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<document id="B6EA3FC2D355B26C7B82FE06034B45FA" ID-CLB-Dataset="157781" ID-DOI="10.11646/zootaxa.5192.1.1" ID-GBIF-Dataset="8e2cc009-d580-4904-a0b7-9ed7a9e3ab9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138639" ID-ZooBank="236A609B-8A6A-47D7-9BB9-E2FDCA5C37E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664794801684" checkinUser="plazi" docAuthor="Kuwahara, Gregory K. &amp; Marshall, Stephen A." docDate="2022" docId="0F519B74FF54ADB2FF2C93E1FB4FFEBA" docLanguage="en" docName="zootaxa.5192.1.1.pdf" docOrigin="Zootaxa 5192 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5192.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Howickia tricolor Kuwahara &amp; Marshall 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="144" masterDocId="F368E30CFFD9AD3DFFBB9376FFA9FFFB" masterDocTitle="A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)" masterLastPageNumber="152" masterPageNumber="1" pageNumber="142" updateTime="1699386905592" updateUser="ExternalLinkService">
<mods:mods id="39BB50AB23A1BFCFA4F87A3472CD74A8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="33A218F1DF62E0A7825368E30A97BACA">
<mods:title id="C7371A6CEA4AFC0D576CBFC860C34DD3">A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)</mods:title>
</mods:titleInfo>
<mods:name id="5257B8FB5DE88814CF58D051F785F3AB" type="personal">
<mods:role id="270FEB1DE5836C3584DC9F65BF39B57A">
<mods:roleTerm id="1D99916C88D80E81A001F0A1A7A75EDB">Author</mods:roleTerm>
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<mods:namePart id="8020938C496B1CA913A633540B2F0341">Kuwahara, Gregory K.</mods:namePart>
<mods:nameIdentifier id="67525A771630DD58C0484CB7A2CE728F" type="ORCID">0000-0002-0319-2672</mods:nameIdentifier>
<mods:nameIdentifier id="BD08A128819A39888349700FDE30DD39" type="email">gkuwahar@uoguelph.ca</mods:nameIdentifier>
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<mods:name id="C07EDE1D566838A90463D2C8EF43E76B" type="personal">
<mods:role id="15DCB9309FC95DDFE52F0FAB94F5120A">
<mods:roleTerm id="21181AABF9942FABD0D679B8B513AF9C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA7761BB05C036B7CC52745D2187924E">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="911C3985B458EBAC11ED23FE794E3FB9" type="ORCID">0000-0002-5732-9718</mods:nameIdentifier>
<mods:nameIdentifier id="9E7D558FA3895F2CB4C56F0EF1EEBE24" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
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<mods:titleInfo id="D58B5A495DF6B14D6AB686FCF635EF0B">
<mods:title id="417BFC7998E9D09903EA1FAC01D8CD83">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1F3C545309FBA4F320A52AFD63E2EEDF">
<mods:date id="F14EBABF973A73DEF6A3594FB881D7E7">2022</mods:date>
<mods:detail id="4CEF2CD3C02934B5125FA5CCE8A61578" type="pubDate">
<mods:number id="55B29E2DB50C3221A90B4A180E3F9427">2022-10-03</mods:number>
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<mods:detail id="6C131040A3B2F48293DFD0E2927111B6" type="volume">
<mods:number id="6831BBA769C818E9600A0BFB64FE9CB1">5192</mods:number>
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<mods:number id="A8688F3751CD901F449CDCC72977CD29">1</mods:number>
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<mods:start id="1F8247B73680D0CA2E920D4BC572A96C">1</mods:start>
<mods:end id="DBFE5245805DAC5CBEC53DEC753EEBD4">152</mods:end>
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<mods:url id="0D5C2E55FED1137E450F3E1CF3203EC6">http://dx.doi.org/10.11646/zootaxa.5192.1.1</mods:url>
</mods:location>
