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<document id="98A46356081233B6441DC4E85D942869" ID-CLB-Dataset="157781" ID-DOI="10.11646/zootaxa.5192.1.1" ID-GBIF-Dataset="8e2cc009-d580-4904-a0b7-9ed7a9e3ab9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138639" ID-ZooBank="236A609B-8A6A-47D7-9BB9-E2FDCA5C37E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664794801684" checkinUser="plazi" docAuthor="Kuwahara, Gregory K. &amp; Marshall, Stephen A." docDate="2022" docId="0F519B74FFCEAD27FF2C92DAFCDBFD42" docLanguage="en" docName="zootaxa.5192.1.1.pdf" docOrigin="Zootaxa 5192 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5192.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Howickia acicula Kuwahara &amp; Marshall 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="F368E30CFFD9AD3DFFBB9376FFA9FFFB" masterDocTitle="A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)" masterLastPageNumber="152" masterPageNumber="1" pageNumber="24" updateTime="1699386905592" updateUser="ExternalLinkService">
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<mods:title id="889CE8786C74EFFE66F5348EFFCADFDA">A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)</mods:title>
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<mods:namePart id="850B592F1917ECC36D3D3FC12D79378E">Kuwahara, Gregory K.</mods:namePart>
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<mods:namePart id="6BA962DECC5A5357F534870EFD8773C5">Marshall, Stephen A.</mods:namePart>
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<treatment id="0F519B74FFCEAD27FF2C92DAFCDBFD42" ID-DOI="http://doi.org/10.5281/zenodo.7138939" ID-GBIF-Taxon="202134580" ID-Zenodo-Dep="7138939" LSID="urn:lsid:plazi:treatment:0F519B74FFCEAD27FF2C92DAFCDBFD42" httpUri="http://treatment.plazi.org/id/0F519B74FFCEAD27FF2C92DAFCDBFD42" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24">
<subSubSection id="CFE279E9FFCEAD2AFF2C92DAFE66FE3C" box="[151,463,428,455]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="87472A62FFCEAD2AFF2C92DAFE66FE3C" blockId="23.[151,463,428,491]" box="[151,463,428,455]" pageId="23" pageNumber="24">
<heading id="DC0F9D0EFFCEAD2AFF2C92DAFE66FE3C" bold="true" box="[151,463,428,455]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<emphasis id="B58CF670FFCEAD2AFF2C92DAFE66FE3C" bold="true" box="[151,463,428,455]" pageId="23" pageNumber="24">
<taxonomicName id="40F851E1FFCEAD2AFF2C92DAFEC3FE3C" ID-CoL="BD3KM" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[151,362,428,455]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="acicula" status="sp. nov.">
<emphasis id="B58CF670FFCEAD2AFF2C92DAFEC3FE3C" bold="true" box="[151,362,428,455]" italics="true" pageId="23" pageNumber="24">Howickia acicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEBF4B0BFFCEAD2AFEC992DBFE66FE3C" box="[370,463,429,455]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="CFE279E9FFCEAD2AFF2C92A6FEBEF808" pageId="23" pageNumber="24" type="description">
<paragraph id="87472A62FFCEAD2AFF2C92A6FEC4FE10" blockId="23.[151,463,428,491]" box="[151,365,464,491]" pageId="23" pageNumber="24">
(
<figureCitation id="1FC336E7FFCEAD2AFF2492A6FEADFE10" box="[159,260,464,491]" captionStart="FIGURES 3.13.22" captionStartId="22.[152,270,1485,1509]" captionTargetBox="[391,1188,189,1455]" captionTargetId="figure-33@22.[386,1202,181,1461]" captionTargetPageId="22" captionText="FIGURES 3.13.22. Mid tibial chaetotaxy of macropterous Australian species of Howickia. 3.1, H. acicula sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.2, H. biantenna sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.3, H. brevior (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.4, H. capitalis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.5, H. erythrocephala sp. nov., a) dorsal, male, b) lateral female; 3.6, H. flaviterga (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.7, H. fuscalis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.8, H. grandisterna sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.9, H. hardyina (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.10, H. kurandensis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.11, H. myrmecophila (Knab &amp; Malloch), a) dorsal, male, b) lateral male, c) lateral, female; 3.12, H. nitidipleura (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.13, H. percostata (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.14, H. pertusa sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.15, H. pseudovittata sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.16, H. subsinuata (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.17, H. symmetrica (Richards), a) dorsal, male, b) lateral male; 3.18, H. trivittata (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.19, H. vernalis (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.20, H. wamini sp. nov., a) dorsal, male, b) lateral male, c) lateral, female; 3.21, H. wilsoni (Richards), a) dorsal, male, b) lateral male, c) lateral, female; 3.22, H. xanthocephala sp. nov., a) dorsal, male, b) lateral male, c) lateral, female. Scale bars indicate 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7138660" httpUri="https://zenodo.org/record/7138660/files/figure.png" pageId="23" pageNumber="24">Figs. 3.1</figureCitation>
