethes Stephens (Col. Nitidulidae). Transactions of the Royal Entomological Society, 105 (16), 373 - 392. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1954. tb 00769. x" type="journal article" year="1954">Easton 1954</bibRefCitation>
;
<bibRefCitation id="F6355C9F7F1DFF87FC483589FB1364F5" author="Audisio" box="[1003,1176,1410,1435]" pageId="2" pageNumber="371" refString="Audisio, P. (1993 b) Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. 32. Calderini ed., Bologna, XVI + 971 pp." type="book" year="1993" yearSuffix="b">Audisio 1993b</bibRefCitation>
) of the Euro-Siberian
<taxonomicName id="55A45AED7F1DFF87FF3435ADFDB264D1" authority="Sturm, 1845" authorityName="Sturm" authorityYear="1845" box="[151,569,1446,1471]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="A0D0FD7C7F1DFF87FF3435ADFE1A64D1" box="[151,401,1446,1471]" italics="true" pageId="2" pageNumber="371">Thymogethes lugubris</emphasis>
(
<bibRefCitation id="F6355C9F7F1DFF87FE0135ACFDBA64D1" author="Sturm" box="[418,561,1446,1471]" pageId="2" pageNumber="371" refString="Sturm, J. (1845) Deutschlands Fauna in Abbildungen nach der Natur mit Beschreibungen. F. Abtheilung. Die Insecten. Zweites Bandchen. Kafer. J. Sturm, Nurnberg, 114 pp., pls. ccciv - cccix." type="book" year="1845">Sturm, 1845</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="55A45AED7F1DFF87FDD735ACFC5264D1" authority="Erichson, 1845" authorityName="Erichson" authorityYear="1845" box="[628,985,1446,1471]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="gagathinus">
<emphasis id="A0D0FD7C7F1DFF87FDD735ACFC8564D1" box="[628,782,1446,1471]" italics="true" pageId="2" pageNumber="371">T. gagathinus</emphasis>
(
<bibRefCitation id="F6355C9F7F1DFF87FC8335ADFC5B64D1" author="Erichson" box="[800,976,1446,1471]" pageId="2" pageNumber="371" refString="Erichson, W. F. (1845) [I., II. Lieferungen]. In: Erichson, W. F. (Ed.), Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Dritter Band. Nikolaische Buchhandlung, Berlin, pp. 1 - 320. [issued in parts: pp. 1 - 320: 1845; pp. 321 - 480: 1846; pp. 481 - 800: 1847; pp. 801 - 968: 1848]." type="journal article" year="1845">Erichson, 1845</bibRefCitation>
)
</taxonomicName>
, both known to occur in N
<collectingCountry id="EAB361FE7F1DFF87FAB935ADFAE164D1" box="[1306,1386,1446,1471]" name="Turkey" pageId="2" pageNumber="371">Turkey</collectingCountry>
and Caucasus (see key to identification of the Near East
<taxonomicName id="55A45AED7F1DFF87FD4835C1FCF4648D" box="[747,895,1482,1507]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="A0D0FD7C7F1DFF87FD4835C1FCF4648D" box="[747,895,1482,1507]" italics="true" pageId="2" pageNumber="371">Thymogethes</emphasis>
</taxonomicName>
below), the presence of these two taxa in some mountain localities of the NW
<collectingCountry id="EAB361FE7F1DFF87FE5235E4FD956768" box="[497,542,1519,1542]" name="Iran" pageId="2" pageNumber="371">Iran</collectingCountry>
cannot be excluded.
</paragraph>
</subSubSection>
</treatment>
</document>