399 lines
52 KiB
XML
399 lines
52 KiB
XML
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<mods:title id="E1938DA1002F9226CFC1DD0B544E7764">New species and records of pollen and sap beetles for Iran (Coleoptera: Kateretidae, Nitidulidae)</mods:title>
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<mods:namePart id="EA0E877116A4EF9649925580FDB445B1">Audisio, Paolo</mods:namePart>
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<mods:namePart id="6E7990C1C9A205B8DB2BA869FAF9D989">Cline, Andrew R.</mods:namePart>
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<mods:namePart id="DD2BCDA4FBB27401F4529FBEC5D3C9D5">Lasoń, Andrzej</mods:namePart>
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<mods:namePart id="FA5612D8DC28785BF621F1D59B97C8F2">Jelínek, Josef</mods:namePart>
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<mods:namePart id="6D3BF00CC4A40B7C2157664C70D9ECAF">Sabatelli, Simone</mods:namePart>
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<mods:namePart id="CE5AA730FF97323C15ACB7F05E0C7B47">Serri, Sayeh</mods:namePart>
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<mods:title id="B6A690BCF7C303F76221B3AF052BBD59">Zootaxa</mods:title>
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<mods:date id="5EEE894D9FF28E8189B0DA0375298AE5">2017</mods:date>
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<treatment id="1A0D90787F1DFF80FF34365BFBC06586" ID-DOI="http://doi.org/10.5281/zenodo.6030359" ID-GBIF-Taxon="127378285" ID-Zenodo-Dep="6030359" LSID="urn:lsid:plazi:treatment:1A0D90787F1DFF80FF34365BFBC06586" httpUri="http://treatment.plazi.org/id/1A0D90787F1DFF80FF34365BFBC06586" lastPageId="5" lastPageNumber="374" pageId="2" pageNumber="371">
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<subSubSection id="DABE72E57F1DFF87FF34365BFEBF67E5" pageId="2" pageNumber="371" type="nomenclature">
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<paragraph id="921B216E7F1DFF87FF34365BFD8A6704" blockId="2.[151,513,1616,1675]" box="[151,513,1616,1642]" pageId="2" pageNumber="371">
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<heading id="C95396027F1DFF87FF34365BFD8A6704" bold="true" box="[151,513,1616,1642]" fontSize="11" level="1" pageId="2" pageNumber="371" reason="1">
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<emphasis id="A0D0FD7C7F1DFF87FF34365BFD8A6704" bold="true" box="[151,513,1616,1642]" pageId="2" pageNumber="371">
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<taxonomicName id="55A45AED7F1DFF87FF34365BFE176704" box="[151,412,1616,1642]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="kassites" status="sp. nov.">
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<emphasis id="A0D0FD7C7F1DFF87FF34365BFE176704" bold="true" box="[151,412,1616,1642]" italics="true" pageId="2" pageNumber="371">Thymogethes kassites</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBE340077F1DFF87FE00365BFD8A6704" box="[419,513,1616,1642]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="921B216E7F1DFF87FF343679FEBF67E5" blockId="2.[151,513,1616,1675]" box="[151,308,1650,1675]" pageId="2" pageNumber="371">
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(
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<figureCitation id="0A9F3DEB7F1DFF87FF3C3679FE8F67E5" box="[159,260,1650,1675]" captionStart="FIGURES 3 – 10" captionStartId="4.[151,264,1624,1647]" captionTargetBox="[226,1362,205,1593]" captionTargetId="figure@4.[212,1374,193,1604]" captionTargetPageId="4" captionText="FIGURES 3 – 10. Male genitalia (median lobe of aedeagus and tegmen) of: 3 – 4. Thymogethes ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 5 – 6. T. klapperichi (Easton, 1957) (male from Pakistan, Madaglasht); 7 – 8. T. kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 9 – 10. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3 – 6); = 0.20 mm (Figs 7 – 10)." httpUri="https://zenodo.org/record/242295/files/figure.png" pageId="2" pageNumber="371">Figs 7–8</figureCitation>
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,
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<figureCitation id="0A9F3DEB7F1DFF87FEAC3678FEA767E5" box="[271,300,1651,1675]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1202,1225]" captionTargetBox="[158,1430,193,1168]" captionTargetId="figure@7.[151,1436,193,1182]" captionTargetPageId="7" captionText="FIGURES 19 – 22. SEM images of tubercles on last abdominal ventrite in males of: 19. Thymogethes khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 20. T. ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 21. T. kassites sp. nov. (male holotype from Iran, Zagros Mts, Mt. Oshtoran); 22. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/242297/files/figure.png" pageId="2" pageNumber="371">21</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DABE72E57F1DFF87FF3436B1FEFB669D" pageId="2" pageNumber="371" type="diagnosis">
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<paragraph id="921B216E7F1DFF87FF3436B1FEFB669D" blockId="2.[151,1437,1722,2035]" pageId="2" pageNumber="371">
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<emphasis id="A0D0FD7C7F1DFF87FF3436B1FE9A67BD" bold="true" box="[151,273,1722,1747]" pageId="2" pageNumber="371">Diagnosis.</emphasis>
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Medium-sized (length 2.48), body shiny black, legs blackish with protibiae chestnut brown to blackish, and antennae chestnut brown with blackish antennal club. Maximum pronotal width near posterior angles. Similar to the widespread European species
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<taxonomicName id="55A45AED7F1DFF87FDE6370FFCF96675" authority="Sturm, 1845" authorityName="Sturm" authorityYear="1845" box="[581,882,1794,1819]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="lugubris">
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<emphasis id="A0D0FD7C7F1DFF87FDE6370FFD496675" box="[581,706,1794,1819]" italics="true" pageId="2" pageNumber="371">T. lugubris</emphasis>
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(
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<bibRefCitation id="F6355C9F7F1DFF87FD753708FCE16675" author="Sturm" box="[726,874,1794,1819]" pageId="2" pageNumber="371" refString="Sturm, J. (1845) Deutschlands Fauna in Abbildungen nach der Natur mit Beschreibungen. F. Abtheilung. Die Insecten. Zweites Bandchen. Kafer. J. Sturm, Nurnberg, 114 pp., pls. ccciv - cccix." type="book" year="1845">Sturm, 1845</bibRefCitation>
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)
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</taxonomicName>
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, revised by