<mods:classification id="8FF7E7924BAD3C22DD100BF87A8E8235">journal article</mods:classification>
<mods:identifier id="330BB7345554F94E415E78E5453C3122" type="CLB-Dataset">157781</mods:identifier>
<mods:identifier id="577A09041A278641A0ECB9F82234374A" type="DOI">10.11646/zootaxa.5192.1.1</mods:identifier>
<mods:identifier id="58734C7C8E241389BD880165BDD73DC0" type="GBIF-Dataset">8e2cc009-d580-4904-a0b7-9ed7a9e3ab9e</mods:identifier>
<mods:identifier id="B595C3A85841F9D7869C4F2BDB8819B4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0F519B74FF54ADB2FF2C93E1FB4FFEBA" ID-DOI="http://doi.org/10.5281/zenodo.7138863" ID-GBIF-Taxon="202134547" ID-Zenodo-Dep="7138863" LSID="urn:lsid:plazi:treatment:0F519B74FF54ADB2FF2C93E1FB4FFEBA" httpUri="http://treatment.plazi.org/id/0F519B74FF54ADB2FF2C93E1FB4FFEBA" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142">
<subSubSection id="CFE279E9FF54ADB0FF2C93E1FE78FF4A" box="[151,465,151,178]" pageId="141" pageNumber="142" type="nomenclature">
<paragraph id="87472A62FF54ADB0FF2C93E1FE78FF4A" blockId="141.[151,465,151,213]" box="[151,465,151,178]" pageId="141" pageNumber="142">
<heading id="DC0F9D0EFF54ADB0FF2C93E1FE78FF4A" bold="true" box="[151,465,151,178]" fontSize="11" level="2" pageId="141" pageNumber="142" reason="4">
<emphasis id="B58CF670FF54ADB0FF2C93E1FE78FF4A" bold="true" box="[151,465,151,178]" pageId="141" pageNumber="142">
<taxonomicName id="40F851E1FF54ADB0FF2C93E1FEC5FF49" ID-CoL="BD3LS" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[151,364,151,178]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="tricolor" status="sp. nov.">
<emphasis id="B58CF670FF54ADB0FF2C93E1FEC5FF49" bold="true" box="[151,364,151,178]" italics="true" pageId="141" pageNumber="142">Howickia tricolor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEBF4B0BFF54ADB0FEC893E1FE78FF4A" box="[371,465,151,177]" pageId="141" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FF54ADB0FF2C93CDFE7AF9F2" pageId="141" pageNumber="142" type="description">
<paragraph id="87472A62FF54ADB0FF2C93CDFEF1FF2E" blockId="141.[151,465,151,213]" box="[151,344,187,213]" pageId="141" pageNumber="142">
(
<figureCitation id="1FC336E7FF54ADB0FF2493CDFEF9FF2E" box="[159,336,187,213]" captionStart="FIGURES 44.1–44.5" captionStartId="142.[152,270,1943,1967]" captionTargetBox="[156,1431,181,1913]" captionTargetId="figure-33@142.[156,1431,181,1919]" captionTargetPageId="142" captionText="FIGURES 44.1–44.5. Howickia tricolor sp. nov. 44.1, male habitus, lateral; 44.2, male head, anterior; 44.3, male S5–8, ventral; 44.4, male terminalia, lateral; 44.5, aedeagus and associated structures, lateral. Body scale bar indicates 0.5 mm, head scale bar indicates 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7138899" httpUri="https://zenodo.org/record/7138899/files/figure.png" pageId="141" pageNumber="142">Figs. 44.1–44.8</figureCitation>
)
</paragraph>
<paragraph id="87472A62FF54ADB0FF2C9275FC3AFEE5" blockId="141.[151,1437,259,2014]" box="[151,915,259,286]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF2C9275FE80FEE6" bold="true" box="[151,297,259,285]" pageId="141" pageNumber="142">Description.</emphasis>
<emphasis id="B58CF670FF54ADB0FE8B9275FE69FEE6" box="[304,448,259,285]" italics="true" pageId="141" pageNumber="142">Body length:</emphasis>
Males 1.2–2.0 mm, females
<quantity id="40008787FF54ADB0FCBE9275FC26FEE5" box="[773,911,259,286]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.4" metricValueMin="1.3" pageId="141" pageNumber="142" unit="mm" value="1.85" valueMax="2.4" valueMin="1.3">1.3–2.4 mm</quantity>
.
