treatments-xml/data/05/4F/DB/054FDB70FFDFFFFAB49C1EC27F372134.xml
2024-06-21 12:28:31 +02:00

1061 lines
160 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FFDFFFFAB49C1EC27F372134" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax Mayr 1861" docType="key" docVersion="4" lastPageNumber="33" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="25" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PeerJ</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>462</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="183542572" LSID="urn:lsid:plazi:treatment:054FDB70FFDFFFFAB49C1EC27F372134" httpUri="http://treatment.plazi.org/id/054FDB70FFDFFFFAB49C1EC27F372134" lastPageId="32" lastPageNumber="33" pageId="24" pageNumber="25">
<keyLead pageId="24" pageNumber="25">
<keyStep pageId="24" pageNumber="25">
<subSubSection pageId="24" pageNumber="25" type="nomenclature">
<paragraph blockId="24.[498,1542,1247,1561]" pageId="24" pageNumber="25">
<heading level="2" pageId="24" pageNumber="25">
Key to the clades of New World
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[981,1178,1247,1277]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Temnothorax</taxonomicName>
based on the worker caste
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="description">
<paragraph blockId="24.[498,1542,1247,1561]" pageId="24" pageNumber="25">
This key is based on the clades of
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[882,1039,1336,1362]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Temnothorax</taxonomicName>
recovered in
<bibRefCitation author="Prebus M." box="[1193,1339,1336,1362]" journalOrPublisher="BMC Evolutionary Biology" pageId="24" pageNumber="25" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">Prebus, 2017</bibRefCitation>
, which was based on phylogenetic analysis of ultraconserved element data. To augment this key, please refer to the list of all described
<taxonomicName box="[806,963,1415,1441]" class="Insecta" family="Formicidae" genus="Temnothorax" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="undetermined">Temnothorax</taxonomicName>
species in the Americas and their clade memberships in Supplemental File 2.
</paragraph>
</subSubSection>
</keyStep>
</keyLead>
<subSubSection lastPageId="32" lastPageNumber="33" pageId="24" pageNumber="25" type="key">
<keyStep pageId="24" pageNumber="25">
<paragraph blockId="24.[498,1542,1247,1561]" box="[498,1542,1495,1522]" pageId="24" pageNumber="25">
<keyLead box="[498,1542,1495,1522]" pageId="24" pageNumber="25">1. Antennae with 11 segments........................................................................................... 2</keyLead>
</paragraph>
<paragraph blockId="24.[498,1542,1247,1561]" box="[498,1542,1535,1561]" pageId="24" pageNumber="25">
<keyLead box="[498,1542,1535,1561]" pageId="24" pageNumber="25">- Antennae with 12 segments ............................................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="24" pageNumber="25">
<paragraph blockId="24.[498,1542,1582,1689]" pageId="24" pageNumber="25">
<keyLead pageId="24" pageNumber="25">
2. Postpetiole broad and campaniform in dorsal view; mesosoma strongly arched and bearing long propodeal spines;
<collectingCountry box="[867,929,1623,1649]" name="Cuba" pageId="24" pageNumber="25">Cuba</collectingCountry>
.............
<taxonomicName authority="(Fontenla)" baseAuthorityName="Fontenla" box="[1042,1349,1622,1650]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cuyaguateje">T. cuyaguateje (Fontenla)</taxonomicName>
(incertae sedis)
</keyLead>
</paragraph>
<paragraph blockId="24.[498,1542,1582,1689]" box="[498,1542,1662,1689]" pageId="24" pageNumber="25">
<keyLead box="[498,1542,1662,1689]" pageId="24" pageNumber="25">- Not matching the above description; broadly distributed .............................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="24" pageNumber="25">
<paragraph blockId="24.[498,1542,1710,1856]" pageId="24" pageNumber="25">
<keyLead pageId="24" pageNumber="25">
3. Antennal scrobes present (
<figureCitation box="[845,944,1710,1736]" captionStart="Figure 18" captionStartId="26.[524,589,1059,1081]" captionTargetBox="[498,1541,234,1046]" captionTargetId="figure-229@26.[498,1295,234,1031]" captionTargetPageId="26" captionText="Figure 18 Comparison of morphology in Temnothorax worker heads in full-face view. (A) T. americanus (CASENT0904772; photo: Will Ericson, from www.antweb.org). (B) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-18" figureDoi="http://doi.org/10.5281/zenodo.5102087" httpUri="https://zenodo.org/record/5102087/files/figure.png" pageId="24" pageNumber="25">Fig. 18A</figureCitation>
); masticatory margin of mandible with four teeth; eastern North America ................
<taxonomicName authority="(Emery)" baseAuthorityName="Emery" baseAuthorityYear="1895" box="[980,1257,1750,1776]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="americanus">T. americanus (Emery)</taxonomicName>
(Palearctic clade, part)
</keyLead>
</paragraph>
<paragraph blockId="24.[498,1542,1710,1856]" pageId="24" pageNumber="25">
<keyLead pageId="24" pageNumber="25">
- Antennal scrobes absent (
<figureCitation box="[822,918,1790,1816]" captionStart="Figure 18" captionStartId="26.[524,589,1059,1081]" captionTargetBox="[498,1541,234,1046]" captionTargetId="figure-229@26.[498,1295,234,1031]" captionTargetPageId="26" captionText="Figure 18 Comparison of morphology in Temnothorax worker heads in full-face view. (A) T. americanus (CASENT0904772; photo: Will Ericson, from www.antweb.org). (B) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-18" figureDoi="http://doi.org/10.5281/zenodo.5102087" httpUri="https://zenodo.org/record/5102087/files/figure.png" pageId="24" pageNumber="25">Fig. 18B</figureCitation>
); masticatory margin of mandible with five teeth; broadly distributed............................................................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102083" ID-Zenodo-Dep="5102083" httpUri="https://zenodo.org/record/5102083/files/figure.png" pageId="25" pageNumber="26" startId="25.[116,181,968,990]" targetBox="[90,1541,234,955]" targetPageId="25">
<paragraph blockId="25.[116,1516,967,1224]" pageId="25" pageNumber="26">
<emphasis bold="true" box="[116,1071,967,990]" pageId="25" pageNumber="26">
Figure 17 Comparison of wings of
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[499,564,967,989]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="salvini">
<emphasis bold="true" box="[499,564,967,989]" italics="true" pageId="25" pageNumber="26">salvini</emphasis>
</taxonomicName>
clade reproductive organized by species group.
</emphasis>
(A)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1129,1365,968,989]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="quercicola" status="sp. nov.">
<emphasis box="[1129,1365,968,989]" italics="true" pageId="25" pageNumber="26">Temnothorax quercicola</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1378,1459,968,989]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
gyne (CASENT0869108). (B)
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[351,454,997,1019]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis box="[351,454,997,1019]" italics="true" pageId="25" pageNumber="26">T. fuscatus</emphasis>
</taxonomicName>
gyne (CASENT0615617). (C)
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[749,852,997,1019]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis box="[749,852,997,1019]" italics="true" pageId="25" pageNumber="26">T. fuscatus</emphasis>
</taxonomicName>
male (CASENT0625205). (D)
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1150,1266,997,1019]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[1150,1266,997,1019]" italics="true" pageId="25" pageNumber="26">T. pastinifer</emphasis>
</taxonomicName>
gyne (CASENT0758861). (E)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[151,282,1026,1048]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rutabulafer" status="sp. nov.">
<emphasis box="[151,282,1026,1048]" italics="true" pageId="25" pageNumber="26">T. rutabulafer</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[288,362,1027,1048]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
gyne (CASENT0758831). (F)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[648,780,1026,1048]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rutabulafer" status="sp. nov.">
<emphasis box="[648,780,1026,1048]" italics="true" pageId="25" pageNumber="26">T. rutabulafer</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[786,860,1027,1048]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
male (CASENT0756085). (G)
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" box="[1151,1268,1026,1048]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis box="[1151,1268,1026,1048]" italics="true" pageId="25" pageNumber="26">T. pergandei</emphasis>
</taxonomicName>
gyne (CASENT0758229). (H)
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" box="[160,281,1055,1077]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pergandei">
<emphasis box="[160,281,1055,1077]" italics="true" pageId="25" pageNumber="26">T. pergandei</emphasis>
</taxonomicName>
male (CASENT0758229). (I)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[580,704,1055,1077]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[580,704,1055,1077]" italics="true" pageId="25" pageNumber="26">T. albispinus</emphasis>
</taxonomicName>
male (LACMENT323246). (J)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1014,1112,1055,1077]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="laticrus" status="sp. nov.">
<emphasis box="[1014,1112,1055,1077]" italics="true" pageId="25" pageNumber="26">T. laticrus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1122,1200,1056,1077]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
gyne (MCZENT00510556). (K)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[116,206,1085,1106]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="wilsoni" status="sp. nov.">
<emphasis box="[116,206,1085,1106]" italics="true" pageId="25" pageNumber="26">T. wilsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[212,286,1085,1106]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
gyne (CASENT0758813). (L)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[570,660,1085,1106]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="wilsoni" status="sp. nov.">
<emphasis box="[570,660,1085,1106]" italics="true" pageId="25" pageNumber="26">T. wilsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[666,740,1085,1106]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
male (CASENT0758813). (M)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1033,1152,1085,1106]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="aztecoides" status="sp. nov.">
<emphasis box="[1033,1152,1085,1106]" italics="true" pageId="25" pageNumber="26">T. aztecoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1157,1231,1085,1106]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
gyne (CASENT0758789). (N)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[116,238,1114,1136]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="aztecoides" status="sp. nov.">
<emphasis box="[116,238,1114,1136]" italics="true" pageId="25" pageNumber="26">T. aztecoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[247,325,1115,1136]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
male (CASENT0758790). (O)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1931" box="[630,726,1115,1136]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="aztecus">
<emphasis box="[630,726,1115,1136]" italics="true" pageId="25" pageNumber="26">T. aztecus</emphasis>
</taxonomicName>
male (CASENT0609756). (P)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1028,1133,1114,1136]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis box="[1028,1133,1114,1136]" italics="true" pageId="25" pageNumber="26">T. longinoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1142,1220,1115,1136]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
gyne (CASENT0619354). (Q)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[116,219,1143,1165]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis box="[116,219,1143,1165]" italics="true" pageId="25" pageNumber="26">T. longinoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[227,303,1144,1165]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
male (CASENT0619355). (R)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[601,758,1143,1165]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="parvidentatus" status="sp. nov.">
<emphasis box="[601,758,1143,1165]" italics="true" pageId="25" pageNumber="26">T. parvidentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[766,842,1144,1165]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
gyne (CASENT0612665). (S)
<taxonomicName box="[1135,1225,1143,1165]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="politus">
<emphasis box="[1135,1225,1143,1165]" italics="true" pageId="25" pageNumber="26">T. politus</emphasis>
</taxonomicName>
gyne (CASENT0758021). (T)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[116,239,1172,1194]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[116,239,1172,1194]" italics="true" pageId="25" pageNumber="26">T. subditivus</emphasis>
</taxonomicName>
gyne (CASENT0758338). (U)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[543,666,1172,1194]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[543,666,1172,1194]" italics="true" pageId="25" pageNumber="26">T. subditivus</emphasis>
</taxonomicName>
male (CASENT0915978). (V)
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[970,1121,1172,1194]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="tenuisculptus">
<emphasis box="[970,1121,1172,1194]" italics="true" pageId="25" pageNumber="26">T. tenuisculptus</emphasis>
</taxonomicName>
gyne (CASENT0756078). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-17
</paragraph>
</caption>
<keyStep pageId="25" pageNumber="26">
<paragraph blockId="25.[498,1542,1279,1425]" pageId="25" pageNumber="26">
<keyLead pageId="25" pageNumber="26">
4. Propodeal spines reduced to small, rounded angles (
<figureCitation box="[1134,1211,1279,1305]" captionStart="Figure 19" captionStartId="27.[524,589,899,921]" captionTargetBox="[498,1541,234,886]" captionTargetId="figure-290@27.[498,1135,234,871]" captionTargetPageId="27" captionText="Figure 19 Temnothorax oxynodis worker propodeum and waist segments in profile view. (CASENT0005443; photo: April Nobile, from www.antweb.org). Scale bar 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-19" figureDoi="http://doi.org/10.5281/zenodo.5102089" httpUri="https://zenodo.org/record/5102089/files/figure.png" pageId="25" pageNumber="26">Fig. 19</figureCitation>
); petiole acutely cuneiform in profile; arboreal species known only from west central
<collectingRegion box="[1164,1278,1319,1345]" country="United States of America" name="California" pageId="25" pageNumber="26">California</collectingRegion>
...............