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<bibRefCitation id="F6355C9F7F1DFF87FBA43708FB216675" author="Easton" box="[1031,1194,1794,1819]" pageId="2" pageNumber="371" refString="Easton, A. M. (1954) A revision of the lugubris complex in the genus Meligethes Stephens (Col. Nitidulidae). Transactions of the Royal Entomological Society, 105 (16), 373 - 392. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1954. tb 00769. x" type="journal article" year="1954">Easton (1954)</bibRefCitation>
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and re-described by
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<bibRefCitation id="F6355C9F7F1DFF87FF34372DFEDD6651" author="Audisio" box="[151,342,1830,1855]" pageId="2" pageNumber="371" refString="Audisio, P. (1993 b) Coleoptera Nitidulidae - Kateretidae. Fauna d'Italia. 32. Calderini ed., Bologna, XVI + 971 pp." type="book" year="1993" yearSuffix="b">Audisio (1993b)</bibRefCitation>
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, but with interspaces between dorsal punctures on pronotum and most of elytra smooth and shining. Differentiated (see key to identification of the Near East
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<taxonomicName id="55A45AED7F1DFF87FC233741FB9F660D" box="[896,1044,1866,1891]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
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<emphasis id="A0D0FD7C7F1DFF87FC233741FB9F660D" box="[896,1044,1866,1891]" italics="true" pageId="2" pageNumber="371">Thymogethes</emphasis>
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</taxonomicName>
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below) from the closely related
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<taxonomicName id="55A45AED7F1DFF87FA243747FE2766E9" authority="Easton, 1957" authorityName="Easton" authorityYear="1957" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="A0D0FD7C7F1DFF87FA243747FE8A66E9" italics="true" pageId="2" pageNumber="371">T. rebmanni</emphasis>
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(
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<bibRefCitation id="F6355C9F7F1DFF87FEAC3764FE2F66E9" author="Easton" box="[271,420,1902,1927]" pageId="2" pageNumber="371" refString="Easton, A. M. (1957) The Meligethes (Col., Nitidulidae) of Afghanistan. Entomologist's Monthly Magazine, 92, 385 - 401." type="journal article" year="1957">Easton, 1957</bibRefCitation>
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)
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</taxonomicName>
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by the comparatively larger male genitalia, the paramera much more extruded laterad, and by the more raised and centrally-placed concave tubercle on male last abdominal ventrite (
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<figureCitation id="0A9F3DEB7F1DFF87FB0C3799FA8866C5" box="[1199,1283,1938,1963]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1202,1225]" captionTargetBox="[158,1430,193,1168]" captionTargetId="figure@7.[151,1436,193,1182]" captionTargetPageId="7" captionText="FIGURES 19 – 22. SEM images of tubercles on last abdominal ventrite in males of: 19. Thymogethes khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 20. T. ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 21. T. kassites sp. nov. (male holotype from Iran, Zagros Mts, Mt. Oshtoran); 22. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/242297/files/figure.png" pageId="2" pageNumber="371">Fig. 21</figureCitation>
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) (less raised, flattened, wider, not concave nor evidently bicuspid, and placed closer to the posterior edge of the ventrite in
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<taxonomicName id="55A45AED7F1DFF87FA2437BCFE8A669D" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="rebmanni">
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<emphasis id="A0D0FD7C7F1DFF87FA2437BCFE8A669D" italics="true" pageId="2" pageNumber="371">T. rebmanni</emphasis>
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</taxonomicName>
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:
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<figureCitation id="0A9F3DEB7F1DFF87FEB337D1FEE8669D" box="[272,355,2010,2035]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1202,1225]" captionTargetBox="[158,1430,193,1168]" captionTargetId="figure@7.[151,1436,193,1182]" captionTargetPageId="7" captionText="FIGURES 19 – 22. SEM images of tubercles on last abdominal ventrite in males of: 19. Thymogethes khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 20. T. ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 21. T. kassites sp. nov. (male holotype from Iran, Zagros Mts, Mt. Oshtoran); 22. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/242297/files/figure.png" pageId="2" pageNumber="371">Fig. 22</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<caption id="C6DB71E67F1CFF86FF34342DFC696532" httpUri="https://zenodo.org/record/242294/files/figure.png" pageId="3" pageNumber="372" targetBox="[164,1424,208,1029]" targetPageId="3">
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<paragraph id="921B216E7F1CFF86FF34342DFC696532" blockId="3.[151,1436,1062,1116]" pageId="3" pageNumber="372">
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||
<emphasis id="A0D0FD7C7F1CFF86FF34342DFED76552" bold="true" box="[151,348,1062,1085]" pageId="3" pageNumber="372">FIGURES 1–2. 1.</emphasis>
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<taxonomicName id="55A45AED7F1CFF86FEC5342CFD6D6553" authority="Sturm, 1845" authorityName="Sturm" authorityYear="1845" box="[358,742,1063,1085]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="lugubris">
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<emphasis id="A0D0FD7C7F1CFF86FEC5342CFDC16553" box="[358,586,1063,1085]" italics="true" pageId="3" pageNumber="372">Thymogethes lugubris</emphasis>
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||
(Sturm, 1845)
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</taxonomicName>
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habitus (male from Poland);
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<emphasis id="A0D0FD7C7F1CFF86FB87342CFBBC6552" bold="true" box="[1060,1079,1063,1084]" pageId="3" pageNumber="372">2.</emphasis>
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||
<taxonomicName id="55A45AED7F1CFF86FBE23423FAD66553" authority="Erichson, 1845" authorityName="Erichson" authorityYear="1845" box="[1089,1373,1063,1085]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="egenus">
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<emphasis id="A0D0FD7C7F1CFF86FBE23423FB2F6553" box="[1089,1188,1064,1085]" italics="true" pageId="3" pageNumber="372">T. egenus</emphasis>
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(Erichson, 1845)
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</taxonomicName>
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(s.l.), habitus (male from Poland). Both specimens ca. 2.2 mm long. Photos L. Borowiec.
