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<document id="AA4DE83006FCAEAEB0B86967F7355B49" ID-CLB-Dataset="8112" ID-DOI="10.37520/aemnp.2020.25" ID-GBIF-Dataset="bd0522ba-e842-4416-82ac-98640acd8e45" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489312" ID-ZooBank="94935C98-664E-4D3C-87FA-F3FC6513512B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612221404353" checkinUser="felipe" docAuthor="Nabozhenko, Maxim, Keskin, Bekir &amp; Papadopoulou, Anna" docDate="2020" docId="0E36BF5DFFF9FF9995B8D21E0F97F8DF" docLanguage="en" docName="ActaEntMusNatPrag.60.2.411-417.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Hedyphanes Fischer von Waldheim 1820" docType="key" docVersion="8" lastPageNumber="413" masterDocId="F20FC725FFFBFF9B9550D5510A0AFFBF" masterDocTitle="Two new species and new records of darkling beetles of the tribe Helopini from Turkey and Cyprus (Coleoptera: Tenebrionidae)" masterLastPageNumber="417" masterPageNumber="411" pageNumber="413" updateTime="1698926180628" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="50F969B5A10D609517D1F4D596ABB774">Two new species and new records of darkling beetles of the tribe Helopini from Turkey and Cyprus (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="15420EA2E62467AA3C83596786F191E5">Nabozhenko, Maxim</mods:namePart>
<mods:affiliation id="D5547D1D15F3EBE5136450C87C3EE848">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan 367000 Russia; e-mail: nalassus @ mail. ru &amp; Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia</mods:affiliation>
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<mods:namePart id="23C6DF1A1CE29921510A94464FE5C51F">Keskin, Bekir</mods:namePart>
<mods:affiliation id="D93936BC18BD15F46D247FC83CD06510">Ege University, Bornova-Izmir 35100 Turkey; e-mail: bekir. keskin. phd @ gmail. com</mods:affiliation>
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<mods:namePart id="1F5C7A6AEAFE833FA1B91396B17F3282">Papadopoulou, Anna</mods:namePart>
<mods:affiliation id="2866DD80DF76B409EF8D3963F01C8BF5">University of Cyprus, 1678 Nicosia P. O. Box 20537 Cyprus; e-mail: papadopoulou. g. anna @ ucy. ac. cy</mods:affiliation>
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<treatment id="0E36BF5DFFF9FF9995B8D21E0F97F8DF" ID-DOI="http://doi.org/10.5281/zenodo.4549657" ID-GBIF-Taxon="177426384" ID-Zenodo-Dep="4549657" LSID="urn:lsid:plazi:treatment:0E36BF5DFFF9FF9995B8D21E0F97F8DF" httpUri="http://treatment.plazi.org/id/0E36BF5DFFF9FF9995B8D21E0F97F8DF" lastPageNumber="413" pageId="2" pageNumber="413">
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<paragraph id="86200E4BFFF9FF9995B8D21E0805F839" blockId="2.[232,678,1870,1926]" pageId="2" pageNumber="413">
<heading id="DD68B927FFF9FF9995B8D21E0805F839" centered="true" fontSize="10" level="2" pageId="2" pageNumber="413" reason="3">
<emphasis id="B4EBD259FFF9FF9995B8D21E0805F839" bold="true" pageId="2" pageNumber="413">
Key to species of the genus
<taxonomicName id="419F75C8FFF9FF99974ED21F08ACF8D9" authorityName="Fischer von Waldheim" authorityYear="1820" box="[542,678,1870,1894]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="genus">
<emphasis id="B4EBD259FFF9FF99974ED21F08ACF8D9" bold="true" box="[542,678,1870,1894]" italics="true" pageId="2" pageNumber="413">Hedyphanes</emphasis>
</taxonomicName>
from
<collectingCountry id="FE884EDBFFF9FF9994EDD23E0805F839" box="[445,527,1903,1926]" name="Turkey" pageId="2" pageNumber="413">Turkey</collectingCountry>
</emphasis>
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<subSubSection id="CE855DC0FFF9FF9995E5D2CF08FFF7AA" pageId="2" pageNumber="413" type="description">
<paragraph id="86200E4BFFF9FF9995E5D2CF08FFF7AA" blockId="2.[151,771,1950,2069]" pageId="2" pageNumber="413">
The key is modified from NΑΒοΖ*üΕΝΚο (2013).