<taxonomicName authority="(Mackay)" baseAuthorityName="Mackay" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="oxynodis">T. oxynodis (Mackay)</taxonomicName>
(
<taxonomicName box="[626,703,1358,1385]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">andrei</taxonomicName>
clade, part)
</keyLead>
</paragraph>
<paragraph blockId="25.[498,1542,1279,1425]" box="[498,1542,1398,1425]" pageId="25" pageNumber="26">
<keyLead box="[498,1542,1398,1425]" pageId="25" pageNumber="26">- Propodeal spines variable; petiole in profile shaped otherwise; broadly distributed...... 5</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="25" pageNumber="26">
<paragraph blockId="25.[498,1542,1446,1831]" pageId="25" pageNumber="26">
<keyLead pageId="25" pageNumber="26">
5. Petiole compact and node evenly rounded in profile (
<figureCitation box="[1155,1332,1446,1472]" captionStart="Figure 20" captionStartId="28.[524,589,1004,1026]" captionTargetBox="[498,1541,234,991]" captionTargetId="figure-140@28.[498,1534,234,976]" captionTargetPageId="28" captionText="Figure 20 Comparison of Temnothorax worker waist segments and heads. (AE) T. obturator clade worker propodeum and waist segments in profile view. (A) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B) T. whitfordi (CASENT0005934; photo: April Nobile, from www. antweb.org). (C) T. mmp09 (CASENT0613041). (D) T. sp. nr. mmp09 (CASENT0758934) (E) T. mmp15 (LACMENT142344). (FJ) T. rugatulus and longispinosus clade worker propodeum and waist segments in profile view. (F) T. rugatulus (CASENT0005690; photo: April Nobile, from www.antweb.org). (G) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (I) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (J) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (KO) T. obturator clade worker heads in full-face view. (K) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L) T. whitfordi (CASENT0005934; photo: April Nobile, from www.antweb.org). (M) T. mmp09 (CASENT0613041). (N) T. sp. nr. mmp09 (CASENT0758934). (O) T. mmp15 (LACMENT142344). (PT) T. rugatulus and longispinosus clade worker heads in full-face view. (P) T. rugatulus (CASENT0005690; photo: April Nobile, from www. antweb.org). (Q) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (S) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (T) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20" figureDoi="http://doi.org/10.5281/zenodo.5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="25" pageNumber="26">Figs. 20A 20E</figureCitation>
) and laterodorsal surfaces of head directly behind compound eyes smooth and shining (
<figureCitation box="[1328,1502,1486,1512]" captionStart="Figure 20" captionStartId="28.[524,589,1004,1026]" captionTargetBox="[498,1541,234,991]" captionTargetId="figure-140@28.[498,1534,234,976]" captionTargetPageId="28" captionText="Figure 20 Comparison of Temnothorax worker waist segments and heads. (AE) T. obturator clade worker propodeum and waist segments in profile view. (A) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B) T. whitfordi (CASENT0005934; photo: April Nobile, from www. antweb.org). (C) T. mmp09 (CASENT0613041). (D) T. sp. nr. mmp09 (CASENT0758934) (E) T. mmp15 (LACMENT142344). (FJ) T. rugatulus and longispinosus clade worker propodeum and waist segments in profile view. (F) T. rugatulus (CASENT0005690; photo: April Nobile, from www.antweb.org). (G) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (I) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (J) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (KO) T. obturator clade worker heads in full-face view. (K) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L) T. whitfordi (CASENT0005934; photo: April Nobile, from www.antweb.org). (M) T. mmp09 (CASENT0613041). (N) T. sp. nr. mmp09 (CASENT0758934). (O) T. mmp15 (LACMENT142344). (PT) T. rugatulus and longispinosus clade worker heads in full-face view. (P) T. rugatulus (CASENT0005690; photo: April Nobile, from www. antweb.org). (Q) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (S) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (T) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20" figureDoi="http://doi.org/10.5281/zenodo.5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="25" pageNumber="26">Figs. 20K20O</figureCitation>
); postpetiole often with an anterior bulge in profile view (except
<taxonomicName box="[1246,1343,1526,1553]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="31" phylum="Arthropoda" rank="species" species="mmp">T. mmp</taxonomicName>
15 from
<collectingCountry name="Puerto Rico" pageId="25" pageNumber="26">Puerto Rico</collectingCountry>
; see
<bibRefCitation author="Prebus M." box="[605,763,1566,1592]" journalOrPublisher="BMC Evolutionary Biology" pageId="25" pageNumber="26" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">Prebus (2017)</bibRefCitation>
and antweb.org) (
<figureCitation box="[975,1142,1566,1592]" captionStart="Figure 20" captionStartId="28.[524,589,1004,1026]" captionTargetBox="[498,1541,234,991]" captionTargetId="figure-140@28.[498,1534,234,976]" captionTargetPageId="28" captionText="Figure 20 Comparison of Temnothorax worker waist segments and heads. (AE) T. obturator clade worker propodeum and waist segments in profile view. (A) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B) T. whitfordi (CASENT0005934; photo: April Nobile, from www. antweb.org). (C) T. mmp09 (CASENT0613041). (D) T. sp. nr. mmp09 (CASENT0758934) (E) T. mmp15 (LACMENT142344). (FJ) T. rugatulus and longispinosus clade worker propodeum and waist segments in profile view. (F) T. rugatulus (CASENT0005690; photo: April Nobile, from www.antweb.org). (G) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (I) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (J) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (KO) T. obturator clade worker heads in full-face view. (K) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L) T. whitfordi (CASENT0005934; photo: April Nobile, from www.antweb.org). (M) T. mmp09 (CASENT0613041). (N) T. sp. nr. mmp09 (CASENT0758934). (O) T. mmp15 (LACMENT142344). (PT) T. rugatulus and longispinosus clade worker heads in full-face view. (P) T. rugatulus (CASENT0005690; photo: April Nobile, from www. antweb.org). (Q) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (S) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (T) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20" figureDoi="http://doi.org/10.5281/zenodo.5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="25" pageNumber="26">Figs. 20A20E</figureCitation>
); arboreal species of the American Southwest through Central America and Greater Antilles ..............
<taxonomicName box="[1290,1405,1605,1632]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="obturator">obturator</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="25.[498,1542,1446,1831]" pageId="25" pageNumber="26">
<keyLead pageId="25" pageNumber="26">
- If petiole is rounded (some species in the
<taxonomicName box="[1012,1121,1645,1671]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rugatulus">rugatulus</taxonomicName>
clade,
<figureCitation box="[1204,1403,1645,1672]" captionStart="Figure 20" captionStartId="28.[524,589,1004,1026]" captionTargetBox="[498,1541,234,991]" captionTargetId="figure-140@28.[498,1534,234,976]" captionTargetPageId="28" captionText="Figure 20 Comparison of Temnothorax worker waist segments and heads. (AE) T. obturator clade worker propodeum and waist segments in profile view. (A) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B) T. whitfordi (CASENT0005934; photo: April Nobile, from www. antweb.org). (C) T. mmp09 (CASENT0613041). (D) T. sp. nr. mmp09 (CASENT0758934) (E) T. mmp15 (LACMENT142344). (FJ) T. rugatulus and longispinosus clade worker propodeum and waist segments in profile view. (F) T. rugatulus (CASENT0005690; photo: April Nobile, from www.antweb.org). (G) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (I) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (J) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (KO) T. obturator clade worker heads in full-face view. (K) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L) T. whitfordi (CASENT0005934; photo: April Nobile, from www.antweb.org). (M) T. mmp09 (CASENT0613041). (N) T. sp. nr. mmp09 (CASENT0758934). (O) T. mmp15 (LACMENT142344). (PT) T. rugatulus and longispinosus clade worker heads in full-face view. (P) T. rugatulus (CASENT0005690; photo: April Nobile, from www. antweb.org). (Q) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (S) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (T) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20" figureDoi="http://doi.org/10.5281/zenodo.5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="25" pageNumber="26">Figs. 20G &amp; 20H</figureCitation>
) then laterodorsal surfaces of head directly behind compound eyes sculptured (
<figureCitation box="[1357,1523,1685,1711]" captionStart="Figure 20" captionStartId="28.[524,589,1004,1026]" captionTargetBox="[498,1541,234,991]" captionTargetId="figure-140@28.[498,1534,234,976]" captionTargetPageId="28" captionText="Figure 20 Comparison of Temnothorax worker waist segments and heads. (AE) T. obturator clade worker propodeum and waist segments in profile view. (A) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B) T. whitfordi (CASENT0005934; photo: April Nobile, from www. antweb.org). (C) T. mmp09 (CASENT0613041). (D) T. sp. nr. mmp09 (CASENT0758934) (E) T. mmp15 (LACMENT142344). (FJ) T. rugatulus and longispinosus clade worker propodeum and waist segments in profile view. (F) T. rugatulus (CASENT0005690; photo: April Nobile, from www.antweb.org). (G) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (I) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (J) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (KO) T. obturator clade worker heads in full-face view. (K) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L) T. whitfordi (CASENT0005934; photo: April Nobile, from www.antweb.org). (M) T. mmp09 (CASENT0613041). (N) T. sp. nr. mmp09 (CASENT0758934). (O) T. mmp15 (LACMENT142344). (PT) T. rugatulus and longispinosus clade worker heads in full-face view. (P) T. rugatulus (CASENT0005690; photo: April Nobile, from www. antweb.org). (Q) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (S) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (T) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20" figureDoi="http://doi.org/10.5281/zenodo.5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="25" pageNumber="26">Figs. 20P20T</figureCitation>
) (sculpture light in
<taxonomicName authority="(Roger)" baseAuthorityName="Roger" baseAuthorityYear="1863" box="[718,954,1725,1751]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="schaumii">T. schaumii (Roger)</taxonomicName>
,
<figureCitation box="[967,1065,1725,1751]" captionStart="Figure 20" captionStartId="28.[524,589,1004,1026]" captionTargetBox="[498,1541,234,991]" captionTargetId="figure-140@28.[498,1534,234,976]" captionTargetPageId="28" captionText="Figure 20 Comparison of Temnothorax worker waist segments and heads. (AE) T. obturator clade worker propodeum and waist segments in profile view. (A) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B) T. whitfordi (CASENT0005934; photo: April Nobile, from www. antweb.org). (C) T. mmp09 (CASENT0613041). (D) T. sp. nr. mmp09 (CASENT0758934) (E) T. mmp15 (LACMENT142344). (FJ) T. rugatulus and longispinosus clade worker propodeum and waist segments in profile view. (F) T. rugatulus (CASENT0005690; photo: April Nobile, from www.antweb.org). (G) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (I) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (J) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (KO) T. obturator clade worker heads in full-face view. (K) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L) T. whitfordi (CASENT0005934; photo: April Nobile, from www.antweb.org). (M) T. mmp09 (CASENT0613041). (N) T. sp. nr. mmp09 (CASENT0758934). (O) T. mmp15 (LACMENT142344). (PT) T. rugatulus and longispinosus clade worker heads in full-face view. (P) T. rugatulus (CASENT0005690; photo: April Nobile, from www. antweb.org). (Q) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (S) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (T) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20" figureDoi="http://doi.org/10.5281/zenodo.5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="25" pageNumber="26">Fig. 20R</figureCitation>
); postpetiole often without an anterior bulge in profile view (except some specimens of
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[1071,1208,1765,1791]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="schaumii">T. schaumii</taxonomicName>
) (
<figureCitation box="[1237,1394,1765,1791]" captionStart="Figure 20" captionStartId="28.[524,589,1004,1026]" captionTargetBox="[498,1541,234,991]" captionTargetId="figure-140@28.[498,1534,234,976]" captionTargetPageId="28" captionText="Figure 20 Comparison of Temnothorax worker waist segments and heads. (AE) T. obturator clade worker propodeum and waist segments in profile view. (A) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B) T. whitfordi (CASENT0005934; photo: April Nobile, from www. antweb.org). (C) T. mmp09 (CASENT0613041). (D) T. sp. nr. mmp09 (CASENT0758934) (E) T. mmp15 (LACMENT142344). (FJ) T. rugatulus and longispinosus clade worker propodeum and waist segments in profile view. (F) T. rugatulus (CASENT0005690; photo: April Nobile, from www.antweb.org). (G) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (I) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (J) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (KO) T. obturator clade worker heads in full-face view. (K) T. emmae (CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L) T. whitfordi (CASENT0005934; photo: April Nobile, from www.antweb.org). (M) T. mmp09 (CASENT0613041). (N) T. sp. nr. mmp09 (CASENT0758934). (O) T. mmp15 (LACMENT142344). (PT) T. rugatulus and longispinosus clade worker heads in full-face view. (P) T. rugatulus (CASENT0005690; photo: April Nobile, from www. antweb.org). (Q) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R) T. schaumii (CASENT0104050; photo: April Nobile, from www.antweb.org). (S) T. curvispinous (CASENT0104040; photo: April Nobile, from www.antweb.org). (T) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20" figureDoi="http://doi.org/10.5281/zenodo.5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="25" pageNumber="26">Figs. 20F20J</figureCitation>
); broadly distributed .......................................................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<paragraph blockId="25.[498,1542,1852,1919]" lastBlockId="26.[498,1542,1223,1529]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<keyLead pageId="25" pageNumber="26">
<keyStep pageId="25" pageNumber="26">
6. Antennal scapes long, separated from the posterior margin of head by about 1x maximum width of the antennal scape or less when fully retracted: SI&gt; 80 (see
<figureCitation box="[1405,1501,1892,1919]" captionStart="Figure 21" captionStartId="29.[524,589,841,863]" captionTargetBox="[498,1541,234,828]" captionTargetId="figure-272@29.[498,1534,234,813]" captionTargetPageId="29" captionText="Figure 21 Comparison of longispinosus and rugatulus clade worker heads in full-face view. (A) Temnothorax curvispinosus (CASENT0104040; photo: April Nobile, from www.antweb.org). (B) T. ambiguus (CASENT0915985; photo: Michele Esposito, from www.antweb.org). (C) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (D) T. longispinosus (CASENT0914508; photo: Dominique Monie, from www.antweb.org). (E) T. rugatulus worker head in full face view (CASENT0005690; photo: April Nobile, from www.antweb.org). (F) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (G) T. schaumii (CASENT0104050; photo: April Nobile, from www. antweb.org). (H) T. bradleyi (CASENT0104011; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-21" figureDoi="http://doi.org/10.5281/zenodo.5102095" httpUri="https://zenodo.org/record/5102095/files/figure.png" pageId="25" pageNumber="26">Fig. 21A</figureCitation>
for
</keyStep>
</keyLead>
<keyStep pageId="26" pageNumber="27">
<keyLead pageId="26" pageNumber="27">
diagram,
<figureCitation box="[610,790,1224,1250]" captionStart="Figure 21" captionStartId="29.[524,589,841,863]" captionTargetBox="[498,1541,234,828]" captionTargetId="figure-272@29.[498,1534,234,813]" captionTargetPageId="29" captionText="Figure 21 Comparison of longispinosus and rugatulus clade worker heads in full-face view. (A) Temnothorax curvispinosus (CASENT0104040; photo: April Nobile, from www.antweb.org). (B) T. ambiguus (CASENT0915985; photo: Michele Esposito, from www.antweb.org). (C) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (D) T. longispinosus (CASENT0914508; photo: Dominique Monie, from www.antweb.org). (E) T. rugatulus worker head in full face view (CASENT0005690; photo: April Nobile, from www.antweb.org). (F) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (G) T. schaumii (CASENT0104050; photo: April Nobile, from www. antweb.org). (H) T. bradleyi (CASENT0104011; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-21" figureDoi="http://doi.org/10.5281/zenodo.5102095" httpUri="https://zenodo.org/record/5102095/files/figure.png" pageId="26" pageNumber="27">Figs. 21B 21D</figureCitation>
for examples); if unclear (some specimens of
<taxonomicName authority="(Emery))" baseAuthorityName="Emery" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ambiguus">T. ambiguus (Emery))</taxonomicName>
then head sculpture areolate-rugulose, without overlying strong costae (but may be costulate) (
<figureCitation box="[665,761,1304,1330]" captionStart="Figure 21" captionStartId="29.[524,589,841,863]" captionTargetBox="[498,1541,234,828]" captionTargetId="figure-272@29.[498,1534,234,813]" captionTargetPageId="29" captionText="Figure 21 Comparison of longispinosus and rugatulus clade worker heads in full-face view. (A) Temnothorax curvispinosus (CASENT0104040; photo: April Nobile, from www.antweb.org). (B) T. ambiguus (CASENT0915985; photo: Michele Esposito, from www.antweb.org). (C) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (D) T. longispinosus (CASENT0914508; photo: Dominique Monie, from www.antweb.org). (E) T. rugatulus worker head in full face view (CASENT0005690; photo: April Nobile, from www.antweb.org). (F) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (G) T. schaumii (CASENT0104050; photo: April Nobile, from www. antweb.org). (H) T. bradleyi (CASENT0104011; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-21" figureDoi="http://doi.org/10.5281/zenodo.5102095" httpUri="https://zenodo.org/record/5102095/files/figure.png" pageId="26" pageNumber="27">Fig. 21B</figureCitation>
); North America, east of the continental divide......