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</paragraph>
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</caption>
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<subSubSection id="DABE72E57F1CFF86FF643488FE4C658A" pageId="3" pageNumber="372" type="materials_examined">
|
||
<paragraph id="921B216E7F1CFF86FF643488FE4C658A" blockId="3.[151,1437,1155,2008]" pageId="3" pageNumber="372">
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||
<materialsCitation id="22CC2B337F1CFF86FF643488FE4C658A" ID-GBIF-Occurrence="1424563911" collectingDate="2014-06-14" collectionCode="CAR" collectorName="Zagros Mts, SE & P. Audisio & Huds." country="Iran" elevation="1695" latitude="33.418056" location="Chamnar" longLatPrecision="20" longitude="49.162224" municipality="Oshtoran Kuh" pageId="3" pageNumber="372" specimenCount="1" specimenCount-male="1" stateProvince="Lorestan Province" typeStatus="holotype">
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<emphasis id="A0D0FD7C7F1CFF86FF643488FEFD65F2" bold="true" box="[199,374,1155,1180]" pageId="3" pageNumber="372">
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<typeStatus id="4D1F9FCC7F1CFF86FF643488FE8965F2" box="[199,258,1155,1180]" pageId="3" pageNumber="372">Type</typeStatus>
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material.
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||
</emphasis>
|
||
<specimenCount id="84A2EAE77F1CFF86FEDD348FFE2765F5" box="[382,428,1156,1180]" pageId="3" pageNumber="372" type="male">1 ♂</specimenCount>
|
||
, male
|
||
<typeStatus id="4D1F9FCC7F1CFF86FE5B3488FDD665F2" box="[504,605,1155,1180]" pageId="3" pageNumber="372" type="holotype">holotype</typeStatus>
|
||
,
|
||
<emphasis id="A0D0FD7C7F1CFF86FDC93488FD3965F2" bold="true" box="[618,690,1155,1180]" pageId="3" pageNumber="372">
|
||
<collectingCountry id="EAB361FE7F1CFF86FDC93488FD3965F2" box="[618,690,1155,1180]" name="Iran" pageId="3" pageNumber="372">IRAN</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="5060EF8C7F1CFF86FD61348FFC1965F2" box="[706,914,1155,1180]" country="Iran" name="Lorestan" pageId="3" pageNumber="372">Lorestan Province</collectingRegion>
|
||
,
|
||
<collectorName id="3F5144B87F1CFF86FC3C348FFBDA65F2" box="[927,1105,1156,1180]" pageId="3" pageNumber="372">Zagros Mts, SE</collectorName>
|
||
Dorud,
|
||
<collectingMunicipality id="727FBB147F1CFF86FB133488FAD865F2" box="[1200,1363,1155,1180]" pageId="3" pageNumber="372">Oshtoran Kuh</collectingMunicipality>
|
||
, small village near
|
||
<location id="977B77B57F1CFF86FE8434ACFE0465AE" LSID="urn:lsid:plazi:treatment:1A0D90787F1DFF80FF34365BFBC06586:977B77B57F1CFF86FE8434ACFE0465AE" box="[295,399,1191,1216]" country="Iran" latitude="33.418056" longLatPrecision="20" longitude="49.162224" municipality="Oshtoran Kuh" name="Chamnar" pageId="3" pageNumber="372" stateProvince="Lorestan Province">Chamnar</location>
|
||
,
|
||
<geoCoordinate id="F79047A97F1CFF86FE3E34ACFD9465AE" box="[413,543,1191,1216]" direction="north" orientation="latitude" pageId="3" pageNumber="372" precision="15" value="33.418056">33°25'05''N</geoCoordinate>
|
||
<geoCoordinate id="F79047A97F1CFF86FD8A34ACFD2765D1" box="[553,684,1191,1215]" direction="east" orientation="longitude" pageId="3" pageNumber="372" precision="15" value="49.162224">49°09'44''E</geoCoordinate>
|
||
,
|
||
<quantity id="555C8C8B7F1CFF86FD1A34ACFC9865D1" box="[697,787,1191,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="3" pageNumber="372" unit="m" value="1695.0">
|
||
<elevation id="1989C65D7F1CFF86FD1A34ACFC9865D1" box="[697,787,1191,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="3" pageNumber="372" unit="m" value="1695.0">1695 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="E61A07AE7F1CFF86FC8134ACFC2565D1" box="[802,942,1191,1215]" pageId="3" pageNumber="372" value="2014-06-14">
|
||
<collectingDate id="F65EFE467F1CFF86FC8134ACFC2565D1" box="[802,942,1191,1215]" pageId="3" pageNumber="372" value="2014-06-14">14.VI. 2014</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="3F5144B87F1CFF86FC4C34A3FBE365AE" box="[1007,1128,1191,1216]" pageId="3" pageNumber="372">P. Audisio</collectorName>
|
||
,
|
||
<collectedFrom id="21B862937F1CFF86FBD534A3FAE365AE" box="[1142,1384,1191,1216]" pageId="3" pageNumber="372">
|
||
on
|
||
<taxonomicName id="55A45AED7F1CFF86FB3834ACFAE365AE" box="[1179,1384,1191,1216]" class="Magnoliopsida" family="Lamiaceae" genus="Mentha" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="372" phylum="Tracheophyta" rank="species" species="longifolia">
|
||
<emphasis id="A0D0FD7C7F1CFF86FB3834ACFAE365AE" box="[1179,1384,1191,1216]" italics="true" pageId="3" pageNumber="372">Mentha longifolia</emphasis>
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
(L.)