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C9251FD4EFE0E" blockId="141.[151,1437,259,2014]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF7C9251FEA4FEBA" box="[199,269,295,321]" italics="true" pageId="141" pageNumber="142">Head:</emphasis>
Reddish-brown, darkest behind level of ocelli; frontal vitta rugose and broad; gena reddish-brown; antennae yellow. Frontal width 1.6x interfrontal height. Two pairs of strong interfrontal bristles, preceded by a slightly smaller pair (0.6–0.7x length of posterior pairs); 1 lateroclinate orbital bristle, preceded by a minute setula. Ocelli present but small. Gena with 2–3 fine setae, very finely rugose, shiny, lower quarter microtrichose; vibrissal angle with 1 smaller seta. Face bulging and evenly convex, somewhat continuous with lunule; palpus clavate with 2 large preapical setae. Eye height 3.0x genal height.
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C9289FE98FD7E" blockId="141.[151,1437,259,2014]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF7C9289FE88FDE2" box="[199,289,511,537]" italics="true" pageId="141" pageNumber="142">Thorax:</emphasis>
Pale orange; scutum shiny despite dense microtrichia, lateral edges yellow; katatergite shining black. One pair of dorsocentral bristles; 7–8 rows of short, fine acrostichal setae. Prosternum narrow, linear, posteriorly expanded. Scutellum rugose, semicircular, its length ~0.4x its width; 4 scutellar bristles long, basal pair 1.8x length of apical pair.
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C91E6FD9AFD0A" blockId="141.[151,1437,259,2014]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF7C91E6FEAFFD52" box="[199,262,656,681]" italics="true" pageId="141" pageNumber="142">Legs:</emphasis>
Yellow; tarsi pale yellow, apical 4 tarsomeres of fore leg white. Male mid tibia with a large apicoventral bristle. Basal half of mid tibia with 3 dorsal setae (2 anterodorsal and 1 posterodorsal), distal half with 3 dorsal setae (2 anterodorsal and 1 posterodorsal).
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C918DFE51FCEE" blockId="141.[151,1437,259,2014]" box="[199,504,763,789]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF7C918DFEFCFCEE" box="[199,341,763,789]" italics="true" pageId="141" pageNumber="142">Wing/halter:</emphasis>
Halter absent.
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C9069FBFAFB85" blockId="141.[151,1437,259,2014]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF7C9069FEC6FCC2" box="[199,367,799,825]" italics="true" pageId="141" pageNumber="142">Male abdomen</emphasis>
: Black, heavily sclerotized, shiny, postabdomen reddish. T2–5 and S2–4 uniformly long-setose along entire surface. S5 broad but short (width 4.0x medial length), asymmetrical (right side 1.7x longer than left), sparsely long-setose, broadly posteromedially emarginate with a posterior row of thickened setae, shortest medially and longest laterally. Epandrium moderate, uniformly long-setose; cercus flat, somewhat bulging ventrolaterally, and with numerous long setae. Hypandrium medially sinuate; ventral lobe small, medially sclerotized and broadly membranous laterally; posterodorsal lobe long but thin, largely membranous. Surstylus bilobed, anterior lobe small and rounded, posterior lobe larger, triangular, slightly twisted and folded inwards with a very thick, curved, apicoventral thorn-like seta and many fine, posterior setulae. Postgonite very long, strongly sinuate, and gradually tapered. Phallapodeme slightly sinuate with a short dorsal ‘fin’ in the apical two-thirds; basiphallus short, rounded posteriorly. Distiphallus short, stout, well-sclerotized with a small membranous apex.
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C97F1FE7AF9F2" blockId="141.[151,1437,259,2014]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF7C97F1FE38FB5A" box="[199,401,1159,1185]" italics="true" pageId="141" pageNumber="142">Female abdomen:</emphasis>
Black, heavily sclerotized, shiny despite dense microtrichia. T2–5 and S2–5 uniformly longsetose along entire surface. Preabdomen slightly longer than length of head + thorax. Postabdomen short, 2.0x length of T5. T6 well-developed and heavily sclerotized, densely microtrichose, and bearing a posterior row of setae; T7 well-developed and heavily sclerotized, shiny with only two lateral patches of microtrichia, with a small posteromedial emargination and only 4 posterior setae. Dorsal sclerite of T8 absent, lateral plates subquadrate. Epiproct small, shield-like, medially desclerotized and dorsomedially bearing a pair of setae. Cercus short, rounded with 4–5 long, sinuate setae (1 apical, 1–2 lateral, 1 dorsal, 1 preapical). S6 and S7 well-developed and heavily sclerotized, bearing a posterior row of setae. S8 elongate, U-shaped, anterior margin deeply emarginate, posterior third heavily microtrichose. Hypoproct small, shield-like, posteriorly microtrichose with a row of 4 stout setulae. Spermathecae stout, hemispherical and entirely lightly grooved; stem elongate (1.5–1.6x length of spermatheca), tapered and swollen basally.