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[1383,1542,1303,1330]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="longispinosus">longispinosus</taxonomicName>
group (Palearctic clade, part)
</keyLead>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102087" ID-Zenodo-Dep="5102087" httpUri="https://zenodo.org/record/5102087/files/figure.png" pageId="26" pageNumber="27" startId="26.[524,589,1059,1081]" targetBox="[498,1541,234,1046]" targetPageId="26">
<paragraph blockId="26.[524,1515,1059,1169]" pageId="26" pageNumber="27">
<emphasis bold="true" box="[524,1386,1059,1081]" pageId="26" pageNumber="27">
Figure 18 Comparison of morphology in
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[943,1076,1059,1081]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[943,1076,1059,1081]" italics="true" pageId="26" pageNumber="27">Temnothorax</emphasis>
</taxonomicName>
worker heads in full-face view.
</emphasis>
(A)
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1895" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="americanus">
<emphasis italics="true" pageId="26" pageNumber="27">T. americanus</emphasis>
</taxonomicName>
(CASENT0904772; photo: Will Ericson, from www.antweb.org). (B)
<taxonomicName baseAuthorityName="Wesson" baseAuthorityYear="1937" box="[1387,1515,1088,1110]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="duloticus">
<emphasis box="[1387,1515,1088,1110]" italics="true" pageId="26" pageNumber="27">T. duloticus</emphasis>
</taxonomicName>
(CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-18
</paragraph>
</caption>
<paragraph blockId="26.[498,1542,1223,1529]" pageId="26" pageNumber="27">
<keyLead pageId="26" pageNumber="27">
- Antennal scapes short, separated from the posterior margin of head by&gt;1x maximum width of the antennal scape when fully retracted: SI &lt;80 (see
<figureCitation box="[1216,1310,1423,1449]" captionStart="Figure 21" captionStartId="29.[524,589,841,863]" captionTargetBox="[498,1541,234,828]" captionTargetId="figure-272@29.[498,1534,234,813]" captionTargetPageId="29" captionText="Figure 21 Comparison of longispinosus and rugatulus clade worker heads in full-face view. (A) Temnothorax curvispinosus (CASENT0104040; photo: April Nobile, from www.antweb.org). (B) T. ambiguus (CASENT0915985; photo: Michele Esposito, from www.antweb.org). (C) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (D) T. longispinosus (CASENT0914508; photo: Dominique Monie, from www.antweb.org). (E) T. rugatulus worker head in full face view (CASENT0005690; photo: April Nobile, from www.antweb.org). (F) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (G) T. schaumii (CASENT0104050; photo: April Nobile, from www. antweb.org). (H) T. bradleyi (CASENT0104011; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-21" figureDoi="http://doi.org/10.5281/zenodo.5102095" httpUri="https://zenodo.org/record/5102095/files/figure.png" pageId="26" pageNumber="27">Fig. 21E</figureCitation>
for diagram,
<figureCitation box="[498,663,1463,1489]" captionStart="Figure 21" captionStartId="29.[524,589,841,863]" captionTargetBox="[498,1541,234,828]" captionTargetId="figure-272@29.[498,1534,234,813]" captionTargetPageId="29" captionText="Figure 21 Comparison of longispinosus and rugatulus clade worker heads in full-face view. (A) Temnothorax curvispinosus (CASENT0104040; photo: April Nobile, from www.antweb.org). (B) T. ambiguus (CASENT0915985; photo: Michele Esposito, from www.antweb.org). (C) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (D) T. longispinosus (CASENT0914508; photo: Dominique Monie, from www.antweb.org). (E) T. rugatulus worker head in full face view (CASENT0005690; photo: April Nobile, from www.antweb.org). (F) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (G) T. schaumii (CASENT0104050; photo: April Nobile, from www. antweb.org). (H) T. bradleyi (CASENT0104011; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-21" figureDoi="http://doi.org/10.5281/zenodo.5102095" httpUri="https://zenodo.org/record/5102095/files/figure.png" pageId="26" pageNumber="27">Figs. 21F21H</figureCitation>
for examples); if unclear (some specimens of
<taxonomicName authority="(Emery))" baseAuthorityName="Emery" box="[1178,1424,1463,1489]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugatulus">T. rugatulus (Emery))</taxonomicName>
then head sculpture rugulose, with overlying strong costae (
<figureCitation box="[1067,1161,1503,1529]" captionStart="Figure 21" captionStartId="29.[524,589,841,863]" captionTargetBox="[498,1541,234,828]" captionTargetId="figure-272@29.[498,1534,234,813]" captionTargetPageId="29" captionText="Figure 21 Comparison of longispinosus and rugatulus clade worker heads in full-face view. (A) Temnothorax curvispinosus (CASENT0104040; photo: April Nobile, from www.antweb.org). (B) T. ambiguus (CASENT0915985; photo: Michele Esposito, from www.antweb.org). (C) T. duloticus (CASENT0103163; photo: April Nobile, from www.antweb.org). (D) T. longispinosus (CASENT0914508; photo: Dominique Monie, from www.antweb.org). (E) T. rugatulus worker head in full face view (CASENT0005690; photo: April Nobile, from www.antweb.org). (F) T. josephi (CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (G) T. schaumii (CASENT0104050; photo: April Nobile, from www. antweb.org). (H) T. bradleyi (CASENT0104011; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-21" figureDoi="http://doi.org/10.5281/zenodo.5102095" httpUri="https://zenodo.org/record/5102095/files/figure.png" pageId="26" pageNumber="27">Fig. 21E</figureCitation>
); widespread ....
<taxonomicName box="[1361,1473,1502,1529]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugatulus">rugatulus</taxonomicName>
clade
</keyLead>
</paragraph>
</keyStep>
</paragraph>
<keyStep pageId="26" pageNumber="27">
<paragraph blockId="26.[498,1542,1551,1816]" pageId="26" pageNumber="27">
<keyLead pageId="26" pageNumber="27">
7. Matching the following description: head, mesosoma, and waist segments areolate and propodeal spines reduced to small teeth or angles and petiolar peduncle short (
<figureCitation box="[508,680,1630,1656]" captionStart="Figure 22" captionStartId="30.[524,589,651,673]" captionTargetBox="[498,1541,234,638]" captionTargetId="figure-97@30.[498,1534,234,623]" captionTargetPageId="30" captionText="Figure 22 Characters distinguishing andersoni clade species from lookalike species. (A) Temnothorax andersoni, dorsal view (CASENT0635924). (B) T. andersoni, waist segments in profile (CASENT0635924). (C) T. punctatissimus (andersoni clade), dorsal view (CASENT0758783). (D) T. punctatissimus (andersoni clade), waist segments in profile (CASENT0758783). (E) T. pilicornis sp. nov. (salvini clade), dorsal view (CASENT0756186). (F) T. pilicornis sp. nov. (salvini clade), waist segments in profile (CASENT0756186). (G) T. terrigena (salvini clade), dorsal view (CASENT0104775; photo: April Nobile, from www.antweb.org) (H) T. terrigena (salvini clade), waist segments in profile (CASENT0104775; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-22" figureDoi="http://doi.org/10.5281/zenodo.5102099" httpUri="https://zenodo.org/record/5102099/files/figure.png" pageId="26" pageNumber="27">Figs. 22A22D</figureCitation>
) and postpetiole moderately broad: PWI 140160; southern
<collectingCountry box="[1387,1542,1630,1656]" name="United States of America" pageId="26" pageNumber="27">United States</collectingCountry>
and Northern Mexico ...........................................................................................
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1353,1471,1670,1697]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andersoni">andersoni</taxonomicName>
clade
</keyLead>
</paragraph>
<paragraph blockId="26.[498,1542,1551,1816]" pageId="26" pageNumber="27">
<keyLead pageId="26" pageNumber="27">
- Not matching the above description; if head, mesosoma, and waist segments areolate, then propodeal spines more developed and petiolar peduncle longer (
<figureCitation box="[1308,1478,1750,1776]" captionStart="Figure 22" captionStartId="30.[524,589,651,673]" captionTargetBox="[498,1541,234,638]" captionTargetId="figure-97@30.[498,1534,234,623]" captionTargetPageId="30" captionText="Figure 22 Characters distinguishing andersoni clade species from lookalike species. (A) Temnothorax andersoni, dorsal view (CASENT0635924). (B) T. andersoni, waist segments in profile (CASENT0635924). (C) T. punctatissimus (andersoni clade), dorsal view (CASENT0758783). (D) T. punctatissimus (andersoni clade), waist segments in profile (CASENT0758783). (E) T. pilicornis sp. nov. (salvini clade), dorsal view (CASENT0756186). (F) T. pilicornis sp. nov. (salvini clade), waist segments in profile (CASENT0756186). (G) T. terrigena (salvini clade), dorsal view (CASENT0104775; photo: April Nobile, from www.antweb.org) (H) T. terrigena (salvini clade), waist segments in profile (CASENT0104775; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-22" figureDoi="http://doi.org/10.5281/zenodo.5102099" httpUri="https://zenodo.org/record/5102099/files/figure.png" pageId="26" pageNumber="27">Figs. 22E22H</figureCitation>
) and postpetiole broader; widespread ......................................................................................................8
</keyLead>
</paragraph>
</keyStep>
<paragraph blockId="26.[498,1494,1837,1904]" lastBlockId="27.[498,1542,1035,1301]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<keyLead pageId="26" pageNumber="27">
<keyStep pageId="26" pageNumber="27">
8. Postpetiole narrow, less than 1.5 times the maximum width of the petiolar caudal cylinder (PEW) in dorsal view: PWI &lt;150 (
<figureCitation box="[1020,1188,1877,1903]" captionStart="Figure 23" captionStartId="30.[116,181,1440,1462]" captionTargetBox="[90,1541,933,1427]" captionTargetId="figure-223@30.[90,1524,933,1412]" captionTargetPageId="30" captionText="Figure 23 Comparison of Temnothorax worker waist segments in dorsal view. (A) T. obturator (CASENT0105860; photo: Dan Kjar, from www. antweb.org). (B) T. acuminatus sp. nov. (JTLC000007439). (C) T. andrei (CASENT0005683; photo: April Nobile, from www.antweb.org). (D) T. adustus (CASENT0105862; photo: Dan Kjar, from www.antweb.org). (E) T. furunculus (UTEPENTO60001; photo: Lindsie McCabe, from www. antweb.org). (F) T. gallae (CASENT0005684; photo: April Nobile, from www.antweb.org). (G) T. poeyi (CASENT0106241; photo: Michael Branstetter, from www.antweb.org). (H) T. mmp11 (CASENT0617101). (I) T. squamifer (CASENT0901793; photo: Will Ericson, from www.antweb.org). (J) T. fuscatus (CASENT0625205). (K) T. tricarinatus (CASENT0102845; photo: Jen Fogarty, from www.antweb.org). (L) T. quercicola sp. nov. (CASENT0105865; photo: Dan Kjar, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-23" figureDoi="http://doi.org/10.5281/zenodo.5102101" httpUri="https://zenodo.org/record/5102101/files/figure.png" pageId="26" pageNumber="27">Figs. 23A23F</figureCitation>
); if PWI is 150155, then
</keyStep>
</keyLead>
<keyStep pageId="27" pageNumber="28">
<keyLead pageId="27" pageNumber="28">
antennal scapes short, when fully retracted, failing to reach the posterior margin of the head by about two times the maximum width of the antennal scape: SI &lt;80 (
<figureCitation box="[1336,1505,1075,1101]" captionStart="Figure 24" captionStartId="31.[524,589,937,959]" captionTargetBox="[498,1541,234,924]" captionTargetId="figure-245@31.[498,1534,234,909]" captionTargetPageId="31" captionText="Figure 24 Synopsis of characters distinguishing workers of obturator clade species with 12 -segmented antennae from other Temnothorax species. (A) T. obturator, full face view (CASENT0104756; photo: April Nobile, from www.antweb.org). (B) T. creolus, full face view (CASENT0755643). (C) T. sp. nr. obturator, full face view (CASENT0733254). (D) T. obturator waist segments in profile (CASENT0104756; photo: April Nobile, from www.antweb.org). (E) T. creolus waist segments in profile (CASENT0755643). (F) T. sp. nr. obturator waist segments in profile (CASENT0733254). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-24" figureDoi="http://doi.org/10.5281/zenodo.5102111" httpUri="https://zenodo.org/record/5102111/files/figure.png" pageId="27" pageNumber="28">Figs. 24A24C</figureCitation>
); NOTE: PWI is not always obvious at first glance: careful measurements should be taken; PEW in this article is measured across the caudal cylinder of the petiole........................... 9
</keyLead>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102089" ID-Zenodo-Dep="5102089" httpUri="https://zenodo.org/record/5102089/files/figure.png" pageId="27" pageNumber="28" startId="27.[524,589,899,921]" targetBox="[498,1541,234,886]" targetPageId="27">
<paragraph blockId="27.[524,1515,899,980]" pageId="27" pageNumber="28">
Figure 19
<taxonomicName box="[641,878,899,921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="oxynodis">
<emphasis bold="true" box="[641,878,899,921]" italics="true" pageId="27" pageNumber="28">Temnothorax oxynodis</emphasis>
</taxonomicName>
worker propodeum and waist segments in profile view. (CASENT0005443; photo: April Nobile, from www.antweb.org). Scale bar 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-19
</paragraph>
</caption>
<paragraph blockId="27.[498,1542,1035,1301]" pageId="27" pageNumber="28">
<keyLead pageId="27" pageNumber="28">
- Postpetiole broad, greater than 1.5 times the maximum width of the petiolar caudal cylinder in dorsal view: PWI&gt; 150 (
<figureCitation box="[930,1099,1234,1261]" captionStart="Figure 23" captionStartId="30.[116,181,1440,1462]" captionTargetBox="[90,1541,933,1427]" captionTargetId="figure-223@30.[90,1524,933,1412]" captionTargetPageId="30" captionText="Figure 23 Comparison of Temnothorax worker waist segments in dorsal view. (A) T. obturator (CASENT0105860; photo: Dan Kjar, from www. antweb.org). (B) T. acuminatus sp. nov. (JTLC000007439). (C) T. andrei (CASENT0005683; photo: April Nobile, from www.antweb.org). (D) T. adustus (CASENT0105862; photo: Dan Kjar, from www.antweb.org). (E) T. furunculus (UTEPENTO60001; photo: Lindsie McCabe, from www. antweb.org). (F) T. gallae (CASENT0005684; photo: April Nobile, from www.antweb.org). (G) T. poeyi (CASENT0106241; photo: Michael Branstetter, from www.antweb.org). (H) T. mmp11 (CASENT0617101). (I) T. squamifer (CASENT0901793; photo: Will Ericson, from www.antweb.org). (J) T. fuscatus (CASENT0625205). (K) T. tricarinatus (CASENT0102845; photo: Jen Fogarty, from www.antweb.org). (L) T. quercicola sp. nov. (CASENT0105865; photo: Dan Kjar, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-23" figureDoi="http://doi.org/10.5281/zenodo.5102101" httpUri="https://zenodo.org/record/5102101/files/figure.png" pageId="27" pageNumber="28">Figs. 23G23L</figureCitation>
); if PWI is 150155, then antennal scapes longer: SI&gt; 80 ...................................................................................................... 14
</keyLead>
</paragraph>
</keyStep>
</paragraph>
<keyStep pageId="27" pageNumber="28">
<paragraph blockId="27.[498,1542,1322,1588]" pageId="27" pageNumber="28">
<keyLead pageId="27" pageNumber="28">
9. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head by about two times the maximum width of the antennal scape: SI &lt;80 (
<figureCitation box="[508,680,1402,1428]" captionStart="Figure 24" captionStartId="31.[524,589,937,959]" captionTargetBox="[498,1541,234,924]" captionTargetId="figure-245@31.[498,1534,234,909]" captionTargetPageId="31" captionText="Figure 24 Synopsis of characters distinguishing workers of obturator clade species with 12 -segmented antennae from other Temnothorax species. (A) T. obturator, full face view (CASENT0104756; photo: April Nobile, from www.antweb.org). (B) T. creolus, full face view (CASENT0755643). (C) T. sp. nr. obturator, full face view (CASENT0733254). (D) T. obturator waist segments in profile (CASENT0104756; photo: April Nobile, from www.antweb.org). (E) T. creolus waist segments in profile (CASENT0755643). (F) T. sp. nr. obturator waist segments in profile (CASENT0733254). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-24" figureDoi="http://doi.org/10.5281/zenodo.5102111" httpUri="https://zenodo.org/record/5102111/files/figure.png" pageId="27" pageNumber="28">Figs. 24A24C</figureCitation>
); petiolar node evenly rounded in profile; caudal cylinder of petiole generally elongate (
<figureCitation box="[722,889,1442,1468]" captionStart="Figure 24" captionStartId="31.[524,589,937,959]" captionTargetBox="[498,1541,234,924]" captionTargetId="figure-245@31.[498,1534,234,909]" captionTargetPageId="31" captionText="Figure 24 Synopsis of characters distinguishing workers of obturator clade species with 12 -segmented antennae from other Temnothorax species. (A) T. obturator, full face view (CASENT0104756; photo: April Nobile, from www.antweb.org). (B) T. creolus, full face view (CASENT0755643). (C) T. sp. nr. obturator, full face view (CASENT0733254). (D) T. obturator waist segments in profile (CASENT0104756; photo: April Nobile, from www.antweb.org). (E) T. creolus waist segments in profile (CASENT0755643). (F) T. sp. nr. obturator waist segments in profile (CASENT0733254). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-24" figureDoi="http://doi.org/10.5281/zenodo.5102111" httpUri="https://zenodo.org/record/5102111/files/figure.png" pageId="27" pageNumber="28">Figs. 24D24F</figureCitation>
); arboreal species from the American southwest through Central America and Greater Antilles..............................................