|
||
<collectorName id="3F5144B87F1CFF86FF3434C0FF51658A" box="[151,218,1227,1252]" pageId="3" pageNumber="372">Huds.</collectorName>
|
||
(
|
||
<taxonomicName id="55A45AED7F1CFF86FF4B34C0FEEE658A" box="[232,357,1227,1252]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="372" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
|
||
) (
|
||
<collectionCode id="F4B5B9AB7F1CFF86FED834C7FE30658A" box="[379,443,1228,1252]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13632" name="Museo de Historia Natural La Salle" pageId="3" pageNumber="372">CAR</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DABE72E57F1CFF80FF6434E4FC2265CE" lastPageId="5" lastPageNumber="374" pageId="3" pageNumber="372" type="description">
|
||
<paragraph id="921B216E7F1CFF86FF6434E4FD64641A" blockId="3.[151,1437,1155,2008]" pageId="3" pageNumber="372">
|
||
<emphasis id="A0D0FD7C7F1CFF86FF6434E4FED26466" bold="true" box="[199,345,1263,1288]" pageId="3" pageNumber="372">Description.</emphasis>
|
||
Male [
|
||
<typeStatus id="4D1F9FCC7F1CFF86FE1334E4FD906466" box="[432,539,1263,1288]" pageId="3" pageNumber="372" type="holotype">holotype</typeStatus>
|
||
]. Length
|
||
<quantity id="555C8C8B7F1CFF86FD2F34E4FD7C6469" box="[652,759,1263,1287]" metricMagnitude="-3" metricUnit="m" metricValue="2.48" pageId="3" pageNumber="372" unit="mm" value="2.48">2.48 mm</quantity>
|
||
; width (at elytral widest point)
|
||
<quantity id="555C8C8B7F1CFF86FBD034E4FB546469" box="[1139,1247,1263,1287]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="3" pageNumber="372" unit="mm" value="1.28">1.28 mm</quantity>
|
||
. Body elongate, narrow, transversely convex and moderately parallel-sided (nearly as in
|
||
<figureCitation id="0A9F3DEB7F1CFF86FC6B3518FB826442" box="[968,1033,1299,1324]" captionStart="FIGURES 1 – 2. 1" captionStartId="3.[151,264,1062,1085]" captionTargetBox="[164,1424,208,1029]" captionTargetId="figure@3.[150,1437,193,1042]" captionTargetPageId="3" captionText="FIGURES 1 – 2. 1. Thymogethes lugubris (Sturm, 1845) habitus (male from Poland); 2. T. egenus (Erichson, 1845) (s. l.), habitus (male from Poland). Both specimens ca. 2.2 mm long. Photos L. Borowiec." httpUri="https://zenodo.org/record/242294/files/figure.png" pageId="3" pageNumber="372">Fig. 1</figureCitation>
|
||
), black and shining, with short and fine silvery to golden-olivaceous pubescence. Legs blackish with protibiae chestnut brown to blackish, and antennae chestnut brown with blackish antennal club.
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1CFF86FF643574FE996746" blockId="3.[151,1437,1155,2008]" pageId="3" pageNumber="372">
|
||
Head with dorsal punctures as large as or slightly smaller than an eye facet, moderately impressed, separated by one diameter or less, surface smooth and shining; front margin of clypeus regularly arcuately emarginate, completely bordered, with rather acute angles. Fronto-genal grooves narrow and shallow, but distinct. Antennae small, of normal size for group (cf.
|
||
<figureCitation id="0A9F3DEB7F1CFF86FD8C35E0FDFA676A" box="[559,625,1515,1540]" captionStart="FIGURES 1 – 2. 1" captionStartId="3.[151,264,1062,1085]" captionTargetBox="[164,1424,208,1029]" captionTargetId="figure@3.[150,1437,193,1042]" captionTargetPageId="3" captionText="FIGURES 1 – 2. 1. Thymogethes lugubris (Sturm, 1845) habitus (male from Poland); 2. T. egenus (Erichson, 1845) (s. l.), habitus (male from Poland). Both specimens ca. 2.2 mm long. Photos L. Borowiec." httpUri="https://zenodo.org/record/242294/files/figure.png" pageId="3" pageNumber="372">Fig. 1</figureCitation>
|
||
), with third antennomere slender, slightly longer than second; antennal club small.
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1CFF86FF64363FFD0B67D6" blockId="3.[151,1437,1155,2008]" pageId="3" pageNumber="372">
|
||
Pronotum markedly narrowed anteriorly, 1.68× as wide as long, broadest close to posterior angles (cf.
|
||
<figureCitation id="0A9F3DEB7F1CFF86FAEA3638FA016722" box="[1353,1418,1587,1612]" captionStart="FIGURES 1 – 2. 1" captionStartId="3.[151,264,1062,1085]" captionTargetBox="[164,1424,208,1029]" captionTargetId="figure@3.[150,1437,193,1042]" captionTargetPageId="3" captionText="FIGURES 1 – 2. 1. Thymogethes lugubris (Sturm, 1845) habitus (male from Poland); 2. T. egenus (Erichson, 1845) (s. l.), habitus (male from Poland). Both specimens ca. 2.2 mm long. Photos L. Borowiec." httpUri="https://zenodo.org/record/242294/files/figure.png" pageId="3" pageNumber="372">Fig. 1</figureCitation>
|
||
); sides narrowly bordered; posterior angles slightly obtuse but markedly distinct. Posterior base faintly sinuate on either side of scutellum; pronotal punctures and surface smooth and shining, each puncture nearly as large as an eye facet and separated by one diameter or less.
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1CFF86FF6436C8FECC666E" blockId="3.[151,1437,1155,2008]" pageId="3" pageNumber="372">Scutellar shield medium-sized, densely and uniformly punctate; surface exhibiting a faint trace of reticulate microsculpture.</paragraph>
|
||
<paragraph id="921B216E7F1CFF86FF643700FEBA66B6" blockId="3.[151,1437,1155,2008]" pageId="3" pageNumber="372">Elytra nearly 1.17× as long as combined width (if length measured from posterior edge of pronotum; 1.04× if measured from posterior apex of scutellar shield), broadest in basal fifth, distinctly wider (1.12×), and approximately twice as long (1.95×) as pronotum; humeral angle rounded, humeral striae absent. Elytral punctures in basal half as on head and pronotum, but slightly coarser and more elongate, exhibiting a feeble transverse rugosity and with a shiny surface between punctures. Elytral punctures becoming finer and shallower towards posterior end.</paragraph>
|
||
<caption id="C6DB71E67F1BFF81FF343653FCA167A5" httpUri="https://zenodo.org/record/242295/files/figure.png" pageId="4" pageNumber="373" targetBox="[226,1362,205,1593]" targetPageId="4">
|
||
<paragraph id="921B216E7F1BFF81FF343653FCA167A5" blockId="4.[151,1436,1624,1739]" pageId="4" pageNumber="373">
|
||