,
<figureCitation id="1FC336E7FFCEAD2AFEAB92A7FECCFE10" box="[272,357,465,491]" captionStart="FIGURES 4.14.5" captionStartId="24.[152,270,1901,1925]" captionTargetBox="[181,1406,181,1864]" captionTargetId="figure-33@24.[181,1406,181,1877]" captionTargetPageId="24" captionText="FIGURES 4.14.5. Howickia acicula sp. nov. 4.1, male habitus, lateral; 4.2, male head, anterolateral; 4.3, male S58, ventral; 4.4, male terminalia, lateral; 4.5, aedeagus and associated structures, lateral. Abbreviations: bas—basiphallus, cer—cercus, dist—distiphallus, epa—epandrium, hyp—hypandrium, phal—phallapodeme, post—postgonite, S—sternite, sur—surstylus. Scale bars indicate 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7138664" httpUri="https://zenodo.org/record/7138664/files/figure.png" pageId="23" pageNumber="24">4.14.9</figureCitation>
)
</paragraph>
<paragraph id="87472A62FFCEAD2AFF2C9165FC3AFDD5" blockId="23.[151,1437,531,2035]" box="[151,915,531,558]" pageId="23" pageNumber="24">
<emphasis id="B58CF670FFCEAD2AFF2C9165FE80FDD6" bold="true" box="[151,297,531,557]" pageId="23" pageNumber="24">Description.</emphasis>
<emphasis id="B58CF670FFCEAD2AFE8B9165FE69FDD6" box="[304,448,531,557]" italics="true" pageId="23" pageNumber="24">Body length:</emphasis>
Males
<quantity id="40008787FFCEAD2AFDA99165FD32FDD5" box="[530,667,531,558]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" pageId="23" pageNumber="24" unit="mm" value="3.05" valueMax="3.4" valueMin="2.7">2.73.4 mm</quantity>
, females
<quantity id="40008787FFCEAD2AFCBE9165FC26FDD5" box="[773,911,531,558]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="23" pageNumber="24" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.73.1 mm</quantity>
.
</paragraph>
<paragraph id="87472A62FFCEAD2AFF7C9140FE49FCD8" blockId="23.[151,1437,531,2035]" pageId="23" pageNumber="24">
<emphasis id="B58CF670FFCEAD2AFF7C9140FEA4FDAB" box="[199,269,566,592]" italics="true" pageId="23" pageNumber="24">Head:</emphasis>
Black, lower margin of frons orange, frontal vitta and orbital plates reddish; frontal vitta, interfrontal plates and orbital plates with pale microtrichia; gena reddish anteriorly; antenna brown. Frontal width 2.1x interfrontal height. Two pairs of large interfrontal bristles preceded by a smaller pair 0.5x length of posterior pair; 2 lateroclinate orbital bristles, anterior bristle 0.5x length of posterior bristle. Gena with strong upcurved bristle and 56 smaller setae, finely and densely striate, shiny, lower third microtrichose; vibrissal angle with 3 subvibrissal setae. Face with a slightly flattened triangle along ventromedial margin; palpus clavate with 1 large apical and 1 large preapical seta. Eye height 2.7x genal height.
</paragraph>
<paragraph id="87472A62FFCEAD2AFF7C905BFB1BFC28" blockId="23.[151,1437,531,2035]" pageId="23" pageNumber="24">
<emphasis id="B58CF670FFCEAD2AFF7C905BFE88FCBC" box="[199,289,813,839]" italics="true" pageId="23" pageNumber="24">Thorax:</emphasis>
Dark brown; scutum shiny despite brown microtrichia, posterolateral edge lighter and with pale spots on posterior notopleural lobe; anepimeron with lighter posteroventral spot. Two pairs of dorsocentral bristles (anterior pair 0.5x length of posterior); 45 rows of long, fine acrostichal setulae. Prosternum narrow, linear, posteriorly expanded. Katepisternum with several small setulae in addition to large posterior seta. Scutellum microtrichose, subtriangular, length 0.7x width; 4 long scutellar bristles, apical pair 2.0x length of scutellum.
</paragraph>
<paragraph id="87472A62FFCEAD2AFF7C90ABFD22FB9B" blockId="23.[151,1437,531,2035]" pageId="23" pageNumber="24">
<emphasis id="B58CF670FFCEAD2AFF7C90ABFEAFFC0D" box="[199,262,989,1014]" italics="true" pageId="23" pageNumber="24">Legs:</emphasis>
Brown, tibiae proximally pale brown; tarsi pale brown. Male mid tibia with a stout apicoventral bristle and 2 rows of stout ventral setae along entire length, mid femur with 2 corresponding rows of strong proximoventral setae. Basal half of mid tibia with 3 dorsal bristles (2 anterodorsal and 1 posterodorsal), distal half with 3 dorsal bristles (2 anterodorsal and 1 posterodorsal).