<taxonomicName box="[1290,1405,1481,1508]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="obturator">obturator</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="27.[498,1542,1322,1588]" pageId="27" pageNumber="28">
<keyLead pageId="27" pageNumber="28">- Petiolar node shaped otherwise; if petiolar node is rounded, then antennal scapes long: SI&gt; 80; arboreal or not; broadly distributed................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="27" pageNumber="28">
<paragraph blockId="27.[498,1542,1609,1755]" pageId="27" pageNumber="28">
<keyLead pageId="27" pageNumber="28">
10. Propodeum without erect setae dorsally, but a single pair may arise from the metanotal groove (
<figureCitation box="[596,789,1649,1676]" captionStart="Figure 25" captionStartId="32.[524,589,928,950]" captionTargetBox="[498,1541,234,915]" captionTargetId="figure-252@32.[498,1534,234,900]" captionTargetPageId="32" captionText="Figure 25 Characters separating workers of salvini clade species with narrow postpetioles from other Temnothorax species, propodeum and waist segments in profile. (A) T. acuminatus sp. nov. (JTLC000007439). (B) T. acutispinosus sp. nov. (JTLC000007447). (C) T. rudis (CASENT0005689; photo: April Nobile, from www.antweb.org). (D) T. gallae (CASENT0005684; photo: April Nobile, from www. antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-25" figureDoi="http://doi.org/10.5281/zenodo.5102113" httpUri="https://zenodo.org/record/5102113/files/figure.png" pageId="27" pageNumber="28">Figs. 25A &amp; 25B</figureCitation>
);
<collectingRegion box="[814,910,1649,1675]" country="Mexico" name="Chiapas" pageId="27" pageNumber="28">Chiapas</collectingRegion>
,
<collectingCountry box="[923,1009,1649,1675]" name="Mexico" pageId="27" pageNumber="28">Mexico</collectingCountry>
..........................................
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1326,1405,1648,1675]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="salvini">salvini</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="27.[498,1542,1609,1755]" pageId="27" pageNumber="28">
<keyLead pageId="27" pageNumber="28">
- Propodeum with at least one pair of erect setae dorsally (
<figureCitation box="[1195,1391,1689,1716]" captionStart="Figure 25" captionStartId="32.[524,589,928,950]" captionTargetBox="[498,1541,234,915]" captionTargetId="figure-252@32.[498,1534,234,900]" captionTargetPageId="32" captionText="Figure 25 Characters separating workers of salvini clade species with narrow postpetioles from other Temnothorax species, propodeum and waist segments in profile. (A) T. acuminatus sp. nov. (JTLC000007439). (B) T. acutispinosus sp. nov. (JTLC000007447). (C) T. rudis (CASENT0005689; photo: April Nobile, from www.antweb.org). (D) T. gallae (CASENT0005684; photo: April Nobile, from www. antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-25" figureDoi="http://doi.org/10.5281/zenodo.5102113" httpUri="https://zenodo.org/record/5102113/files/figure.png" pageId="27" pageNumber="28">Figs. 25C &amp; 25D</figureCitation>
); North America, west of the Great Plains ................................................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="27" pageNumber="28">
<paragraph blockId="27.[498,1541,1776,1843]" box="[498,1541,1776,1803]" pageId="27" pageNumber="28">
<keyLead box="[498,1541,1776,1803]" pageId="27" pageNumber="28">
11. Petiolar node not cuneiform (
<figureCitation box="[896,1071,1776,1803]" captionStart="Figure 26" captionStartId="33.[524,589,778,800]" captionTargetBox="[498,1541,234,766]" captionTargetId="figure-235@33.[498,1534,234,750]" captionTargetPageId="33" captionText="Figure 26 Petioles of Temnothorax workers in profile, comparing non-cuneiform and cuneiform. (A) T. rudis (CASENT0005689; photo: April Nobile, from www.antweb.org). (B) T. andrei (CASENT0005683; photo: April Nobile, from www.antweb.org). (C) T. myrmiciformis (CASENT0339323; photo Marek Borowiec, from www.antweb.org). (D) T. paiute (CASENT0005932; photo Marek Borowiec, from www.antweb.org). (E) T. gallae (CASENT0005684; photo: April Nobile, from www.antweb.org). (F) T. nitens (CASENT0005686; photo: April Nobile, from www.antweb.org). (G) T. furunculus (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (H) T. arboreus (CASENT0005933; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-26" figureDoi="http://doi.org/10.5281/zenodo.5102117" httpUri="https://zenodo.org/record/5102117/files/figure.png" pageId="27" pageNumber="28">Figs. 26A26D</figureCitation>
)................................
<taxonomicName box="[1326,1404,1776,1803]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="andrei">andrei</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="27.[498,1541,1776,1843]" box="[498,1541,1816,1843]" pageId="27" pageNumber="28">
<keyLead box="[498,1541,1816,1843]" pageId="27" pageNumber="28">
- Petiolar node cuneiform (
<figureCitation box="[822,993,1816,1842]" captionStart="Figure 26" captionStartId="33.[524,589,778,800]" captionTargetBox="[498,1541,234,766]" captionTargetId="figure-235@33.[498,1534,234,750]" captionTargetPageId="33" captionText="Figure 26 Petioles of Temnothorax workers in profile, comparing non-cuneiform and cuneiform. (A) T. rudis (CASENT0005689; photo: April Nobile, from www.antweb.org). (B) T. andrei (CASENT0005683; photo: April Nobile, from www.antweb.org). (C) T. myrmiciformis (CASENT0339323; photo Marek Borowiec, from www.antweb.org). (D) T. paiute (CASENT0005932; photo Marek Borowiec, from www.antweb.org). (E) T. gallae (CASENT0005684; photo: April Nobile, from www.antweb.org). (F) T. nitens (CASENT0005686; photo: April Nobile, from www.antweb.org). (G) T. furunculus (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (H) T. arboreus (CASENT0005933; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-26" figureDoi="http://doi.org/10.5281/zenodo.5102117" httpUri="https://zenodo.org/record/5102117/files/figure.png" pageId="27" pageNumber="28">Figs. 26E26H</figureCitation>
) .................................................................... 12
</keyLead>
</paragraph>
</keyStep>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102091" ID-Zenodo-Dep="5102091" httpUri="https://zenodo.org/record/5102091/files/figure.png" pageId="28" pageNumber="29" startId="28.[524,589,1004,1026]" targetBox="[498,1541,234,991]" targetPageId="28">
<paragraph blockId="28.[524,1516,1004,1552]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[524,1269,1004,1026]" pageId="28" pageNumber="29">
Figure 20 Comparison of
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[789,922,1004,1026]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[789,922,1004,1026]" italics="true" pageId="28" pageNumber="29">Temnothorax</emphasis>
</taxonomicName>
worker waist segments and heads.