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FF54ADB0FF7C9565FAA4F826" pageId="141" pageNumber="142" type="materials_examined">
<paragraph id="87472A62FF54ADB0FF7C9565FE8DF98E" blockId="141.[151,1437,259,2014]" pageId="141" pageNumber="142">
<emphasis id="B58CF670FF54ADB0FF7C9565FE1DF9D6" bold="true" box="[199,436,1555,1581]" pageId="141" pageNumber="142">Material examined.</emphasis>
<materialsCitation id="3790203FFF54ADB0FE069565FE8DF98E" ID-GBIF-Occurrence="3920718335" collectingDate="2005-03-21" collectingDateMax="2005-03-31" collectingDateMin="2005-03-21" collectionCode="AMSA" collectorName="Site G &amp; N. Velez" country="Australia" elevation="660" location="Mount Gower" municipality="Lord Howe Island" pageId="141" pageNumber="142" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="holotype">
<typeStatus id="584394C0FF54ADB0FE069565FD83F9D6" box="[445,554,1555,1581]" pageId="141" pageNumber="142" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="FFEF6AF2FF54ADB0FD8D9565FD70F9D6" box="[566,729,1555,1581]" name="Australia" pageId="141" pageNumber="142">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FF54ADB0FD5D9565FC6FF9D6" bold="true" box="[742,966,1555,1581]" pageId="141" pageNumber="142">
<collectingRegion id="453CE480FF54ADB0FD5D9565FC68F9D6" box="[742,961,1555,1581]" country="Australia" name="New South Wales" pageId="141" pageNumber="142">New South Wales</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6723B018FF54ADB0FC6B9565FB0DF9D6" box="[976,1188,1555,1582]" pageId="141" pageNumber="142">Lord Howe Island</collectingMunicipality>
,
<location id="82277CB9FF54ADB0FB099562FAFAF9D6" LSID="urn:lsid:plazi:treatment:0F519B74FF54ADB2FF2C93E1FB4FFEBA:82277CB9FF54ADB0FB099562FAFAF9D6" box="[1202,1363,1555,1582]" country="Australia" municipality="Lord Howe Island" name="Mount Gower" pageId="141" pageNumber="142" stateProvince="New South Wales">Mount Gower</location>
track,
<collectorName id="2A0D4FB4FF54ADB0FF2C9541FF48F9AA" box="[151,225,1591,1617]" pageId="141" pageNumber="142">Site G</collectorName>
26, 31˚34’58”S 159˚05’03”E,
<quantity id="40008787FF54ADB0FD829541FD2DF9A9" box="[569,644,1591,1618]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="141" pageNumber="142" unit="m" value="660.0">
<elevation id="0CD5CD51FF54ADB0FD829541FD2DF9A9" box="[569,644,1591,1618]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="141" pageNumber="142" unit="m" value="660.0">660 m</elevation>
</quantity>
,
<taxonomicName id="40F851E1FF54ADB0FD299541FC10F9AA" box="[658,953,1591,1617]" class="Liliopsida" family="Arecaceae" genus="Hedyscepe" kingdom="Plantae" order="Arecales" pageId="141" pageNumber="142" phylum="Tracheophyta" rank="species" species="canterburyana">
<emphasis id="B58CF670FF54ADB0FD299541FC10F9AA" box="[658,953,1591,1617]" italics="true" pageId="141" pageNumber="142">Hedyscepe canterburyana</emphasis>
</taxonomicName>
forest, leaf litter,
<date id="F3460CA2FF54ADB0FB3C9541FA87F9A9" box="[1159,1326,1591,1618]" pageId="141" pageNumber="142" value="2005-03-21" valueMax="2005-03-31" valueMin="2005-03-21">
<collectingDate id="E302F54AFF54ADB0FB3C9541FA87F9A9" box="[1159,1326,1591,1618]" pageId="141" pageNumber="142" value="2005-03-21" valueMax="2005-03-31" valueMin="2005-03-21">21–31.iii.2005</collectingDate>
</date>
,
<collectorName id="2A0D4FB4FF54ADB0FA81954EFA35F9AA" box="[1338,1436,1591,1618]" pageId="141" pageNumber="142">N. Velez</collectorName>