</paragraph>
<paragraph id="87472A62FFCEAD2AFF7C971FFC1EFB31" blockId="23.[151,1437,531,2035]" pageId="23" pageNumber="24">
<emphasis id="B58CF670FFCEAD2AFF7C971FFEFCFB78" box="[199,341,1129,1155]" italics="true" pageId="23" pageNumber="24">Wing/halter:</emphasis>
Strongly infuscate; ventral surface of vein R
<subScript id="1B7C2827FFCEAD2AFCDB970AFCC1FB71" attach="left" box="[864,872,1148,1162]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
white, other veins light brown. CS2 subequal to CS3. R
<subScript id="1B7C2827FFCEAD2AFF5297E9FEABFB56" attach="left" box="[233,258,1183,1197]" fontSize="6" pageId="23" pageNumber="24">2+3</subScript>
sinuate, distal curve stronger than basal curve; R
<subScript id="1B7C2827FFCEAD2AFC8B97E9FCE3FB56" attach="left" box="[816,842,1183,1197]" fontSize="6" pageId="23" pageNumber="24">4+5</subScript>
upcurved, meeting costa well before wing tip; costa extending well past apex of R
<subScript id="1B7C2827FFCEAD2AFE5E97B4FE56FB2B" attach="left" box="[485,511,1218,1232]" fontSize="6" pageId="23" pageNumber="24">4+5</subScript>
(5x costal width). Halter yellow-white.
</paragraph>
<paragraph id="87472A62FFCEAD2AFF7C97A5FCA2F968" blockId="23.[151,1437,531,2035]" pageId="23" pageNumber="24">
<emphasis id="B58CF670FFCEAD2AFF7C97A5FED8FB16" box="[199,369,1235,1261]" italics="true" pageId="23" pageNumber="24">Male abdomen</emphasis>
: Dark brown, almost black, shiny despite microtrichia. T25 and S24 slightly desclerotized along edges, otherwise heavily sclerotized, uniformly long-setose along posterior half with large posterolateral setae. S5 (
<figureCitation id="1FC336E7FFCEAD2AFEAB966FFEC2FACF" box="[272,363,1305,1332]" captionStart="FIGURES 4.14.5" captionStartId="24.[152,270,1901,1925]" captionTargetBox="[181,1406,181,1864]" captionTargetId="figure-33@24.[181,1406,181,1877]" captionTargetPageId="24" captionText="FIGURES 4.14.5. Howickia acicula sp. nov. 4.1, male habitus, lateral; 4.2, male head, anterolateral; 4.3, male S58, ventral; 4.4, male terminalia, lateral; 4.5, aedeagus and associated structures, lateral. Abbreviations: bas—basiphallus, cer—cercus, dist—distiphallus, epa—epandrium, hyp—hypandrium, phal—phallapodeme, post—postgonite, S—sternite, sur—surstylus. Scale bars indicate 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7138664" httpUri="https://zenodo.org/record/7138664/files/figure.png" pageId="23" pageNumber="24">Fig. 4.3</figureCitation>
) large, shiny, broad but short (width 3.0x length), posteromedially desclerotized and bulging with dense patch of short setae, laterally with much longer setae. Epandrium large, hemispherical, and uniformly long-setose; cercus flattened, rectangular with 2 very long setae. Hypandrium medially strongly sinuate; ventral lobe thin, tapered, slightly sinuate with a thin sheet connecting to medial part; posterodorsal lobe short. Surstylus subtriangular and largely bare, with a short, triangular, laterobasal lobe, apically with an extremely large, thickened ventral thorn-like seta and some smaller lateral setae. Postgonite elongate, sinuate, distal third with anterior edge serrate with numerous minute setulae, apex curved forwards into hook. Phallapodeme sinuate with large anterodorsal fin; basiphallus with short, rectangular arms. Distiphallus divided into two distinct sections: basal half heavily sclerotized with a large saddle-like dorsal sclerite and a flattened ventral sclerite articulating to form a tube; distal half with a V-shaped dorsal sclerite and a thin ventral sclerite fused laterally into a tube supporting a membranous apex with numerous dorsoapical rows of small spicules.
</paragraph>
<paragraph id="87472A62FFCEAD2AFF7C95EBFEBEF808" blockId="23.[151,1437,531,2035]" pageId="23" pageNumber="24">
<emphasis id="B58CF670FFCEAD2AFF7C95EBFE3FF94C" box="[199,406,1693,1719]" italics="true" pageId="23" pageNumber="24">Female abdomen:</emphasis>
Dark brown, sometimes desclerotized, shiny despite microtrichia. T25 uniformly longsetose in posterior two-thirds with large posterolateral setae, S25 uniformly long-setose along entire surface. Preabdomen slightly shorter than length of head + thorax. Postabdomen elongate, narrow, 2.6x length of T5. T6 and T7 reduced to a pair of L-shaped lateral plates, each bearing a posterior row of 4 setae. Epiproct rectangular with a triangular posteromedial point and paired dorsomedial setae. Cercus subtriangular with 4 long sinuate setae (1 apical, 2 lateral and 1 dorsal) and several smaller setulae. S67 relatively well-developed, entirely microtrichose, medially desclerotized, bell-shaped plates bearing a posterior row of 10 setae. S8 reduced to an ovoid, microtrichose plate with a preapical pair of setae. Hypoproct largely desclerotized, rounded, microtrichose with a patch of setae. Spermathecae short, stout, conical and entirely lightly grooved; stem subequal in length to spermatheca and cylindrical.