</emphasis>
(AE)
<taxonomicName box="[1342,1459,1004,1026]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="obturator">
<emphasis box="[1342,1459,1004,1026]" italics="true" pageId="28" pageNumber="29">T. obturator</emphasis>
</taxonomicName>
clade worker propodeum and waist segments in profile view. (A)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1117,1214,1034,1055]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="emmae">
<emphasis box="[1117,1214,1034,1055]" italics="true" pageId="28" pageNumber="29">T. emmae</emphasis>
</taxonomicName>
(CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (B)
<taxonomicName box="[886,1001,1063,1084]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="whitfordi">
<emphasis box="[886,1001,1063,1084]" italics="true" pageId="28" pageNumber="29">T. whitfordi</emphasis>
</taxonomicName>
(CASENT0005934; photo: April Nobile, from www. antweb.org). (C)
<taxonomicName box="[685,760,1092,1114]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="31" phylum="Arthropoda" rank="species" species="mmp">
<emphasis box="[685,705,1093,1114]" italics="true" pageId="28" pageNumber="29">T.</emphasis>
mmp
</taxonomicName>
09 (CASENT0613041). (D)
<emphasis box="[1023,1044,1093,1114]" italics="true" pageId="28" pageNumber="29">T.</emphasis>
sp. nr. mmp09 (CASENT0758934) (E)
<taxonomicName box="[1416,1494,1092,1114]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="31" phylum="Arthropoda" rank="species" species="mmp">
<emphasis box="[1416,1437,1093,1114]" italics="true" pageId="28" pageNumber="29">T.</emphasis>
mmp
</taxonomicName>
15 (LACMENT142344). (FJ)
<taxonomicName box="[788,903,1121,1143]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rugatulus">
<emphasis box="[788,903,1121,1143]" italics="true" pageId="28" pageNumber="29">T. rugatulus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[952,1076,1121,1142]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis box="[952,1076,1121,1142]" italics="true" pageId="28" pageNumber="29">longispinosus</emphasis>
</taxonomicName>
clade worker propodeum and waist segments in profile view. (F)
<taxonomicName box="[733,855,1150,1172]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rugatulus">
<emphasis box="[733,855,1150,1172]" italics="true" pageId="28" pageNumber="29">T. rugatulus</emphasis>
</taxonomicName>
(CASENT0005690; photo: April Nobile, from www.antweb.org). (G)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[572,671,1180,1201]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="josephi">
<emphasis box="[572,671,1180,1201]" italics="true" pageId="28" pageNumber="29">T. josephi</emphasis>
</taxonomicName>
(CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (H)
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[1396,1515,1180,1201]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis box="[1396,1515,1180,1201]" italics="true" pageId="28" pageNumber="29">T. schaumii</emphasis>
</taxonomicName>
(CASENT0104050; photo: April Nobile, from www.antweb.org). (I)
<taxonomicName box="[1181,1325,1209,1231]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="curvispinous">
<emphasis box="[1181,1325,1209,1231]" italics="true" pageId="28" pageNumber="29">T. curvispinous</emphasis>
</taxonomicName>
(CASENT0104040; photo: April Nobile, from www.antweb.org). (J)
<taxonomicName baseAuthorityName="Wesson" baseAuthorityYear="1937" box="[1002,1116,1238,1260]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="duloticus">
<emphasis box="[1002,1116,1238,1260]" italics="true" pageId="28" pageNumber="29">T. duloticus</emphasis>
</taxonomicName>
(CASENT0103163; photo: April Nobile, from www.antweb.org). (KO)
<taxonomicName box="[844,966,1267,1289]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="obturator">
<emphasis box="[844,966,1267,1289]" italics="true" pageId="28" pageNumber="29">T. obturator</emphasis>
</taxonomicName>
clade worker heads in full-face view. (K)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1415,1515,1268,1289]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="emmae">
<emphasis box="[1415,1515,1268,1289]" italics="true" pageId="28" pageNumber="29">T. emmae</emphasis>
</taxonomicName>
(CASENT0102836; photo: Jen Fogarty, from www.antweb.org). (L)
<taxonomicName box="[1204,1322,1296,1318]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="whitfordi">
<emphasis box="[1204,1322,1296,1318]" italics="true" pageId="28" pageNumber="29">T. whitfordi</emphasis>
</taxonomicName>
(CASENT0005934; photo: April Nobile, from www.antweb.org). (M)
<taxonomicName box="[1002,1079,1326,1348]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="31" phylum="Arthropoda" rank="species" species="mmp">
<emphasis box="[1002,1022,1327,1348]" italics="true" pageId="28" pageNumber="29">T.</emphasis>
mmp
</taxonomicName>
09 (CASENT0613041). (N)
<emphasis box="[1348,1368,1327,1348]" italics="true" pageId="28" pageNumber="29">T.</emphasis>
sp. nr. mmp09 (CASENT0758934). (O)
<taxonomicName box="[768,850,1355,1377]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="31" phylum="Arthropoda" rank="species" species="mmp">
<emphasis box="[768,788,1356,1377]" italics="true" pageId="28" pageNumber="29">T.</emphasis>
mmp
</taxonomicName>
15 (LACMENT142344). (PT)
<taxonomicName box="[1159,1277,1355,1377]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rugatulus">
<emphasis box="[1159,1277,1355,1377]" italics="true" pageId="28" pageNumber="29">T. rugatulus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[1332,1456,1355,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis box="[1332,1456,1355,1376]" italics="true" pageId="28" pageNumber="29">longispinosus</emphasis>
</taxonomicName>
clade worker heads in full-face view. (P)
<taxonomicName box="[876,994,1384,1406]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="rugatulus">
<emphasis box="[876,994,1384,1406]" italics="true" pageId="28" pageNumber="29">T. rugatulus</emphasis>
</taxonomicName>
(CASENT0005690; photo: April Nobile, from www. antweb.org). (Q)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[707,806,1413,1435]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="josephi">
<emphasis box="[707,806,1413,1435]" italics="true" pageId="28" pageNumber="29">T. josephi</emphasis>
</taxonomicName>
(CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (R)
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[524,642,1443,1464]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis box="[524,642,1443,1464]" italics="true" pageId="28" pageNumber="29">T. schaumii</emphasis>
</taxonomicName>
(CASENT0104050; photo: April Nobile, from www.antweb.org). (S)
<taxonomicName box="[1364,1515,1443,1464]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="curvispinous">
<emphasis box="[1364,1515,1443,1464]" italics="true" pageId="28" pageNumber="29">T. curvispinous</emphasis>
</taxonomicName>
(CASENT0104040; photo: April Nobile, from www.antweb.org). (T)
<taxonomicName baseAuthorityName="Wesson" baseAuthorityYear="1937" box="[1208,1322,1472,1494]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="duloticus">
<emphasis box="[1208,1322,1472,1494]" italics="true" pageId="28" pageNumber="29">T. duloticus</emphasis>
</taxonomicName>
(CASENT0103163; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-20
</paragraph>
</caption>
<keyStep pageId="28" pageNumber="29">
<paragraph blockId="28.[498,1542,1607,1833]" pageId="28" pageNumber="29">
<keyLead pageId="28" pageNumber="29">
12. Anterior face of petiolar node straight, not concave in profile view (
<figureCitation box="[1316,1502,1607,1634]" captionStart="Figure 27" captionStartId="34.[524,589,951,973]" captionTargetBox="[498,1541,234,938]" captionTargetId="figure-45@34.[498,1534,234,923]" captionTargetPageId="34" captionText="Figure 27 Characters separating workers of sallei clade species with cuneiform petioles from other Temnothorax species. (A) T. arboreus waist segments in profile (CASENT0005933; photo: April Nobile, from www.antweb.org). (B) T. adustus waist segments in profile (CASENT0172590; photo: April Nobile, from www.antweb.org). (C) T. adustus head in full-face view (CASENT0105862; photo: Dan Kjar, from www.antweb.org). (D) T. nitens waist segments in profile (CASENT0005686; photo: April Nobile, from www.antweb.org). (E) T. furunculus waist segments in profile (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (F) T. gallae head in full-face view (CASENT0005684; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-27" figureDoi="http://doi.org/10.5281/zenodo.5102119" httpUri="https://zenodo.org/record/5102119/files/figure.png" pageId="28" pageNumber="29">Figs. 27A &amp; 27B</figureCitation>
); if unclear, posterior margin of head with ten or more erect setae and medial lobe clypeus with three distinct carinae (
<figureCitation box="[825,924,1687,1713]" captionStart="Figure 27" captionStartId="34.[524,589,951,973]" captionTargetBox="[498,1541,234,938]" captionTargetId="figure-45@34.[498,1534,234,923]" captionTargetPageId="34" captionText="Figure 27 Characters separating workers of sallei clade species with cuneiform petioles from other Temnothorax species. (A) T. arboreus waist segments in profile (CASENT0005933; photo: April Nobile, from www.antweb.org). (B) T. adustus waist segments in profile (CASENT0172590; photo: April Nobile, from www.antweb.org). (C) T. adustus head in full-face view (CASENT0105862; photo: Dan Kjar, from www.antweb.org). (D) T. nitens waist segments in profile (CASENT0005686; photo: April Nobile, from www.antweb.org). (E) T. furunculus waist segments in profile (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (F) T. gallae head in full-face view (CASENT0005684; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-27" figureDoi="http://doi.org/10.5281/zenodo.5102119" httpUri="https://zenodo.org/record/5102119/files/figure.png" pageId="28" pageNumber="29">Fig. 27C</figureCitation>
)......................................................
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1343,1404,1687,1714]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sallei">sallei</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="28.[498,1542,1607,1833]" pageId="28" pageNumber="29">
<keyLead pageId="28" pageNumber="29">
- Anterior of petiolar node concave in profile view (
<figureCitation box="[1112,1306,1727,1753]" captionStart="Figure 27" captionStartId="34.[524,589,951,973]" captionTargetBox="[498,1541,234,938]" captionTargetId="figure-45@34.[498,1534,234,923]" captionTargetPageId="34" captionText="Figure 27 Characters separating workers of sallei clade species with cuneiform petioles from other Temnothorax species. (A) T. arboreus waist segments in profile (CASENT0005933; photo: April Nobile, from www.antweb.org). (B) T. adustus waist segments in profile (CASENT0172590; photo: April Nobile, from www.antweb.org). (C) T. adustus head in full-face view (CASENT0105862; photo: Dan Kjar, from www.antweb.org). (D) T. nitens waist segments in profile (CASENT0005686; photo: April Nobile, from www.antweb.org). (E) T. furunculus waist segments in profile (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (F) T. gallae head in full-face view (CASENT0005684; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-27" figureDoi="http://doi.org/10.5281/zenodo.5102119" httpUri="https://zenodo.org/record/5102119/files/figure.png" pageId="28" pageNumber="29">Figs. 27D &amp; 27E</figureCitation>
); if unclear, posterior of head with 8 or fewer erect setae and medial lobe of clypeus without three distinct carinae (
<figureCitation box="[696,791,1807,1833]" captionStart="Figure 27" captionStartId="34.[524,589,951,973]" captionTargetBox="[498,1541,234,938]" captionTargetId="figure-45@34.[498,1534,234,923]" captionTargetPageId="34" captionText="Figure 27 Characters separating workers of sallei clade species with cuneiform petioles from other Temnothorax species. (A) T. arboreus waist segments in profile (CASENT0005933; photo: April Nobile, from www.antweb.org). (B) T. adustus waist segments in profile (CASENT0172590; photo: April Nobile, from www.antweb.org). (C) T. adustus head in full-face view (CASENT0105862; photo: Dan Kjar, from www.antweb.org). (D) T. nitens waist segments in profile (CASENT0005686; photo: April Nobile, from www.antweb.org). (E) T. furunculus waist segments in profile (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (F) T. gallae head in full-face view (CASENT0005684; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-27" figureDoi="http://doi.org/10.5281/zenodo.5102119" httpUri="https://zenodo.org/record/5102119/files/figure.png" pageId="28" pageNumber="29">Fig. 27F</figureCitation>
) ................................................................................................ 13
</keyLead>
</paragraph>
</keyStep>
<keyStep lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<paragraph blockId="28.[498,1542,1854,1921]" pageId="28" pageNumber="29">
<keyLead pageId="28" pageNumber="29">
13. Medial lobe of clypeus convex anteriorly and bearing three distinct carinae (
<figureCitation box="[1427,1526,1854,1881]" captionStart="Figure 28" captionStartId="34.[524,589,1608,1630]" captionTargetBox="[498,1541,1234,1595]" captionTargetId="figure-172@34.[498,1534,1234,1580]" captionTargetPageId="34" captionText="Figure 28 Characters separating workers of Temnothorax furunculus from workers of andrei clade workers with cuneiform petioles, heads in full-face view. (A) T. furunculus (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (B) T. gallae (CASENT0005684; photo: April Nobile, from www.antweb.org). (C) T. nitens (CASENT0005686; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-28" figureDoi="http://doi.org/10.5281/zenodo.5102125" httpUri="https://zenodo.org/record/5102125/files/figure.png" pageId="28" pageNumber="29">Fig. 28A</figureCitation>
); North America, Rocky Mountains .......
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" box="[968,1257,1894,1921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="furunculus">T. furunculus (Wheeler)</taxonomicName>
(Palearctic clade, part)
</keyLead>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102095" ID-Zenodo-Dep="5102095" httpUri="https://zenodo.org/record/5102095/files/figure.png" pageId="29" pageNumber="30" startId="29.[524,589,841,863]" targetBox="[498,1541,234,828]" targetPageId="29">
<paragraph blockId="29.[524,1516,841,1097]" pageId="29" pageNumber="30">
<emphasis bold="true" box="[524,1472,841,863]" pageId="29" pageNumber="30">
Figure 21 Comparison of
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[799,929,841,863]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis bold="true" box="[799,929,841,863]" italics="true" pageId="29" pageNumber="30">longispinosus</emphasis>
</taxonomicName>
and
<taxonomicName box="[988,1081,841,863]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="rugatulus">
<emphasis bold="true" box="[988,1081,841,863]" italics="true" pageId="29" pageNumber="30">rugatulus</emphasis>
</taxonomicName>
clade worker heads in full-face view.
</emphasis>
(A)
<taxonomicName baseAuthorityName="Mayr" baseAuthorityYear="1866" box="[524,794,870,892]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="curvispinosus">
<emphasis box="[524,794,870,892]" italics="true" pageId="29" pageNumber="30">Temnothorax curvispinosus</emphasis>
</taxonomicName>
(CASENT0104040; photo: April Nobile, from www.antweb.org). (B)
<taxonomicName box="[524,647,899,921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis box="[524,647,899,921]" italics="true" pageId="29" pageNumber="30">T. ambiguus</emphasis>
</taxonomicName>
(CASENT0915985; photo: Michele Esposito, from www.antweb.org). (C)
<taxonomicName baseAuthorityName="Wesson" baseAuthorityYear="1937" box="[1399,1515,899,921]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="duloticus">
<emphasis box="[1399,1515,899,921]" italics="true" pageId="29" pageNumber="30">T. duloticus</emphasis>
</taxonomicName>
(CASENT0103163; photo: April Nobile, from www.antweb.org). (D)
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[1178,1326,929,951]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis box="[1178,1326,929,951]" italics="true" pageId="29" pageNumber="30">T. longispinosus</emphasis>
</taxonomicName>
(CASENT0914508; photo: Dominique Monie, from www.antweb.org). (E)
<taxonomicName box="[1081,1201,958,980]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="rugatulus">
<emphasis box="[1081,1201,958,980]" italics="true" pageId="29" pageNumber="30">T. rugatulus</emphasis>
</taxonomicName>
worker head in full face view (CASENT0005690; photo: April Nobile, from www.antweb.org). (F)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1171,1261,987,1009]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="josephi">
<emphasis box="[1171,1261,987,1009]" italics="true" pageId="29" pageNumber="30">T. josephi</emphasis>
</taxonomicName>
(CASENT0102837; photo: Jen Fogarty, from www.antweb.org). (G)
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[911,1020,1016,1038]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis box="[911,1020,1016,1038]" italics="true" pageId="29" pageNumber="30">T. schaumii</emphasis>
</taxonomicName>
(CASENT0104050; photo: April Nobile, from www. antweb.org). (H)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1913" box="[690,792,1046,1067]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bradleyi">
<emphasis box="[690,792,1046,1067]" italics="true" pageId="29" pageNumber="30">T. bradleyi</emphasis>
</taxonomicName>
(CASENT0104011; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-21
</paragraph>
</caption>
<paragraph blockId="29.[498,1542,1152,1298]" pageId="29" pageNumber="30">
<keyLead pageId="29" pageNumber="30">
- Medial lobe of clypeus flat to emarginate anteriorly and not bearing three distinct carinae (several to many indistinct, but median carina not distinct along the majority of the median clypeal lobe;
<figureCitation box="[747,940,1232,1258]" captionStart="Figure 28" captionStartId="34.[524,589,1608,1630]" captionTargetBox="[498,1541,1234,1595]" captionTargetId="figure-172@34.[498,1534,1234,1580]" captionTargetPageId="34" captionText="Figure 28 Characters separating workers of Temnothorax furunculus from workers of andrei clade workers with cuneiform petioles, heads in full-face view. (A) T. furunculus (CASENT0105615; photo: Dan Kjar, from www.antweb.org). (B) T. gallae (CASENT0005684; photo: April Nobile, from www.antweb.org). (C) T. nitens (CASENT0005686; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-28" figureDoi="http://doi.org/10.5281/zenodo.5102125" httpUri="https://zenodo.org/record/5102125/files/figure.png" pageId="29" pageNumber="30">Figs. 28B &amp; 28C</figureCitation>
); North America, west of the Great Plains ............ ................................................................................................................