(
<specimenCount id="91FEE1EBFF54ADB0FF1A952DFF11F98E" box="[161,184,1627,1653]" pageId="141" pageNumber="142" type="male">♂</specimenCount>
,
<collectionCode id="E1E9B2A7FF54ADB0FF79952DFEB1F98E" box="[194,280,1627,1653]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="141" pageNumber="142">AMSA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C95F6FB0DF842" blockId="141.[151,1437,259,2014]" pageId="141" pageNumber="142">
<typeStatus id="584394C0FF54ADB0FF7C95F6FE91F961" box="[199,312,1664,1690]" pageId="141" pageNumber="142" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="FFEF6AF2FF54ADB0FEF89509FE4FF962" box="[323,486,1663,1689]" name="Australia" pageId="141" pageNumber="142">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FF54ADB0FE499509FD67F962" bold="true" box="[498,718,1663,1689]" pageId="141" pageNumber="142">
<collectingRegion id="453CE480FF54ADB0FE499509FD60F962" box="[498,713,1663,1689]" country="Australia" name="New South Wales" pageId="141" pageNumber="142">New South Wales</collectingRegion>
:
</emphasis>
Lord Howe Island, Mount Gower track, Site G27,
<geoCoordinate id="E2CC4CA5FF54ADB0FAA39509FA35F961" box="[1304,1436,1663,1690]" degrees="31" direction="south" minutes="35" orientation="latitude" pageId="141" pageNumber="142" precision="15" seconds="00" value="-31.583334">31°35’00”S</geoCoordinate>
<geoCoordinate id="E2CC4CA5FF54ADB0FF2C95D5FE87F945" box="[151,302,1699,1726]" degrees="159" direction="east" minutes="05" orientation="longitude" pageId="141" pageNumber="142" precision="15" seconds="01" value="159.0836">159°05’01”E</geoCoordinate>
,
<quantity id="40008787FF54ADB0FE8795D5FE2EF945" box="[316,391,1699,1726]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="141" pageNumber="142" unit="m" value="690.0">690 m</quantity>
,
<taxonomicName id="40F851E1FF54ADB0FE2E95D5FD14F946" box="[405,701,1699,1725]" class="Liliopsida" family="Arecaceae" genus="Hedyscepe" kingdom="Plantae" order="Arecales" pageId="141" pageNumber="142" phylum="Tracheophyta" rank="species" species="canterburyana">
<emphasis id="B58CF670FF54ADB0FE2E95D5FD14F946" box="[405,701,1699,1725]" italics="true" pageId="141" pageNumber="142">Hedyscepe canterburyana</emphasis>
</taxonomicName>
forest, litter,
<date id="F3460CA2FF54ADB0FCE195D5FBA9F945" box="[858,1024,1699,1726]" pageId="141" pageNumber="142" value="2005-03-21" valueMax="2005-03-31" valueMin="2005-03-21">21–31.iii.2005</date>
, N. Velez (
<specimenCount id="91FEE1EBFF54ADB0FB3F95D5FB18F945" box="[1156,1201,1699,1726]" pageId="141" pageNumber="142" type="male">1 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFF54ADB0FB0495D5FB45F945" box="[1215,1260,1699,1726]" pageId="141" pageNumber="142" type="female">1 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FF54ADB0FB4195D5FAF9F946" box="[1274,1360,1699,1725]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="141" pageNumber="142">AMSA</collectionCode>
); Lord Howe Island, Mount Gower track, Site G30, 31˚35’03”S 159˚04’57”E,
<quantity id="40008787FF54ADB0FC7395B1FBBAF919" box="[968,1043,1735,1762]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="141" pageNumber="142" unit="m" value="760.0">760 m</quantity>
,
<emphasis id="B58CF670FF54ADB0FB9A95B1FAE5F91A" box="[1057,1356,1735,1761]" italics="true" pageId="141" pageNumber="142">
