treatments-xml/data/05/4F/DB/054FDB70FECFFECBB49C1FF97D91228F.xml

<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FECFFECBB49C1FF97D91228F" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax albispinus" docType="treatment" docVersion="4" lastPageNumber="273" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="265" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
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<mods:title>e 11514</mods:title>
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<mods:number>2021-06-30</mods:number>
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<mods:number>9</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102574" ID-GBIF-Taxon="183542656" ID-Zenodo-Dep="5102574" LSID="urn:lsid:plazi:treatment:054FDB70FECFFECBB49C1FF97D91228F" httpUri="http://treatment.plazi.org/id/054FDB70FECFFECBB49C1FF97D91228F" lastPageId="273" lastPageNumber="273" pageId="264" pageNumber="265">
<subSubSection box="[498,973,1508,1534]" pageId="264" pageNumber="265" type="nomenclature">
<paragraph blockId="264.[498,1383,1508,1574]" box="[498,973,1508,1534]" pageId="264" pageNumber="265">
<heading box="[498,973,1508,1534]" level="2" pageId="264" pageNumber="265">
<taxonomicName authority="(Wheeler, 1908)" baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[498,973,1508,1534]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[498,780,1508,1534]" italics="true" pageId="264" pageNumber="265">Temnothorax albispinus</emphasis>
(
<bibRefCitation author="Wheeler WM" box="[798,964,1508,1534]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="264" pageNumber="265" pagination="117 - 158" part="24" refId="ref273343" refString="Wheeler WM. 1908. The ants of Porto Rico and the Virgin Islands. Bulletin of the American Museum of Natural History 24: 117 - 158." title="The ants of Porto Rico and the Virgin Islands" type="journal article" year="1908">
<emphasis box="[798,964,1508,1534]" italics="true" pageId="264" pageNumber="265">Wheeler, 1908</emphasis>
</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,1383,1547,1574]" pageId="264" pageNumber="265" type="distribution">
<paragraph blockId="264.[498,1383,1508,1574]" box="[498,1383,1547,1574]" pageId="264" pageNumber="265">
Distribution: 
<figureCitation box="[657,767,1548,1574]" captionStart="Figure 133" captionStartId="258.[116,181,1036,1058]" captionTargetBox="[90,1541,234,1023]" captionTargetId="figure-184@258.[90,1524,234,1008]" captionTargetPageId="258" captionText="Figure 133 Geographical and elevational distribution of the pulchellus group. (A) Temnothorax agavicola sp. nov. (B) T. albispinus. (C) T. bahoruco sp. nov. (D) T. balaclava sp. nov. (E) T. ciferrii (F) T. flavidulus (G) T. harlequina sp. nov. (H) T. hippolytus sp. nov. (I) T. laticrus sp. nov. (J) T. magnabulla sp. nov. (K) T. pulchellus (L) T. terricola (M) T. torrei (N) T. wettereri sp. nov. (O) T. wilsoni sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-133" figureDoi="http://doi.org/10.5281/zenodo.5102387" httpUri="https://zenodo.org/record/5102387/files/figure.png" pageId="264" pageNumber="265">Fig. 133B</figureCitation>
; worker, gyne &amp; male: 
<figureCitation box="[1043,1137,1547,1574]" captionStart="Figure 135" captionStartId="264.[524,589,1001,1023]" captionTargetBox="[498,1541,234,988]" captionTargetId="figure-200@264.[498,1534,234,973]" captionTargetPageId="264" captionText="Figure 135 Temnothorax albispinus. (A–C) Worker (CASENT0756097). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Gyne (MCZENT00505431). (D) Full-face view. (E) Profile view. (F) Dorsal view. (G–J) Male (CASENT0756098). (G) Full-face view. (H) Profile view. (I) Forewing. (J) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-135" figureDoi="http://doi.org/10.5281/zenodo.5102396" httpUri="https://zenodo.org/record/5102396/files/figure.png" pageId="264" pageNumber="265">Fig. 135</figureCitation>
; variability: 
<figureCitation box="[1284,1377,1548,1574]" captionStart="Figure 136" captionStartId="265.[524,589,1001,1023]" captionTargetBox="[498,1541,234,988]" captionTargetId="figure-196@265.[498,1534,234,973]" captionTargetPageId="265" captionText="Figure 136 Light form of Temnothorax albispinus. (A–C) Worker (LACMENT323245). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Gyne (LACMENT323245). (D) Full-face view. (E) Profile view. (F) Dorsal view.(G–J) Male (LACMENT323246). (G) Full-face view.(H) Profile view. (I) Wings. (J) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-136" figureDoi="http://doi.org/10.5281/zenodo.5102398" httpUri="https://zenodo.org/record/5102398/files/figure.png" pageId="264" pageNumber="265">Fig. 136</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="265" lastPageNumber="266" pageId="264" pageNumber="265" type="reference_group">
<paragraph blockId="264.[498,1542,1610,1876]" pageId="264" pageNumber="265">
<treatmentCitationGroup pageId="264" pageNumber="265">
<treatmentCitation author="Wheeler WM" box="[498,1194,1610,1637]" page="139" pageId="264" pageNumber="265" year="1908">
<taxonomicName authority="Wheeler, 1908: 139" authorityName="Wheeler" authorityPageNumber="139" authorityYear="1908" box="[498,941,1610,1636]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="species" species="albispina">
<emphasis box="[498,941,1610,1636]" italics="true" pageId="264" pageNumber="265">
Macromischa albispina 
<bibRefCitation author="Wheeler WM" box="[776,1000,1610,1637]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="264" pageNumber="265" pagination="117 - 158" part="24" refId="ref273343" refString="Wheeler WM. 1908. The ants of Porto Rico and the Virgin Islands. Bulletin of the American Museum of Natural History 24: 117 - 158." title="The ants of Porto Rico and the Virgin Islands" type="journal article" year="1908">Wheeler, 1908: 139</bibRefCitation>
</emphasis>
</taxonomicName>
, pl. 11, figs. 3, 4
</treatmentCitation>
. 
<typeStatus box="[1207,1299,1610,1636]" pageId="264" pageNumber="265">Syntype</typeStatus>
workers and gyne. 
<collectingCountry box="[498,574,1650,1676]" name="United States of America" pageId="264" pageNumber="265">U.S.A.</collectingCountry>
, Puerto Rico. 
<location LSID="urn:lsid:plazi:treatment:054FDB70FECFFECBB49C1FF97D91228F:88393CBDFECFFED2B7881C6F7A2A21DB" box="[742,792,1650,1676]" country="United States of America" name="One" pageId="264" pageNumber="265">One</location>
<typeStatus box="[800,889,1651,1677]" pageId="264" pageNumber="265">syntype</typeStatus>
worker here designated 
<emphasis bold="true" box="[1181,1290,1650,1676]" pageId="264" pageNumber="265">
<typeStatus box="[1181,1290,1650,1676]" pageId="264" pageNumber="265">lectotype</typeStatus>
</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="264.[498,1542,1610,1876]" pageId="264" pageNumber="265">
<treatmentCitationGroup pageId="264" pageNumber="265">
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" box="[498,1109,1690,1717]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="species" species="albispina" subGenus="Antillaemyrmex">
<emphasis box="[498,980,1690,1716]" italics="true" pageId="264" pageNumber="265">Macromischa (Antillaemyrmex) albispina</emphasis>
(Wheeler)
</taxonomicName>
: 
<treatmentCitation author="Mann WM" box="[1121,1321,1690,1716]" page="408" pageId="264" pageNumber="265" year="1920">
<bibRefCitation author="Mann WM" box="[1121,1321,1690,1716]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="264" pageNumber="265" pagination="403 - 439" part="42" refId="ref271797" refString="Mann WM. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403 - 439." title="Additions to the ant fauna of the West Indies and Central America" type="journal article" year="1920">
<emphasis box="[1121,1262,1690,1716]" italics="true" pageId="264" pageNumber="265">
<author box="[1121,1194,1691,1716]" fns="*" inm="true" npo="*" pageId="264" pageNumber="265">Mann</author>
, 
<year box="[1207,1262,1690,1716]" pageId="264" pageNumber="265">1920</year>
</emphasis>
: 408
</bibRefCitation>
</treatmentCitation>
. First combination in 
<taxonomicName authority="" box="[530,899,1730,1756]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="subGenus" subGenus="Antillaemyrmex">
<emphasis box="[530,899,1730,1756]" italics="true" pageId="264" pageNumber="265">Macromischa (Antillaemyrmex)</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<treatmentCitationGroup pageId="264" pageNumber="265">
<paragraph blockId="264.[498,1542,1610,1876]" box="[498,1532,1769,1796]" pageId="264" pageNumber="265">
<treatmentCitation author="Mann WM" box="[498,1337,1769,1796]" page="424" pageId="264" pageNumber="265" year="1920">
<taxonomicName authority="Mann, 1920: 424" authorityName="Mann" authorityPageNumber="424" authorityYear="1920" box="[498,1277,1769,1796]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="variety" species="albispina" subGenus="Antillaemyrmex" variety="pallipes">
<emphasis box="[498,981,1769,1795]" italics="true" pageId="264" pageNumber="265">Macromischa (Antillaemyrmex) albispina</emphasis>
var. 
<emphasis box="[1041,1277,1769,1796]" italics="true" pageId="264" pageNumber="265">
pallipes 
<bibRefCitation author="Mann WM" box="[1136,1337,1770,1796]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="264" pageNumber="265" pagination="403 - 439" part="42" refId="ref271797" refString="Mann WM. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403 - 439." title="Additions to the ant fauna of the West Indies and Central America" type="journal article" year="1920">Mann, 1920: 424</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
. 
<emphasis box="[1349,1526,1769,1795]" italics="true" pageId="264" pageNumber="265">Nomen nudum</emphasis>
.
</paragraph>
<paragraph blockId="264.[498,1542,1610,1876]" pageId="264" pageNumber="265">
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[498,927,1809,1836]" class="Insecta" family="Formicidae" genus="Antillaemyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="species" species="albispina">
<emphasis box="[498,798,1809,1835]" italics="true" pageId="264" pageNumber="265">Antillaemyrmex albispina</emphasis>
(Wheeler)
</taxonomicName>
: 
<treatmentCitation author="Wheeler WM" box="[939,1150,1809,1836]" page="27" pageId="264" pageNumber="265" year="1931">
<bibRefCitation author="Wheeler WM" box="[939,1150,1809,1836]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="264" pageNumber="265" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
<emphasis box="[939,1105,1809,1835]" italics="true" pageId="264" pageNumber="265">
<author box="[939,1036,1809,1835]" fns="*" inm="true" npo="*" pageId="264" pageNumber="265">Wheeler</author>
, 
<year box="[1050,1105,1809,1835]" pageId="264" pageNumber="265">1931</year>
</emphasis>
: 27
</bibRefCitation>
</treatmentCitation>
. First combination in 
<taxonomicName authorityName="Mann" authorityYear="1920" box="[498,683,1849,1875]" class="Insecta" family="Formicidae" genus="Antillaemyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="264" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis box="[498,683,1849,1875]" italics="true" pageId="264" pageNumber="265">Antillaemyrmex</emphasis>
</taxonomicName>
.
</paragraph>
</treatmentCitationGroup>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102398" ID-Zenodo-Dep="5102398" httpUri="https://zenodo.org/record/5102398/files/figure.png" pageId="265" pageNumber="266" startId="265.[524,589,1001,1023]" targetBox="[498,1541,234,988]" targetPageId="265">
<paragraph blockId="265.[524,1515,1001,1111]" pageId="265" pageNumber="266">
<emphasis bold="true" box="[524,1029,1001,1023]" pageId="265" pageNumber="266">
Figure 136 Light form of 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[785,1023,1001,1023]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis bold="true" box="[785,1023,1001,1023]" italics="true" pageId="265" pageNumber="266">Temnothorax albispinus</emphasis>
</taxonomicName>
.
</emphasis>
(A–C) Worker (LACMENT323245). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Gyne (LACMENT323245). (D) Full-face view. (E) Profile view. (F) Dorsal view. (G–J) Male (LACMENT323246). (G) Full-face view. (H) Profile view. (I) Wings. (J) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-136
</paragraph>
</caption>
<paragraph blockId="265.[498,1542,1166,1910]" pageId="265" pageNumber="266">
<treatmentCitationGroup pageId="265" pageNumber="266">
<taxonomicName authority="(Mann)" baseAuthorityName="Mann" box="[498,1045,1166,1193]" class="Insecta" family="Formicidae" genus="Antillaemyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="subSpecies" species="albispina" subSpecies="pallipes">
<emphasis box="[498,798,1166,1192]" italics="true" pageId="265" pageNumber="266">Antillaemyrmex albispina</emphasis>
ssp. 