<taxonomicName box="[1326,1404,1271,1298]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="andrei">andrei</taxonomicName>
clade, part
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="29" pageNumber="30">
<paragraph blockId="29.[498,1542,1319,1585]" pageId="29" pageNumber="30">
<keyLead pageId="29" pageNumber="30">
14. Petiole without tubercles united by a distinct carina anterodorsally: profile of dorsal surface of petiole, when viewed from a ventrolateral angle, not interrupted by a tubercle anteriorly (
<figureCitation box="[631,731,1399,1425]" captionStart="Figure 29" captionStartId="35.[524,589,1035,1057]" captionTargetBox="[498,1541,234,1022]" captionTargetId="figure-197@35.[498,1534,234,1007]" captionTargetPageId="35" captionText="Figure 29 Characters separating workers of several clades of Temnothorax. (A) T. aztecoides sp. nov. petiole in ventrolateral view (CASENT0733319). (B) T. agavicola sp. nov. worker waist segments and gaster in dorsal view (CASENT0758360). (C) T. tricarinatus worker petiole in ventrolateral view (LACMENT323609). (D) T. texanus worker waist segments and gaster in dorsal view (CASENT0104063; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-29" figureDoi="http://doi.org/10.5281/zenodo.5102127" httpUri="https://zenodo.org/record/5102127/files/figure.png" pageId="29" pageNumber="30">Fig. 29A</figureCitation>
); or, if present, postpetiole is broadly attached to anterior margin of gaster (
<figureCitation box="[617,714,1439,1465]" captionStart="Figure 29" captionStartId="35.[524,589,1035,1057]" captionTargetBox="[498,1541,234,1022]" captionTargetId="figure-197@35.[498,1534,234,1007]" captionTargetPageId="35" captionText="Figure 29 Characters separating workers of several clades of Temnothorax. (A) T. aztecoides sp. nov. petiole in ventrolateral view (CASENT0733319). (B) T. agavicola sp. nov. worker waist segments and gaster in dorsal view (CASENT0758360). (C) T. tricarinatus worker petiole in ventrolateral view (LACMENT323609). (D) T. texanus worker waist segments and gaster in dorsal view (CASENT0104063; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-29" figureDoi="http://doi.org/10.5281/zenodo.5102127" httpUri="https://zenodo.org/record/5102127/files/figure.png" pageId="29" pageNumber="30">Fig. 29B</figureCitation>
) .......................................................................................................................... 15
</keyLead>
</paragraph>
<paragraph blockId="29.[498,1542,1319,1585]" pageId="29" pageNumber="30">
<keyLead pageId="29" pageNumber="30">
- Petiole with tubercles united by a distinct carina anterodorsally: profile of dorsal surface of petiole, when viewed from a ventrolateral angle, interrupted by a tubercle anteriorly (
<figureCitation box="[508,607,1558,1585]" captionStart="Figure 29" captionStartId="35.[524,589,1035,1057]" captionTargetBox="[498,1541,234,1022]" captionTargetId="figure-197@35.[498,1534,234,1007]" captionTargetPageId="35" captionText="Figure 29 Characters separating workers of several clades of Temnothorax. (A) T. aztecoides sp. nov. petiole in ventrolateral view (CASENT0733319). (B) T. agavicola sp. nov. worker waist segments and gaster in dorsal view (CASENT0758360). (C) T. tricarinatus worker petiole in ventrolateral view (LACMENT323609). (D) T. texanus worker waist segments and gaster in dorsal view (CASENT0104063; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-29" figureDoi="http://doi.org/10.5281/zenodo.5102127" httpUri="https://zenodo.org/record/5102127/files/figure.png" pageId="29" pageNumber="30">Fig. 29C</figureCitation>
); postpetiole not broadly attached to anterior margin of gaster (
<figureCitation box="[1366,1467,1558,1585]" captionStart="Figure 29" captionStartId="35.[524,589,1035,1057]" captionTargetBox="[498,1541,234,1022]" captionTargetId="figure-197@35.[498,1534,234,1007]" captionTargetPageId="35" captionText="Figure 29 Characters separating workers of several clades of Temnothorax. (A) T. aztecoides sp. nov. petiole in ventrolateral view (CASENT0733319). (B) T. agavicola sp. nov. worker waist segments and gaster in dorsal view (CASENT0758360). (C) T. tricarinatus worker petiole in ventrolateral view (LACMENT323609). (D) T. texanus worker waist segments and gaster in dorsal view (CASENT0104063; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-29" figureDoi="http://doi.org/10.5281/zenodo.5102127" httpUri="https://zenodo.org/record/5102127/files/figure.png" pageId="29" pageNumber="30">Fig. 29D</figureCitation>
) .... 19
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="29" pageNumber="30">
<paragraph blockId="29.[498,1541,1606,1912]" pageId="29" pageNumber="30">
<keyLead pageId="29" pageNumber="30">
15. Dorsum of mesosoma evenly convex in profile; dorsum of propodeum rounding evenly into the propodeal declivity; propodeum either entirely without teeth or spines (
<figureCitation box="[508,681,1686,1712]" captionStart="Figure 30" captionStartId="36.[524,589,1068,1090]" captionTargetBox="[498,1541,234,1055]" captionTargetId="figure-218@36.[498,1534,234,1040]" captionTargetPageId="36" captionText="Figure 30 Propodeum and waist segments workers of sallei clade species with rounded petioles in profile view. (A) Temnothorax striatulus (FOCOL2038; photo: Christiana Klingenberg, from www. antweb.org). (B) T. gundlachi (CASENT0916006; photo: Michele Esposito, from www.antweb.org). (C) T. versicolor (CASENT0914180; photo: Ziv Lieberman, from www.antweb.org). (D) T. isabellae (CASENT0915380; photo: Will Ericson, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-30" figureDoi="http://doi.org/10.5281/zenodo.5102131" httpUri="https://zenodo.org/record/5102131/files/figure.png" pageId="29" pageNumber="30">Figs. 30A30C</figureCitation>
) or with small teeth, in which case the teeth do not mark an angulate transition between the dorsum and declivity of the propodeum (
<figureCitation box="[1260,1361,1726,1752]" captionStart="Figure 30" captionStartId="36.[524,589,1068,1090]" captionTargetBox="[498,1541,234,1055]" captionTargetId="figure-218@36.[498,1534,234,1040]" captionTargetPageId="36" captionText="Figure 30 Propodeum and waist segments workers of sallei clade species with rounded petioles in profile view. (A) Temnothorax striatulus (FOCOL2038; photo: Christiana Klingenberg, from www. antweb.org). (B) T. gundlachi (CASENT0916006; photo: Michele Esposito, from www.antweb.org). (C) T. versicolor (CASENT0914180; photo: Ziv Lieberman, from www.antweb.org). (D) T. isabellae (CASENT0915380; photo: Will Ericson, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-30" figureDoi="http://doi.org/10.5281/zenodo.5102131" httpUri="https://zenodo.org/record/5102131/files/figure.png" pageId="29" pageNumber="30">Fig. 30D</figureCitation>
); Central America,
<collectingCountry box="[613,677,1766,1792]" name="Cuba" pageId="29" pageNumber="30">Cuba</collectingCountry>
, and
<collectingCountry box="[742,881,1766,1792]" name="Puerto Rico" pageId="29" pageNumber="30">Puerto Rico</collectingCountry>
.............................................................
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1343,1404,1765,1792]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sallei">sallei</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="29.[498,1541,1606,1912]" pageId="29" pageNumber="30">
<keyLead pageId="29" pageNumber="30">- Dorsum of mesosoma in profile variable: evenly convex or not; propodeum bearing teeth or spines or, if uncertain, transition from dorsum to declivity of propodeum is marked by an angle, not rounded as above; broadly distributed ................................................. 16</keyLead>
</paragraph>
</keyStep>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102099" ID-Zenodo-Dep="5102099" httpUri="https://zenodo.org/record/5102099/files/figure.png" pageId="30" pageNumber="31" startId="30.[524,589,651,673]" targetBox="[498,1541,234,638]" targetPageId="30">
<paragraph blockId="30.[524,1517,651,878]" pageId="30" pageNumber="31">
<emphasis bold="true" box="[524,1343,651,673]" pageId="30" pageNumber="31">
Figure 22 Characters distinguishing
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[889,984,651,673]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis bold="true" box="[889,984,651,673]" italics="true" pageId="30" pageNumber="31">andersoni</emphasis>
</taxonomicName>
clade species from lookalike species.
</emphasis>
(A)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis italics="true" pageId="30" pageNumber="31">Temnothorax andersoni</emphasis>
</taxonomicName>
, dorsal view (CASENT0635924). (B)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[965,1081,680,702]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis box="[965,1081,680,702]" italics="true" pageId="30" pageNumber="31">T. andersoni</emphasis>
</taxonomicName>
, waist segments in profile (CASENT0635924). (C)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[561,724,710,731]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis box="[561,724,710,731]" italics="true" pageId="30" pageNumber="31">T. punctatissimus</emphasis>
</taxonomicName>
(
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[738,829,709,730]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis box="[738,829,709,730]" italics="true" pageId="30" pageNumber="31">andersoni</emphasis>
</taxonomicName>
clade), dorsal view (CASENT0758783). (D)
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1247,1410,710,731]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis box="[1247,1410,710,731]" italics="true" pageId="30" pageNumber="31">T. punctatissimus</emphasis>
</taxonomicName>
(
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1424,1515,709,730]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis box="[1424,1515,709,730]" italics="true" pageId="30" pageNumber="31">andersoni</emphasis>
</taxonomicName>
clade), waist segments in profile (CASENT0758783). (E)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1066,1177,739,760]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pilicornis" status="sp. nov.">
<emphasis box="[1066,1177,739,760]" italics="true" pageId="30" pageNumber="31">T. pilicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1184,1259,739,760]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1273,1333,739,760]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[1273,1333,739,760]" italics="true" pageId="30" pageNumber="31">salvini</emphasis>
</taxonomicName>
clade), dorsal view (CASENT0756186). (F)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[752,862,768,790]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pilicornis" status="sp. nov.">
<emphasis box="[752,862,768,790]" italics="true" pageId="30" pageNumber="31">T. pilicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[868,942,768,789]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[955,1015,768,789]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[955,1015,768,789]" italics="true" pageId="30" pageNumber="31">salvini</emphasis>
</taxonomicName>
clade), waist segments in profile (CASENT0756186). (G)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[562,668,798,819]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[562,668,798,819]" italics="true" pageId="30" pageNumber="31">T. terrigena</emphasis>
</taxonomicName>
(
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[682,742,797,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[682,742,797,818]" italics="true" pageId="30" pageNumber="31">salvini</emphasis>
</taxonomicName>
clade), dorsal view (CASENT0104775; photo: April Nobile, from www.antweb.org) (H)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[566,676,827,848]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="terrigena">
<emphasis box="[566,676,827,848]" italics="true" pageId="30" pageNumber="31">T. terrigena</emphasis>
</taxonomicName>
(
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[693,753,826,847]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[693,753,826,847]" italics="true" pageId="30" pageNumber="31">salvini</emphasis>
</taxonomicName>
clade), waist segments in profile (CASENT0104775; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-22
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102101" ID-Zenodo-Dep="5102101" httpUri="https://zenodo.org/record/5102101/files/figure.png" pageId="30" pageNumber="31" startId="30.[116,181,1440,1462]" targetBox="[90,1541,933,1427]" targetPageId="30">
<paragraph blockId="30.[115,1516,1440,1638]" pageId="30" pageNumber="31">
<emphasis bold="true" box="[116,888,1440,1462]" pageId="30" pageNumber="31">
Figure 23 Comparison of
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[377,510,1440,1462]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[377,510,1440,1462]" italics="true" pageId="30" pageNumber="31">Temnothorax</emphasis>
</taxonomicName>
worker waist segments in dorsal view.