</paragraph>
</subSubSection>
<caption id="D3877AEAFFC1AD25FF23941BFE19F80A" ID-DOI="http://doi.org/10.5281/zenodo.7138664" ID-Zenodo-Dep="7138664" httpUri="https://zenodo.org/record/7138664/files/figure.png" pageId="24" pageNumber="25" startId="24.[152,270,1901,1925]" targetBox="[181,1406,181,1864]" targetPageId="24">
<paragraph id="87472A62FFC1AD25FF23941BFE19F80A" blockId="24.[151,1437,1901,2033]" pageId="24" pageNumber="25">
<emphasis id="B58CF670FFC1AD25FF23941BFECEF87E" bold="true" box="[152,359,1901,1925]" pageId="24" pageNumber="25">FIGURES 4.14.5.</emphasis>
<taxonomicName id="40F851E1FFC1AD25FED7941BFDB6F87E" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[364,543,1901,1925]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="acicula" status="sp. nov.">
<emphasis id="B58CF670FFC1AD25FED7941BFDB6F87E" box="[364,543,1901,1925]" italics="true" pageId="24" pageNumber="25">Howickia acicula</emphasis>
</taxonomicName>
<emphasis id="B58CF670FFC1AD25FD9F941BFDDDF87E" bold="true" box="[548,628,1901,1925]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="AEBF4B0BFFC1AD25FD9F941BFDDDF87E" box="[548,628,1901,1925]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
4.1, male habitus, lateral; 4.2, male head, anterolateral; 4.3, male S58, ventral; 4.4, male terminalia, lateral; 4.5, aedeagus and associated structures, lateral. Abbreviations: bas—basiphallus, cer—cercus, dist—distiphallus, epa—epandrium, hyp—hypandrium, phal—phallapodeme, post—postgonite, S—sternite, sur—surstylus. Scale bars indicate 0.5 mm.
</paragraph>
</caption>
<caption id="D3877AEAFFC0AD24FF2394EEFCB8F803" ID-DOI="http://doi.org/10.5281/zenodo.7138666" ID-Zenodo-Dep="7138666" httpUri="https://zenodo.org/record/7138666/files/figure.png" pageId="25" pageNumber="26" startId="25.[152,270,1944,1968]" targetBox="[258,1330,181,1913]" targetPageId="25">
<paragraph id="87472A62FFC0AD24FF2394EEFCB8F803" blockId="25.[151,1436,1944,2040]" pageId="25" pageNumber="26">
<emphasis id="B58CF670FFC0AD24FF2394EEFECFF84B" bold="true" box="[152,358,1944,1968]" pageId="25" pageNumber="26">FIGURES 4.64.9.</emphasis>
<taxonomicName id="40F851E1FFC0AD24FED094EEFDB4F84B" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[363,541,1944,1968]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="acicula" status="sp. nov.">
<emphasis id="B58CF670FFC0AD24FED094EEFDB4F84B" box="[363,541,1944,1968]" italics="true" pageId="25" pageNumber="26">Howickia acicula</emphasis>
</taxonomicName>
<emphasis id="B58CF670FFC0AD24FD9994EEFDD8F84B" bold="true" box="[546,625,1944,1968]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="AEBF4B0BFFC0AD24FD9994EEFDD8F84B" box="[546,625,1944,1968]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
4.6, left wing; 4.7, female terminalia, dorsal; 4.8, female terminalia, ventral; 4.9, spermathecae. Abbreviations: epi—epiproct, hypo—hypoproct, S—sternite, sprm—spermatheca, T—tergite. Female cerci were excluded from ventral illustration. Scale bar indicates 0.5 mm.