<emphasis id="A0D0FD7C7F1BFF81FF343653FEC36700" bold="true" box="[151,328,1624,1647]" pageId="4" pageNumber="373">FIGURES 3–10.</emphasis>
|
||
Male genitalia (median lobe of aedeagus and tegmen) of:
|
||
<emphasis id="A0D0FD7C7F1BFF81FC003652FC5B6700" bold="true" box="[931,976,1625,1646]" pageId="4" pageNumber="373">3–4.</emphasis>
|
||
<taxonomicName id="55A45AED7F1BFF81FC743652FAD16701" authority="Jelinek, 1981" authorityName="Jelinek" authorityYear="1981" box="[983,1370,1625,1647]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="ahriman">
|
||
<emphasis id="A0D0FD7C7F1BFF81FC743652FB326701" box="[983,1209,1625,1647]" italics="true" pageId="4" pageNumber="373">Thymogethes ahriman</emphasis>
|
||
(Jelínek, 1981)
|
||
</taxonomicName>
|
||
(male from Iran, Zagros Mts., Mt. Oshtoran);
|
||
<emphasis id="A0D0FD7C7F1BFF81FD8D3673FDD067E3" bold="true" box="[558,603,1656,1677]" pageId="4" pageNumber="373">5–6.</emphasis>
|
||
<taxonomicName id="55A45AED7F1BFF81FDC13672FC0667E0" authority="Easton, 1957" authorityName="Easton" authorityYear="1957" box="[610,909,1656,1678]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="klapperichi">
|
||
<emphasis id="A0D0FD7C7F1BFF81FDC13672FD6467E0" box="[610,751,1656,1678]" italics="true" pageId="4" pageNumber="373">T. klapperichi</emphasis>
|
||
(Easton, 1957)
|
||
</taxonomicName>
|
||
(male from Pakistan, Madaglasht);
|
||
<emphasis id="A0D0FD7C7F1BFF81FB5C3673FAA767E3" bold="true" box="[1279,1324,1656,1677]" pageId="4" pageNumber="373">7–8.</emphasis>
|
||
<taxonomicName id="55A45AED7F1BFF81FA903672FA1767E0" box="[1331,1436,1656,1678]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="kassites" status="sp. nov.">
|
||
<emphasis id="A0D0FD7C7F1BFF81FA903672FA1767E0" box="[1331,1436,1656,1678]" italics="true" pageId="4" pageNumber="373">T. kassites</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0D0FD7C7F1BFF81FF34369CFF6167C2" bold="true" box="[151,234,1687,1708]" pageId="4" pageNumber="373">
|
||
<taxonomicNameLabel id="BBE340077F1BFF81FF34369CFF6167C2" box="[151,234,1687,1708]" pageId="4" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(male holotype from Iran, Zagros Mts., Mt. Oshtoran);
|
||
<emphasis id="A0D0FD7C7F1BFF81FC94369DFCFB67C5" bold="true" box="[823,880,1686,1707]" pageId="4" pageNumber="373">9–10.</emphasis>
|
||
<taxonomicName id="55A45AED7F1BFF81FCDA369CFB1F67C2" authority="Easton, 1957" authorityName="Easton" authorityYear="1957" box="[889,1172,1686,1708]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="rebmanni">
|
||
<emphasis id="A0D0FD7C7F1BFF81FCDA369CFC7F67C2" box="[889,1012,1686,1708]" italics="true" pageId="4" pageNumber="373">T. rebmanni</emphasis>
|
||
(Easton, 1957)
|
||
</taxonomicName>
|
||
(male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3–6); = 0.20 mm (Figs 7–10).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="921B216E7F1BFF81FF6436F9FBAC6619" blockId="4.[151,1437,1778,2019]" pageId="4" pageNumber="373">
|
||
Ventral surface black, with sparse fine silvery pubescence. Prosternal antennal furrows (at notosternal sutures) strongly raised, reaching near the middle point of the hypomeron. Prosternal process moderately long, subtruncate at apex, ventral surface (excluding predistal lateral expansions) only moderately wider before apex (as in
|
||
<figureCitation id="0A9F3DEB7F1BFF81FA9F3731FA04663D" box="[1340,1423,1850,1875]" captionStart="FIGURES 23 – 24" captionStartId="7.[151,264,1914,1937]" captionTargetBox="[157,1429,1373,1884]" captionTargetId="figure@7.[151,1435,1367,1893]" captionTargetPageId="7" captionText="FIGURES 23 – 24. SEM images of prosternal process of: 23. Thymogethes kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 24. T. otini (Easton, 1954) (male from Morocco, near Taza). Scale bar = 0.24 mm." httpUri="https://zenodo.org/record/242298/files/figure.png" pageId="4" pageNumber="373">Fig. 23</figureCitation>
|
||
), widest subdistal portion approximately 1.28× wider than narrowest basal portion.
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1BFF81FF643789FDBB668D" blockId="4.[151,1437,1778,2019]" pageId="4" pageNumber="373">Metaventrite moderately convex (punctures as on head and pronotum, surface shiny), with a barely distinct and posteriorly widened longitudinal impression on posterior two-thirds, and a slightly raised elongate medial tubercle. Posterior angles only slightly raised.</paragraph>
|
||
<paragraph id="921B216E7F1AFF80FF64309CFC186199" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="374">
|
||
Caudal marginal line of metacoxal cavity closely following posterior edge, turning back just before outer end. Last visible abdominal ventrite with a markedly raised obtuse projection nearly in middle, distally concave and bicuspid (
|
||
<figureCitation id="0A9F3DEB7F1AFF80FEA630D4FED36196" box="[261,344,223,248]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1202,1225]" captionTargetBox="[158,1430,193,1168]" captionTargetId="figure@7.[151,1436,193,1182]" captionTargetPageId="7" captionText="FIGURES 19 – 22. SEM images of tubercles on last abdominal ventrite in males of: 19. Thymogethes khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 20. T. ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 21. T. kassites sp. nov. (male holotype from Iran, Zagros Mts, Mt. Oshtoran); 22. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/242297/files/figure.png" pageId="5" pageNumber="374">Fig. 21</figureCitation>
|
||
), similar to males of
|
||
<taxonomicName id="55A45AED7F1AFF80FDE630EAFD366196" box="[581,701,223,248]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="A0D0FD7C7F1AFF80FDE630EAFD366196" box="[581,701,223,248]" italics="true" pageId="5" pageNumber="374">T. lugubris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="55A45AED7F1AFF80FD5730EAFC076196" box="[756,908,223,248]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="gagathinus">
|
||
<emphasis id="A0D0FD7C7F1AFF80FD5730EAFC076196" box="[756,908,223,248]" italics="true" pageId="5" pageNumber="374">T. gagathinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1AFF80FF64310FFAEA600B" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="374">
|
||
Protibiae (cf.