<taxonomicName id="40F851E1FF54ADB0FB9A95B1FB0BF91A" box="[1057,1186,1735,1761]" class="Magnoliopsida" family="Winteraceae" genus="Zygogynum" kingdom="Plantae" order="Canellales" pageId="141" pageNumber="142" phylum="Tracheophyta" rank="genus">Zygogynum</taxonomicName>
/Dracophyllum
</emphasis>
forest, leaf litter,
<date id="F3460CA2FF54ADB0FEB5959DFE0DF8FE" box="[270,420,1771,1797]" pageId="141" pageNumber="142" value="2004-11-01" valueMax="2004-11-14" valueMin="2004-11-01">1–14.xi.2004</date>
, N. Velez (
<specimenCount id="91FEE1EBFF54ADB0FD9E959DFDFDF8FD" box="[549,596,1771,1798]" pageId="141" pageNumber="142" type="male">7 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFF54ADB0FDD9959DFD32F8FE" box="[610,667,1771,1797]" pageId="141" pageNumber="142" type="female">13 ♀</specimenCount>
); Lord Howe Island, Mount Gower, bottom of southernmost gully, 31˚35’17”S 159˚04’23”E,
<quantity id="40008787FF54ADB0FE729479FDBFF8D1" box="[457,534,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="141" pageNumber="142" unit="m" value="670.0">670 m</quantity>
, pitfall trap,
<date id="F3460CA2FF54ADB0FD099479FCE1F8D2" box="[690,840,1807,1833]" pageId="141" pageNumber="142" value="2002-01-18" valueMax="2002-01-31" valueMin="2002-01-18">18–31.i.2002</date>
, I. Hutton (
<specimenCount id="91FEE1EBFF54ADB0FC6E9479FBACF8D1" box="[981,1029,1807,1834]" pageId="141" pageNumber="142" type="male">3 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFF54ADB0FBAE9479FBEDF8D1" box="[1045,1092,1807,1834]" pageId="141" pageNumber="142" type="female">5 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FF54ADB0FBEF9479FB03F8D2" box="[1108,1194,1807,1833]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="141" pageNumber="142">AMSA</collectionCode>
); Lord Howe Island, Mount Lidgbird, west side of valley between Pimple and summit,
<geoCoordinate id="E2CC4CA5FF54ADB0FC2D9445FBB2F8B6" box="[918,1051,1843,1869]" degrees="31" direction="south" minutes="33" orientation="latitude" pageId="141" pageNumber="142" precision="15" seconds="18" value="-31.554998">31°33’18”S</geoCoordinate>
<geoCoordinate id="E2CC4CA5FF54ADB0FB9E9445FB12F8B5" box="[1061,1211,1843,1870]" degrees="159" direction="east" minutes="05" orientation="longitude" pageId="141" pageNumber="142" precision="15" seconds="02" value="159.08388">159°05’02”E</geoCoordinate>
,
<quantity id="40008787FF54ADB0FB729445FABCF8B5" box="[1225,1301,1843,1870]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="141" pageNumber="142" unit="m" value="650.0">650 m</quantity>
,
<taxonomicName id="40F851E1FF54ADB0FA989445FE95F88A" class="Liliopsida" family="Arecaceae" genus="Hedyscepe" kingdom="Plantae" order="Arecales" pageId="141" pageNumber="142" phylum="Tracheophyta" rank="species" species="canterburyana">
<emphasis id="B58CF670FF54ADB0FA989445FE95F88A" italics="true" pageId="141" pageNumber="142">Hedyscepe canterburyana</emphasis>
</taxonomicName>
,
<taxonomicName id="40F851E1FF54ADB0FEF2942EFDE4F88A" box="[329,589,1879,1905]" class="Magnoliopsida" family="Piperaceae" genus="Macropiper" kingdom="Plantae" order="Piperales" pageId="141" pageNumber="142" phylum="Tracheophyta" rank="species" species="hooglandii">
<emphasis id="B58CF670FF54ADB0FEF2942EFDE4F88A" box="[329,589,1879,1905]" italics="true" pageId="141" pageNumber="142">Macropiper hooglandii</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="40F851E1FF54ADB0FDD4942EFCCBF88A" box="[623,866,1879,1906]" class="Magnoliopsida" family="Rubiaceae" genus="Coprosma" kingdom="Plantae" order="Gentianales" pageId="141" pageNumber="142" phylum="Tracheophyta" rank="species" species="huttoniana">