</paragraph>
</caption>
<subSubSection id="CFE279E9FFC3AD27FF7C93E0FB40FE7A" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="87472A62FFC3AD27FF7C93E0FBF6FF28" blockId="26.[151,1437,150,697]" pageId="26" pageNumber="27">
<emphasis id="B58CF670FFC3AD27FF7C93E0FE1BFF4B" bold="true" box="[199,434,150,176]" pageId="26" pageNumber="27">Material examined.</emphasis>
<materialsCitation id="3790203FFFC3AD27FE0193E1FBF6FF28" ID-GBIF-Occurrence="3920718444" collectingDate="2003-12-14" collectionCode="TMAG" collectorName="S. A. Marshall" country="Australia" location="Falls Rd." municipality="Cuckoo Falls" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="584394C0FFC3AD27FE0193E1FD8EFF4A" box="[442,551,151,177]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="FFEF6AF2FFC3AD27FD8A93E1FD7DFF4A" box="[561,724,151,177]" name="Australia" pageId="26" pageNumber="27">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FFC3AD27FD5B93E0FCF4FF4B" bold="true" box="[736,861,150,176]" pageId="26" pageNumber="27">
<collectingRegion id="453CE480FFC3AD27FD5B93E0FCF1FF4B" box="[736,856,150,176]" country="Australia" name="Tasmania" pageId="26" pageNumber="27">Tasmania</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6723B018FFC3AD27FCDE93E1FC54FF4A" box="[869,1021,151,177]" pageId="26" pageNumber="27">Cuckoo Falls</collectingMunicipality>
,
<locationDeviation id="96F53340FFC3AD27FBB193E1FB75FF4A" box="[1034,1244,151,177]" location="Scottsdale" pageId="26" pageNumber="27">
<quantity id="40008787FFC3AD27FBB193E1FBEDFF4A" box="[1034,1092,151,177]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="km" value="4.0">4 km</quantity>
E Scottsdale
</locationDeviation>
, along
<location id="82277CB9FFC3AD27FA9493E1FA31FF4A" LSID="urn:lsid:plazi:treatment:0F519B74FFCEAD27FF2C92DAFCDBFD42:82277CB9FFC3AD27FA9493E1FA31FF4A" box="[1327,1432,151,177]" country="Australia" municipality="Cuckoo Falls" name="Falls Rd." pageId="26" pageNumber="27" stateProvince="Tasmania">Falls Rd.</location>
, along creek, pans/some dung,
<date id="F3460CA2FFC3AD27FE5193CFFDC5FF28" box="[490,620,185,211]" pageId="26" pageNumber="27" value="2003-12-14">
<collectingDate id="E302F54AFFC3AD27FE5193CFFDC5FF28" box="[490,620,185,211]" pageId="26" pageNumber="27" value="2003-12-14">14.xii.2003</collectingDate>
</date>
,
<collectorName id="2A0D4FB4FFC3AD27FDC393CFFCBEFF28" box="[632,791,185,211]" pageId="26" pageNumber="27">S.A. Marshall</collectorName>
, debu00248893 (
<specimenCount id="91FEE1EBFFC3AD27FC6293CFFC59FF28" box="[985,1008,185,211]" pageId="26" pageNumber="27" type="male"></specimenCount>
,
<collectionCode id="E1E9B2A7FFC3AD27FC4093CFFBFAFF28" box="[1019,1107,185,211]" country="Australia" httpUri="http://grbio.org/cool/88vw-h26m" name="Tasmanian Museum and Art Gallery" pageId="26" pageNumber="27">TMAG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="87472A62FFC3AD27FF7C93ABFADDFEC0" blockId="26.[151,1437,150,697]" pageId="26" pageNumber="27">
<materialsCitation id="3790203FFFC3AD27FF7C93ABFD7AFEE2" ID-GBIF-Occurrence="3920718304" collectingDate="2003-12-12" collectingDateMax="2003-12-13" collectingDateMin="2003-12-12" collectionCode="DEBU" collectorName="S. A. Marshall" country="Australia" county="Mount Williams National Park" location="Stumpys Bay" municipality="Stumpys Bay Campground" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Tasmania" typeStatus="paratype">
<typeStatus id="584394C0FFC3AD27FF7C93ABFE91FF0C" box="[199,312,221,247]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="FFEF6AF2FFC3AD27FEF893AAFE4FFF0D" box="[323,486,220,246]" name="Australia" pageId="26" pageNumber="27">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FFC3AD27FE4893AAFDD9FF0D" bold="true" box="[499,624,220,246]" pageId="26" pageNumber="27">
<collectingRegion id="453CE480FFC3AD27FE4893AAFDC2FF0D" box="[499,619,220,246]" country="Australia" name="Tasmania" pageId="26" pageNumber="27">Tasmania</collectingRegion>
:
</emphasis>
<collectingCounty id="6E2652EEFFC3AD27FDC293ABFC73FF0D" box="[633,986,220,247]" pageId="26" pageNumber="27">Mount Williams National Park</collectingCounty>
,
<collectingMunicipality id="6723B018FFC3AD27FC5C93AAFAB4FF0D" box="[999,1309,220,247]" pageId="26" pageNumber="27">
<location id="82277CB9FFC3AD27FC5C93AAFBD6FF0C" LSID="urn:lsid:plazi:treatment:0F519B74FFCEAD27FF2C92DAFCDBFD42:82277CB9FFC3AD27FC5C93AAFBD6FF0C" box="[999,1151,220,247]" country="Australia" county="Mount Williams National Park" municipality="Stumpys Bay Campground" name="Stumpys Bay" pageId="26" pageNumber="27" stateProvince="Tasmania">Stumpys Bay</location>
Campground
</collectingMunicipality>
, dry heath, pans,
<date id="F3460CA2FFC3AD27FF629389FE2FFEE2" box="[217,390,255,281]" pageId="26" pageNumber="27" value="2003-12-12" valueMax="2003-12-13" valueMin="2003-12-12">