|
||
<figureCitation id="0A9F3DEB7F1AFF80FEC2310FFE2A6072" box="[353,417,260,285]" captionStart="FIGURES 1 – 2. 1" captionStartId="3.[151,264,1062,1085]" captionTargetBox="[164,1424,208,1029]" captionTargetId="figure@3.[150,1437,193,1042]" captionTargetPageId="3" captionText="FIGURES 1 – 2. 1. Thymogethes lugubris (Sturm, 1845) habitus (male from Poland); 2. T. egenus (Erichson, 1845) (s. l.), habitus (male from Poland). Both specimens ca. 2.2 mm long. Photos L. Borowiec." httpUri="https://zenodo.org/record/242294/files/figure.png" pageId="5" pageNumber="374">Fig. 1</figureCitation>
|
||
) with outer edges finely crenulate from basal third, with subapical group of 5 sharp teeth, the first and penultimate tooth markedly bigger than the other teeth; protarsi distinctly wider than antennal club, ratio WFTA/LFTA = 0.35–0.36; metatibiae narrow, inner edges not sinuate (as in
|
||
<figureCitation id="0A9F3DEB7F1AFF80FB893147FBE0600A" box="[1066,1131,332,357]" captionStart="FIGURES 1 – 2. 1" captionStartId="3.[151,264,1062,1085]" captionTargetBox="[164,1424,208,1029]" captionTargetId="figure@3.[150,1437,193,1042]" captionTargetPageId="3" captionText="FIGURES 1 – 2. 1. Thymogethes lugubris (Sturm, 1845) habitus (male from Poland); 2. T. egenus (Erichson, 1845) (s. l.), habitus (male from Poland). Both specimens ca. 2.2 mm long. Photos L. Borowiec." httpUri="https://zenodo.org/record/242294/files/figure.png" pageId="5" pageNumber="374">Fig. 1</figureCitation>
|
||
); tarsal claws simple.
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1AFF80FF643164FB53609B" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="374">
|
||
Genitalia. Tegmen as in
|
||
<figureCitation id="0A9F3DEB7F1AFF80FE4D3164FDBC60E9" box="[494,567,367,392]" captionStart="FIGURES 3 – 10" captionStartId="4.[151,264,1624,1647]" captionTargetBox="[226,1362,205,1593]" captionTargetId="figure@4.[212,1374,193,1604]" captionTargetPageId="4" captionText="FIGURES 3 – 10. Male genitalia (median lobe of aedeagus and tegmen) of: 3 – 4. Thymogethes ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 5 – 6. T. klapperichi (Easton, 1957) (male from Pakistan, Madaglasht); 7 – 8. T. kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 9 – 10. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3 – 6); = 0.20 mm (Figs 7 – 10)." httpUri="https://zenodo.org/record/242295/files/figure.png" pageId="5" pageNumber="374">Fig. 7</figureCitation>
|
||
, rather strongly sclerotized, brownish, with a moderately deep, U-shaped median excision, and paramera markedly arcuately protruded at sides in anterior half; median lobe of aedeagus elongate, narrow, about 2.4× as long as wide (
|
||
<figureCitation id="0A9F3DEB7F1AFF80FD3A31BCFD5760BE" box="[665,732,439,464]" captionStart="FIGURES 3 – 10" captionStartId="4.[151,264,1624,1647]" captionTargetBox="[226,1362,205,1593]" captionTargetId="figure@4.[212,1374,193,1604]" captionTargetPageId="4" captionText="FIGURES 3 – 10. Male genitalia (median lobe of aedeagus and tegmen) of: 3 – 4. Thymogethes ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 5 – 6. T. klapperichi (Easton, 1957) (male from Pakistan, Madaglasht); 7 – 8. T. kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 9 – 10. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3 – 6); = 0.20 mm (Figs 7 – 10)." httpUri="https://zenodo.org/record/242295/files/figure.png" pageId="5" pageNumber="374">Fig. 8</figureCitation>
|
||
), nearly as sclerotized and coloured as tegmen, widest at distal three-fifths, subparallel-sided proximad, narrowed distad, apex subtruncate and minutely incised.
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1AFF80FF6431F4FE156376" blockId="5.[151,1437,151,1257]" box="[199,414,511,536]" pageId="5" pageNumber="374">Female. Unknown.</paragraph>
|
||
<paragraph id="921B216E7F1AFF80FF64322FFBF36232" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="374">
|
||
<emphasis id="A0D0FD7C7F1AFF80FF64322FFE396353" bold="true" box="[199,434,548,573]" pageId="5" pageNumber="374">Comparative notes.</emphasis>
|
||
<taxonomicName id="55A45AED7F1AFF80FE19322FFD266353" box="[442,685,548,573]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="kassites" status="sp. nov.">
|
||
<emphasis id="A0D0FD7C7F1AFF80FE19322FFD266353" box="[442,685,548,573]" italics="true" pageId="5" pageNumber="374">Thymogethes kassites</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0D0FD7C7F1AFF80FD15322FFC9A6353" bold="true" box="[694,785,548,573]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="BBE340077F1AFF80FD15322FFC9A6353" box="[694,785,548,573]" pageId="5" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
exhibits a dorsal habitus similar to large specimens of
|
||
<taxonomicName id="55A45AED7F1AFF80FA24322EFF7F630E" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="A0D0FD7C7F1AFF80FA24322EFF7F630E" italics="true" pageId="5" pageNumber="374">T. lugubris</emphasis>
|
||
</taxonomicName>
|
||
(a rare but widespread species in Europe and N
|
||
<collectingCountry id="EAB361FE7F1AFF80FCB9324CFCF4630E" box="[794,895,583,608]" name="Turkey" pageId="5" pageNumber="374">Anatolia</collectingCountry>
|
||
), but with larger and slightly less elongate body size, and a much shinier dorsal surface, similar to the recently described
|
||
<taxonomicName id="55A45AED7F1AFF80FC6C3266FF4463C6" authority="Audisio, De Biase & Trizzino, 2009" authorityName="Audisio, De Biase & Trizzino" authorityYear="2009" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="foddaii">
|
||
<emphasis id="A0D0FD7C7F1AFF80FC6C3266FBB063EB" box="[975,1083,620,645]" italics="true" pageId="5" pageNumber="374">T. foddaii</emphasis>
|
||
Audisio, De Biase & Trizzino, 2009
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="5060EF8C7F1AFF80FEB03284FEFB63C6" box="[275,368,655,680]" country="Italy" name="Sardegna" pageId="5" pageNumber="374">Sardinia</collectingRegion>
|
||
and
|
||
<collectingCountry id="EAB361FE7F1AFF80FE053284FE7063C6" box="[422,507,655,680]" name="France" pageId="5" pageNumber="374">
|
||
<collectingRegion id="5060EF8C7F1AFF80FE053284FE7063C6" box="[422,507,655,680]" country="France" name="Corse" pageId="5" pageNumber="374">Corsica</collectingRegion>
|
||
</collectingCountry>
|
||
(
|
||