<emphasis id="B58CF670FF54ADB0FDD4942EFCCBF88A" box="[623,866,1879,1906]" italics="true" pageId="141" pageNumber="142">Coprosma huttoniana</emphasis>
</taxonomicName>
leaf litter,
<date id="F3460CA2FF54ADB0FC619421FBF8F88A" box="[986,1105,1879,1905]" pageId="141" pageNumber="142" value="2002-04-11">11.iv.2002</date>
, I. Hutton (
<specimenCount id="91FEE1EBFF54ADB0FB749421FB53F889" box="[1231,1274,1879,1906]" pageId="141" pageNumber="142" type="male">3 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFF54ADB0FABD9421FA86F889" box="[1286,1327,1879,1906]" pageId="141" pageNumber="142" type="female">2 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FF54ADB0FA819421FA39F88A" box="[1338,1424,1879,1905]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="141" pageNumber="142">AMSA</collectionCode>
); Lord Howe Islands, Mount Gower, midway down ridge south of igloo,
<geoCoordinate id="E2CC4CA5FF54ADB0FC75940DFBFAF86D" box="[974,1107,1915,1942]" degrees="31" direction="south" minutes="35" orientation="latitude" pageId="141" pageNumber="142" precision="15" seconds="10" value="-31.586111">31°35’10”S</geoCoordinate>
<geoCoordinate id="E2CC4CA5FF54ADB0FBE7940DFB5BF86D" box="[1116,1266,1915,1942]" degrees="159" direction="east" minutes="04" orientation="longitude" pageId="141" pageNumber="142" precision="15" seconds="30" value="159.075">159°04’30”E</geoCoordinate>
,
<quantity id="40008787FF54ADB0FABB940DFAE2F86D" box="[1280,1355,1915,1942]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="141" pageNumber="142" unit="m" value="838.0">838 m</quantity>
, pitfall trap,
<date id="F3460CA2FF54ADB0FF6A94E9FECEF842" box="[209,359,1951,1977]" pageId="141" pageNumber="142" value="2002-01-18" valueMax="2002-01-31" valueMin="2002-01-18">18–31.i.2002</date>
, I. Hutton (
<specimenCount id="91FEE1EBFF54ADB0FE5C94E9FDBDF841" box="[487,532,1951,1978]" pageId="141" pageNumber="142" type="male">4 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFF54ADB0FD9B94E9FDF1F841" box="[544,600,1951,1978]" pageId="141" pageNumber="142" type="female">17 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FF54ADB0FDDF94E9FD13F842" box="[612,698,1951,1977]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="141" pageNumber="142">AMSA</collectionCode>
); same data as
<typeStatus id="584394C0FF54ADB0FCD994E9FC6DF842" box="[866,964,1951,1977]" pageId="141" pageNumber="142" type="holotype">holotype</typeStatus>
(
<specimenCount id="91FEE1EBFF54ADB0FC6894E9FBA9F841" box="[979,1024,1951,1978]" pageId="141" pageNumber="142" type="male">2 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFF54ADB0FBB794E9FB9FF841" box="[1036,1078,1951,1978]" pageId="141" pageNumber="142" type="female">5 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FF54ADB0FBF894E9FB30F842" box="[1091,1177,1951,1977]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="141" pageNumber="142">AMSA</collectionCode>
).
</paragraph>
<paragraph id="87472A62FF54ADB0FF7C94B5FAA4F826" blockId="141.[151,1437,259,2014]" box="[199,1293,1987,2014]" pageId="141" pageNumber="142">
Other material examined: 233 additional specimens from Lord Howe Island, deposited in
<collectionCode id="E1E9B2A7FF54ADB0FB0E94B5FAA3F826" box="[1205,1290,1987,2013]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="141" pageNumber="142">AMSA</collectionCode>
.