<collectingDate id="E302F54AFFC3AD27FF629389FE2FFEE2" box="[217,390,255,281]" pageId="26" pageNumber="27" value="2003-12-12" valueMax="2003-12-13" valueMin="2003-12-12">1213.xii.2003</collectingDate>
</date>
,
<collectorName id="2A0D4FB4FFC3AD27FE299389FD99FEE2" box="[402,560,255,281]" pageId="26" pageNumber="27">S.A. Marshall</collectorName>
(
<specimenCount id="91FEE1EBFFC3AD27FDF99389FDC4FEE2" box="[578,621,255,281]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
,
<collectionCode id="E1E9B2A7FFC3AD27FDC19389FD62FEE2" box="[634,715,255,281]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="26" pageNumber="27">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3790203FFFC3AD27FD649389FABEFEE2" ID-GBIF-Occurrence="3920718646" box="[735,1303,255,281]" collectingDate="2003-12-14" collectionCode="TMAG, DEBU" collectorName="S. A. Marshall" country="Australia" location="Falls Rd." municipality="Cuckoo Falls" pageId="26" pageNumber="27" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Tasmania" typeStatus="paratype">
same data as holotype (
<specimenCount id="91FEE1EBFFC3AD27FC529389FBBEFEE2" box="[1001,1047,255,281]" pageId="26" pageNumber="27" type="male">9 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFFC3AD27FB989389FBE7FEE2" box="[1059,1102,255,281]" pageId="26" pageNumber="27" type="female">6 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FFC3AD27FBE69389FB1BFEE2" box="[1117,1202,255,281]" country="Australia" httpUri="http://grbio.org/cool/88vw-h26m" name="Tasmanian Museum and Art Gallery" pageId="26" pageNumber="27">TMAG</collectionCode>
,
<collectionCode id="E1E9B2A7FFC3AD27FB059389FAA6FEE2" box="[1214,1295,255,281]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="26" pageNumber="27">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3790203FFFC3AD27FA989389FAD9FEC0" ID-GBIF-Occurrence="3920718382" collectingDate="2003-12-20" collectingDateMax="2003-12-21" collectingDateMin="2003-12-20" collectionCode="TMAG, DEBU" collectorName="S. A. Marshall" country="Australia" location="Waratah" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="2" stateProvince="Tasmania" typeStatus="paratype">
<location id="82277CB9FFC3AD27FA989389FA2AFEE2" LSID="urn:lsid:plazi:treatment:0F519B74FFCEAD27FF2C92DAFCDBFD42:82277CB9FFC3AD27FA989389FA2AFEE2" box="[1315,1411,255,281]" country="Australia" name="Waratah" pageId="26" pageNumber="27" stateProvince="Tasmania">Waratah</location>
,
<quantity id="40008787FFC3AD27FA359389FF12FEC0" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="27" unit="km" value="8.0">8 km</quantity>
SW, 41˚2904”S 145˚2741”E, rainforest, dung pans,
<date id="F3460CA2FFC3AD27FCA09257FC6EFEC0" box="[795,967,289,315]" pageId="26" pageNumber="27" value="2003-12-20" valueMax="2003-12-21" valueMin="2003-12-20">
<collectingDate id="E302F54AFFC3AD27FCA09257FC6EFEC0" box="[795,967,289,315]" pageId="26" pageNumber="27" value="2003-12-20" valueMax="2003-12-21" valueMin="2003-12-20">2021.xii.2003</collectingDate>
</date>
,
<collectorName id="2A0D4FB4FFC3AD27FC699257FBC6FEC0" box="[978,1135,289,315]" pageId="26" pageNumber="27">S.A. Marshall</collectorName>
(
<specimenCount id="91FEE1EBFFC3AD27FBC59257FB00FEC7" box="[1150,1193,289,316]" pageId="26" pageNumber="27" type="female">2 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FFC3AD27FB0C9257FAA5FEC0" box="[1207,1292,289,315]" country="Australia" httpUri="http://grbio.org/cool/88vw-h26m" name="Tasmanian Museum and Art Gallery" pageId="26" pageNumber="27">TMAG</collectionCode>
,
<collectionCode id="E1E9B2A7FFC3AD27FAAC9257FAC1FEC0" box="[1303,1384,289,315]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="26" pageNumber="27">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="87472A62FFC3AD27FF7C9232FB40FE7A" blockId="26.[151,1437,150,697]" pageId="26" pageNumber="27">
<materialsCitation id="3790203FFFC3AD27FF7C9232FB4FFE7A" ID-GBIF-Occurrence="3920718688" collectingDate="2003-12-20" collectingDateMax="2003-12-21" collectingDateMin="2003-12-20" collectionCode="DEBU" collectorName="S. A. Marshall" country="Australia" location="Waratah" municipality="Cuckoo Falls" pageId="26" pageNumber="27" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Tasmania">
Other material examined:
<collectingCountry id="FFEF6AF2FFC3AD27FE509232FD27FEA5" box="[491,654,324,350]" name="Australia" pageId="26" pageNumber="27">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FFC3AD27FD229232FCBFFEA5" bold="true" box="[665,790,324,350]" pageId="26" pageNumber="27">
<collectingRegion id="453CE480FFC3AD27FD229232FCB8FEA5" box="[665,785,324,350]" country="Australia" name="Tasmania" pageId="26" pageNumber="27">Tasmania</collectingRegion>
:
</emphasis>
same data as holotype (
<specimenCount id="91FEE1EBFFC3AD27FB9D9232FBF9FEA4" box="[1062,1104,324,351]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFFC3AD27FBE79232FB2EFEA4" box="[1116,1159,324,351]" pageId="26" pageNumber="27" type="female">4 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FFC3AD27FB2E9232FB4FFEA5" box="[1173,1254,324,350]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="26" pageNumber="27">DEBU</collectionCode>