<bibRefCitation id="F6355C9F7F1AFF80FDA93284FD7963C6" author="Audisio" box="[522,754,655,680]" pageId="5" pageNumber="374" refString="Audisio, P., De Biase, A., Trizzino, M., Mancini, E. & Antonini, G. (2009 a) A new species of Meligethes (Coleoptera: Nitidulidae: Meligethinae) of the M. lugubris complex from Sardinia. Zootaxa, 2318, 386 - 393." type="journal article" year="2009" yearSuffix="a">
|
||
Audisio
|
||
<emphasis id="A0D0FD7C7F1AFF80FDC8329AFD2863C6" box="[619,675,655,680]" italics="true" pageId="5" pageNumber="374">et al.</emphasis>
|
||
2009a
|
||
</bibRefCitation>
|
||
). This new species is recognizable from the closely related
|
||
<taxonomicName id="55A45AED7F1AFF80FA24329AFE8A63A3" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="rebmanni">
|
||
<emphasis id="A0D0FD7C7F1AFF80FA24329AFE8A63A3" italics="true" pageId="5" pageNumber="374">T. rebmanni</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="EAB361FE7F1AFF80FEEA32BFFE5963A3" box="[329,466,692,717]" name="Afghanistan" pageId="5" pageNumber="374">Afghanistan</collectingCountry>
|
||
chiefly by parameres more protruding laterad in distal portion of tegmen (
|
||
<figureCitation id="0A9F3DEB7F1AFF80FA8532BFFA0563A2" box="[1318,1422,692,717]" captionStart="FIGURES 3 – 10" captionStartId="4.[151,264,1624,1647]" captionTargetBox="[226,1362,205,1593]" captionTargetId="figure@4.[212,1374,193,1604]" captionTargetPageId="4" captionText="FIGURES 3 – 10. Male genitalia (median lobe of aedeagus and tegmen) of: 3 – 4. Thymogethes ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 5 – 6. T. klapperichi (Easton, 1957) (male from Pakistan, Madaglasht); 7 – 8. T. kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 9 – 10. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3 – 6); = 0.20 mm (Figs 7 – 10)." httpUri="https://zenodo.org/record/242295/files/figure.png" pageId="5" pageNumber="374">Figs 7, 9</figureCitation>
|
||
), and the more raised and concave (bicuspidate) tubercle on the ventral side of last abdominal ventrite (
|
||
<figureCitation id="0A9F3DEB7F1AFF80FAAE32DCFA1B639E" box="[1293,1424,727,752]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1202,1225]" captionTargetBox="[158,1430,193,1168]" captionTargetId="figure@7.[151,1436,193,1182]" captionTargetPageId="7" captionText="FIGURES 19 – 22. SEM images of tubercles on last abdominal ventrite in males of: 19. Thymogethes khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 20. T. ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 21. T. kassites sp. nov. (male holotype from Iran, Zagros Mts, Mt. Oshtoran); 22. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/242297/files/figure.png" pageId="5" pageNumber="374">Figs 21, 22</figureCitation>
|
||
). The partial morphological convergence observed in tegmen shape between
|
||
<taxonomicName id="55A45AED7F1AFF80FC5632F7FB62627B" box="[1013,1257,764,789]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="kassites" status="sp. nov.">
|
||
<emphasis id="A0D0FD7C7F1AFF80FC5632F7FB62627B" box="[1013,1257,764,789]" italics="true" pageId="5" pageNumber="374">Thymogethes kassites</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0D0FD7C7F1AFF80FB5132F7FAC6627B" bold="true" box="[1266,1357,764,789]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="BBE340077F1AFF80FB5132F7FAC6627B" box="[1266,1357,764,789]" pageId="5" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="55A45AED7F1AFF80FA2432F6FEA76256" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="khorasanicus" status="sp. nov.">
|
||
<emphasis id="A0D0FD7C7F1AFF80FA2432F6FEA76256" italics="true" pageId="5" pageNumber="374">T. khorasanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0D0FD7C7F1AFF80FE903314FE066256" bold="true" box="[307,397,799,824]" pageId="5" pageNumber="374">
|
||
<taxonomicNameLabel id="BBE340077F1AFF80FE903314FE066256" box="[307,397,799,824]" pageId="5" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is remarkable (
|
||
<figureCitation id="0A9F3DEB7F1AFF80FD993314FD096259" box="[570,642,799,824]" captionStart="FIGURES 3 – 10" captionStartId="4.[151,264,1624,1647]" captionTargetBox="[226,1362,205,1593]" captionTargetId="figure@4.[212,1374,193,1604]" captionTargetPageId="4" captionText="FIGURES 3 – 10. Male genitalia (median lobe of aedeagus and tegmen) of: 3 – 4. Thymogethes ahriman (Jelínek, 1981) (male from Iran, Zagros Mts., Mt. Oshtoran); 5 – 6. T. klapperichi (Easton, 1957) (male from Pakistan, Madaglasht); 7 – 8. T. kassites sp. nov. (male holotype from Iran, Zagros Mts., Mt. Oshtoran); 9 – 10. T. rebmanni (Easton, 1957) (male from Afghanistan, Kabul). Scale bar = 0.17 mm (Figs 3 – 6); = 0.20 mm (Figs 7 – 10)." httpUri="https://zenodo.org/record/242295/files/figure.png" pageId="5" pageNumber="374">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="0A9F3DEB7F1AFF80FD2D332BFD276256" box="[654,684,800,824]" captionStart="FIGURES 11 – 18" captionStartId="6.[151,264,1565,1588]" captionTargetBox="[262,1351,179,1540]" captionTargetId="figure@6.[236,1352,169,1544]" captionTargetPageId="6" captionText="FIGURES 11 – 18. Male genitalia (median lobe of aedeagus and tegmen) of: 11 – 12, 15 – 16, Thymogethes egenus (Erichson, 1845) s. l. (11 – 12: male from Italy, Lazio, Tivoli; 1 5 – 16: male from N Iran, Kadkan); 13 – 14. T. khorasanicus sp. nov. (male paratype from N Iran, Zoshk Valley above Shandiz); 17 – 18. T. otini (Easton, 1954) (male from Morocco, near Taza). Scale bar = 0.24 mm." httpUri="https://zenodo.org/record/242296/files/figure.png" pageId="5" pageNumber="374">13</figureCitation>
|
||
); however, these two Iranian species obviously belong to two quite distinct groups of species (former in the
|
||
<taxonomicName id="55A45AED7F1AFF80FDFF334EFD5E6233" box="[604,725,836,861]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="lugubris">
|
||
<emphasis id="A0D0FD7C7F1AFF80FDFF334EFD5E6233" box="[604,725,836,861]" italics="true" pageId="5" pageNumber="374">T. lugubris</emphasis>
|
||
</taxonomicName>
|
||
-group, latter in the
|
||
<taxonomicName id="55A45AED7F1AFF80FC17334EFB966232" box="[948,1053,837,860]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="egenus">
|
||
<emphasis id="A0D0FD7C7F1AFF80FC17334EFB966232" box="[948,1053,837,860]" italics="true" pageId="5" pageNumber="374">T. egenus</emphasis>
|
||
</taxonomicName>
|
||
-group).