<taxonomicName id="419F75C8FFF9FF999780D2CF0861F86A" authority="(Nabozhenko, 2008)" baseAuthorityName="Nabozhenko" baseAuthorityYear="2008" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="roznerorum">
<emphasis id="B4EBD259FFF9FF999780D2CF0B88F86A" italics="true" pageId="2" pageNumber="413">Hedyphanes roznerorum</emphasis>
(Nabozhenko, 2008)
</taxonomicName>
represents a junior synonym of
<taxonomicName id="419F75C8FFF9FF999430D28F085FF84A" authority="Allard, 1877" authorityName="Allard" authorityYear="1877" box="[352,597,2014,2037]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="lutosus">
<emphasis id="B4EBD259FFF9FF999430D28F0BC6F84A" box="[352,460,2014,2037]" italics="true" pageId="2" pageNumber="413">H. lutosus</emphasis>
Allard, 1877
</taxonomicName>
(NΑΒοΖ*üΕΝΚο 2020); therefore, we use only the senior name in the key.
</paragraph>
</subSubSection>
<subSubSection id="CE855DC0FFF9FF99966CD1990F97F8DF" pageId="2" pageNumber="413" type="key">
<keyStep id="3D6B16EEFFF9FF99966CD1990F96FA9F" pageId="2" pageNumber="413">
<paragraph id="86200E4BFFF9FF99966CD1990F96FABF" blockId="2.[828,1437,1224,1888]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD1990F96FABF" pageId="2" pageNumber="413">
1 Elytral intervals covered with small granules and short strong setae. ..............................
<taxonomicName id="419F75C8FFF9FF9991F6D1B90F96FABF" authority="Allard, 1877" authorityName="Allard" authorityYear="1877" box="[1190,1436,1256,1280]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="lutosus">
<emphasis id="B4EBD259FFF9FF9991F6D1B90F1EFABF" bold="true" box="[1190,1300,1256,1280]" italics="true" pageId="2" pageNumber="413">H. lutosus</emphasis>
Allard, 1877
</taxonomicName>
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</paragraph>
<paragraph id="86200E4BFFF9FF99966CD0580F96FA9F" blockId="2.[828,1437,1224,1888]" box="[828,1436,1288,1312]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD0580F96FA9F" box="[828,1436,1288,1312]" pageId="2" pageNumber="413"> Elytral intervals punctured. ........................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D6B16EEFFF9FF99966CD0790F96F9BF" pageId="2" pageNumber="413">
<paragraph id="86200E4BFFF9FF99966CD0790F96FA1F" blockId="2.[828,1437,1224,1888]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD0790F96FA1F" pageId="2" pageNumber="413">
2 Prothoracic hypomera, metaventrite and abdominal ventrites 13 covered with dense granules, epipleura with sparse small granules. Elytra with very small humeral angles. .....................
<taxonomicName id="419F75C8FFF9FF999127D0D90F96FA1F" authority="Seidlitz, 1896" authorityName="Seidlitz" authorityYear="1896" box="[1143,1436,1416,1440]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="cordicollis">
<emphasis id="B4EBD259FFF9FF999127D0D90F02FA1F" bold="true" box="[1143,1288,1416,1440]" italics="true" pageId="2" pageNumber="413">H. cordicollis</emphasis>
Seidlitz, 1896
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="86200E4BFFF9FF99966CD0F80F96F9BF" blockId="2.[828,1437,1224,1888]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD0F80F96F9BF" pageId="2" pageNumber="413"> Prothoracic hypomera, metaventrite and abdominal ventrites 13 punctated with simple or asperate punctures. Humeral angles absent. ..................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D6B16EEFFF9FF99966CD3590F96F91F" pageId="2" pageNumber="413">
<paragraph id="86200E4BFFF9FF99966CD3590F96F9DF" blockId="2.[828,1437,1224,1888]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD3590F96F9DF" pageId="2" pageNumber="413">
3 Prothoracic hypomera with simple round punctures. Body length
<quantity id="4167A3AEFFF9FF9996B2D3790E28F9FF" box="[994,1058,1576,1600]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="413" unit="mm" value="8.0">8 mm</quantity>
. ............................................................ ...............................