);
<location id="82277CB9FFC3AD27FB429232FAF0FEA5" LSID="urn:lsid:plazi:treatment:0F519B74FFCEAD27FF2C92DAFCDBFD42:82277CB9FFC3AD27FB429232FAF0FEA5" box="[1273,1369,324,350]" country="Australia" name="Waratah" pageId="26" pageNumber="27" stateProvince="Tasmania">Waratah</location>
,
<quantity id="40008787FFC3AD27FAD89232FA34FEA5" box="[1379,1437,324,350]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="27" unit="km" value="8.0">8 km</quantity>
SW, 41˚2904”S 145˚2741”E, rainforest, dung pans,
<date id="F3460CA2FFC3AD27FD4B9211FC35FE7A" box="[752,924,359,385]" pageId="26" pageNumber="27" value="2003-12-20" valueMax="2003-12-21" valueMin="2003-12-20">
<collectingDate id="E302F54AFFC3AD27FD4B9211FC35FE7A" box="[752,924,359,385]" pageId="26" pageNumber="27" value="2003-12-20" valueMax="2003-12-21" valueMin="2003-12-20">2021.xii.2003</collectingDate>
</date>
,
<collectorName id="2A0D4FB4FFC3AD27FC139211FBEDFE7A" box="[936,1092,359,385]" pageId="26" pageNumber="27">S.A. Marshall</collectorName>
(
<specimenCount id="91FEE1EBFFC3AD27FBED9211FBD7FE7A" box="[1110,1150,359,385]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FFC3AD27FB379211FB74FE7A" box="[1164,1245,359,385]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="26" pageNumber="27">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFC3AD27FF7C92FFFD78FE3D" pageId="26" pageNumber="27" type="etymology">
<paragraph id="87472A62FFC3AD27FF7C92FFFD78FE3D" blockId="26.[151,1437,150,697]" pageId="26" pageNumber="27">
<emphasis id="B58CF670FFC3AD27FF7C92FFFEE7FE58" bold="true" box="[199,334,393,419]" pageId="26" pageNumber="27">Etymology.</emphasis>
This name is in reference to the elongate, pin- or needle-like postgonite and surstylar seta (Latin
<taxonomicName id="40F851E1FFC3AD27FF2C92DAFF40FE3D" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[151,233,428,454]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="acicula">
<emphasis id="B58CF670FFC3AD27FF2C92DAFF40FE3D" box="[151,233,428,454]" italics="true" pageId="26" pageNumber="27">acicula</emphasis>
</taxonomicName>
: “a needle-like spine or bristle; a hairpin”).
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFC3AD27FF7C92B8FCDBFD42" pageId="26" pageNumber="27" type="discussion">
<paragraph id="87472A62FFC3AD27FF7C92B8FCDBFD42" blockId="26.[151,1437,150,697]" pageId="26" pageNumber="27">
<emphasis id="B58CF670FFC3AD27FF7C92B8FEE6FE13" bold="true" box="[199,335,462,488]" pageId="26" pageNumber="27">Comments.</emphasis>
<taxonomicName id="40F851E1FFC3AD27FEEC92B9FDB5FE12" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[343,540,463,489]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="acicula">
<emphasis id="B58CF670FFC3AD27FEEC92B9FDB5FE12" box="[343,540,463,489]" italics="true" pageId="26" pageNumber="27">Howickia acicula</emphasis>
</taxonomicName>
is part of the
<taxonomicName id="40F851E1FFC3AD27FD0092B9FCE0FE12" authorityName="ACT, NSW, TAS, VIC" baseAuthorityName="Richards" baseAuthorityYear="1973" box="[699,841,463,489]" class="Insecta" family="Sphaeroceridae" genus="Australimosina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="flaviterga">
<emphasis id="B58CF670FFC3AD27FD0092B9FCE0FE12" box="[699,841,463,489]" italics="true" pageId="26" pageNumber="27">H. flaviterga</emphasis>
</taxonomicName>
species group, which also includes
<taxonomicName id="40F851E1FFC3AD27FB6492B9FADBFE12" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[1247,1394,463,489]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="biantenna">
<emphasis id="B58CF670FFC3AD27FB6492B9FADBFE12" box="[1247,1394,463,489]" italics="true" pageId="26" pageNumber="27">H. biantenna</emphasis>
</taxonomicName>
,
<taxonomicName id="40F851E1FFC3AD27FA3A92B9FEAAFDF0" authorityName="ACT, NSW, TAS, VIC" baseAuthorityName="Richards" baseAuthorityYear="1973" class="Insecta" family="Sphaeroceridae" genus="Australimosina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="flaviterga">
<emphasis id="B58CF670FFC3AD27FA3A92B9FEAAFDF0" italics="true" pageId="26" pageNumber="27">H. flaviterga</emphasis>
</taxonomicName>
,
<taxonomicName id="40F851E1FFC3AD27FEAB9284FE6BFDF0" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[272,450,497,523]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="grandisterna">
<emphasis id="B58CF670FFC3AD27FEAB9284FE6BFDF0" box="[272,450,497,523]" italics="true" pageId="26" pageNumber="27">H. grandisterna</emphasis>
</taxonomicName>
, and
<emphasis id="B58CF670FFC3AD27FE469284FD32FDF0" box="[509,667,498,523]" italics="true" pageId="26" pageNumber="27">
<taxonomicName id="40F851E1FFC3AD27FE469284FD3EFDF0" box="[509,663,498,523]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="percostata">H. percostata</taxonomicName>
.