</paragraph>
</subSubSection>
<caption id="D3877AEAFF57ADB3FF2394E1FE10F80C" ID-DOI="http://doi.org/10.5281/zenodo.7138899" ID-Zenodo-Dep="7138899" httpUri="https://zenodo.org/record/7138899/files/figure.png" pageId="142" pageNumber="143" startId="142.[152,270,1943,1967]" targetBox="[156,1431,181,1913]" targetPageId="142">
<paragraph id="87472A62FF57ADB3FF2394E1FE10F80C" blockId="142.[151,1436,1943,2040]" pageId="142" pageNumber="143">
<emphasis id="B58CF670FF57ADB3FF2394E1FE2AF854" bold="true" box="[152,387,1943,1967]" pageId="142" pageNumber="143">FIGURES 44.1–44.5.</emphasis>
<taxonomicName id="40F851E1FF57ADB3FE3594EEFDEFF84B" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[398,582,1944,1968]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="tricolor" status="sp. nov.">
<emphasis id="B58CF670FF57ADB3FE3594EEFDEFF84B" box="[398,582,1944,1968]" italics="true" pageId="142" pageNumber="143">Howickia tricolor</emphasis>
</taxonomicName>
<emphasis id="B58CF670FF57ADB3FDEA94EEFD0CF84B" bold="true" box="[593,677,1944,1968]" pageId="142" pageNumber="143">
<taxonomicNameLabel id="AEBF4B0BFF57ADB3FDEA94EEFD0CF84B" box="[593,677,1944,1968]" pageId="142" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
44.1, male habitus, lateral; 44.2, male head, anterior; 44.3, male S5–8, ventral; 44.4, male terminalia, lateral; 44.5, aedeagus and associated structures, lateral. Body scale bar indicates 0.5 mm, head scale bar indicates 0.25 mm.
</paragraph>
</caption>
<subSubSection id="CFE279E9FF56ADB2FF7C93E1FC17FF2E" pageId="143" pageNumber="144" type="etymology">
<paragraph id="87472A62FF56ADB2FF7C93E1FC17FF2E" blockId="143.[151,1436,151,321]" pageId="143" pageNumber="144">
<emphasis id="B58CF670FF56ADB2FF7C93E1FEE7FF4A" bold="true" box="[199,334,151,177]" pageId="143" pageNumber="144">Etymology.</emphasis>
This name refers to the three different colours of the body tagmata: the orange-brown head, the orange thorax, and the black abdomen (Latin ‘tricolor’: three-coloured).
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FF56ADB2FF7C93A8FB4FFEBA" pageId="143" pageNumber="144" type="discussion">
<paragraph id="87472A62FF56ADB2FF7C93A8FB4FFEBA" blockId="143.[151,1436,151,321]" pageId="143" pageNumber="144">
<emphasis id="B58CF670FF56ADB2FF7C93A8FEE6FF03" bold="true" box="[199,335,222,248]" pageId="143" pageNumber="144">Comments.</emphasis>
The three species of
<taxonomicName id="40F851E1FF56ADB2FDFA93A9FD05FF02" authorityName="Richards" authorityYear="1951" box="[577,684,223,249]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FF56ADB2FDFA93A9FD05FF02" box="[577,684,223,249]" italics="true" pageId="143" pageNumber="144">Howickia</emphasis>
</taxonomicName>
found on Lord Howe Island are distinct and easily separated from one another.
<taxonomicName id="40F851E1FF56ADB2FE949275FE52FEE6" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[303,507,259,285]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis id="B58CF670FF56ADB2FE949275FE52FEE6" box="[303,507,259,285]" italics="true" pageId="143" pageNumber="144">Howickia tricolor</emphasis>
</taxonomicName>
differs from
<taxonomicName id="40F851E1FF56ADB2FD209275FD50FEE6" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[667,761,259,285]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="B58CF670FF56ADB2FD209275FD50FEE6" box="[667,761,259,285]" italics="true" pageId="143" pageNumber="144">H. nuda</emphasis>
</taxonomicName>
and
<taxonomicName id="40F851E1FF56ADB2FC839275FC49FEE6" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[824,992,259,285]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="robustaseta">
<emphasis id="B58CF670FF56ADB2FC839275FC49FEE6" box="[824,992,259,285]" italics="true" pageId="143" pageNumber="144">H. robustaseta</emphasis>
</taxonomicName>
in the characteristic tricoloured body, having ocelli, large eyes, and a bulging face; no other species of
<taxonomicName id="40F851E1FF56ADB2FCD09251FC7FFEBA" authorityName="Richards" authorityYear="1951" box="[875,982,295,321]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FF56ADB2FCD09251FC7FFEBA" box="[875,982,295,321]" italics="true" pageId="143" pageNumber="144">Howickia</emphasis>
</taxonomicName>
has such a convex face.
</paragraph>
</subSubSection>
</treatment>
</document>
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