</emphasis>
(A)
<taxonomicName box="[933,1048,1440,1462]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="obturator">
<emphasis box="[933,1048,1440,1462]" italics="true" pageId="30" pageNumber="31">T. obturator</emphasis>
</taxonomicName>
(CASENT0105860; photo: Dan Kjar, from www. antweb.org). (B)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[283,423,1470,1491]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis box="[283,423,1470,1491]" italics="true" pageId="30" pageNumber="31">T. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[432,510,1470,1491]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(JTLC000007439). (C)
<taxonomicName box="[744,834,1469,1491]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="andrei">
<emphasis box="[744,834,1469,1491]" italics="true" pageId="30" pageNumber="31">T. andrei</emphasis>
</taxonomicName>
(CASENT0005683; photo: April Nobile, from www.antweb.org). (D)
<taxonomicName box="[116,213,1498,1520]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="adustus">
<emphasis box="[116,213,1498,1520]" italics="true" pageId="30" pageNumber="31">T. adustus</emphasis>
</taxonomicName>
(CASENT0105862; photo: Dan Kjar, from www.antweb.org). (E)
<taxonomicName box="[843,969,1498,1520]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="furunculus">
<emphasis box="[843,969,1498,1520]" italics="true" pageId="30" pageNumber="31">T. furunculus</emphasis>
</taxonomicName>
(UTEPENTO60001; photo: Lindsie McCabe, from www. antweb.org). (F)
<taxonomicName box="[278,359,1528,1549]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="gallae">
<emphasis box="[278,359,1528,1549]" italics="true" pageId="30" pageNumber="31">T. gallae</emphasis>
</taxonomicName>
(CASENT0005684; photo: April Nobile, from www.antweb.org). (G)
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1913" box="[1034,1110,1528,1549]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="poeyi">
<emphasis box="[1034,1110,1528,1549]" italics="true" pageId="30" pageNumber="31">T. poeyi</emphasis>
</taxonomicName>
(CASENT0106241; photo: Michael Branstetter, from www.antweb.org). (H)
<emphasis box="[455,551,1557,1579]" italics="true" pageId="30" pageNumber="31">
<taxonomicName box="[455,529,1557,1579]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="mmp">T. mmp</taxonomicName>
11
</emphasis>
(CASENT0617101). (I)
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[783,901,1557,1579]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="squamifer">
<emphasis box="[783,901,1557,1579]" italics="true" pageId="30" pageNumber="31">T. squamifer</emphasis>
</taxonomicName>
(CASENT0901793; photo: Will Ericson, from www.antweb.org). (J)
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[148,252,1586,1608]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis box="[148,252,1586,1608]" italics="true" pageId="30" pageNumber="31">T. fuscatus</emphasis>
</taxonomicName>
(CASENT0625205). (K)
<taxonomicName box="[500,638,1587,1608]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis box="[500,638,1587,1608]" italics="true" pageId="30" pageNumber="31">T. tricarinatus</emphasis>
</taxonomicName>
(CASENT0102845; photo: Jen Fogarty, from www.antweb.org). (L)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1308,1431,1586,1608]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="quercicola" status="sp. nov.">
<emphasis box="[1308,1431,1586,1608]" italics="true" pageId="30" pageNumber="31">T. quercicola</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1439,1516,1587,1608]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(CASENT0105865; photo: Dan Kjar, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-23
</paragraph>
</caption>
<keyStep pageId="30" pageNumber="31">
<paragraph blockId="30.[498,1542,1692,1918]" pageId="30" pageNumber="31">
<keyLead pageId="30" pageNumber="31">
16. Matching the following description: propodeal teeth reduced, or transition from dorsum to declivity of propodeum is marked by an angle (
<figureCitation box="[1192,1385,1732,1759]" captionStart="Figure 31" captionStartId="37.[524,589,1082,1104]" captionTargetBox="[498,1541,234,1069]" captionTargetId="figure-209@37.[498,1534,234,1054]" captionTargetPageId="37" captionText="Figure 31 Temnothorax sallei clade workers with reduced propodeal teeth. Propodeum and waist segments in profile view. (A) T. mmp11 (CASENT0617101). (B) T. sp. nr. mmp49 (CASENT0733208); legs in dorsal view. (C) T. mmp11 (CASENT0617101). (D) T. sp. nr. mmp49 (CASENT0733208). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-31" figureDoi="http://doi.org/10.5281/zenodo.5102133" httpUri="https://zenodo.org/record/5102133/files/figure.png" pageId="30" pageNumber="31">Figs. 31A &amp; 31B</figureCitation>
); legs moderately incrassate (FI &lt;300), and without abundant erect to suberect setae (
<figureCitation captionStart="Figure 31" captionStartId="37.[524,589,1082,1104]" captionTargetBox="[498,1541,234,1069]" captionTargetId="figure-209@37.[498,1534,234,1054]" captionTargetPageId="37" captionText="Figure 31 Temnothorax sallei clade workers with reduced propodeal teeth. Propodeum and waist segments in profile view. (A) T. mmp11 (CASENT0617101). (B) T. sp. nr. mmp49 (CASENT0733208); legs in dorsal view. (C) T. mmp11 (CASENT0617101). (D) T. sp. nr. mmp49 (CASENT0733208). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-31" figureDoi="http://doi.org/10.5281/zenodo.5102133" httpUri="https://zenodo.org/record/5102133/files/figure.png" pageId="30" pageNumber="31">
Figs. 31C &amp;
<keyLead pageId="30" pageNumber="31">
31D); petiolar node evenly rounded in profile (
<figureCitation box="[1043,1233,1812,1838]" captionStart="Figure 31" captionStartId="37.[524,589,1082,1104]" captionTargetBox="[498,1541,234,1069]" captionTargetId="figure-209@37.[498,1534,234,1054]" captionTargetPageId="37" captionText="Figure 31 Temnothorax sallei clade workers with reduced propodeal teeth. Propodeum and waist segments in profile view. (A) T. mmp11 (CASENT0617101). (B) T. sp. nr. mmp49 (CASENT0733208); legs in dorsal view. (C) T. mmp11 (CASENT0617101). (D) T. sp. nr. mmp49 (CASENT0733208). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-31" figureDoi="http://doi.org/10.5281/zenodo.5102133" httpUri="https://zenodo.org/record/5102133/files/figure.png" pageId="30" pageNumber="31">Figs. 31A &amp; 31B</figureCitation>
); mid to high elevations, from central
<collectingCountry box="[650,735,1852,1878]" name="Mexico" pageId="30" pageNumber="31">Mexico</collectingCountry>
to the Nicaraguan depression ...................................
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1348,1408,1851,1878]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="sallei">sallei</taxonomicName>
clade, part
</keyLead>
</figureCitation>
</keyLead>
</paragraph>
<paragraph blockId="30.[498,1542,1692,1918]" box="[498,1541,1892,1918]" pageId="30" pageNumber="31">
<keyLead box="[498,1541,1892,1918]" pageId="30" pageNumber="31">- Not matching the above description; broadly distributed ............................................ 17</keyLead>
</paragraph>
</keyStep>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102111" ID-Zenodo-Dep="5102111" httpUri="https://zenodo.org/record/5102111/files/figure.png" pageId="31" pageNumber="32" startId="31.[524,589,937,959]" targetBox="[498,1541,234,924]" targetPageId="31">
<paragraph blockId="31.[524,1515,937,1135]" pageId="31" pageNumber="32">
<emphasis bold="true" pageId="31" pageNumber="32">
Figure 24 Synopsis of characters distinguishing workers of
<taxonomicName box="[1176,1270,937,959]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obturator">
<emphasis bold="true" box="[1176,1270,937,959]" italics="true" pageId="31" pageNumber="32">obturator</emphasis>
</taxonomicName>
clade species with 12 -segmented antennae from other
<taxonomicName box="[910,1136,966,988]" class="Insecta" family="Formicidae" genus="Temnothorax" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis bold="true" box="[910,1043,966,988]" italics="true" pageId="31" pageNumber="32">Temnothorax</emphasis>
species.
</taxonomicName>
</emphasis>
(A)
<taxonomicName box="[1209,1338,966,988]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obturator">
<emphasis box="[1209,1338,966,988]" italics="true" pageId="31" pageNumber="32">T. obturator</emphasis>
</taxonomicName>
, full face view (CASENT0104756; photo: April Nobile, from www.antweb.org). (B)
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1249,1350,995,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="creolus">
<emphasis box="[1249,1350,995,1017]" italics="true" pageId="31" pageNumber="32">T. creolus</emphasis>
</taxonomicName>
, full face view (CASENT0755643). (C)
<emphasis box="[763,783,1026,1047]" italics="true" pageId="31" pageNumber="32">T.</emphasis>
sp. nr.
<taxonomicName box="[860,950,1025,1046]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obturator">
<emphasis box="[860,950,1025,1046]" italics="true" pageId="31" pageNumber="32">obturator</emphasis>
</taxonomicName>
, full face view (CASENT0733254). (D)
<taxonomicName box="[1340,1458,1025,1047]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obturator">
<emphasis box="[1340,1458,1025,1047]" italics="true" pageId="31" pageNumber="32">T. obturator</emphasis>
</taxonomicName>
waist segments in profile (CASENT0104756; photo: April Nobile, from www.antweb.org). (E)
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1369,1460,1054,1076]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="creolus">
<emphasis box="[1369,1460,1054,1076]" italics="true" pageId="31" pageNumber="32">T. creolus</emphasis>
</taxonomicName>
waist segments in profile (CASENT0755643). (F)
<emphasis box="[1005,1025,1084,1105]" italics="true" pageId="31" pageNumber="32">T.</emphasis>
sp. nr.
<taxonomicName box="[1133,1223,1083,1104]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obturator">
<emphasis box="[1133,1223,1083,1104]" italics="true" pageId="31" pageNumber="32">obturator</emphasis>
</taxonomicName>
waist segments in profile (CASENT0733254). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-24
</paragraph>
</caption>
<keyStep pageId="31" pageNumber="32">
<paragraph blockId="31.[498,1542,1189,1534]" pageId="31" pageNumber="32">
<keyLead pageId="31" pageNumber="32">
17. Matching the following description: clypeus forming wall between antennal insertions and mandibles (
<figureCitation box="[687,859,1229,1256]" captionStart="Figure 32" captionStartId="38.[116,181,724,746]" captionTargetBox="[90,1541,234,711]" captionTargetId="figure-67@38.[90,1524,234,696]" captionTargetPageId="38" captionText="Figure 32 Temnothorax sallei clade and salvini clade workers. Heads in full-face view. (A) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (B) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (C) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (D) T. quercicola sp. nov. (LACMENT323216). (E) T. obtusigaster sp. nov. (CASENT0758654). (F) T. balnearius sp. nov. (LACMENT323199); legs in dorsal view. (G) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (H) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (I) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (J) T. quercicola sp. nov. (LACMENT323216). (K) T. obtusigaster sp. nov. (CASENT0758654). (L) T. balnearius sp. nov. (LACMENT323199). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-32" figureDoi="http://doi.org/10.5281/zenodo.5102135" httpUri="https://zenodo.org/record/5102135/files/figure.png" pageId="31" pageNumber="32">Figs. 32A32C</figureCitation>
); anterior margin of clypeus flat; posterior margin of head rounding evenly into lateral margins; legs not incrassate (
<figureCitation box="[1177,1340,1269,1295]" captionStart="Figure 32" captionStartId="38.[116,181,724,746]" captionTargetBox="[90,1541,234,711]" captionTargetId="figure-67@38.[90,1524,234,696]" captionTargetPageId="38" captionText="Figure 32 Temnothorax sallei clade and salvini clade workers. Heads in full-face view. (A) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (B) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (C) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (D) T. quercicola sp. nov. (LACMENT323216). (E) T. obtusigaster sp. nov. (CASENT0758654). (F) T. balnearius sp. nov. (LACMENT323199); legs in dorsal view. (G) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (H) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (I) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (J) T. quercicola sp. nov. (LACMENT323216). (K) T. obtusigaster sp. nov. (CASENT0758654). (L) T. balnearius sp. nov. (LACMENT323199). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-32" figureDoi="http://doi.org/10.5281/zenodo.5102135" httpUri="https://zenodo.org/record/5102135/files/figure.png" pageId="31" pageNumber="32">Figs. 32G32I</figureCitation>
); southwestern
<collectingCountry box="[498,655,1309,1335]" name="United States of America" pageId="31" pageNumber="32">United States</collectingCountry>
and northwestern
<collectingCountry box="[880,966,1309,1335]" name="Mexico" pageId="31" pageNumber="32">Mexico</collectingCountry>
........................................................
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1343,1404,1309,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sallei">sallei</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="31.[498,1542,1189,1534]" pageId="31" pageNumber="32">
<keyLead pageId="31" pageNumber="32">
- Not matching the above description. If specimen is from southwestern
<collectingCountry box="[1337,1491,1349,1375]" name="United States of America" pageId="31" pageNumber="32">United States</collectingCountry>
and northwestern
<collectingCountry box="[662,748,1389,1415]" name="Mexico" pageId="31" pageNumber="32">Mexico</collectingCountry>
and the clypeal character is unclear, then the anterior margin of clypeus emarginated, posterior margin of head transitions to lateral margins through a rounded angle (
<figureCitation box="[686,856,1468,1495]" captionStart="Figure 32" captionStartId="38.[116,181,724,746]" captionTargetBox="[90,1541,234,711]" captionTargetId="figure-67@38.[90,1524,234,696]" captionTargetPageId="38" captionText="Figure 32 Temnothorax sallei clade and salvini clade workers. Heads in full-face view. (A) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (B) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (C) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (D) T. quercicola sp. nov. (LACMENT323216). (E) T. obtusigaster sp. nov. (CASENT0758654). (F) T. balnearius sp. nov. (LACMENT323199); legs in dorsal view. (G) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (H) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (I) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (J) T. quercicola sp. nov. (LACMENT323216). (K) T. obtusigaster sp. nov. (CASENT0758654). (L) T. balnearius sp. nov. (LACMENT323199). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-32" figureDoi="http://doi.org/10.5281/zenodo.5102135" httpUri="https://zenodo.org/record/5102135/files/figure.png" pageId="31" pageNumber="32">Figs. 32D32F</figureCitation>
), and legs strongly incrassate (
<figureCitation box="[1222,1379,1468,1495]" captionStart="Figure 32" captionStartId="38.[116,181,724,746]" captionTargetBox="[90,1541,234,711]" captionTargetId="figure-67@38.[90,1524,234,696]" captionTargetPageId="38" captionText="Figure 32 Temnothorax sallei clade and salvini clade workers. Heads in full-face view. (A) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (B) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (C) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (D) T. quercicola sp. nov. (LACMENT323216). (E) T. obtusigaster sp. nov. (CASENT0758654). (F) T. balnearius sp. nov. (LACMENT323199); legs in dorsal view. (G) T. silvestrii (CASENT0913000; photo: Ziv Lieberman, from www.antweb.org). (H) T. morongo (CASENT0103108; photo: April Nobile, from www.antweb.org). (I) T. bca07 (CASENT0922107; photo: Michele Esposito, from www.antweb.org). (J) T. quercicola sp. nov. (LACMENT323216). (K) T. obtusigaster sp. nov. (CASENT0758654). (L) T. balnearius sp. nov. (LACMENT323199). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-32" figureDoi="http://doi.org/10.5281/zenodo.5102135" httpUri="https://zenodo.org/record/5102135/files/figure.png" pageId="31" pageNumber="32">Figs. 32J32L</figureCitation>
); broadly distributed.......................................................................................................................................... 18
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="31" pageNumber="32">
<paragraph blockId="31.[498,1541,1556,1862]" pageId="31" pageNumber="32">
<keyLead pageId="31" pageNumber="32">
18. Postpetiole campaniform in dorsal view (
<figureCitation box="[1031,1130,1556,1582]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Fig. 33A</figureCitation>
); legs with abundant suberect to erect setae (
<figureCitation box="[638,735,1596,1622]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Fig. 33B</figureCitation>
); if petiole squamiform and increasing in width toward the apex (
<figureCitation box="[508,608,1636,1662]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Fig. 33A</figureCitation>
), medial lobe of clypeus with 3 or fewer distinct carinae (
<figureCitation box="[1292,1488,1636,1662]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Figs. 33C &amp; 33D</figureCitation>
); southern Florida,
<collectingCountry box="[709,816,1676,1702]" name="Bahamas" pageId="31" pageNumber="32">Bahamas</collectingCountry>
,
<collectingCountry box="[829,893,1676,1702]" name="Cuba" pageId="31" pageNumber="32">Cuba</collectingCountry>
, Hispaniola ........................................
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1343,1404,1675,1702]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sallei">sallei</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="31.[498,1541,1556,1862]" pageId="31" pageNumber="32">
<keyLead pageId="31" pageNumber="32">
- Postpetiole variable; legs usually without abundant suberect to erect setae; if present (
<figureCitation box="[508,603,1755,1782]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Fig. 33F</figureCitation>
) and petiolar node squamiform and increasing in width toward the apex (
<figureCitation box="[508,603,1795,1822]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Fig. 33E</figureCitation>
) then medial lobe of clypeus with many fine carinae (
<figureCitation box="[1231,1329,1795,1822]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Fig. 33G</figureCitation>
) or propodeum is strongly depressed (
<figureCitation box="[734,836,1835,1861]" captionStart="Figure 33" captionStartId="38.[524,589,1461,1483]" captionTargetBox="[498,1541,1012,1448]" captionTargetId="figure-202@38.[498,1534,1012,1432]" captionTargetPageId="38" captionText="Figure 33 Characters separating some members of the sallei and salvini clades. (AC) Temnothorax squamifer worker (CASENT0901793; photos: Will Ericson, from www.antweb.org). (A) waist segments in dorsal view. (B) legs in dorsal view. (C) clypeus and mandibles in full-face view. (D) T. creightoni worker, clypeus and mandibles in full-face view (CASENT0901794; photo: Will Ericson, from www. antweb.org) (EG) T. aztecoides sp. nov. worker (CASENT0733319). (E) waist segments in posterodorsal view. (F) legs in dorsal view. (G) clypeus and mandibles in full-face view. (H) T. bison sp. nov. worker, propodeum and petiole in profile view (CASENT0636947). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-33" figureDoi="http://doi.org/10.5281/zenodo.5102137" httpUri="https://zenodo.org/record/5102137/files/figure.png" pageId="31" pageNumber="32">Fig. 33H</figureCitation>
); broadly distributed ...............................