<emphasis box="[857,942,1166,1192]" italics="true" pageId="265" pageNumber="266">pallipes</emphasis>
(Mann)
</taxonomicName>
: 
<treatmentCitation author="Wheeler WM" box="[1057,1268,1166,1193]" page="27" pageId="265" pageNumber="266" year="1931">
<bibRefCitation author="Wheeler WM" box="[1057,1268,1166,1193]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="265" pageNumber="266" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
<emphasis box="[1057,1223,1166,1192]" italics="true" pageId="265" pageNumber="266">
<author box="[1057,1154,1166,1192]" fns="*" inm="true" npo="*" pageId="265" pageNumber="266">Wheeler</author>
, 
<year box="[1168,1223,1166,1192]" pageId="265" pageNumber="266">1931</year>
</emphasis>
: 27
</bibRefCitation>
</treatmentCitation>
. 
<typeStatus box="[1281,1373,1166,1192]" pageId="265" pageNumber="266">Syntype</typeStatus>
workers and gyne; junior synonym of 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[796,1078,1206,1232]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[796,1078,1206,1232]" italics="true" pageId="265" pageNumber="266">Temnothorax albispinus</emphasis>
</taxonomicName>
by 
<treatmentCitation author="Baroni Urbani C." box="[1123,1423,1206,1233]" page="413" pageId="265" pageNumber="266" year="1978">
<bibRefCitation author="Baroni Urbani C." box="[1123,1423,1206,1233]" journalOrPublisher="Entomologica Basiliensia" pageId="265" pageNumber="266" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis box="[1123,1363,1206,1232]" italics="true" pageId="265" pageNumber="266">
<author box="[1123,1295,1206,1232]" case="TC" fns="*" inm="true" npo="*" pageId="265" pageNumber="266">Baroni Urbani</author>
, 
<year box="[1308,1363,1206,1232]" pageId="265" pageNumber="266">1978</year>
</emphasis>
: 413
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection box="[498,1052,1246,1272]" pageId="265" pageNumber="266" type="materials_examined">
<paragraph blockId="265.[498,1542,1166,1910]" box="[498,1052,1246,1272]" pageId="265" pageNumber="266">
One 
<typeStatus box="[557,646,1246,1272]" pageId="265" pageNumber="266">syntype</typeStatus>
worker here designated 
<emphasis bold="true" box="[937,1046,1246,1272]" pageId="265" pageNumber="266">
<typeStatus box="[937,1046,1246,1272]" pageId="265" pageNumber="266">lectotype</typeStatus>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="265" pageNumber="266" type="reference_group">
<paragraph blockId="265.[498,1542,1166,1910]" pageId="265" pageNumber="266">
<treatmentCitationGroup pageId="265" pageNumber="266">
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[530,913,1286,1313]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="species" species="albispina">
<emphasis box="[530,784,1286,1312]" italics="true" pageId="265" pageNumber="266">Leptothorax albispina</emphasis>
(Wheeler)
</taxonomicName>
: 
<treatmentCitation author="Baroni Urbani C." box="[925,1226,1286,1312]" page="413" pageId="265" pageNumber="266" year="1978">
<bibRefCitation author="Baroni Urbani C." box="[925,1226,1286,1312]" journalOrPublisher="Entomologica Basiliensia" pageId="265" pageNumber="266" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis box="[925,1166,1286,1312]" italics="true" pageId="265" pageNumber="266">Baroni Urbani, 1978</emphasis>
: 413
</bibRefCitation>
</treatmentCitation>
. First combination in 
<taxonomicName authority="Mayr" box="[498,638,1325,1351]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis box="[498,638,1325,1351]" italics="true" pageId="265" pageNumber="266">Leptothorax</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="265.[498,1542,1166,1910]" pageId="265" pageNumber="266">
<treatmentCitationGroup pageId="265" pageNumber="266">
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[530,941,1365,1392]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[530,812,1365,1391]" italics="true" pageId="265" pageNumber="266">Temnothorax albispinus</emphasis>
(Wheeler)
</taxonomicName>
: 
<treatmentCitation author="Bolton B." box="[954,1159,1365,1392]" page="271" pageId="265" pageNumber="266" year="2003">
<bibRefCitation author="Bolton B." box="[954,1159,1365,1392]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="265" pageNumber="266" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
<emphasis box="[954,1100,1365,1392]" italics="true" pageId="265" pageNumber="266">
<collectingRegion box="[954,1032,1365,1391]" country="United Kingdom" name="Bolton" pageId="265" pageNumber="266">Bolton</collectingRegion>
, 2003
</emphasis>
: 271
</bibRefCitation>
</treatmentCitation>
. First combination in 
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1405,1431]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="genus">
<emphasis box="[498,655,1405,1431]" italics="true" pageId="265" pageNumber="266">Temnothorax</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="266" lastPageNumber="267" pageId="265" pageNumber="266" type="materials_examined">
<paragraph blockId="265.[498,1542,1166,1910]" pageId="265" pageNumber="266">
<emphasis bold="true" box="[530,834,1445,1471]" pageId="265" pageNumber="266">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861492" collectionCode="MCZC" collectorName="W. M. Wheeler" country="Puerto Rico" location="Culebra" pageId="265" pageNumber="266" specimenCount="1" stateProvince="Culebra" typeStatus="lectotype">
<emphasis box="[843,1069,1445,1471]" italics="true" pageId="265" pageNumber="266">
<typeStatus box="[843,951,1446,1471]" pageId="265" pageNumber="266">Lectotype</typeStatus>
worker of
</emphasis>
<taxonomicName box="[1078,1352,1445,1471]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="species" species="albispina">Macromischa albispina</taxonomicName>
: U.S.A.: 
<collectingCountry name="Puerto Rico" pageId="265" pageNumber="266">Puerto Rico</collectingCountry>
: 
<collectingRegion box="[566,658,1485,1511]" country="Puerto Rico" name="Culebra" pageId="265" pageNumber="266">Culebra</collectingRegion>
<collectorName box="[666,816,1485,1512]" pageId="265" pageNumber="266">Island, W.M</collectorName>
. Wheeler, TYPE 
<collectorName box="[1020,1188,1485,1512]" pageId="265" pageNumber="266">No. A.M.N.H.</collectorName>
, specimen furthest from pin (images of 
<collectorName box="[629,778,1525,1551]" pageId="265" pageNumber="266">M.C.Z. Type</collectorName>
1-3 21013 examined on the MCZ Type Database website) [
<collectionCode box="[505,596,1564,1591]" pageId="265" pageNumber="266">MCZC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="265.[498,1542,1166,1910]" pageId="265" pageNumber="266">
<materialsCitation ID-GBIF-Occurrence="3321861585" collectionCode="MCZC" collectorName="de Mona" country="Puerto Rico" location="Puerto Rico" pageId="265" pageNumber="266" specimenCount="2" typeStatus="lectotype">
<emphasis box="[530,757,1605,1631]" italics="true" pageId="265" pageNumber="266">
<typeStatus box="[530,638,1605,1630]" pageId="265" pageNumber="266">Lectotype</typeStatus>
worker of
</emphasis>
<taxonomicName authority="" box="[765,1359,1605,1632]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="265" pageNumber="266" phylum="Arthropoda" rank="subGenus" subGenus="Antillaemyrmex">
Macromischa (Antillaemyrmex) alibispina pallipe 
<emphasis box="[1349,1365,1605,1630]" italics="true" pageId="265" pageNumber="266">s:</emphasis>
</taxonomicName>
U.S.A.: 
<collectingCountry name="Puerto Rico" pageId="265" pageNumber="266">Puerto Rico</collectingCountry>
: Isla 
<collectorName box="[611,718,1644,1671]" pageId="265" pageNumber="266">de Mona</collectorName>
: Mona Passage, top specimen (images of MCZ Type 1-2 21014 
<typeStatus box="[1452,1541,1645,1671]" pageId="265" pageNumber="266">syntype</typeStatus>
worker examined on the MCZ Type Database website) [
<collectionCode box="[1160,1251,1684,1711]" pageId="265" pageNumber="266">MCZC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="265.[498,1542,1166,1910]" lastBlockId="266.[498,1542,230,1934]" lastPageId="266" lastPageNumber="267" pageId="265" pageNumber="266">
<emphasis bold="true" box="[530,885,1724,1750]" pageId="265" pageNumber="266">Non-type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861602" lastPageId="265" lastPageNumber="267" pageId="265" pageNumber="266">
U.S.A.: 
<collectingCountry box="[979,1119,1724,1751]" name="Puerto Rico" pageId="265" pageNumber="266">Puerto Rico</collectingCountry>
: 
<collectingRegion box="[1130,1215,1724,1750]" country="Puerto Rico" name="Coamo" pageId="265" pageNumber="266">Coamo</collectingRegion>
Springs, 
<specimenCount box="[1323,1428,1724,1750]" pageId="265" pageNumber="266" type="worker">1 worker</specimenCount>
, 1 dealate gyne (LACMENT323175) [
<collectionCode box="[818,907,1763,1790]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="265" pageNumber="266" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861773" lastPageId="265" lastPageNumber="267" pageId="265" pageNumber="266">
<collectingRegion box="[927,1024,1764,1790]" country="Puerto Rico" name="Guanica" pageId="265" pageNumber="266">Guánica</collectingRegion>
Forest 
<geoCoordinate box="[1113,1265,1759,1791]" degrees="17.960000" direction="north" orientation="latitude" pageId="265" pageNumber="266" precision="1" value="17.96">
17.960000 
<emphasis box="[1232,1242,1759,1777]" italics="true" pageId="265" pageNumber="266">
<superScript attach="right" box="[1232,1242,1759,1777]" fontSize="8" pageId="265" pageNumber="266">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[1274,1433,1759,1791]" degrees="66.870000" direction="west" orientation="longitude" pageId="265" pageNumber="266" precision="1" value="-66.87">
66.870000 
<emphasis box="[1392,1402,1759,1777]" italics="true" pageId="265" pageNumber="266">
<superScript attach="right" box="[1392,1402,1759,1777]" fontSize="8" pageId="265" pageNumber="266">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<date pageId="265" pageNumber="266" value="2000-08-28">28 August 2000</date>
, J.A. Torres, ex sifted leaf litter, 1 dealate gyne &amp; 
<specimenCount box="[1236,1352,1804,1830]" pageId="265" pageNumber="266" type="worker">2 workers</specimenCount>
(LACMENT323245) [
<collectionCode box="[754,843,1843,1870]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="265" pageNumber="266" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861501" lastPageId="265" lastPageNumber="267" pageId="265" pageNumber="266">
same data as previous, except: 
<date box="[1223,1400,1844,1870]" pageId="265" pageNumber="266" value="2000-01-01">1 January 2000</date>
J.A. Torres, ex sifted leaf litter, 
<specimenCount box="[718,832,1883,1910]" pageId="265" pageNumber="266" type="worker">4 workers</specimenCount>
(LACMENT323247) [
<collectionCode box="[1093,1182,1883,1910]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="265" pageNumber="266" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861856" lastPageId="266" lastPageNumber="267" pageId="265" pageNumber="266">
<collectingRegion box="[1201,1298,1883,1909]" country="Puerto Rico" name="Guanica" pageId="265" pageNumber="266">Guánica</collectingRegion>
Forest, 
<geoCoordinate box="[1390,1542,1879,1910]" degrees="17.840000" direction="north" orientation="latitude" pageId="265" pageNumber="266" precision="1" value="17.84">
17.840000 
<emphasis box="[1508,1518,1879,1897]" italics="true" pageId="265" pageNumber="266">
<superScript attach="right" box="[1508,1518,1879,1897]" fontSize="8" pageId="265" pageNumber="266">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[498,658,230,261]" degrees="66.860000" direction="west" orientation="longitude" pageId="266" pageNumber="267" precision="1" value="-66.86">
66.860000 
<emphasis box="[617,627,230,248]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[617,627,230,248]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<date box="[672,828,234,261]" pageId="266" pageNumber="267" value="1997-06-14">14 June 1997</date>
, M. Canals, lite trap, 
<specimenCount box="[1082,1161,234,260]" pageId="266" pageNumber="267" type="male">1 male</specimenCount>
(LACMENT323241) [
<collectionCode box="[1425,1513,234,261]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
] 
<specimenCount pageId="266" pageNumber="267" type="male">1 male</specimenCount>
(LACMENT323242) [
<collectionCode box="[819,907,273,300]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[922,1000,274,300]" pageId="266" pageNumber="267" type="male">1 male</specimenCount>
(LACMENT323246) [
<collectionCode box="[1265,1354,273,300]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861367" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<geoCoordinate box="[1374,1526,269,301]" degrees="17.972000" direction="north" orientation="latitude" pageId="266" pageNumber="267" precision="1" value="17.972">
17.972000 
<emphasis box="[1492,1502,269,287]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[1492,1502,269,287]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[498,658,309,340]" degrees="66.868000" direction="west" orientation="longitude" pageId="266" pageNumber="267" precision="1" value="-66.868">
66.868000 
<emphasis box="[617,627,309,327]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[617,627,309,327]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<quantity box="[672,747,314,340]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="266" pageNumber="267" unit="m" value="120.0">120 m</quantity>
, 
<date box="[761,988,314,340]" pageId="266" pageNumber="267" value="1998-09-20">20 September 1998</date>
, R.R. Snelling#98-239, #1998-239, subtropical deciduous forest, ex rotten stick in leaf litter, 
<specimenCount box="[1034,1150,354,380]" pageId="266" pageNumber="267" type="worker">2 workers</specimenCount>
(LACMENT323243) [
<collectionCode box="[1414,1503,353,380]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861493" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<geoCoordinate box="[498,650,389,420]" degrees="17.966667" direction="north" orientation="latitude" pageId="266" pageNumber="267" precision="1" value="17.966667">
17.966667 
<emphasis box="[617,627,389,407]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[617,627,389,407]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[659,818,389,420]" degrees="66.883333" direction="west" orientation="longitude" pageId="266" pageNumber="267" precision="1" value="-66.88333">
66.883333 
<emphasis box="[777,787,389,407]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[777,787,389,407]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<quantity box="[832,922,393,420]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="266" pageNumber="267" unit="m" value="25.0" valueMax="50.0" valueMin="0.0">0–50 m</quantity>
, 
<date box="[936,1092,394,420]" pageId="266" pageNumber="267" value="1993-05-30">30 May 1993</date>
, J. Longino#JTL3493-s, dry forest, sifted leaf litter sample, taken from moist litter in ravine, 
<specimenCount box="[1177,1282,433,460]" pageId="266" pageNumber="267" type="worker">1 worker</specimenCount>
(LACMENT002310) [
<collectionCode box="[505,593,473,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
] 
<specimenCount box="[608,724,473,499]" pageId="266" pageNumber="267" type="worker">2 workers</specimenCount>
(LACMENT002311) [
<collectionCode box="[988,1077,473,500]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="266" pageNumber="267" type="Museum">UCDC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861612" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
same data as previous, except: J. Longino#JTL3494-s, dry forest, strays from first site, 
<specimenCount box="[1124,1228,513,539]" pageId="266" pageNumber="267" type="worker">1 worker</specimenCount>
(LACMENT002269) [
<collectionCode box="[505,595,552,579]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
].
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861580" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingRegion box="[614,711,553,579]" country="Puerto Rico" name="Guanica" pageId="266" pageNumber="267">Guánica</collectingRegion>
Forest, 
<date box="[807,949,553,580]" pageId="266" pageNumber="267" value="1991-06-07">7 June 1991</date>
, J. Torres#592, 
<specimenCount box="[1135,1252,553,579]" pageId="266" pageNumber="267" type="worker">2 workers</specimenCount>
, 1 dealate gyne (LACMENT323244) 
<collectingCountry box="[748,788,593,619]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona: 
<date box="[917,1073,593,619]" pageId="266" pageNumber="267" value="1982-06-30">30 June 1982</date>
, J.A. Torres#65, 
<specimenCount box="[1271,1387,593,619]" pageId="266" pageNumber="267" type="worker">3 workers</specimenCount>
(LACMENT323169) [
<collectionCode box="[754,843,632,659]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861485" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
Maygüez: 
<collectingCountry box="[983,1023,633,659]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona, 
<date box="[1153,1288,633,659]" pageId="266" pageNumber="267" value="1982-07-04">4 July 1982</date>
, J.A. Torres#255, 
<specimenCount pageId="266" pageNumber="267" type="worker">2 workers</specimenCount>
(LACMENT323170) [
<collectionCode box="[856,945,672,699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861613" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingCountry box="[965,1005,673,699]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona, 5 min E of antenna, 
<geoCoordinate box="[1379,1531,668,699]" degrees="18.100000" direction="north" orientation="latitude" pageId="266" pageNumber="267" precision="1" value="18.1">
18.100000 
<emphasis box="[1497,1507,668,686]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[1497,1507,668,686]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[498,658,708,739]" degrees="67.927000" direction="west" orientation="longitude" pageId="266" pageNumber="267" precision="1" value="-67.927">
67.927000 
<emphasis box="[617,627,708,726]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[617,627,708,726]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<date box="[671,811,712,739]" pageId="266" pageNumber="267" value="2006-06-03">3 June 2006</date>
, J.K. Wetterer#272, 
<specimenCount box="[1047,1161,712,738]" pageId="266" pageNumber="267" type="worker">2 workers</specimenCount>
(MCZENT00505426) 
<specimenCount box="[1427,1542,712,738]" pageId="266" pageNumber="267" type="worker">2 workers</specimenCount>
(MCZENT00505427) [
<collectionCode box="[764,854,752,779]" pageId="266" pageNumber="267">MCZC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861438" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingCountry box="[874,914,752,778]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona, 50 min E of antenna, 
<geoCoordinate box="[1302,1454,748,779]" degrees="18.090000" direction="north" orientation="latitude" pageId="266" pageNumber="267" precision="1" value="18.09">
18.090000 
<emphasis box="[1420,1430,748,766]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[1420,1430,748,766]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[498,658,787,819]" degrees="67.910000" direction="west" orientation="longitude" pageId="266" pageNumber="267" precision="1" value="-67.91">
67.910000 
<emphasis box="[617,627,787,805]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[617,627,787,805]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<date box="[672,814,792,819]" pageId="266" pageNumber="267" value="2006-06-03">3 June 2006</date>
, J.K. Wetterer#282, 1 dealate gyne (MCZENT00505431) [
<collectionCode box="[505,595,831,858]" pageId="266" pageNumber="267">MCZC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861801" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingCountry box="[615,655,832,858]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona, Los Cerezos, 
<geoCoordinate box="[941,1093,827,858]" degrees="18.089000" direction="north" orientation="latitude" pageId="266" pageNumber="267" precision="1" value="18.089">
18.089000 
<emphasis box="[1059,1069,827,845]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[1059,1069,827,845]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[1101,1261,827,858]" degrees="67.900000" direction="west" orientation="longitude" pageId="266" pageNumber="267" precision="1" value="-67.9">
67.900000 
<emphasis box="[1219,1229,827,845]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[1219,1229,827,845]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<date box="[1274,1416,832,858]" pageId="266" pageNumber="267" value="2006-06-03">3 June 2006</date>
, James K. Wetterer#277, forest, 
<specimenCount box="[754,858,872,898]" pageId="266" pageNumber="267" type="worker">1 worker</specimenCount>
(CASENT0756097) [
<collectionCode box="[1110,1204,871,898]" pageId="266" pageNumber="267">MMPC</collectionCode>
] 
<specimenCount box="[1219,1298,872,898]" pageId="266" pageNumber="267" type="male">1 male</specimenCount>
(CASENT0756098) [
<collectionCode box="[505,601,911,938]" pageId="266" pageNumber="267">MMPC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861748" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingCountry box="[621,661,912,938]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona, near antenna, 
<geoCoordinate box="[960,1111,907,938]" degrees="18.910000" direction="north" orientation="latitude" pageId="266" pageNumber="267" precision="1" value="18.91">
18.910000 
<emphasis box="[1078,1088,907,925]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[1078,1088,907,925]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[1120,1280,907,938]" degrees="67.936000" direction="west" orientation="longitude" pageId="266" pageNumber="267" precision="1" value="-67.936">
67.936000 
<emphasis box="[1238,1248,907,925]" italics="true" pageId="266" pageNumber="267">
<superScript attach="right" box="[1238,1248,907,925]" fontSize="8" pageId="266" pageNumber="267">°</superScript>
</emphasis>
W
</geoCoordinate>
, 
<quantity box="[1293,1355,912,938]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="266" pageNumber="267" unit="m" value="40.0">40 m</quantity>
, 
<date box="[1368,1511,911,938]" pageId="266" pageNumber="267" value="2006-06-02">2 June 2006</date>
, J.K. Wetterer#JKW-2006-250, 
<specimenCount box="[854,957,951,978]" pageId="266" pageNumber="267" type="worker">1 worker</specimenCount>
(CASENT0758674) [
<collectionCode box="[1204,1293,951,978]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="266" pageNumber="267" type="Museum">UCDC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861504" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingCountry box="[1312,1352,951,977]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona, 
<date pageId="266" pageNumber="267" value="2014-02-21" valueMax="2014-02-26" valueMin="2014-02-21">21-26 February 2014</date>
, 
<specimenCount box="[680,794,991,1018]" pageId="266" pageNumber="267" type="worker">3 workers</specimenCount>
(CASENT0758358) [
<collectionCode box="[1041,1132,991,1018]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="266" pageNumber="267" type="Museum">USNM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861769" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingCountry box="[1151,1191,991,1017]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
de Mona, 
<date box="[1315,1537,992,1018]" pageId="266" pageNumber="267" value="1944-08-11" valueMax="1944-08-31" valueMin="1944-08-11">11-31 August 1944</date>
, Harry A. Beatty, 
<specimenCount box="[700,805,1031,1057]" pageId="266" pageNumber="267" type="worker">1 worker</specimenCount>
(CASENT0758359) [
<collectionCode box="[1056,1147,1031,1058]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="266" pageNumber="267" type="Museum">USNM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861722" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
Cayo Luis Peña, 
<date box="[1365,1519,1031,1057]" pageId="266" pageNumber="267" value="1982-09-10">10 Sept 1982</date>
, J.A. Torres#51b, 
<specimenCount box="[698,814,1071,1097]" pageId="266" pageNumber="267" type="worker">3 workers</specimenCount>
(LACMENT323171) [
<collectionCode box="[1078,1167,1070,1097]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861618" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
<collectingCountry box="[1187,1227,1071,1097]" name="Iceland" pageId="266" pageNumber="267">Isla</collectingCountry>
Piñeras, Humacao 
<date pageId="266" pageNumber="267" value="1982-03-11">11 March 1982</date>
, J.A. Torres#109, 
<specimenCount box="[854,970,1111,1137]" pageId="266" pageNumber="267" type="worker">3 workers</specimenCount>
(LACMENT323172) [
<collectionCode box="[1234,1323,1110,1137]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861578" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
Cayo Ratones, 
<date box="[498,647,1151,1177]" pageId="266" pageNumber="267" value="1982-07-17">17 July 1982</date>
, J.A. Torres#252, 
<specimenCount box="[859,964,1151,1177]" pageId="266" pageNumber="267" type="worker">1 worker</specimenCount>
(LACMENT323173) [
<collectionCode box="[1228,1317,1150,1177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861644" lastPageId="266" lastPageNumber="267" pageId="266" pageNumber="266">
Caja de Muertos, 
<date box="[498,725,1191,1217]" pageId="266" pageNumber="267" value="1980-09-27">27 September 1980</date>
, coll. J.A. Torres#51, 
<specimenCount box="[979,1095,1191,1217]" pageId="266" pageNumber="267" type="worker">3 workers</specimenCount>
(LACMENT323174) [
<collectionCode box="[1359,1447,1190,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="266" pageNumber="267" type="Museum">LACM</collectionCode>
] 
<specimenCount pageId="266" pageNumber="267" type="worker">3 workers</specimenCount>
(CASENT0759975) [
<collectionCode box="[842,920,1230,1257]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="266" pageNumber="267" type="Museum">FSCA</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="266" pageNumber="267" type="distribution">
<paragraph blockId="266.[498,1542,230,1934]" pageId="266" pageNumber="267">
<emphasis bold="true" box="[530,755,1270,1296]" pageId="266" pageNumber="267">Geographic range:</emphasis>
Low elevations of 
<collectingCountry box="[978,1116,1270,1297]" name="Puerto Rico" pageId="266" pageNumber="267">Puerto Rico</collectingCountry>
and its small outlying islands (
<figureCitation box="[508,619,1310,1336]" captionStart="Figure 133" captionStartId="258.[116,181,1036,1058]" captionTargetBox="[90,1541,234,1023]" captionTargetId="figure-184@258.[90,1524,234,1008]" captionTargetPageId="258" captionText="Figure 133 Geographical and elevational distribution of the pulchellus group. (A) Temnothorax agavicola sp. nov. (B) T. albispinus. (C) T. bahoruco sp. nov. (D) T. balaclava sp. nov. (E) T. ciferrii (F) T. flavidulus (G) T. harlequina sp. nov. (H) T. hippolytus sp. nov. (I) T. laticrus sp. nov. (J) T. magnabulla sp. nov. (K) T. pulchellus (L) T. terricola (M) T. torrei (N) T. wettereri sp. nov. (O) T. wilsoni sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-133" figureDoi="http://doi.org/10.5281/zenodo.5102387" httpUri="https://zenodo.org/record/5102387/files/figure.png" pageId="266" pageNumber="267">Fig. 133B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="267" lastPageNumber="268" pageId="266" pageNumber="267" type="diagnosis">
<paragraph blockId="266.[498,1542,230,1934]" lastBlockId="267.[498,1542,234,1934]" lastPageId="267" lastPageNumber="268" pageId="266" pageNumber="267">
<emphasis bold="true" box="[530,754,1350,1376]" pageId="266" pageNumber="267">Worker diagnosis:</emphasis>
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[763,1044,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="266" pageNumber="267" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[763,1044,1350,1376]" italics="true" pageId="266" pageNumber="267">Temnothorax albispinus</emphasis>
</taxonomicName>
can be separated from all other species in the 
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[544,619,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="266" pageNumber="267" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[544,619,1390,1416]" italics="true" pageId="266" pageNumber="267">salvini</emphasis>
</taxonomicName>
clade by the following character combination: head in full face view with posterior margin flat to slightly convex; mesosoma compact; in profile view, dorsum of mesosoma evenly convex; metanotal groove not impressed; propodeum not depressed below the level of the promesonotum; propodeum bearing standing setae dorsally; propodeal spines about as long as the propodeal declivity, directed posteriorly, and straight to downcurved; in dorsal view, propodeal spines broadly approximated basally, the negative space between them “U” shaped; hind femora moderately to strongly incrassate; petiolar node rounded dorsally; in dorsal view, apex of petiolar node narrower or slightly wider than the caudal cylinder of the petiole; petiolar node with four erect setae dorsally; postpetiole very broad: greater than 2.5 times the width of the petiole; median clypeal carina well-defined; dorsum of head uniformly areolate; mesosoma bearing more than ten erect setae dorsally; setae on head, mesosoma, waist segments and gaster erect, moderately long, sparse and blunt (never long and tapering); integument variously colored: either predominantly dark brown, with light yellow antennal funiculus, tibiae, tarsi, and propodeal spines, or femora and antennal scapes yellow in addition to the preceding body parts, or uniformly yellow.
</paragraph>
<paragraph blockId="267.[498,1542,234,1934]" pageId="267" pageNumber="268">
<emphasis bold="true" box="[530,719,314,340]" pageId="267" pageNumber="268">Similar species:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[727,1000,314,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="agavicola" status="sp. nov.">
<emphasis box="[727,1000,314,340]" italics="true" pageId="267" pageNumber="268">Temnothorax agavicola</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1009,1102,314,340]" pageId="267" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1115,1233,314,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="laticrus" status="sp. nov.">
<emphasis box="[1115,1233,314,340]" italics="true" pageId="267" pageNumber="268">T. laticrus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1241,1335,314,340]" pageId="267" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
, 
<emphasis italics="true" pageId="267" pageNumber="268">
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1348,1447,315,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="torrei">T. torrei</taxonomicName>
, 
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,631,355,380]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="wettereri" status="sp. nov.">T. wettereri</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[638,730,354,380]" pageId="267" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
, and members of the 
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[982,1090,354,380]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[982,1090,354,380]" italics="true" pageId="267" pageNumber="268">pastinifer</emphasis>
</taxonomicName>
group. 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[1180,1460,354,380]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[1180,1460,354,380]" italics="true" pageId="267" pageNumber="268">Temnothorax albispinus</emphasis>
</taxonomicName>
can be separated from most members of the 
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[944,1052,394,420]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[944,1052,394,420]" italics="true" pageId="267" pageNumber="268">pastinifer</emphasis>
</taxonomicName>
group, by the petiolar node, which not squamiform and relatively narrow in dorsal view: 0.9 to 1.1 times the width of the caudal cylinder of the petiole in 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[797,946,473,499]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[797,946,473,499]" italics="true" pageId="267" pageNumber="268">T. albispinus</emphasis>
</taxonomicName>
vs. greater than 1.3 times in the 
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1337,1445,473,499]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[1337,1445,473,499]" italics="true" pageId="267" pageNumber="268">pastinifer</emphasis>
</taxonomicName>
group. The petiolar node shape, in addition to the long propodeal spines and relatively narrow hind femora will distinguish 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[832,978,553,579]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[832,978,553,579]" italics="true" pageId="267" pageNumber="268">T. albispinus</emphasis>
</taxonomicName>
from 
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1049,1165,553,579]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="laticrus" status="sp. nov.">
<emphasis box="[1049,1165,553,579]" italics="true" pageId="267" pageNumber="268">T. laticrus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1171,1261,554,580]" pageId="267" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
The erect setae count of four on the dorsum of the petiolar node differentiates 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[1141,1290,593,619]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[1141,1290,593,619]" italics="true" pageId="267" pageNumber="268">T. albispinus</emphasis>
</taxonomicName>
from 
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1364,1460,594,619]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1364,1460,594,619]" italics="true" pageId="267" pageNumber="268">T. torrei</emphasis>
</taxonomicName>
, which has only two along the posterior margin. The flat to weakly convex posterior margin of the head of 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[660,806,673,699]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[660,806,673,699]" italics="true" pageId="267" pageNumber="268">T. albispinus</emphasis>
</taxonomicName>
contrasts with 
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[980,1117,673,699]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="agavicola" status="sp. nov.">
<emphasis box="[980,1117,673,699]" italics="true" pageId="267" pageNumber="268">T. agavicola</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1123,1213,673,699]" pageId="267" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
, which has a weakly concave posterior head margin. The body of 
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[928,1062,713,738]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="wettereri" status="sp. nov.">
<emphasis box="[928,1062,713,738]" italics="true" pageId="267" pageNumber="268">T. wettereri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1071,1162,713,739]" pageId="267" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
is mostly devoid of erect setae, whereas 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[601,750,752,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="267" pageNumber="268" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[601,750,752,778]" italics="true" pageId="267" pageNumber="268">T. albispinus</emphasis>
</taxonomicName>
has more than ten erect setae on the dorsum of the mesosoma.
</paragraph>
</subSubSection>
<subSubSection lastPageId="272" lastPageNumber="273" pageId="267" pageNumber="268" type="description">
<paragraph blockId="267.[498,1542,234,1934]" pageId="267" pageNumber="268">
<emphasis bold="true" box="[530,1047,792,819]" pageId="267" pageNumber="268">
Worker measurements &amp; indices (
<emphasis bold="true" box="[950,966,793,819]" italics="true" pageId="267" pageNumber="268">n</emphasis>
= 28):
</emphasis>
SL = 
<geoCoordinate box="[1119,1181,792,818]" degrees="0.432" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.432">0.432</geoCoordinate>
<geoCoordinate box="[1181,1258,792,819]" degrees="0.530" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.53">–0.530</geoCoordinate>
(0.487); FRS = 
<geoCoordinate box="[579,641,832,858]" degrees="0.167" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.167">0.167</geoCoordinate>
<geoCoordinate box="[642,719,832,858]" degrees="0.216" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.216">–0.216</geoCoordinate>
(0.190); CW = 
<geoCoordinate box="[907,969,832,859]" degrees="0.518" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.518">0.518</geoCoordinate>
<geoCoordinate box="[969,1046,832,858]" degrees="0.641" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.641">–0.641</geoCoordinate>
(0.578); CWb = 
<geoCoordinate box="[1249,1311,832,859]" degrees="0.456" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.456">0.456</geoCoordinate>
<geoCoordinate box="[1311,1388,832,859]" degrees="0.590" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.59">–0.590</geoCoordinate>
(0.525); PoOC = 
<geoCoordinate box="[604,666,872,898]" degrees="0.233" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.233">0.233</geoCoordinate>
<geoCoordinate box="[667,744,872,898]" degrees="0.294" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.294">–0.294</geoCoordinate>
(0.265); CL = 
<geoCoordinate box="[919,981,872,899]" degrees="0.563" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.563">0.563</geoCoordinate>
<geoCoordinate box="[981,1058,872,898]" degrees="0.694" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.694">–0.694</geoCoordinate>
(0.637); EL = 
<geoCoordinate box="[1231,1293,872,898]" degrees="0.120" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.12">0.120</geoCoordinate>
<geoCoordinate box="[1293,1370,872,899]" degrees="0.156" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.156">–0.156</geoCoordinate>
(0.139); EW = 
<geoCoordinate box="[577,639,912,938]" degrees="0.091" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.091">0.091</geoCoordinate>
<geoCoordinate box="[640,716,911,938]" degrees="0.116" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.116">–0.116</geoCoordinate>
(0.103); MD = 
<geoCoordinate box="[905,967,912,938]" degrees="0.120" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.12">0.120</geoCoordinate>
<geoCoordinate box="[967,1044,912,938]" degrees="0.179" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.179">–0.179</geoCoordinate>
(0.147); WL = 
<geoCoordinate box="[1228,1290,911,937]" degrees="0.608" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.608">0.608</geoCoordinate>
<geoCoordinate box="[1291,1368,911,938]" degrees="0.763" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.763">–0.763</geoCoordinate>
(0.691); SPST = 
<geoCoordinate box="[594,656,952,978]" degrees="0.190" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.19">0.190</geoCoordinate>
<geoCoordinate box="[657,734,951,978]" degrees="0.276" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.276">–0.276</geoCoordinate>
(0.245); MPST = 
<geoCoordinate box="[948,1010,952,978]" degrees="0.184" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.184">0.184</geoCoordinate>
<geoCoordinate box="[1011,1088,951,978]" degrees="0.250" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.25">–0.250</geoCoordinate>
(0.224); PEL = 
<geoCoordinate box="[1277,1339,951,978]" degrees="0.259" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.259">0.259</geoCoordinate>
<geoCoordinate box="[1339,1416,951,978]" degrees="0.350" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.35">–0.350</geoCoordinate>
(0.317); NOL = 
<geoCoordinate box="[591,653,991,1017]" degrees="0.161" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.161">0.161</geoCoordinate>
<geoCoordinate box="[654,731,991,1018]" degrees="0.231" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.231">–0.231</geoCoordinate>
(0.195); NOH = 
<geoCoordinate box="[937,999,992,1018]" degrees="0.093" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.093">0.093</geoCoordinate>
<geoCoordinate box="[999,1076,991,1018]" degrees="0.147" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.147">–0.147</geoCoordinate>
(0.120); PEH = 
<geoCoordinate box="[1272,1334,992,1018]" degrees="0.172" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.172">0.172</geoCoordinate>
<geoCoordinate box="[1334,1411,991,1018]" degrees="0.238" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.238">–0.238</geoCoordinate>
(0.214); PPL = 
<geoCoordinate box="[581,643,1031,1057]" degrees="0.171" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.171">0.171</geoCoordinate>
<geoCoordinate box="[643,720,1031,1057]" degrees="0.240" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.24">–0.240</geoCoordinate>
(0.208); PPH = 
<geoCoordinate box="[916,978,1031,1057]" degrees="0.209" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.209">0.209</geoCoordinate>
<geoCoordinate box="[978,1055,1031,1057]" degrees="0.297" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.297">–0.297</geoCoordinate>
(0.246); PW = 
<geoCoordinate box="[1241,1303,1031,1057]" degrees="0.332" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.332">0.332</geoCoordinate>
<geoCoordinate box="[1303,1380,1031,1057]" degrees="0.416" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.416">–0.416</geoCoordinate>
(0.386); SBPA = 
<geoCoordinate box="[600,662,1071,1098]" degrees="0.125" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.125">0.125</geoCoordinate>
<geoCoordinate box="[662,739,1071,1097]" degrees="0.180" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.18">–0.180</geoCoordinate>
(0.154); SPTI = 
<geoCoordinate box="[938,1000,1071,1097]" degrees="0.171" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.171">0.171</geoCoordinate>
<geoCoordinate box="[1000,1077,1071,1098]" degrees="0.259" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.259">–0.259</geoCoordinate>
(0.218); PEW = 
<geoCoordinate box="[1279,1341,1071,1097]" degrees="0.119" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.119">0.119</geoCoordinate>
<geoCoordinate box="[1341,1418,1071,1097]" degrees="0.169" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.169">–0.169</geoCoordinate>
(0.146); PNW = 
<geoCoordinate box="[599,661,1111,1137]" degrees="0.116" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.116">0.116</geoCoordinate>
<geoCoordinate box="[661,738,1111,1137]" degrees="0.176" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.176">–0.176</geoCoordinate>
(0.148); PPW = 
<geoCoordinate box="[940,1002,1111,1137]" degrees="0.334" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.334">0.334</geoCoordinate>
<geoCoordinate box="[1002,1079,1111,1137]" degrees="0.434" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.434">–0.434</geoCoordinate>
(0.385); HFL = 
<geoCoordinate box="[1273,1335,1111,1137]" degrees="0.431" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.431">0.431</geoCoordinate>
<geoCoordinate box="[1335,1412,1111,1138]" degrees="0.534" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.534">–0.534</geoCoordinate>
(0.492); HFWmax = 
<geoCoordinate box="[648,710,1151,1177]" degrees="0.120" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.12">0.120</geoCoordinate>
<geoCoordinate box="[711,788,1151,1178]" degrees="0.157" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.157">–0.157</geoCoordinate>
(0.141); HFWmin = 
<geoCoordinate box="[1041,1103,1151,1177]" degrees="0.044" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.044">0.044</geoCoordinate>
<geoCoordinate box="[1104,1181,1151,1178]" degrees="0.058" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.058">–0.058</geoCoordinate>
(0.050); CS = 
<geoCoordinate box="[1354,1416,1151,1177]" degrees="0.738" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.738">0.738</geoCoordinate>
<geoCoordinate box="[1417,1494,1151,1177]" degrees="0.937" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.937">–0.937</geoCoordinate>
(0.844); ES = 
<geoCoordinate box="[660,722,1191,1217]" degrees="0.169" direction="north" orientation="latitude" pageId="267" pageNumber="268" precision="55" value="0.169">0.169</geoCoordinate>
<geoCoordinate box="[723,799,1191,1217]" degrees="0.212" direction="west" orientation="longitude" pageId="267" pageNumber="268" precision="55" value="-0.212">–0.212</geoCoordinate>
(0.191); SI = 86–97 (93); OI = 22–24 (23); CI = 79–87 (82); WLI = 123–138 (132); SBI = 25–33 (29); PSI = 31–39 (36); PWI = 247–281 (265); PLI = 121–178 (153); NI = 141–216 (164); PNWI = 85–117 (102); NLI = 56–78 (62); FI = 245–320 (283).
</paragraph>
<paragraph blockId="267.[498,1542,234,1934]" pageId="267" pageNumber="268">
<emphasis bold="true" box="[530,777,1350,1376]" pageId="267" pageNumber="268">Worker description:</emphasis>
In full–face view, head subquadrate, longer than broad (CI 79–87). Mandibles weakly striate, shining, and armed with five teeth: the apical-most well developed, followed by a four roughly equally developed smaller teeth. Anterior clypeal margin entire to very slightly concave medially. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 86–97). Antennae 12-segmented; antennal club of three segments, with the apical-most segment longer than the preceding two in combination. Frontal carinae very short, extending past the antennal toruli by the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, converging evenly to the mandibular insertions. Posterior head margin flat to very slightly convex, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="267.[498,1542,234,1934]" lastBlockId="268.[498,1542,234,1934]" lastPageId="268" lastPageNumber="269" pageId="267" pageNumber="268">
In profile view, compound eyes ovular and moderately large (OI 22–24), with 11 ommatidia in longest row. Pronotal declivity indistinct, neck and anterior face of pronotum forming a rounded ~120 
<emphasis box="[910,920,1863,1881]" italics="true" pageId="267" pageNumber="268">
<superScript attach="left" box="[910,920,1863,1881]" fontSize="8" pageId="267" pageNumber="268">°</superScript>
</emphasis>
angle. Mesosoma evenly, but weakly, convex dorsally from where it joins the pronotal neck to the propodeal spines. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which is well developed. Metanotal groove nearly entirely absent: only visible as a faint disruption in the surface sculpture between meso- and metacoxal insertions to the minute metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about three spiracle diameters. Propodeal spines well developed and moderately long (PSI 31–39), about as long as the propodeal declivity, tapering evenly from the base, evenly downcurved, and acute. Propodeal declivity straight and flat, forming a rounded ~100 
<emphasis box="[789,799,548,566]" italics="true" pageId="268" pageNumber="269">
<superScript attach="left" box="[789,799,548,566]" fontSize="8" pageId="268" pageNumber="269">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and very weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 121–178), without tubercles anterodorsally. Subpetiolar process in the form of a weakly developed, blunt tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle short, comprising about one third of the total length of the petiole. Petiolar node robust and erect: transition between peduncle and node marked by a rounded angle of ~120 
<emphasis box="[825,835,827,845]" italics="true" pageId="268" pageNumber="269">
<superScript attach="none" box="[825,835,827,845]" fontSize="8" pageId="268" pageNumber="269">°</superScript>
</emphasis>
, resulting in a strongly concave anterior node face; anterior face forming a rounded ~90 
<emphasis box="[830,840,867,885]" italics="true" pageId="268" pageNumber="269">
<superScript attach="left" box="[830,840,867,885]" fontSize="8" pageId="268" pageNumber="269">°</superScript>
</emphasis>
angle with the dorsal face, which is evenly convex; dorsal face rounding evenly into the posterior face, which is straight and forms a ~100 
<emphasis box="[1405,1415,907,925]" italics="true" pageId="268" pageNumber="269">
<superScript attach="left" box="[1405,1415,907,925]" fontSize="8" pageId="268" pageNumber="269">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole evenly rounded anterodorsally, before flattening posterodorsally; weakly lobed ventrally.
</paragraph>
<paragraph blockId="268.[498,1542,234,1934]" pageId="268" pageNumber="269">In dorsal view, humeri weakly developed: rounded and only slightly wider than the rest of the mesosoma; mesothoracic spiracles not protruding past the lateral margins of the mesosoma. Metanotal groove absent: mesonotum and propodeum completely fused and converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by slightly more than their length; negative space between them “U” shaped. Petiolar peduncle with spiracles slightly protruding past the lateral margins, the peduncle subtly constricted anterior to them. Petiolar node campaniform: rounded anteriorly, flatter posteriorly, slightly wider than the peduncle, and evenly grading into the caudal cylinder, which is slightly wider than the node. Postpetiole very broad (PWI 247–281) and campaniform, articulating with the entire anterior margin of the gaster. Anterior margin of the postpetiole broadly convex, with corners evenly rounding into the lateral margins, which evenly diverge to the angulate posterior corners; posterior margin broadly concave. Metafemur moderately to strongly incrassate (FI 245–320).</paragraph>
<paragraph blockId="268.[498,1542,234,1934]" lastBlockId="269.[498,1542,234,1934]" lastPageId="269" lastPageNumber="270" pageId="268" pageNumber="269">Sculpture: median clypeal carina present, flanked by additional faint carinae over weakly areolate ground sculpture. Antennal scapes weakly areolate and dull. Cephalic dorsum and lateral surfaces of head densely areolate; ventral surface shining through weaker areolate sculpture. Mesosoma with uniformly areolate sculpture, with the areolae subtly arranged into longitudinal rows on the dorsal faces of the pronotum, meso- and metapleurae; area directly anterior to the propodeal spiracle, as well as the area between the propodeal spiracle and base of the propodeal spines shining through weaker sculpture. Femora shining through weak areolate sculpture. Petiole smooth and shining ventrally, with shallow areolate sculpture surrounding the base of the petiolar node, which is mostly smooth and shining dorsally. Dorsal surface of postpetiole dull with weak costulae and shallow areolate sculpture. Gaster smooth and shining, with extensive spectral iridescence on the first tergite. Surface of the first gastral sternite smooth and shining, with weaker spectral iridescence.</paragraph>
<paragraph blockId="269.[498,1542,234,1934]" pageId="269" pageNumber="270">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of head, mesosoma, waist segments and gaster with short, erect, blunt-tipped setae, the longest of which are roughly the width of the compound eye. The head bears ~28, mesosoma ~24, petiole 4, postpetiole ~12, and first gastral tergite ~44 setae. Sparse, adpressed pubescence present on the entire body, but only apparent on the gaster due to the dense ground sculpture on the rest of the body.</paragraph>
<paragraph blockId="269.[498,1542,234,1934]" pageId="269" pageNumber="270">Color: head, mesosoma (except for propodeal spines), waist segments (except for strigil), and gaster are uniformly dark brown. Mandibles, extremities, and strigil are pale yellow.</paragraph>
<paragraph blockId="269.[498,1542,234,1934]" pageId="269" pageNumber="270">
<emphasis bold="true" box="[530,1001,752,779]" pageId="269" pageNumber="270">
Gyne measurements &amp; indices (
<emphasis bold="true" box="[919,935,753,779]" italics="true" pageId="269" pageNumber="270">n</emphasis>
= 4):
</emphasis>
SL = 
<geoCoordinate box="[1073,1135,752,779]" degrees="0.561" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.561">0.561</geoCoordinate>
<geoCoordinate box="[1136,1213,752,778]" degrees="0.613" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.613">–0.613</geoCoordinate>
(0.591); FRS = 
<geoCoordinate box="[1400,1462,752,778]" degrees="0.224" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.224">0.224</geoCoordinate>
<geoCoordinate box="[1463,1539,752,779]" degrees="0.254" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.254">–0.254</geoCoordinate>
(0.243); CW = 
<geoCoordinate box="[678,740,792,818]" degrees="0.776" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.776">0.776</geoCoordinate>
<geoCoordinate box="[740,817,792,818]" degrees="0.807" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.807">–0.807</geoCoordinate>
(0.787); CWb = 
<geoCoordinate box="[1020,1082,792,819]" degrees="0.715" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.715">0.715</geoCoordinate>
<geoCoordinate box="[1082,1159,792,819]" degrees="0.758" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.758">–0.758</geoCoordinate>
(0.731); PoOC = 
<geoCoordinate box="[1372,1434,792,818]" degrees="0.278" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.278">0.278</geoCoordinate>
<geoCoordinate box="[1434,1511,792,819]" degrees="0.315" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.315">–0.315</geoCoordinate>
(0.301); CL = 
<geoCoordinate box="[665,727,832,858]" degrees="0.716" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.716">0.716</geoCoordinate>
<geoCoordinate box="[727,804,832,859]" degrees="0.775" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.775">–0.775</geoCoordinate>
(0.756); EL = 
<geoCoordinate box="[977,1039,832,858]" degrees="0.210" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.21">0.210</geoCoordinate>
<geoCoordinate box="[1039,1116,832,858]" degrees="0.224" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.224">–0.224</geoCoordinate>
(0.217); EW = 
<geoCoordinate box="[1302,1364,832,858]" degrees="0.160" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.16">0.160</geoCoordinate>
<geoCoordinate box="[1364,1441,832,859]" degrees="0.185" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.185">–0.185</geoCoordinate>
(0.169); MD = 
<geoCoordinate box="[580,642,872,898]" degrees="0.146" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.146">0.146</geoCoordinate>
<geoCoordinate box="[642,719,872,899]" degrees="0.154" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.154">–0.154</geoCoordinate>
(0.150); WL = 
<geoCoordinate box="[904,966,872,899]" degrees="1.153" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="1.153">1.153</geoCoordinate>
<geoCoordinate box="[966,1043,872,898]" degrees="1.262" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-1.262">–1.262</geoCoordinate>
(1.206); SPST = 
<geoCoordinate box="[1245,1307,872,898]" degrees="0.282" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.282">0.282</geoCoordinate>
<geoCoordinate box="[1308,1385,872,898]" degrees="0.328" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.328">–0.328</geoCoordinate>
(0.302); MPST = 
<geoCoordinate box="[607,669,912,938]" degrees="0.304" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.304">0.304</geoCoordinate>
<geoCoordinate box="[669,746,912,938]" degrees="0.342" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.342">–0.342</geoCoordinate>
(0.321); PEL = 
<geoCoordinate box="[935,997,912,938]" degrees="0.384" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.384">0.384</geoCoordinate>
<geoCoordinate box="[997,1074,911,938]" degrees="0.466" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.466">–0.466</geoCoordinate>
(0.438); NOL = 
<geoCoordinate box="[1274,1336,912,938]" degrees="0.212" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.212">0.212</geoCoordinate>
<geoCoordinate box="[1336,1413,911,938]" degrees="0.265" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.265">–0.265</geoCoordinate>
(0.244); NOH = 
<geoCoordinate box="[598,660,952,978]" degrees="0.131" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.131">0.131</geoCoordinate>
<geoCoordinate box="[660,737,951,978]" degrees="0.176" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.176">–0.176</geoCoordinate>
(0.159); PEH = 
<geoCoordinate box="[933,995,951,978]" degrees="0.256" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.256">0.256</geoCoordinate>
<geoCoordinate box="[995,1072,952,978]" degrees="0.310" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.31">–0.310</geoCoordinate>
(0.288); PPL = 
<geoCoordinate box="[1262,1324,952,978]" degrees="0.210" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.21">0.210</geoCoordinate>
<geoCoordinate box="[1324,1401,952,978]" degrees="0.297" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.297">–0.297</geoCoordinate>
(0.251); PPH = 
<geoCoordinate box="[588,650,991,1017]" degrees="0.316" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.316">0.316</geoCoordinate>
<geoCoordinate box="[650,727,991,1018]" degrees="0.359" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.359">–0.359</geoCoordinate>
(0.336); PW = 
<geoCoordinate box="[912,974,991,1018]" degrees="0.715" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.715">0.715</geoCoordinate>
<geoCoordinate box="[975,1052,991,1018]" degrees="0.779" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.779">–0.779</geoCoordinate>
(0.757); SBPA = 
<geoCoordinate box="[1260,1322,991,1018]" degrees="0.354" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.354">0.354</geoCoordinate>
<geoCoordinate box="[1322,1399,991,1018]" degrees="0.374" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.374">–0.374</geoCoordinate>
(0.363); SPTI = 
<geoCoordinate box="[590,652,1031,1057]" degrees="0.293" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.293">0.293</geoCoordinate>
<geoCoordinate box="[653,730,1031,1058]" degrees="0.345" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.345">–0.345</geoCoordinate>
(0.327); PEW = 
<geoCoordinate box="[932,994,1031,1057]" degrees="0.182" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.182">0.182</geoCoordinate>
<geoCoordinate box="[994,1071,1031,1057]" degrees="0.206" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.206">–0.206</geoCoordinate>
(0.196); PNW = 
<geoCoordinate box="[1278,1340,1031,1057]" degrees="0.189" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.189">0.189</geoCoordinate>
<geoCoordinate box="[1340,1417,1031,1057]" degrees="0.212" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.212">–0.212</geoCoordinate>
(0.201); PPW = 
<geoCoordinate box="[590,652,1071,1097]" degrees="0.492" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.492">0.492</geoCoordinate>
<geoCoordinate box="[653,730,1071,1098]" degrees="0.553" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.553">–0.553</geoCoordinate>
(0.533); HFL = 
<geoCoordinate box="[917,979,1071,1097]" degrees="0.629" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.629">0.629</geoCoordinate>
<geoCoordinate box="[979,1056,1071,1097]" degrees="0.674" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.674">–0.674</geoCoordinate>
(0.655); HFWmax = 
<geoCoordinate box="[1306,1368,1071,1097]" degrees="0.122" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.122">0.122</geoCoordinate>
<geoCoordinate box="[1368,1445,1071,1098]" degrees="0.151" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.151">–0.151</geoCoordinate>
(0.139); HFWmin = 
<geoCoordinate box="[645,707,1111,1138]" degrees="0.051" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.051">0.051</geoCoordinate>
<geoCoordinate box="[708,785,1111,1137]" degrees="0.061" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.061">–0.061</geoCoordinate>
(0.057); CS = 
<geoCoordinate box="[958,1020,1111,1137]" degrees="1.073" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="1.073">1.073</geoCoordinate>
<geoCoordinate box="[1021,1098,1111,1137]" degrees="1.142" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-1.142">–1.142</geoCoordinate>
(1.109); ES = 
<geoCoordinate box="[1269,1331,1111,1137]" degrees="0.293" direction="north" orientation="latitude" pageId="269" pageNumber="270" precision="55" value="0.293">0.293</geoCoordinate>
<geoCoordinate box="[1331,1408,1111,1137]" degrees="0.313" direction="west" orientation="longitude" pageId="269" pageNumber="270" precision="55" value="-0.313">–0.313</geoCoordinate>
(0.301); SI = 78–86 (81); OI = 26–28 (27); CI = 92–100 (97); WLI = 161–172 (165); SBI = 48–52 (50); PSI = 23–28 (25); PWI = 260–284 (273); PLI = 148–189 (177); NI = 151–162 (154); PNWI = 100–105 (103); NLI = 55–57 (56); FI = 234–256 (245).
</paragraph>
<paragraph blockId="269.[498,1542,234,1934]" pageId="269" pageNumber="270">
<emphasis bold="true" box="[530,747,1270,1296]" pageId="269" pageNumber="270">Gyne description:</emphasis>
In full-face view, head subquadrate, roughly as long as broad (CI 92–100). Mandibles weakly striate, shining, and armed with five teeth: the apical-most well developed, followed by a four roughly equally developed smaller teeth. Anterior clypeal very slightly concave medially. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 78–86). Antennae 12-segmented; antennal club of three segments, with the apical-most segment as long as the preceding two in combination. Frontal carinae very short, extending past the antennal toruli by one and a half times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head slightly convex, converging evenly to the mandibular insertions. Posterior head margin flat to very slightly convex, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="269.[498,1542,234,1934]" lastBlockId="270.[498,1542,234,1934]" lastPageId="270" lastPageNumber="271" pageId="269" pageNumber="270">
In profile view, compound eyes teardrop-ovular and large (OI 26–28), with 16 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, covering the dorsal surface of the pronotum, and flat dorsally. Mesoscutellum depressed slightly below the level of the mesoscutum; posterior margin of mesoscutellum even with the posterior margin of the metanotum. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines stout and well developed, but short (PSI 23–28), about half as long as the propodeal declivity, tapering evenly from the base, straight, and directed posteriorly. Propodeal declivity straight and flat, forming a rounded ~90 
<emphasis box="[1083,1093,309,327]" italics="true" pageId="270" pageNumber="271">
<superScript attach="left" box="[1083,1093,309,327]" fontSize="8" pageId="270" pageNumber="271">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and very weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 148–189), without tubercles anterodorsally. Subpetiolar process in the form of a weakly developed, blunt triangular tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle short; comprising about one third of the total length of the petiole. Petiolar node erect: transition between peduncle and node evenly rounded, resulting in a very slightly concave anterior node face; anterior face forming a sharp ~90 
<emphasis box="[796,806,628,646]" italics="true" pageId="270" pageNumber="271">
<superScript attach="left" box="[796,806,628,646]" fontSize="8" pageId="270" pageNumber="271">°</superScript>
</emphasis>
angle with the dorsal face, which is evenly convex and short; dorsal face rounding evenly into the posterior face, which forms a ~130 
<emphasis box="[1344,1354,668,686]" italics="true" pageId="270" pageNumber="271">
<superScript attach="left" box="[1344,1354,668,686]" fontSize="8" pageId="270" pageNumber="271">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole evenly rounded anteriorly, dorsal face weakly bulging anterodorsally before flattening posteriorly; ventral surface lobed.
</paragraph>
<paragraph blockId="270.[498,1542,234,1934]" pageId="270" pageNumber="271">In dorsal view, mesoscutum covering pronotum anteriorly, but humeri visible laterally as rounded sclerites. Propodeal spines weakly diverging proximally, but converging distally, their apices separated from each other by about one and a half times their length. Petiolar peduncle with spiracles slightly protruding past the lateral margins, the peduncle broadened where they arise. Petiolar node weakly convex anteriorly, its shape roughly trapezoidal; posterior face of node indistinct. Petiolar node slightly wider than the peduncle, and evenly grading into the caudal cylinder, which is roughly the same width as the node. Postpetiole very broad (PWI 260–284), anteroposteriorly compressed, and campaniform, articulating with most of the anterior margin of the gaster, leaving small, angulate margins on each side of the gaster exposed. Anterior margin of postpetiole weakly convex, with corners evenly rounding into the lateral margins, which evenly diverge to the angulate posterior corners; posterior margin broadly concave. Metafemur weakly to moderately incrassate (FI 234–256).</paragraph>
<paragraph blockId="270.[498,1542,234,1934]" pageId="270" pageNumber="271">Sculpture: median clypeal carina present, flanked by additional equally strong carinae over weakly areolate ground sculpture. Antennal scapes weakly areolate and moderately shining. Cephalic dorsum densely areolate, with weak longitudinal striae. Ventral surface of head weakly, but densely, areolate. Pronotal neck areolate. Pronotum, propodeum, and posterior halves of the anepisternum and katepisternum sculptured like the cephalic dorsum; anterior halves of anepisternum and katepisternum smooth and shining. Mesoscutum costulate with weak areolate sculpture; a small patch of smooth and shining sculpture present anteromedially. Mesoscutellum densely areolate, with a central strip of weaker, shining sculpture. Femora smooth and shining, with traces weak areolate sculpture on the distal quarter. Petiole covered with shallow areolate sculpture, which is weaker ventrally and on the angulate dorsal surface of the node. Dorsal surface of postpetiole dull, with weak costulae and shallow areolate sculpture. Gaster smooth and shining, with extensive spectral iridescence on the first, second, and third tergites. Surface of the and first gastral sternite smooth and shining, with weaker spectral iridescence.</paragraph>
<paragraph blockId="270.[498,1542,234,1934]" lastBlockId="271.[498,1542,234,1895]" lastPageId="271" lastPageNumber="272" pageId="270" pageNumber="271">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of head, mesosoma, waist segments and gaster with short, erect, blunt-tipped setae, the longest of which are roughly a third of the width of the compound eye. Sparse, adpressed pubescence present on the entire body, but only apparent on the gaster due to the dense ground sculpture on the rest of the body.</paragraph>
<paragraph blockId="271.[498,1542,234,1895]" pageId="271" pageNumber="272">Color: head, mesosoma (except for propodeal spines), waist segments (except for strigil), and gaster are uniformly dark brown. Mandibles, extremities, and strigil are pale yellow.</paragraph>
<paragraph blockId="271.[498,1542,234,1895]" pageId="271" pageNumber="272">
<emphasis bold="true" box="[530,997,473,500]" pageId="271" pageNumber="272">
Male measurements &amp; indices (
<emphasis bold="true" box="[916,932,474,500]" italics="true" pageId="271" pageNumber="272">n</emphasis>
= 4):
</emphasis>
SL = 
<geoCoordinate box="[1070,1132,473,499]" degrees="0.312" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.312">0.312</geoCoordinate>
<geoCoordinate box="[1132,1209,473,499]" degrees="0.338" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.338">–0.338</geoCoordinate>
(0.322); FRS = 
<geoCoordinate box="[1397,1459,473,499]" degrees="0.137" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.137">0.137</geoCoordinate>
<geoCoordinate box="[1459,1536,473,499]" degrees="0.177" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.177">–0.177</geoCoordinate>
(0.152); CW = 
<geoCoordinate box="[678,740,513,540]" degrees="0.504" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.504">0.504</geoCoordinate>
<geoCoordinate box="[740,817,513,540]" degrees="0.574" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.574">–0.574</geoCoordinate>
(0.534); CWb = 
<geoCoordinate box="[1020,1082,513,539]" degrees="0.449" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.449">0.449</geoCoordinate>
<geoCoordinate box="[1082,1159,513,539]" degrees="0.494" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.494">–0.494</geoCoordinate>
(0.470); PoOC = 
<geoCoordinate box="[1372,1434,513,539]" degrees="0.190" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.19">0.190</geoCoordinate>
<geoCoordinate box="[1434,1511,513,539]" degrees="0.202" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.202">–0.202</geoCoordinate>
(0.196); CL = 
<geoCoordinate box="[665,727,553,579]" degrees="0.479" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.479">0.479</geoCoordinate>
<geoCoordinate box="[727,804,553,580]" degrees="0.516" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.516">–0.516</geoCoordinate>
(0.499); EL = 
<geoCoordinate box="[977,1039,553,579]" degrees="0.209" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.209">0.209</geoCoordinate>
<geoCoordinate box="[1039,1116,553,579]" degrees="0.228" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.228">–0.228</geoCoordinate>
(0.217); EW = 
<geoCoordinate box="[1302,1364,553,579]" degrees="0.172" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.172">0.172</geoCoordinate>
<geoCoordinate box="[1364,1441,553,579]" degrees="0.182" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.182">–0.182</geoCoordinate>
(0.177); MD = 
<geoCoordinate box="[579,641,593,620]" degrees="0.056" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.056">0.056</geoCoordinate>
<geoCoordinate box="[641,718,593,619]" degrees="0.069" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.069">–0.069</geoCoordinate>
(0.063); WL = 
<geoCoordinate box="[901,963,593,619]" degrees="0.698" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.698">0.698</geoCoordinate>
<geoCoordinate box="[964,1041,593,620]" degrees="0.765" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.765">–0.765</geoCoordinate>
(0.737); SPST = n/a; MPST = 
<geoCoordinate box="[1402,1464,593,619]" degrees="0.141" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.141">0.141</geoCoordinate>
<geoCoordinate box="[1464,1541,593,619]" degrees="0.172" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.172">–0.172</geoCoordinate>
(0.156); PEL = 
<geoCoordinate box="[678,740,632,659]" degrees="0.295" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.295">0.295</geoCoordinate>
<geoCoordinate box="[741,818,633,659]" degrees="0.349" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.349">–0.349</geoCoordinate>
(0.322); NOL = 
<geoCoordinate box="[1017,1079,633,659]" degrees="0.191" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.191">0.191</geoCoordinate>
<geoCoordinate box="[1080,1157,633,659]" degrees="0.217" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.217">–0.217</geoCoordinate>
(0.203); NOH = 
<geoCoordinate box="[1363,1425,633,659]" degrees="0.042" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.042">0.042</geoCoordinate>
<geoCoordinate box="[1425,1502,632,659]" degrees="0.058" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.058">–0.058</geoCoordinate>
(0.050); PEH = 
<geoCoordinate box="[685,747,673,699]" degrees="0.120" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.12">0.120</geoCoordinate>
<geoCoordinate box="[747,824,672,699]" degrees="0.150" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.15">–0.150</geoCoordinate>
(0.130); PPL = 
<geoCoordinate box="[1014,1076,672,699]" degrees="0.154" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.154">0.154</geoCoordinate>
<geoCoordinate box="[1076,1153,673,699]" degrees="0.193" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.193">–0.193</geoCoordinate>
(0.174); PPH = 
<geoCoordinate box="[1349,1411,672,699]" degrees="0.158" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.158">0.158</geoCoordinate>
<geoCoordinate box="[1411,1488,673,699]" degrees="0.198" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.198">–0.198</geoCoordinate>
(0.181); PW = 
<geoCoordinate box="[672,734,712,739]" degrees="0.445" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.445">0.445</geoCoordinate>
<geoCoordinate box="[735,812,712,739]" degrees="0.519" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.519">–0.519</geoCoordinate>
(0.467); SBPA = n/a; SPTI = n/a; PEW = 
<geoCoordinate box="[1305,1367,712,739]" degrees="0.115" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.115">0.115</geoCoordinate>
<geoCoordinate box="[1368,1445,712,738]" degrees="0.130" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.13">–0.130</geoCoordinate>
(0.122); PNW = 
<geoCoordinate box="[599,661,752,778]" degrees="0.123" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.123">0.123</geoCoordinate>
<geoCoordinate box="[661,738,752,779]" degrees="0.155" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.155">–0.155</geoCoordinate>
(0.137); PPW = 
<geoCoordinate box="[940,1002,752,778]" degrees="0.262" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.262">0.262</geoCoordinate>
<geoCoordinate box="[1002,1079,752,778]" degrees="0.339" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.339">–0.339</geoCoordinate>
(0.313); HFL = 
<geoCoordinate box="[1273,1335,752,779]" degrees="0.523" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.523">0.523</geoCoordinate>
<geoCoordinate box="[1335,1412,752,778]" degrees="0.649" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.649">–0.649</geoCoordinate>
(0.580); HFWmax = 
<geoCoordinate box="[648,710,792,818]" degrees="0.069" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.069">0.069</geoCoordinate>
<geoCoordinate box="[711,788,792,818]" degrees="0.084" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.084">–0.084</geoCoordinate>
(0.078); HFWmin = 
<geoCoordinate box="[1041,1103,792,818]" degrees="0.036" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.036">0.036</geoCoordinate>
<geoCoordinate box="[1104,1181,792,818]" degrees="0.039" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.039">–0.039</geoCoordinate>
(0.038); CS = 
<geoCoordinate box="[1354,1416,792,818]" degrees="0.693" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.693">0.693</geoCoordinate>
<geoCoordinate box="[1417,1494,792,819]" degrees="0.752" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.752">–0.752</geoCoordinate>
(0.719); ES = 
<geoCoordinate box="[660,722,832,859]" degrees="0.295" direction="north" orientation="latitude" pageId="271" pageNumber="272" precision="55" value="0.295">0.295</geoCoordinate>
<geoCoordinate box="[723,799,832,859]" degrees="0.315" direction="west" orientation="longitude" pageId="271" pageNumber="272" precision="55" value="-0.315">–0.315</geoCoordinate>
(0.305); SI = 67–70 (69); OI = 42–43 (42); CI = 90–96 (94); WLI = 152–163 (157); SBI = n/a; PSI= n/a; PWI = 228–277 (256); PLI = 153–213 (186); NI = 341–481 (412); PNWI = 99–119 (112); NLI = 59–68 (63); FI = 177–221 (204).
</paragraph>
<paragraph blockId="271.[498,1542,234,1895]" pageId="271" pageNumber="272">
<emphasis bold="true" box="[530,743,951,978]" pageId="271" pageNumber="272">Male description:</emphasis>
In full-face view, head subovate, slightly longer than broad (CI 90-96). Mandibles weakly striate, shining, and armed with five teeth: the apical-most well developed, followed by a four roughly equally developed smaller teeth, and the basal-most two separated from the rest by a brief diastema. Anterior clypeal margin entire and evenly convex. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 67–70). Antennae 13-segmented; antennal club of four segments, with the apical-most segment as long as the preceding two in combination. Frontal carinae very short, extending past the antennal toruli by one and a half times the maximum width of the antennal scape. Compound eyes strongly protruding past the lateral margins of the head capsule. Lateral margin of head convex, margin between the anterior margin of the compound eye and the mandibular insertions straight. Posterior head margin very slightly convex, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="271.[498,1542,234,1895]" lastBlockId="272.[498,1542,234,1934]" lastPageId="272" lastPageNumber="273" pageId="271" pageNumber="272">In profile view, compound eyes ovular and large (OI 42–43), with 19 ommatidia in the longest row. Mesoscutum bulging anteriorly, covering the dorsal surface of the pronotum, flat dorsally, and rounded posteriorly. Mesoscutellum depressed slightly below the level of the mesoscutum, overhanging the very small metanotum. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about three spiracle diameters. Propodeal spines absent, indicated by weak carinae on the dorsal and declivitous faces of the propodeum, which converge into a rounded angle. Propodeal lobes rounded and weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole long (PLI 153–213), without tubercles anterodorsally where it articulates with the mesosoma. Subpetiolar process absent. Petiolar peduncle short: comprising about one third of the total length of the petiole. Petiolar node low and rounded, the convergence of the anterior and dorsal faces marked by a rounded angle. Postpetiole evenly rounded anterodorsally, flattened dorsally, and with a lobed, concave ventral surface.</paragraph>
<paragraph blockId="272.[498,1542,234,1934]" pageId="272" pageNumber="273">In dorsal view, mesoscutum covering pronotum anteriorly, humeri barely visible laterally as slivers of rounded sclerites. Petiolar peduncle with spiracles slightly protruding past the lateral margins, the peduncle broadened where they arise. Petiolar node slightly wider than the peduncle; petiole narrowed posterior to the node, before widening again to the caudal cylinder, which is slightly narrower than the node. Postpetiole very broad (PWI 228–277) and campaniform, articulating with most of the anterior margin of the gaster, leaving small, rounded margins on each side exposed. Anterior margin of postpetiole weakly convex, with the anterior corners evenly rounding into the lateral margins, which evenly diverge to the angulate posterior corners; posterior margin of postpetiole broadly concave. Metafemur not incrassate (FI 177–221).</paragraph>
<paragraph blockId="272.[498,1542,234,1934]" pageId="272" pageNumber="273">Sculpture: median clypeal carina present, flanked by additional weak carinae over weakly areolate ground sculpture. Antennal scapes weakly shining through shallow areolate sculpture. Head uniformly densely areolate. Lateral surfaces of pronotum and propodeum moderately shining through weak areolate sculpture. Anepisternum and katepisternum smooth and shining with weak areolate sculpture on their posterior third. Dorsally, mesoscutum weakly areolate between the Mayrian furrows, with a small patch of smooth sculpture anteromedially; otherwise smooth and shining. Mesoscutellum weakly areolate. Femora smooth and shining, with traces weak areolate sculpture along the entire length. Petiole covered with shallow areolate sculpture, with the dorsal surface of the node smooth and shining. Dorsal surface of postpetiole shining, with traces of shallow areolate sculpture, especially on the posterior third. Gaster smooth and shining, with weak spectral iridescence on the first gastral tergite.</paragraph>
<paragraph blockId="272.[498,1542,234,1934]" pageId="272" pageNumber="273">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of head, mesosoma, waist segments and gaster with short, erect, blunt-tipped setae, the longest of which are roughly a quarter of the width of the compound eye. Sparse, adpressed pubescence present on the entire body, but only apparent on the gaster due to the dense ground sculpture on the rest of the body.</paragraph>
<paragraph blockId="272.[498,1542,234,1934]" pageId="272" pageNumber="273">Color: head, mesosoma, waist segments, and gaster are uniformly dark brown. Mandibles and extremities are pale yellow.</paragraph>
</subSubSection>
<subSubSection pageId="272" pageNumber="273" type="etymology">
<paragraph blockId="272.[498,1542,234,1934]" pageId="272" pageNumber="273">
<emphasis bold="true" box="[530,668,1470,1496]" pageId="272" pageNumber="273">Etymology:</emphasis>
Morphological, from the Latin ‘albus’ (= white) + ‘spinus’ (= spined), in reference to the lightly colored integument of the propodeal spines.
</paragraph>
</subSubSection>
<subSubSection lastPageId="273" lastPageNumber="274" pageId="272" pageNumber="273" type="discussion">
<paragraph blockId="272.[498,1542,234,1934]" lastBlockId="273.[498,1542,234,1497]" lastPageId="273" lastPageNumber="274" pageId="272" pageNumber="273">
<emphasis bold="true" box="[530,670,1549,1575]" pageId="272" pageNumber="273">Comments:</emphasis>
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[679,961,1549,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="272" pageNumber="273" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[679,961,1549,1575]" italics="true" pageId="272" pageNumber="273">Temnothorax albispinus</emphasis>
</taxonomicName>
is known mostly from collections made on xerophytic islands surrounding 
<collectingCountry box="[867,1007,1589,1616]" name="Puerto Rico" pageId="272" pageNumber="273">Puerto Rico</collectingCountry>
; known only from a couple of localities on the main island: the vicinity of 
<collectingRegion box="[824,909,1629,1655]" country="Puerto Rico" name="Coamo" pageId="272" pageNumber="273">Coamo</collectingRegion>
Springs, and a light 
<taxonomicName box="[1155,1278,1629,1655]" form="from" pageId="272" pageNumber="273" rank="form">form from</taxonomicName>
<collectingRegion box="[1287,1449,1629,1655]" country="Puerto Rico" name="Guanica" pageId="272" pageNumber="273">Guánica State</collectingRegion>
Forest. Only one record of their nesting habits has appeared in the literature: 
<bibRefCitation author="Wheeler WM" box="[1329,1506,1669,1695]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="272" pageNumber="273" pagination="117 - 158" part="24" refId="ref273343" refString="Wheeler WM. 1908. The ants of Porto Rico and the Virgin Islands. Bulletin of the American Museum of Natural History 24: 117 - 158." title="The ants of Porto Rico and the Virgin Islands" type="journal article" year="1908">
<emphasis box="[1329,1506,1669,1695]" italics="true" pageId="272" pageNumber="273">Wheeler (1908)</emphasis>
</bibRefCitation>
, when describing the 
<typeStatus box="[737,787,1709,1735]" pageId="272" pageNumber="273">type</typeStatus>
series, noted: “a single colony which was found nesting in a small cavity in the ground in the shade of a thicket, where some of the workers were moving about slowly over the dead leaves and twigs.” Additional collections of the light 
<taxonomicName box="[1423,1542,1788,1814]" form="have" pageId="272" pageNumber="273" rank="form">form have</taxonomicName>
been from either from soil beneath a stone, from rotten sticks in the leaf litter, or from Winkler extraction of leaf litter. An unpublished study by Snelling &amp; Torres notes that it is attracted to tuna fish baits placed on the soil but has never been observed at baits placed on vegetation, even in localities where it is common. Apparently heat tolerant, workers have been observed foraging at baits during the midday hours, when soil surface temperatures probably exceed 45 
<emphasis box="[733,743,309,327]" italics="true" pageId="273" pageNumber="274">
<superScript attach="right" box="[733,743,309,327]" fontSize="8" pageId="273" pageNumber="274">°</superScript>
</emphasis>
C (R. R. Snelling &amp; J.A. Torres, 2001, unpublished data). Males have been collected at light traps in 
<collectingRegion box="[926,1088,354,380]" country="Puerto Rico" name="Guanica" pageId="273" pageNumber="274">Guánica State</collectingRegion>
Forest, but queens have only been collected from nests or from sifting leaf litter. Based on the presence of multiple dealate gynes in nest series, 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[740,889,433,459]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[740,889,433,459]" italics="true" pageId="273" pageNumber="274">T. albispinus</emphasis>
</taxonomicName>
may be polygynous, but only dissections of the gynes can rule out functional monogyny. This species, like other members of the 
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1387,1500,473,499]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[1387,1500,473,499]" italics="true" pageId="273" pageNumber="274">pulchellus</emphasis>
</taxonomicName>
group, is probably ground or leaf litter nesting. Populations from Mona Island, which correspond to subspecies 
<emphasis box="[800,885,553,579]" italics="true" pageId="273" pageNumber="274">pallipes</emphasis>
Wheeler, appear to be highly variable in the amount of infuscation on their appendages, ranging from the entirety of the legs being light yellow to the femora and antennal scapes being medium brown. However, another difference between 
<taxonomicName box="[600,837,673,699]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="subSpecies" species="albispinus" subSpecies="pallipes">
<emphasis box="[600,837,673,699]" italics="true" pageId="273" pageNumber="274">T. albispinus pallipes</emphasis>
</taxonomicName>
and 
<emphasis box="[894,1072,673,699]" italics="true" pageId="273" pageNumber="274">
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[894,1040,673,699]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="albispinus">T. albispinus</taxonomicName>
<taxonomicNameLabel box="[1046,1078,673,699]" pageId="273" pageNumber="274" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
noted by 
<bibRefCitation author="Wheeler WM" box="[1192,1366,673,699]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="273" pageNumber="274" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
<emphasis box="[1192,1366,673,699]" italics="true" pageId="273" pageNumber="274">Wheeler (1931)</emphasis>
</bibRefCitation>
, the absence of sculpture on the dorsal surface of the petiolar node, appears to occur only in the 
<collectingCountry box="[1459,1499,712,738]" name="Iceland" pageId="273" pageNumber="274">Isla</collectingCountry>
de Mona populations. The amount of sculpture on the first gastral tergite appears to vary throughout the range of 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1908" box="[790,939,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="albispinus">
<emphasis box="[790,939,792,818]" italics="true" pageId="273" pageNumber="274">T. albispinus</emphasis>
</taxonomicName>
as well: at the extreme end, the anterior 2/3 of the tergite is weakly sculptured in the Coamo Springs worker. Nest series of workers indicate that the extent of femoral incrassation is at least somewhat dependent on worker size. The light form of this species has been determined as 
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1134,1229,912,937]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1134,1229,912,937]" italics="true" pageId="273" pageNumber="274">T. torrei</emphasis>
</taxonomicName>
in 
<bibRefCitation author="Torres JA &amp; Snelling RR" box="[1270,1542,912,938]" journalOrPublisher="Biodiversity &amp; Conservation" pageId="273" pageNumber="274" pagination="1103 - 1121" part="6" refId="ref272973" refString="Torres JA, Snelling RR. 1997. Biogeography of Puerto Rican ants: a non-equilibrium case? Biodiversity &amp; Conservation 6 (8): 1103 - 1121 DOI 10.1023 / A: 1018332117719." title="Biogeography of Puerto Rican ants: a non-equilibrium case?" type="journal article" year="1997">
<emphasis box="[1270,1542,912,938]" italics="true" pageId="273" pageNumber="274">Torres &amp; Snelling, 1997</emphasis>
</bibRefCitation>
and in the unpublished manuscript by Snelling &amp; Torres, which is a review of the ants of 
<collectingCountry box="[528,664,991,1018]" name="Puerto Rico" pageId="273" pageNumber="274">Puerto Rico</collectingCountry>
(
<figureCitation box="[680,771,991,1017]" captionStart="Figure 136" captionStartId="265.[524,589,1001,1023]" captionTargetBox="[498,1541,234,988]" captionTargetId="figure-196@265.[498,1534,234,973]" captionTargetPageId="265" captionText="Figure 136 Light form of Temnothorax albispinus. (A–C) Worker (LACMENT323245). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Gyne (LACMENT323245). (D) Full-face view. (E) Profile view. (F) Dorsal view.(G–J) Male (LACMENT323246). (G) Full-face view.(H) Profile view. (I) Wings. (J) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-136" figureDoi="http://doi.org/10.5281/zenodo.5102398" httpUri="https://zenodo.org/record/5102398/files/figure.png" pageId="273" pageNumber="274">Fig. 136</figureCitation>
). It is easily separated from 
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1093,1186,992,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1093,1186,992,1017]" italics="true" pageId="273" pageNumber="274">T. torrei</emphasis>
</taxonomicName>
by the larger size and presence of four erect setae on the dorsum of the petiole. Recognition of this species also explains what was interpreted as a disjunct distribution of 
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1087,1183,1072,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1087,1183,1072,1097]" italics="true" pageId="273" pageNumber="274">T. torrei</emphasis>
</taxonomicName>
in 
<bibRefCitation author="Torres JA &amp; Snelling RR" box="[1224,1498,1071,1098]" journalOrPublisher="Biodiversity &amp; Conservation" pageId="273" pageNumber="274" pagination="1103 - 1121" part="6" refId="ref272973" refString="Torres JA, Snelling RR. 1997. Biogeography of Puerto Rican ants: a non-equilibrium case? Biodiversity &amp; Conservation 6 (8): 1103 - 1121 DOI 10.1023 / A: 1018332117719." title="Biogeography of Puerto Rican ants: a non-equilibrium case?" type="journal article" year="1997">
<emphasis box="[1224,1498,1071,1098]" italics="true" pageId="273" pageNumber="274">Torres &amp; Snelling, 1997</emphasis>
</bibRefCitation>
. 
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[498,727,1111,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[498,727,1111,1137]" italics="true" pageId="273" pageNumber="274">Temnothorax torrei</emphasis>
</taxonomicName>
, like many other 
<taxonomicName box="[937,1186,1111,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[937,1094,1111,1137]" italics="true" pageId="273" pageNumber="274">Temnothorax</emphasis>
species
</taxonomicName>
, has a Bahamian-Cuban distribution; it has never been collected from the island of Hispaniola. This species has long been suspected to have close relationships to other members of the 
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1297,1372,1191,1217]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[1297,1372,1191,1217]" italics="true" pageId="273" pageNumber="274">salvini</emphasis>
</taxonomicName>
clade: specifically, 
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1905" box="[638,802,1230,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="androsanus">
<emphasis box="[638,802,1230,1256]" italics="true" pageId="273" pageNumber="274">T. androsanus</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[817,956,1231,1257]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[817,956,1231,1257]" italics="true" pageId="273" pageNumber="274">T. pastinifer</emphasis>
</taxonomicName>
, and 
<taxonomicName authority="(Wheeler, 1908)" baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1021,1360,1230,1257]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1021,1168,1230,1256]" italics="true" pageId="273" pageNumber="274">T. subditivus</emphasis>
(
<bibRefCitation author="Wheeler WM" box="[1186,1350,1230,1257]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="273" pageNumber="274" pagination="117 - 158" part="24" refId="ref273343" refString="Wheeler WM. 1908. The ants of Porto Rico and the Virgin Islands. Bulletin of the American Museum of Natural History 24: 117 - 158." title="The ants of Porto Rico and the Virgin Islands" type="journal article" year="1908">
<emphasis box="[1186,1350,1230,1257]" italics="true" pageId="273" pageNumber="274">Wheeler, 1908</emphasis>
</bibRefCitation>
)
</taxonomicName>
. 
<bibRefCitation author="Baroni Urbani C." journalOrPublisher="Entomologica Basiliensia" pageId="273" pageNumber="274" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis italics="true" pageId="273" pageNumber="274">Baroni Urbani (1978)</emphasis>
</bibRefCitation>
lumped them into the heterogeneous 
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1025,1138,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[1025,1138,1270,1296]" italics="true" pageId="273" pageNumber="274">pulchellus</emphasis>
</taxonomicName>
group. In the first formal morphology-based phylogenetic work including this species, 
<bibRefCitation author="Fontenla Rizo JL" box="[1216,1461,1310,1336]" journalOrPublisher="Avicennia" pageId="273" pageNumber="274" pagination="35 - 44" part="13" refId="ref270970" refString="Fontenla Rizo JL. 2000. Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax). Avicennia 13: 35 - 44." title="Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax)" type="journal article" year="2000">
<emphasis box="[1216,1461,1310,1336]" italics="true" pageId="273" pageNumber="274">Fontenla Rizo (2000)</emphasis>
</bibRefCitation>
, while not including all of the taxa in Baroni Urbani’ s 
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1063,1176,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[1063,1176,1350,1376]" italics="true" pageId="273" pageNumber="274">pulchellus</emphasis>
</taxonomicName>
group, corroborated these relationships to some degree, with the caveat that 
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1090,1218,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[1090,1218,1390,1416]" italics="true" pageId="273" pageNumber="274">T. terricola</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Aguayo" baseAuthorityYear="1931" box="[1280,1376,1391,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="273" pageNumber="274" phylum="Arthropoda" rank="species" species="torrei">
<emphasis box="[1280,1376,1391,1416]" italics="true" pageId="273" pageNumber="274">T. torrei</emphasis>
</taxonomicName>
fell outside of this group. Combined molecular and morphological analyses (M. 
<bibRefCitation author="Prebus M." box="[1308,1461,1430,1456]" journalOrPublisher="Systematic Entomology" pageId="273" pageNumber="274" pagination="307 - 326" part="46" refId="ref272244" refString="Prebus M. 2021. Phylogenomic species delimitation in the ants of the Temnothorax salvini group (Hymenoptera: Formicidae): an integrative approach. Systematic Entomology 46 (2): 307 - 326 DOI 10.1111 / syen. 12463." title="Phylogenomic species delimitation in the ants of the Temnothorax salvini group (Hymenoptera: Formicidae): an integrative approach" type="journal article" year="2021">Prebus, 2021</bibRefCitation>
, unpublished data) mostly agree with 
<bibRefCitation author="Fontenla Rizo JL" box="[937,1181,1470,1496]" journalOrPublisher="Avicennia" pageId="273" pageNumber="274" pagination="35 - 44" part="13" refId="ref270970" refString="Fontenla Rizo JL. 2000. Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax). Avicennia 13: 35 - 44." title="Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax)" type="journal article" year="2000">
<emphasis box="[937,1181,1470,1496]" italics="true" pageId="273" pageNumber="274">Fontenla Rizo (2000)</emphasis>
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>