<taxonomicName id="419F75C8FFF9FF999167D3190F96F9DF" authority="Nabozhenko, 2013" authorityName="Nabozhenko" authorityYear="2013" box="[1079,1436,1608,1632]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="khachikovi">
<emphasis id="B4EBD259FFF9FF999167D3190EDAF9DF" bold="true" box="[1079,1232,1608,1632]" italics="true" pageId="2" pageNumber="413">H. khachikovi</emphasis>
<bibRefCitation id="E20E73BAFFF9FF999186D3190F96F9DF" author="NABOZHENKO M. V." box="[1238,1436,1608,1632]" pageId="2" pageNumber="413" pagination="1 - 9" refId="ref5426" refString="NABOZHENKO M. V. 2013: Taxonomic notes on the genera Hedyphanes Fischer von Waldheim, 1820 and Entomogonus Solier, 1848 (Coleoptera: Tenebrionidae) of Turkey. Journal of Insect Biodiversity 1: 1 - 9." type="journal article" year="2013">Nabozhenko, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="86200E4BFFF9FF99966CD3380F96F91F" blockId="2.[828,1437,1224,1888]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD3380F96F91F" pageId="2" pageNumber="413">
Prothoracic hypomera with aspirate punctures. Body length more than
<quantity id="4167A3AEFFF9FF999148D3D90E6CF91F" box="[1048,1126,1672,1696]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="413" unit="mm" value="10.0">10 mm</quantity>
. .............................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D6B16EEFFF9FF99966CD3F90F97F8DF" pageId="2" pageNumber="413">
<paragraph id="86200E4BFFF9FF99966CD3F90F96F8BF" blockId="2.[828,1437,1224,1888]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD3F90F96F8BF" pageId="2" pageNumber="413">
4 Elytra oval, sides of elytral base bare. Lateral margins of pronotum shortly sinuate near base. Body with bluish tint. ......................
<taxonomicName id="419F75C8FFF9FF999170D3B90F96F8BF" authority="Faldermann, 1837" authorityName="Faldermann" authorityYear="1837" box="[1056,1436,1768,1792]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="mannerheimi">
<emphasis id="B4EBD259FFF9FF999170D3B90EDFF8BF" bold="true" box="[1056,1237,1768,1792]" italics="true" pageId="2" pageNumber="413">H. mannerheimi</emphasis>
Faldermann, 1837
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="86200E4BFFF9FF99966CD2580F97F8DF" blockId="2.[828,1437,1224,1888]" pageId="2" pageNumber="413">
<keyLead id="3D6EAD7EFFF9FF99966CD2580F97F8DF" pageId="2" pageNumber="413">
Elytra subparallel, sides of elytral base weakly pubescent. Lateral margins of pronotum evenly rounded. Body black, without bluish tint. .....
<emphasis id="B4EBD259FFF9FF999186D2190F97F8DF" bold="true" box="[1238,1437,1864,1888]" pageId="2" pageNumber="413">
<taxonomicName id="419F75C8FFF9FF999186D2190F4EF8DF" authority="Nabozhenko &amp; Keskin &amp; Papadopoulou, 2020" authorityName="Nabozhenko &amp; Keskin &amp; Papadopoulou" authorityYear="2020" box="[1238,1348,1864,1888]" class="Insecta" family="Tenebrionidae" genus="Hedyphanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="kmenti" status="sp. nov.">
<emphasis id="B4EBD259FFF9FF999186D2190F4EF8DF" bold="true" box="[1238,1348,1864,1888]" italics="true" pageId="2" pageNumber="413">H. kmenti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFD86F22FFF9FF999019D2190F97F8DF" box="[1353,1437,1864,1888]" pageId="2" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
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