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1326,1405,1835,1862]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="salvini">salvini</taxonomicName>
clade, part
</keyLead>
</paragraph>
</keyStep>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102113" ID-Zenodo-Dep="5102113" httpUri="https://zenodo.org/record/5102113/files/figure.png" pageId="32" pageNumber="33" startId="32.[524,589,928,950]" targetBox="[498,1541,234,915]" targetPageId="32">
<paragraph blockId="32.[524,1516,928,1068]" pageId="32" pageNumber="33">
<emphasis bold="true" pageId="32" pageNumber="33">
Figure 25 Characters separating workers of
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[959,1024,928,950]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="salvini">
<emphasis bold="true" box="[959,1024,928,950]" italics="true" pageId="32" pageNumber="33">salvini</emphasis>
</taxonomicName>
clade species with narrow postpetioles from other
<taxonomicName box="[524,740,957,979]" class="Insecta" family="Formicidae" genus="Temnothorax" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis bold="true" box="[524,657,957,979]" italics="true" pageId="32" pageNumber="33">Temnothorax</emphasis>
species
</taxonomicName>
, propodeum and waist segments in profile.
</emphasis>
(A)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1276,1420,958,979]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="acuminatus" status="sp. nov.">
<emphasis box="[1276,1420,958,979]" italics="true" pageId="32" pageNumber="33">T. acuminatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1434,1516,958,979]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
(JTLC000007439). (B)
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[739,889,988,1009]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[739,889,988,1009]" italics="true" pageId="32" pageNumber="33">T. acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[894,967,987,1008]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
(JTLC000007447). (C)
<taxonomicName box="[1189,1261,987,1009]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rudis">
<emphasis box="[1189,1261,987,1009]" italics="true" pageId="32" pageNumber="33">T. rudis</emphasis>
</taxonomicName>
(CASENT0005689; photo: April Nobile, from www.antweb.org). (D)
<taxonomicName box="[930,1010,1016,1038]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gallae">
<emphasis box="[930,1010,1016,1038]" italics="true" pageId="32" pageNumber="33">T. gallae</emphasis>
</taxonomicName>
(CASENT0005684; photo: April Nobile, from www. antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-25
</paragraph>
</caption>
<keyStep pageId="32" pageNumber="33">
<paragraph blockId="32.[498,1542,1115,1261]" pageId="32" pageNumber="33">
<keyLead pageId="32" pageNumber="33">
19. Postpetiole with a distinctive trapezoidal shape in dorsal view: anteriorly broad, with angulate anterolateral corners (
<figureCitation box="[866,966,1155,1181]" captionStart="Figure 34" captionStartId="39.[524,589,1059,1081]" captionTargetBox="[498,1541,234,1046]" captionTargetId="figure-10@39.[498,1295,234,1031]" captionTargetPageId="39" captionText="Figure 34 Characters separating Temnothorax quasimodo from sallei and salvini clade species, worker waist segments in dorsal view. (A) T. quasimodo (CASENT0005694; photo: Marek Borowiec, from www.antweb.org). (B) T. neomexicanus (sallei clade) (CASENT0010695; photo: April Nobile, from www.antweb.org). (C) T. obtusigaster sp. nov. (salvini clade) (CASENT0758654). (D) T. morongo (sallei clade) (CASENT0103108; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-34" figureDoi="http://doi.org/10.5281/zenodo.5102139" httpUri="https://zenodo.org/record/5102139/files/figure.png" pageId="32" pageNumber="33">Fig. 34A</figureCitation>
); southwestern North America ....
<taxonomicName authority="Snelling et al." authorityName="Snelling" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="quasimodo">T. quasimodo Snelling et al.</taxonomicName>
(
<taxonomicName box="[684,762,1194,1221]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="andrei">andrei</taxonomicName>
clade, part)
</keyLead>
</paragraph>
<paragraph blockId="32.[498,1542,1115,1261]" box="[498,1541,1234,1261]" pageId="32" pageNumber="33">
<keyLead box="[498,1541,1234,1261]" pageId="32" pageNumber="33">
- Postpetiole in dorsal view not as above (
<figureCitation box="[994,1166,1234,1261]" captionStart="Figure 34" captionStartId="39.[524,589,1059,1081]" captionTargetBox="[498,1541,234,1046]" captionTargetId="figure-10@39.[498,1295,234,1031]" captionTargetPageId="39" captionText="Figure 34 Characters separating Temnothorax quasimodo from sallei and salvini clade species, worker waist segments in dorsal view. (A) T. quasimodo (CASENT0005694; photo: Marek Borowiec, from www.antweb.org). (B) T. neomexicanus (sallei clade) (CASENT0010695; photo: April Nobile, from www.antweb.org). (C) T. obtusigaster sp. nov. (salvini clade) (CASENT0758654). (D) T. morongo (sallei clade) (CASENT0103108; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-34" figureDoi="http://doi.org/10.5281/zenodo.5102139" httpUri="https://zenodo.org/record/5102139/files/figure.png" pageId="32" pageNumber="33">Figs. 34B34D</figureCitation>
); broadly distributed ............ 20
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="32" pageNumber="33">
<paragraph blockId="32.[498,1542,1282,1468]" pageId="32" pageNumber="33">
<keyLead pageId="32" pageNumber="33">
20. Medial lobe of clypeus smooth medially or bearing three distinct carinae, although these may be accompanied by additional weaker, incomplete carinae (
<figureCitation box="[1308,1476,1322,1349]" captionStart="Figure 35" captionStartId="39.[524,589,1719,1741]" captionTargetBox="[498,1541,1282,1706]" captionTargetId="figure-94@39.[498,1534,1282,1691]" captionTargetPageId="39" captionText="Figure 35 Characters separating the workers of some species of the sallei and salvini clades, clypeus and mandibles in full-face view. (A) T. Temnothorax texanus (sallei clade) (CASENT0104062; photo: April Nobile, from www.antweb.org). (B) T. brevispinosus (sallei clade) (CASENT0911187; photo: Ziv Lieberman, from www.antweb.org). (C) T. stenotyle (sallei clade) (CASENT0922576; photo: Wade Lee, from www.antweb.org). (D) T. acuminatus sp. nov. (salvini clade) (JTLC000007468). (E) T. morongo (sallei clade) (CASENT0103108; photo: April Nobile, from www.antweb.org). (F) T. quercicola sp. nov. (salvini clade) (LACMENT323216). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-35" figureDoi="http://doi.org/10.5281/zenodo.5102141" httpUri="https://zenodo.org/record/5102141/files/figure.png" pageId="32" pageNumber="33">Figs. 35A35C</figureCitation>
) ..... .......................................................................................................................
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1348,1408,1361,1388]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sallei">sallei</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="32.[498,1542,1282,1468]" pageId="32" pageNumber="33">
<keyLead pageId="32" pageNumber="33">
- Medial lobe of clypeus bearing many fine carinae (
<figureCitation box="[1116,1285,1402,1429]" captionStart="Figure 35" captionStartId="39.[524,589,1719,1741]" captionTargetBox="[498,1541,1282,1706]" captionTargetId="figure-94@39.[498,1534,1282,1691]" captionTargetPageId="39" captionText="Figure 35 Characters separating the workers of some species of the sallei and salvini clades, clypeus and mandibles in full-face view. (A) T. Temnothorax texanus (sallei clade) (CASENT0104062; photo: April Nobile, from www.antweb.org). (B) T. brevispinosus (sallei clade) (CASENT0911187; photo: Ziv Lieberman, from www.antweb.org). (C) T. stenotyle (sallei clade) (CASENT0922576; photo: Wade Lee, from www.antweb.org). (D) T. acuminatus sp. nov. (salvini clade) (JTLC000007468). (E) T. morongo (sallei clade) (CASENT0103108; photo: April Nobile, from www.antweb.org). (F) T. quercicola sp. nov. (salvini clade) (LACMENT323216). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-35" figureDoi="http://doi.org/10.5281/zenodo.5102141" httpUri="https://zenodo.org/record/5102141/files/figure.png" pageId="32" pageNumber="33">Figs. 35D35F</figureCitation>
), although these may be in addition to the three strong carinae (
<figureCitation box="[1062,1156,1441,1468]" captionStart="Figure 35" captionStartId="39.[524,589,1719,1741]" captionTargetBox="[498,1541,1282,1706]" captionTargetId="figure-94@39.[498,1534,1282,1691]" captionTargetPageId="39" captionText="Figure 35 Characters separating the workers of some species of the sallei and salvini clades, clypeus and mandibles in full-face view. (A) T. Temnothorax texanus (sallei clade) (CASENT0104062; photo: April Nobile, from www.antweb.org). (B) T. brevispinosus (sallei clade) (CASENT0911187; photo: Ziv Lieberman, from www.antweb.org). (C) T. stenotyle (sallei clade) (CASENT0922576; photo: Wade Lee, from www.antweb.org). (D) T. acuminatus sp. nov. (salvini clade) (JTLC000007468). (E) T. morongo (sallei clade) (CASENT0103108; photo: April Nobile, from www.antweb.org). (F) T. quercicola sp. nov. (salvini clade) (LACMENT323216). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-35" figureDoi="http://doi.org/10.5281/zenodo.5102141" httpUri="https://zenodo.org/record/5102141/files/figure.png" pageId="32" pageNumber="33">Fig. 35F</figureCitation>
)..................................................... 21
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="32" pageNumber="33">
<paragraph blockId="32.[498,1541,1489,1636]" pageId="32" pageNumber="33">
<keyLead pageId="32" pageNumber="33">
21. Metafemora incrassate: FI&gt; 325 (
<figureCitation box="[927,1024,1489,1515]" captionStart="Figure 36" captionStartId="40.[524,589,1135,1157]" captionTargetBox="[498,1541,234,1122]" captionTargetId="figure-179@40.[498,1295,234,1107]" captionTargetPageId="40" captionText="Figure 36 Characters separating the workers of some species of the sallei and salvini clades, legs in dorsal view. (A) Temnothorax quercicola sp. nov. (salvini clade) worker legs in dorsal view (CASENT0105865; photo: Dan Kjar, from www.antweb.org). (B) T. hispidus (sallei clade) worker legs in dorsal view (CASENT0172594; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-36" figureDoi="http://doi.org/10.5281/zenodo.5102143" httpUri="https://zenodo.org/record/5102143/files/figure.png" pageId="32" pageNumber="33">Fig. 36A</figureCitation>
); or, if not incrassate, erect setae absent from the propodeal dorsum (
<figureCitation box="[779,972,1529,1556]" captionStart="Figure 25" captionStartId="32.[524,589,928,950]" captionTargetBox="[498,1541,234,915]" captionTargetId="figure-252@32.[498,1534,234,900]" captionTargetPageId="32" captionText="Figure 25 Characters separating workers of salvini clade species with narrow postpetioles from other Temnothorax species, propodeum and waist segments in profile. (A) T. acuminatus sp. nov. (JTLC000007439). (B) T. acutispinosus sp. nov. (JTLC000007447). (C) T. rudis (CASENT0005689; photo: April Nobile, from www.antweb.org). (D) T. gallae (CASENT0005684; photo: April Nobile, from www. antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-25" figureDoi="http://doi.org/10.5281/zenodo.5102113" httpUri="https://zenodo.org/record/5102113/files/figure.png" pageId="32" pageNumber="33">Figs. 25A &amp; 25B</figureCitation>
) .............................................
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1326,1405,1529,1556]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="salvini">salvini</taxonomicName>
clade, part
</keyLead>
</paragraph>
<paragraph blockId="32.[498,1541,1489,1636]" pageId="32" pageNumber="33">
<keyLead pageId="32" pageNumber="33">
- Metafemora not incrassate FI &lt;325 (
<figureCitation box="[957,1053,1569,1595]" captionStart="Figure 36" captionStartId="40.[524,589,1135,1157]" captionTargetBox="[498,1541,234,1122]" captionTargetId="figure-179@40.[498,1295,234,1107]" captionTargetPageId="40" captionText="Figure 36 Characters separating the workers of some species of the sallei and salvini clades, legs in dorsal view. (A) Temnothorax quercicola sp. nov. (salvini clade) worker legs in dorsal view (CASENT0105865; photo: Dan Kjar, from www.antweb.org). (B) T. hispidus (sallei clade) worker legs in dorsal view (CASENT0172594; photo: April Nobile, from www.antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-36" figureDoi="http://doi.org/10.5281/zenodo.5102143" httpUri="https://zenodo.org/record/5102143/files/figure.png" pageId="32" pageNumber="33">Fig. 36B</figureCitation>
); erect setae always present on the propodeal dorsum (
<figureCitation box="[734,930,1609,1636]" captionStart="Figure 25" captionStartId="32.[524,589,928,950]" captionTargetBox="[498,1541,234,915]" captionTargetId="figure-252@32.[498,1534,234,900]" captionTargetPageId="32" captionText="Figure 25 Characters separating workers of salvini clade species with narrow postpetioles from other Temnothorax species, propodeum and waist segments in profile. (A) T. acuminatus sp. nov. (JTLC000007439). (B) T. acutispinosus sp. nov. (JTLC000007447). (C) T. rudis (CASENT0005689; photo: April Nobile, from www.antweb.org). (D) T. gallae (CASENT0005684; photo: April Nobile, from www. antweb.org). Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-25" figureDoi="http://doi.org/10.5281/zenodo.5102113" httpUri="https://zenodo.org/record/5102113/files/figure.png" pageId="32" pageNumber="33">Figs. 25C &amp; 25D</figureCitation>
) .....................................................
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1343,1404,1608,1635]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sallei">sallei</taxonomicName>
clade, part
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>