</emphasis>
It differs from
<taxonomicName id="40F851E1FFC3AD27FCF99284FC5DFDF0" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[834,1012,497,523]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="grandisterna">
<emphasis id="B58CF670FFC3AD27FCF99284FC5DFDF0" box="[834,1012,497,523]" italics="true" pageId="26" pageNumber="27">H. grandisterna</emphasis>
</taxonomicName>
and
<taxonomicName id="40F851E1FFC3AD27FB9C9284FB17FDF0" box="[1063,1214,498,523]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="percostata">
<emphasis id="B58CF670FFC3AD27FB9C9284FB17FDF0" box="[1063,1214,498,523]" italics="true" pageId="26" pageNumber="27">H. percostata</emphasis>
</taxonomicName>
in having four rows of acrostichal setulae and vein R
<subScript id="1B7C2827FFC3AD27FDBA9150FDB3FDCF" attach="left" box="[513,538,550,564]" fontSize="6" pageId="26" pageNumber="27">2+3</subScript>
separated from the costa by 2.5x the costal width, and can be distinguished from
<taxonomicName id="40F851E1FFC3AD27FF2C9141FE82FDAA" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[151,299,567,593]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="biantenna">
<emphasis id="B58CF670FFC3AD27FF2C9141FE82FDAA" box="[151,299,567,593]" italics="true" pageId="26" pageNumber="27">H. biantenna</emphasis>
</taxonomicName>
by the presence of two pairs of dorsocentral bristles.
<taxonomicName id="40F851E1FFC3AD27FC289141FBF0FDAA" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[915,1113,567,593]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="acicula">
<emphasis id="B58CF670FFC3AD27FC289141FBF0FDAA" box="[915,1113,567,593]" italics="true" pageId="26" pageNumber="27">Howickia acicula</emphasis>
</taxonomicName>
is externally very similar to
<taxonomicName id="40F851E1FFC3AD27FF2C912CFE8CFD88" authorityName="ACT, NSW, TAS, VIC" baseAuthorityName="Richards" baseAuthorityYear="1973" box="[151,293,601,627]" class="Insecta" family="Sphaeroceridae" genus="Australimosina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="flaviterga">
<emphasis id="B58CF670FFC3AD27FF2C912CFE8CFD88" box="[151,293,601,627]" italics="true" pageId="26" pageNumber="27">H. flaviterga</emphasis>
</taxonomicName>
, though it differs in only having four rows of acrostichal setulae and a slightly longer scutellum. Male genitalia also differ significantly.
<taxonomicName id="40F851E1FFC3AD27FDB0910AFD67FD6D" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[523,718,636,662]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="acicula">
<emphasis id="B58CF670FFC3AD27FDB0910AFD67FD6D" box="[523,718,636,662]" italics="true" pageId="26" pageNumber="27">Howickia acicula</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="40F851E1FFC3AD27FC23910BFB80FD6D" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[920,1065,636,662]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="biantenna">
<emphasis id="B58CF670FFC3AD27FC23910BFB80FD6D" box="[920,1065,636,662]" italics="true" pageId="26" pageNumber="27">H. biantenna</emphasis>
</taxonomicName>
and
<taxonomicName id="40F851E1FFC3AD27FBE6910BFAA6FD6D" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[1117,1295,636,662]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="grandisterna">
<emphasis id="B58CF670FFC3AD27FBE6910BFAA6FD6D" box="[1117,1295,636,662]" italics="true" pageId="26" pageNumber="27">H. grandisterna</emphasis>
</taxonomicName>
, which were also collected at the same time in Mount Williams National Park.
</paragraph>
</subSubSection>
</treatment>
</document>