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1AFF80FF64336CFC986283" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="374">
|
||
<materialsCitation id="22CC2B337F1AFF80FF64336CFC986283" ID-GBIF-Occurrence="1424563916" country="Iran" county="Geographic" location="This" municipality="The" pageId="5" pageNumber="374" specimenCount="10" stateProvince="Central">
|
||
<emphasis id="A0D0FD7C7F1AFF80FF64336CFE7B62EE" bold="true" box="[199,496,871,896]" pageId="5" pageNumber="374">
|
||
<collectingCounty id="7B7A59E27F1AFF80FF64336CFED862EE" box="[199,339,871,896]" pageId="5" pageNumber="374">Geographic</collectingCounty>
|
||
distribution.
|
||
</emphasis>
|
||
<collectingMunicipality id="727FBB147F1AFF80FE5A336CFDAE62EE" box="[505,549,871,896]" pageId="5" pageNumber="374">The</collectingMunicipality>
|
||
single known male specimen is from SW
|
||
<collectingCountry id="EAB361FE7F1AFF80FBAE3363FBB16211" box="[1037,1082,872,895]" name="Iran" pageId="5" pageNumber="374">Iran</collectingCountry>
|
||
(
|
||
<collectingRegion id="5060EF8C7F1AFF80FBE8336CFB1462EE" box="[1099,1183,871,896]" country="Israel" name="Central" pageId="5" pageNumber="374">Central</collectingRegion>
|
||
Zagros Mts; Fig. 25).
|
||
<location id="977B77B57F1AFF80FF343387FF4262CB" LSID="urn:lsid:plazi:treatment:1A0D90787F1DFF80FF34365BFBC06586:977B77B57F1AFF80FF343387FF4262CB" box="[151,201,908,933]" country="Iran" county="Geographic" municipality="The" name="This" pageId="5" pageNumber="374" stateProvince="Central">This</location>
|
||
species is likely rare (a single male specimen collected in company with a series of about
|
||
<specimenCount id="84A2EAE77F1AFF80FB1E3387FADE62CB" box="[1213,1365,908,933]" pageId="5" pageNumber="374" type="generic">10 specimens</specimenCount>
|
||
of the related
|
||
<taxonomicName id="55A45AED7F1AFF80FF4C33BAFEE662A6" box="[239,365,943,968]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="ahriman">
|
||
<emphasis id="A0D0FD7C7F1AFF80FF4C33BAFEE662A6" box="[239,365,943,968]" italics="true" pageId="5" pageNumber="374">T. ahriman</emphasis>
|
||
</taxonomicName>
|
||
), but could be more widely distributed, in suitable habitats, throughout the Zagros Chain. Likewise, the host plant is also widespread in the region.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="921B216E7F1AFF80FF6433FCFC2265CE" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="374">
|
||
<emphasis id="A0D0FD7C7F1AFF80FF6433FCFE05657E" bold="true" box="[199,398,1015,1040]" pageId="5" pageNumber="374">Biological notes.</emphasis>
|
||
The
|
||
<typeStatus id="4D1F9FCC7F1AFF80FE7033F3FD88657E" box="[467,515,1016,1040]" pageId="5" pageNumber="374">type</typeStatus>
|
||
specimen was collected from flowering apical stems of the common
|
||
<taxonomicName id="55A45AED7F1AFF80FAE533FCFE8F655B" class="Magnoliopsida" family="Lamiaceae" genus="Mentha" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="374" phylum="Tracheophyta" rank="species" species="longifolia">
|
||
<emphasis id="A0D0FD7C7F1AFF80FAE533FCFE8F655B" italics="true" pageId="5" pageNumber="374">Mentha longifolia</emphasis>
|
||
</taxonomicName>
|
||
(L.) Huds. (
|
||
<taxonomicName id="55A45AED7F1AFF80FE3B3417FD9C655B" box="[408,535,1052,1077]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="374" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
|
||
), a widespread species known to occur in most West Palearctic areas, in wet places near lakes and ponds, and along the edges of river banks and small springs, from sea level up to
|
||
<quantity id="555C8C8B7F1AFF80FA9C344BFA136539" box="[1343,1432,1088,1112]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="374" unit="m" value="2000.0">2000 m</quantity>
|
||
. Based on the local host-plant flowering period, adults are probably active on mint flowers from May to September, but reproduction likely occurs mainly between early June and August.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DABE72E57F1AFF80FF6434A7FBC06586" pageId="5" pageNumber="374" type="etymology">
|
||
<paragraph id="921B216E7F1AFF80FF6434A7FBC06586" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="374">
|
||
<emphasis id="A0D0FD7C7F1AFF80FF6434A7FEC565AB" bold="true" box="[199,334,1196,1221]" pageId="5" pageNumber="374">Etymology.</emphasis>
|
||
This species is named after the
|
||
<taxonomicName id="55A45AED7F1AFF80FD0C34A7FC8465AB" box="[687,783,1196,1221]" class="Insecta" family="Nitidulidae" genus="Thymogethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="381" phylum="Arthropoda" rank="species" species="kassites" status="sp. nov.">Kassites</taxonomicName>
|
||
, an ancient Near Eastern people who controlled Babylonia and the Zagros Mts after the fall of the Old Babylonian Empire (ca. 1531–1155 BC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |