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<document id="991C432AFF58C8E6AA66D43029989383" ID-DOI="10.1093/zoolinnean/zlad118" ID-ISSN="0024-4082" ID-Zenodo-Dep="11241048" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716363684985" checkinUser="plazi" docAuthor="Sánchez-García, Alba, Sendra, Alberto, Davis, Steven R. &amp; Grimaldi, David A." docDate="2024" docId="039787F42D1AFFB3EBE3F90DFBBFF8D6" docLanguage="en" docName="zlad118.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad118" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Litocampa Silvestri 1933" docType="treatment" docVersion="2" lastPageNumber="149" masterDocId="FFAEFF8C2D12FFBEE826FFA0FFF8F917" masterDocTitle="‘ Dawn’ hexapods in Cenozoic ambers (Diplura: Campodeoidea)" masterLastPageNumber="158" masterPageNumber="136" pageNumber="144" updateTime="1716364893493" updateUser="ExternalLinkService">
<mods:mods id="67063CB821A2867100B736563235C83E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="314D4B8888E73025B38ADB0F4DF0221C">
<mods:title id="F5D36BFC7D05ACB862DFBE1B02A2A7EE">‘ Dawn’ hexapods in Cenozoic ambers (Diplura: Campodeoidea)</mods:title>
</mods:titleInfo>
<mods:name id="FA98F52C51230907682A544370B4B5F6" type="personal">
<mods:role id="259B1362EAA4DCD2DFCE2AEADC443BBB">
<mods:roleTerm id="331C6039CD38266AF81233C881D040FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC407507291D6FAB8C158192C8F4F94D">Sánchez-García, Alba</mods:namePart>
<mods:affiliation id="933586634DB72AFE9899508AE1184E16">Instituto Geológico y Minero de España (IGME), CSIC, València, Spain</mods:affiliation>
<mods:nameIdentifier id="353C51F430019B5E47A3D449B5382F40" type="email">alba.sanchez@csic.es</mods:nameIdentifier>
</mods:name>
<mods:name id="F652E9F06995467D273B6A3AED052AA1" type="personal">
<mods:role id="D54F33E23238E4919EDC7E7F6313C2EA">
<mods:roleTerm id="1AB7429C5B35FD6DDE5DA75C10D8BEC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF9C0D7B40F5E6DB5055394FF2A93169">Sendra, Alberto</mods:namePart>
<mods:affiliation id="1D3B695EFC8EA46A114FDC6ACF014D8F">Coleccions Torres-Sala i Siro de Fez, Servei de Patrimoni Històric, Ajuntament de València, València, Spain &amp; Departament de Didàctica de les Ciències Experimentals i Socials, Facultat de Magisteri, Universitat de València, València, Spain</mods:affiliation>
</mods:name>
<mods:name id="F14291ECDC4A3D280C5079A11DB62CF2" type="personal">
<mods:role id="95EE04144F5B77EB247E2D2284EA959D">
<mods:roleTerm id="485BE4E3FE3D932A42EBC8E282E06F67">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7AC1B212BE6B799136DAD8F5E5060BDE">Davis, Steven R.</mods:namePart>
<mods:affiliation id="7B7E83F26D8428C215FB09679DD29B2E">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA</mods:affiliation>
</mods:name>
<mods:name id="2D18BA9484F97BF93170439B644328D8" type="personal">
<mods:role id="26AB8A5364CC507AC3A4D77549334350">
<mods:roleTerm id="57567E19E922061132398ACC17C101CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32B1B011DB91B1F81EA077EAE77273D6">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="7F562B97366BF6F2871B3A1BBFECA68D">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1C01A595FD8F597AE93C058B8F5D5371">text</mods:typeOfResource>
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<mods:title id="F30A6B6B9D19E5010FE4F6D0F02314AC">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="87B98F78B21D228BC1360F382BD7C2A3">
<mods:date id="439A4DA4FC519714692A4D5BB3B42676">2024</mods:date>
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<mods:number id="83498F5155E9B50D18B96902866EC213">2023-09-30</mods:number>
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<mods:number id="F20C14754EF6F8A6C659C97587C1D97A">201</mods:number>
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<mods:number id="8A4518A38ABE8DB21817445954BAA8A4">1</mods:number>
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<mods:classification id="18068EB38DD0286E679B2D619E5840CB">journal article</mods:classification>
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<mods:identifier id="EA2E0F262CCF4E4DE9883D62E098433B" type="ISSN">0024-4082</mods:identifier>
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<treatment id="039787F42D1AFFB3EBE3F90DFBBFF8D6" ID-DOI="http://doi.org/10.5281/zenodo.11241054" ID-Zenodo-Dep="11241054" LSID="urn:lsid:plazi:treatment:039787F42D1AFFB3EBE3F90DFBBFF8D6" httpUri="http://treatment.plazi.org/id/039787F42D1AFFB3EBE3F90DFBBFF8D6" lastPageId="13" lastPageNumber="149" pageId="8" pageNumber="144">
<subSubSection id="C32465692D1AFFB6EBE3F90DFAE0FFD0" box="[965,1304,1709,1735]" pageId="8" pageNumber="144" type="nomenclature">
<paragraph id="8B8136E22D1AFFB6EBE3F90DFAE0FFD0" blockId="8.[965,1304,1709,1735]" box="[965,1304,1709,1735]" pageId="8" pageNumber="144">
<heading id="D0C9818E2D1AFFB6EBE3F90DFAE0FFD0" bold="true" box="[965,1304,1709,1735]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="144" reason="3">
<emphasis id="B94AEAF02D1AFFB6EBE3F90DFAE0FFD0" bold="true" box="[965,1304,1709,1735]" pageId="8" pageNumber="144">
Genus
<taxonomicName id="4C3E4D612D1AFFB6EC29F90DFAE0FFD0" ID-CoL="8NQZP" authority="Silvestri, 1933" authorityName="Silvestri" authorityYear="1933" box="[1039,1304,1709,1735]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="8" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAF02D1AFFB6EC29F90DFB85FFD0" bold="true" box="[1039,1149,1709,1735]" italics="true" pageId="8" pageNumber="144">Litocampa</emphasis>
Silvestri, 1933
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32465692D1AFFB6EB0CF97FFB6DFE57" pageId="8" pageNumber="144" type="materials_examined">
<paragraph id="8B8136E22D1AFFB6EB0CF97FFAE7FFE0" blockId="8.[810,1311,1759,1783]" box="[810,1311,1759,1783]" pageId="8" pageNumber="144">
<emphasis id="B94AEAF02D1AFFB6EB0CF97FFC5EFFE1" box="[810,934,1759,1782]" italics="true" pageId="8" pageNumber="144">Type species:</emphasis>
<taxonomicName id="4C3E4D612D1AFFB6EB96F97FFAE3FFE0" authority="Silvestri, 1933" authorityName="Silvestri" authorityYear="1933" box="[944,1307,1759,1783]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="8" pageNumber="144" phylum="Arthropoda" rank="species" species="neotropica">
<emphasis id="B94AEAF02D1AFFB6EB96F97FFB7BFFE1" box="[944,1155,1759,1782]" italics="true" pageId="8" pageNumber="144">Litocampa neotropica</emphasis>
Silvestri, 1933
</taxonomicName>
.
</paragraph>
<paragraph id="8B8136E22D1AFFB6EB11F8A7FB6DFE57" blockId="8.[823,1445,1798,1856]" pageId="8" pageNumber="144">
<emphasis id="B94AEAF02D1AFFB6EB11F8A7FB6DFE57" bold="true" pageId="8" pageNumber="144">
<taxonomicName id="4C3E4D612D1AFFB6EB11F8A7FA5DFE36" authority="Sanchez-Garcia, Sendra and Grimaldi" authorityName="Sanchez-Garcia, Sendra and Grimaldi" authorityYear="2024" box="[823,1445,1798,1825]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="8" pageNumber="144" phylum="Arthropoda" rank="species" species="eobaltica" status="sp. nov.">
<emphasis id="B94AEAF02D1AFFB6EB11F8A7FBF1FE36" bold="true" box="[823,1033,1798,1825]" italics="true" pageId="8" pageNumber="144">Litocampa eobaltica</emphasis>
Sánchez-García, Sendra and Grimaldi
</taxonomicName>
<taxonomicNameLabel id="A279578B2D1AFFB6EC61F886FB6DFE57" box="[1095,1173,1830,1856]" pageId="8" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32465692D1AFFB3EC16F8F5FBBFF8D6" lastPageId="13" lastPageNumber="149" pageId="8" pageNumber="144" type="description">
<paragraph id="8B8136E22D1AFFB6EC16F8F5FB55FE78" blockId="8.[1072,1197,1877,1903]" box="[1072,1197,1877,1903]" pageId="8" pageNumber="144">
<emphasis id="B94AEAF02D1AFFB6EC16F8F5FB55FE78" bold="true" box="[1072,1197,1877,1903]" pageId="8" pageNumber="144">
(
<figureCitation id="13052A672D1AFFB6EC1CF8F5FB5AFE78" box="[1082,1186,1877,1903]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStart-3="Figure 10" captionStartId-0="9.[129,194,1954,1978]" captionStartId-1="10.[113,178,1836,1860]" captionStartId-2="11.[129,194,1794,1818]" captionStartId-3="12.[113,178,1836,1860]" captionTargetBox-0="[148,1454,984,1923]" captionTargetBox-1="[169,1403,147,1805]" captionTargetBox-2="[132,1470,148,1763]" captionTargetBox-3="[141,1431,147,1805]" captionTargetId-0="figure-427@9.[145,1457,981,1926]" captionTargetId-1="figure-8@10.[166,1406,144,1808]" captionTargetId-2="figure-9@11.[129,1473,144,1766]" captionTargetId-3="figure-7@12.[138,1434,144,1808]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="Figure 7. Dorsal photomicrograph of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber." captionText-1="Figure 8. Camera lucida drawings of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of thorax, showing the distribution of macrosetae.B, detail of basal antennomeres.C, dorsal habitus.D, detail of stylus.E, detail of pretarsus and tibia of leg II; asterisk indicates the ventral macroseta on the tibia. F, detail of pretarsus and tibia of leg III; asterisk indicates the ventral macroseta on the tibia. Abbreviation:c, calcar." captionText-2="Figure 9. Photomicrographs of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of left leg II; asterisk indicates the ventral macroseta on the tibia. B, detail of pretarsus in A. C, detail of left leg III; asterisk indicates the ventral macroseta on the tibia. D, detail of pretarsus in C. E, distribution of macrosetae on pronotum and mesonotum. F, distribution of macrosetae on metanotum. Abbreviation: c, calcar." captionText-3="Figure 10. Photomicrograph of paratype of Litocampa eobaltica, AMNH Ba-JVe277, in Eocene Baltic amber. A, dorsal habitus; arrows point to the spider web.B, thorax in dorsal view, showing the distribution of macrosetae; the box labelled C corresponds to panel C. C, detail of microdenticles of the cuticle.D, head and thorax in ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.11241064" figureDoi-1="http://doi.org/10.5281/zenodo.11241066" figureDoi-2="http://doi.org/10.5281/zenodo.11241068" figureDoi-3="http://doi.org/10.5281/zenodo.11241070" httpUri-0="https://zenodo.org/record/11241064/files/figure.png" httpUri-1="https://zenodo.org/record/11241066/files/figure.png" httpUri-2="https://zenodo.org/record/11241068/files/figure.png" httpUri-3="https://zenodo.org/record/11241070/files/figure.png" pageId="8" pageNumber="144">Figs 7–10</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B8136E22D1AFFB6EB0CF836FC37FEDA" blockId="8.[810,1459,1942,1997]" pageId="8" pageNumber="144">
<emphasis id="B94AEAF02D1AFFB6EB0CF836FC91FEBA" box="[810,873,1942,1965]" italics="true" pageId="8" pageNumber="144">LSID:</emphasis>
<uri id="FFAF3AE02D1AFFB6EB5AF836FC37FEDA" pageId="8" pageNumber="144">
urn:lsid:zoobank.org:act:
<uuid id="FF980C372D1AFFB6EC5EF836FC37FEDA" pageId="8" pageNumber="144">687BF785-5BE6-488A-B531- 147DF9F51906</uuid>
</uri>
</paragraph>
<paragraph id="8B8136E22D1BFFB7E8A7FF30FD6DF9D0" blockId="9.[129,777,144,199]" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7E8A7FF30FF08F9BF" box="[129,240,144,168]" italics="true" pageId="9" pageNumber="145">Etymology:</emphasis>
The specific epithet is in reference to the geological epoch (Eocene) and amber of origin (Baltic amber).
</paragraph>
<paragraph id="8B8136E22D1BFFB7E8A7FF49FD8FF8AA" blockId="9.[128,778,233,695]" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7E8A7FF49FF24F816" box="[129,220,233,257]" italics="true" pageId="9" pageNumber="145">Material:</emphasis>
<materialsCitation id="3B563CBF2D1BFFB7E8C5FF49FD8FF8AA" collectionCode="AMNH" pageId="9" pageNumber="145" specimenCount="1" typeStatus="holotype">
<typeStatus id="548588402D1BFFB7E8C5FF49FEBEF816" box="[227,326,233,257]" pageId="9" pageNumber="145" type="holotype">Holotype</typeStatus>
<emphasis id="B94AEAF02D1BFFB7E96AFF49FE06F816" box="[332,510,233,257]" italics="true" pageId="9" pageNumber="145">
<collectionCode id="ED2FAE272D1BFFB7E96AFF49FE63F817" box="[332,411,233,256]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="145">AMNH</collectionCode>
Ba-JVe66
</emphasis>
(
<figureCitation id="13052A672D1BFFB7EA28FF49FD9EF816" box="[526,614,233,257]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStartId-0="9.[129,194,1954,1978]" captionStartId-1="10.[113,178,1836,1860]" captionStartId-2="11.[129,194,1794,1818]" captionTargetBox-0="[148,1454,984,1923]" captionTargetBox-1="[169,1403,147,1805]" captionTargetBox-2="[132,1470,148,1763]" captionTargetId-0="figure-427@9.[145,1457,981,1926]" captionTargetId-1="figure-8@10.[166,1406,144,1808]" captionTargetId-2="figure-9@11.[129,1473,144,1766]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 7. Dorsal photomicrograph of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber." captionText-1="Figure 8. Camera lucida drawings of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of thorax, showing the distribution of macrosetae.B, detail of basal antennomeres.C, dorsal habitus.D, detail of stylus.E, detail of pretarsus and tibia of leg II; asterisk indicates the ventral macroseta on the tibia. F, detail of pretarsus and tibia of leg III; asterisk indicates the ventral macroseta on the tibia. Abbreviation:c, calcar." captionText-2="Figure 9. Photomicrographs of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of left leg II; asterisk indicates the ventral macroseta on the tibia. B, detail of pretarsus in A. C, detail of left leg III; asterisk indicates the ventral macroseta on the tibia. D, detail of pretarsus in C. E, distribution of macrosetae on pronotum and mesonotum. F, distribution of macrosetae on metanotum. Abbreviation: c, calcar." figureDoi-0="http://doi.org/10.5281/zenodo.11241064" figureDoi-1="http://doi.org/10.5281/zenodo.11241066" figureDoi-2="http://doi.org/10.5281/zenodo.11241068" httpUri-0="https://zenodo.org/record/11241064/files/figure.png" httpUri-1="https://zenodo.org/record/11241066/files/figure.png" httpUri-2="https://zenodo.org/record/11241068/files/figure.png" pageId="9" pageNumber="145">Figs 7–9</figureCitation>
), sex unknown, adult; held in the
<collectionCode id="ED2FAE272D1BFFB7E916FEA8FE7CF837" box="[304,388,264,288]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="145">AMNH</collectionCode>
; specimen virtually complete, with the left antenna broken; a reddish halo is around most of the specimen; observable ventrally (with high distortion) and dorsally; the piece is very transparent, orange in colour, embedded in a block of EpoTek resin trimmed to
<quantity id="4CC69B072D1BFFB7E9D7FE25FDC1F88A" box="[497,569,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="145" unit="cm" value="1.9">1.9 cm</quantity>
×
<quantity id="4CC69B072D1BFFB7EA73FE25FD66F88A" box="[597,670,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="145" unit="cm" value="1.4">1.4 cm</quantity>
×
<quantity id="4CC69B072D1BFFB7EA9CFE25FCFDF88A" box="[698,773,389,413]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="145" unit="cm" value="0.7">0.7 cm</quantity>
; syninclusions include an ant and plant trichomes.
</materialsCitation>
</paragraph>
<paragraph id="8B8136E22D1BFFB7E8BAFE64FCF8FBA0" blockId="9.[128,778,233,695]" pageId="9" pageNumber="145">
<materialsCitation id="3B563CBF2D1BFFB7E8BAFE64FF0AF8EC" collectionCode="AMNH" pageId="9" pageNumber="145" specimenCount="1" typeStatus="paratype">
<typeStatus id="548588402D1BFFB7E8BAFE64FF0EF8CB" box="[156,246,452,476]" pageId="9" pageNumber="145" type="paratype">Paratype</typeStatus>
<collectionCode id="ED2FAE272D1BFFB7E8D8FE64FEB7F8CB" box="[254,335,452,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="145">AMNH</collectionCode>
Ba-JVe277 (
<figureCitation id="13052A672D1BFFB7E9F0FE64FDE6F8CB" box="[470,542,452,476]" captionStart="Figure 10" captionStartId="12.[113,178,1836,1860]" captionTargetBox="[141,1431,147,1805]" captionTargetId="figure-7@12.[138,1434,144,1808]" captionTargetPageId="12" captionText="Figure 10. Photomicrograph of paratype of Litocampa eobaltica, AMNH Ba-JVe277, in Eocene Baltic amber. A, dorsal habitus; arrows point to the spider web.B, thorax in dorsal view, showing the distribution of macrosetae; the box labelled C corresponds to panel C. C, detail of microdenticles of the cuticle.D, head and thorax in ventral view." figureDoi="http://doi.org/10.5281/zenodo.11241070" httpUri="https://zenodo.org/record/11241070/files/figure.png" pageId="9" pageNumber="145">Fig. 10</figureCitation>
), sex unknown, adult; held in the
</materialsCitation>
AMNH; specimen virtually complete, with the left cercus broken; observable ventrally (with high distortion) and dorsally; the fossilized position of the specimen precludes several measurements and hinders examination of legs and ventral structures; the piece is very transparent, orange in colour, embedded in a block of EpoTek resin trimmed to
<quantity id="4CC69B072D1BFFB7EAA1FD20FD35FB80" box="[647,717,640,664]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="145" unit="cm" value="2.7">2.7 cm</quantity>
×
<quantity id="4CC69B072D1BFFB7EACCFD20FF58FBA0" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="145" unit="cm" value="1.6">1.6 cm</quantity>
×
<quantity id="4CC69B072D1BFFB7E89BFD3FFF0AFBA0" box="[189,242,671,695]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="145" unit="cm" value="1.0">1 cm</quantity>
; in spider webbing, syninclusions include one acari.
</paragraph>
<paragraph id="8B8136E22D1BFFB7E8A6FD79FE2EFBE7" blockId="9.[128,470,728,752]" box="[128,470,728,752]" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7E8A6FD79FF0CFBE7" box="[128,244,729,752]" italics="true" pageId="9" pageNumber="145">Occurrence:</emphasis>
Eocene, Baltic amber.
</paragraph>
<paragraph id="8B8136E22D1BFFB7E8A6FCB3FBFEF8B5" blockId="9.[128,778,786,904]" lastBlockId="9.[825,1474,144,418]" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7E8A6FCB3FF1EFA3D" box="[128,230,787,810]" italics="true" pageId="9" pageNumber="145">Diagnosis:</emphasis>
Cuticle with microdenticles. Frontal process not protruding. Antenna three-fifths length of body, with 25 antennomeres, moniliform. Distribution of thoracic macrosetae: 1 + 1
<emphasis id="B94AEAF02D1BFFB7E898FCD1FF25FA9F" box="[190,221,881,904]" italics="true" pageId="9" pageNumber="145">ma</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1BFFB7E901FCD0FEC1FA9F" box="[295,313,880,904]" italics="true" pageId="9" pageNumber="145">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1BFFB7E9A5FCD0FE6EFA9F" box="[387,406,880,904]" italics="true" pageId="9" pageNumber="145">lp</emphasis>
on pronotum and mesonotum, and 1 + 1
<emphasis id="B94AEAF02D1BFFB7EB5CFF31FC61F9BF" box="[890,921,145,168]" italics="true" pageId="9" pageNumber="145">ma</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1BFFB7EBCCFF30FC05F9BF" box="[1002,1021,144,168]" italics="true" pageId="9" pageNumber="145">lp</emphasis>
on metanotum;
<emphasis id="B94AEAF02D1BFFB7EC90FF30FB31F9BF" box="[1206,1225,144,168]" italics="true" pageId="9" pageNumber="145">lp</emphasis>
and marginal setae robust. Metathoracic leg reaching abdominal segment VI; femur without dorsal macrosetae and with one ventral macroseta; tibia with one ventral macroseta; claws subequal, elbowed, with small and smooth lateral crests; pretarsal lateral processes setiform and smooth. Distribution of abdominal macrosetae: 1 + 1
<emphasis id="B94AEAF02D1BFFB7ED8CFE8CFA44F853" box="[1450,1468,300,324]" italics="true" pageId="9" pageNumber="145">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1BFFB7EB5EFEECFC73F873" box="[888,907,332,356]" italics="true" pageId="9" pageNumber="145">lp</emphasis>
on VI–VII, and 3 + 3
<emphasis id="B94AEAF02D1BFFB7EC5FFEECFB74F873" box="[1145,1164,332,356]" italics="true" pageId="9" pageNumber="145">lp</emphasis>
on VIII. Cerci slightly longer than body, with five primary articles, with whorls of setae and macrosetae smooth.
</paragraph>
<paragraph id="8B8136E22D1BFFB7EB1FFE6EFAB3F8F2" blockId="9.[825,1355,461,485]" box="[825,1355,461,485]" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7EB1FFE6EFB8FF8F2" box="[825,1143,461,485]" italics="true" pageId="9" pageNumber="145">
Description (based on
<typeStatus id="548588402D1BFFB7EC34FE6DFB92F8F2" box="[1042,1130,461,485]" pageId="9" pageNumber="145" type="holotype">holotype</typeStatus>
):
</emphasis>
Sex unknown, adult.
</paragraph>
<paragraph id="8B8136E22D1BFFB7EB1FFDB0FB38FB71" blockId="9.[825,1474,528,615]" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7EB1FFDB0FC8AFB3F" box="[825,882,528,552]" italics="true" pageId="9" pageNumber="145">Body:</emphasis>
Length
<quantity id="4CC69B072D1BFFB7EBEEFDB0FBDEFB3F" box="[968,1062,528,552]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="9" pageNumber="145" unit="mm" value="2.35">2.35 mm</quantity>
. Cuticle with microdenticles perceptible under compound microscope (
<figureCitation id="13052A672D1BFFB7ECA5FD8FFB25FB50" box="[1155,1245,559,583]" captionStart="Figure 10" captionStartId="12.[113,178,1836,1860]" captionTargetBox="[141,1431,147,1805]" captionTargetId="figure-7@12.[138,1434,144,1808]" captionTargetPageId="12" captionText="Figure 10. Photomicrograph of paratype of Litocampa eobaltica, AMNH Ba-JVe277, in Eocene Baltic amber. A, dorsal habitus; arrows point to the spider web.B, thorax in dorsal view, showing the distribution of macrosetae; the box labelled C corresponds to panel C. C, detail of microdenticles of the cuticle.D, head and thorax in ventral view." figureDoi="http://doi.org/10.5281/zenodo.11241070" httpUri="https://zenodo.org/record/11241070/files/figure.png" pageId="9" pageNumber="145">Fig. 10C</figureCitation>
); body highly setose, with long, thin, smooth clothing setae.
</paragraph>
<paragraph id="8B8136E22D1BFFB3EB1FFD31FEF5F9D0" blockId="9.[825,1474,657,901]" lastBlockId="13.[129,777,144,199]" lastPageId="13" lastPageNumber="149" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7EB1FFD31FC8EFBBE" box="[825,886,657,681]" italics="true" pageId="9" pageNumber="145">Head:</emphasis>
Length
<quantity id="4CC69B072D1BFFB7EBE3FD32FBE7FBBE" box="[965,1055,658,682]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="145" unit="mm" value="0.39">0.39mm</quantity>
,0.16 × length of body.Frontal process not protruding. Labial palp not visible. Antenna (non-regenerated) length
<quantity id="4CC69B072D1BFFB7EBA4FD70FC1AFBFF" box="[898,994,720,744]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="145" unit="mm" value="1.42">1.42 mm</quantity>
, 0.6 × length of body, with 25 antennomeres; antennomeres I and II subrectangular; antennomere III onwards moniliform, as long as wide (length and width
<quantity id="4CC69B072D1BFFB7ED70FCAFFA4EFA30" box="[1366,1462,783,807]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="9" pageNumber="145" unit="mm" value="0.07">0.07 mm</quantity>
); apical antennomere length
<quantity id="4CC69B072D1BFFB7EC7DFC8EFB45FA51" box="[1115,1213,814,838]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="145" unit="mm" value="0.09">0.09 mm</quantity>
, width
<quantity id="4CC69B072D1BFFB7ED36FC8EFA89FA51" box="[1296,1393,814,838]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="9" pageNumber="145" unit="mm" value="0.07">0.07 mm</quantity>
, with a cupuliform organ occupying 1/17th of its length, with sensilla not visible; trichobotria visible in antennomeres III–V (
<figureCitation id="13052A672D1BFFB7ED4CFCCDFA57FA92" box="[1386,1455,877,901]" captionStart="Figure 8" captionStartId="10.[113,178,1836,1860]" captionTargetBox="[169,1403,147,1805]" captionTargetId="figure-8@10.[166,1406,144,1808]" captionTargetPageId="10" captionText="Figure 8. Camera lucida drawings of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of thorax, showing the distribution of macrosetae.B, detail of basal antennomeres.C, dorsal habitus.D, detail of stylus.E, detail of pretarsus and tibia of leg II; asterisk indicates the ventral macroseta on the tibia. F, detail of pretarsus and tibia of leg III; asterisk indicates the ventral macroseta on the tibia. Abbreviation:c, calcar." figureDoi="http://doi.org/10.5281/zenodo.11241066" httpUri="https://zenodo.org/record/11241066/files/figure.png" pageId="9" pageNumber="145">Fig. 8B</figureCitation>
); medial and distal antennomeres with whorls of short, thin and smooth setae.
</paragraph>
<caption id="DF41666A2D1BFFB7E8A7F802FB4DFEAC" ID-DOI="http://doi.org/10.5281/zenodo.11241064" ID-Zenodo-Dep="11241064" box="[129,1205,1954,1979]" httpUri="https://zenodo.org/record/11241064/files/figure.png" pageId="9" pageNumber="145" startId="9.[129,194,1954,1978]" targetBox="[148,1454,984,1923]" targetPageId="9" targetType="figure">
<paragraph id="8B8136E22D1BFFB7E8A7F802FB4DFEAC" blockId="9.[129,1205,1954,1979]" box="[129,1205,1954,1979]" pageId="9" pageNumber="145">
<emphasis id="B94AEAF02D1BFFB7E8A7F802FF20FEAC" bold="true" box="[129,216,1954,1979]" pageId="9" pageNumber="145">Figure 7.</emphasis>
Dorsal photomicrograph of holotype of
<taxonomicName id="4C3E4D612D1BFFB7EA71F803FC88FEAD" authority=", AMNH" authorityName="AMNH" box="[599,880,1954,1979]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="9" pageNumber="145" phylum="Arthropoda" rank="species" species="eobaltica">
<emphasis id="B94AEAF02D1BFFB7EA71F803FCE5FEAD" bold="true" box="[599,797,1954,1979]" italics="true" pageId="9" pageNumber="145">Litocampa eobaltica</emphasis>
,
<emphasis id="B94AEAF02D1BFFB7EB01F803FC88FEAD" box="[807,880,1955,1978]" italics="true" pageId="9" pageNumber="145">AMNH</emphasis>
</taxonomicName>
<emphasis id="B94AEAF02D1BFFB7EB53F803FC34FEAC" box="[885,972,1955,1979]" italics="true" pageId="9" pageNumber="145">Ba-JVe66</emphasis>
, in Eocene Baltic amber.
</paragraph>
</caption>
<caption id="DF41666A2D18FFB4E857F88CFE87FE8F" ID-DOI="http://doi.org/10.5281/zenodo.11241066" ID-Zenodo-Dep="11241066" httpUri="https://zenodo.org/record/11241066/files/figure.png" pageId="10" pageNumber="146" startId="10.[113,178,1836,1860]" targetBox="[169,1403,147,1805]" targetPageId="10" targetType="figure">
<paragraph id="8B8136E22D18FFB4E857F88CFE87FE8F" blockId="10.[113,1446,1836,1944]" pageId="10" pageNumber="146">
<emphasis id="B94AEAF02D18FFB4E857F88CFF31FE53" bold="true" box="[113,201,1836,1860]" pageId="10" pageNumber="146">Figure 8.</emphasis>
Camera lucida drawings of holotype of
<taxonomicName id="4C3E4D612D18FFB4EA66F88CFCA2FE54" authority=", AMNH" authorityName="AMNH" box="[576,858,1836,1860]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="10" pageNumber="146" phylum="Arthropoda" rank="species" species="eobaltica">
<emphasis id="B94AEAF02D18FFB4EA66F88CFCFEFE53" bold="true" box="[576,774,1836,1860]" italics="true" pageId="10" pageNumber="146">Litocampa eobaltica</emphasis>
,
<emphasis id="B94AEAF02D18FFB4EB37F88CFCA2FE54" box="[785,858,1836,1859]" italics="true" pageId="10" pageNumber="146">AMNH</emphasis>
</taxonomicName>
<emphasis id="B94AEAF02D18FFB4EB78F88CFC4DFE53" box="[862,949,1836,1860]" italics="true" pageId="10" pageNumber="146">Ba-JVe66</emphasis>
, in Eocene Baltic amber. A, detail of thorax, showing the distribution of macrosetae. B, detail of basal antennomeres. C, dorsal habitus. D, detail of stylus. E, detail of pretarsus and tibia of leg II; asterisk indicates the ventral macroseta on the tibia. F, detail of pretarsus and tibia of leg III; asterisk indicates the ventral macroseta on the tibia. Abbreviation: c, calcar.
</paragraph>
</caption>
<caption id="DF41666A2D19FFB5E8A7F8A2FEA1FE79" ID-DOI="http://doi.org/10.5281/zenodo.11241068" ID-Zenodo-Dep="11241068" httpUri="https://zenodo.org/record/11241068/files/figure.png" pageId="11" pageNumber="147" startId="11.[129,194,1794,1818]" targetBox="[132,1470,148,1763]" targetPageId="11" targetType="figure">
<paragraph id="8B8136E22D19FFB5E8A7F8A2FEA1FE79" blockId="11.[129,1433,1794,1902]" pageId="11" pageNumber="147">
<emphasis id="B94AEAF02D19FFB5E8A7F8A2FF20FE0D" bold="true" box="[129,216,1794,1818]" pageId="11" pageNumber="147">Figure 9.</emphasis>
Photomicrographs of holotype of
<taxonomicName id="4C3E4D612D19FFB5EA3AF8A2FCCEFE0D" authority=", AMNH" authorityName="AMNH" box="[540,822,1794,1818]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="11" pageNumber="147" phylum="Arthropoda" rank="species" species="eobaltica">
<emphasis id="B94AEAF02D19FFB5EA3AF8A2FD1BFE0D" bold="true" box="[540,739,1794,1818]" italics="true" pageId="11" pageNumber="147">Litocampa eobaltica</emphasis>
,
<emphasis id="B94AEAF02D19FFB5EACBF8A3FCCEFE0D" box="[749,822,1795,1818]" italics="true" pageId="11" pageNumber="147">AMNH</emphasis>
</taxonomicName>
<emphasis id="B94AEAF02D19FFB5EB1DF8A2FC6AFE0D" box="[827,914,1794,1818]" italics="true" pageId="11" pageNumber="147">Ba-JVe66</emphasis>
, in Eocene Baltic amber. A, detail of left leg II; asterisk indicates the ventral macroseta on the tibia. B, detail of pretarsus in A. C, detail of left leg III; asterisk indicates the ventral macroseta on the tibia. D, detail of pretarsus in C. E, distribution of macrosetae on pronotum and mesonotum. F, distribution of macrosetae on metanotum. Abbreviation: c, calcar.
</paragraph>
</caption>
<caption id="DF41666A2D1EFFB2E857F88CFD35FE6B" ID-DOI="http://doi.org/10.5281/zenodo.11241070" ID-Zenodo-Dep="11241070" httpUri="https://zenodo.org/record/11241070/files/figure.png" pageId="12" pageNumber="148" startId="12.[113,178,1836,1860]" targetBox="[141,1431,147,1805]" targetPageId="12" targetType="figure">
<paragraph id="8B8136E22D1EFFB2E857F88CFD35FE6B" blockId="12.[113,1442,1836,1916]" pageId="12" pageNumber="148">
<emphasis id="B94AEAF02D1EFFB2E857F88CFF2EFE53" bold="true" box="[113,214,1836,1860]" pageId="12" pageNumber="148">Figure 10.</emphasis>
Photomicrograph of paratype of
<taxonomicName id="4C3E4D612D1EFFB2EA28F88CFCD2FE53" authority=", AMNH" authorityName="AMNH" box="[526,810,1836,1860]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="12" pageNumber="148" phylum="Arthropoda" rank="species" species="eobaltica">
<emphasis id="B94AEAF02D1EFFB2EA28F88CFD2DFE53" bold="true" box="[526,725,1836,1860]" italics="true" pageId="12" pageNumber="148">Litocampa eobaltica</emphasis>
, AMNH
</taxonomicName>
Ba-JVe277, in Eocene Baltic amber. A, dorsal habitus; arrows point to the spider web. B, thorax in dorsal view, showing the distribution of macrosetae; the box labelled C corresponds to panel C. C, detail of microdenticles of the cuticle. D, head and thorax in ventral view.
</paragraph>
</caption>
<paragraph id="8B8136E22D1FFFB3E8A7FF49FE3AFBE1" blockId="13.[128,778,233,758]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3E8A7FF49FF36F816" box="[129,206,233,257]" italics="true" pageId="13" pageNumber="149">Thorax:</emphasis>
Length
<quantity id="4CC69B072D1FFFB3E90DFF49FE73F817" box="[299,395,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="13" pageNumber="149" unit="mm" value="0.66">0.66 mm</quantity>
, 0.28 × length of body. Distribution of macrosetae: pronotum with 1 + 1
<emphasis id="B94AEAF02D1FFFB3EA31FEA9FDCEF837" box="[535,566,265,288]" italics="true" pageId="13" pageNumber="149">ma</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3EAAFFEA8FD63F837" box="[649,667,264,288]" italics="true" pageId="13" pageNumber="149">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3EAC9FEA8FCFAF837" box="[751,770,264,288]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; mesonotum with 1 + 1
<emphasis id="B94AEAF02D1FFFB3E95EFE88FE6FF828" box="[376,407,296,319]" italics="true" pageId="13" pageNumber="149">ma</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3E9C5FE87FE0DF828" box="[483,501,295,319]" italics="true" pageId="13" pageNumber="149">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3EA64FE87FDADF828" box="[578,597,295,319]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; and metanotum with 1 + 1
<emphasis id="B94AEAF02D1FFFB3E8D8FEE7FEE5F849" box="[254,285,327,350]" italics="true" pageId="13" pageNumber="149">ma</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3E955FEE7FE7EF848" box="[371,390,327,351]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
(
<figureCitation id="13052A672D1FFFB3E9BBFEE7FE09F848" box="[413,497,327,351]" captionStart="Figure 8" captionStartId="10.[113,178,1836,1860]" captionTargetBox="[169,1403,147,1805]" captionTargetId="figure-8@10.[166,1406,144,1808]" captionTargetPageId="10" captionText="Figure 8. Camera lucida drawings of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of thorax, showing the distribution of macrosetae.B, detail of basal antennomeres.C, dorsal habitus.D, detail of stylus.E, detail of pretarsus and tibia of leg II; asterisk indicates the ventral macroseta on the tibia. F, detail of pretarsus and tibia of leg III; asterisk indicates the ventral macroseta on the tibia. Abbreviation:c, calcar." figureDoi="http://doi.org/10.5281/zenodo.11241066" httpUri="https://zenodo.org/record/11241066/files/figure.png" pageId="13" pageNumber="149">Figs 8A</figureCitation>
,
<figureCitation id="13052A672D1FFFB3EA22FEE7FDC7F849" box="[516,575,327,351]" captionStart="Figure 9" captionStartId="11.[129,194,1794,1818]" captionTargetBox="[132,1470,148,1763]" captionTargetId="figure-9@11.[129,1473,144,1766]" captionTargetPageId="11" captionText="Figure 9. Photomicrographs of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of left leg II; asterisk indicates the ventral macroseta on the tibia. B, detail of pretarsus in A. C, detail of left leg III; asterisk indicates the ventral macroseta on the tibia. D, detail of pretarsus in C. E, distribution of macrosetae on pronotum and mesonotum. F, distribution of macrosetae on metanotum. Abbreviation: c, calcar." figureDoi="http://doi.org/10.5281/zenodo.11241068" httpUri="https://zenodo.org/record/11241068/files/figure.png" pageId="13" pageNumber="149">9E, F</figureCitation>
,
<figureCitation id="13052A672D1FFFB3EA77FEE7FD81F848" box="[593,633,327,351]" captionStart="Figure 10" captionStartId="12.[113,178,1836,1860]" captionTargetBox="[141,1431,147,1805]" captionTargetId="figure-7@12.[138,1434,144,1808]" captionTargetPageId="12" captionText="Figure 10. Photomicrograph of paratype of Litocampa eobaltica, AMNH Ba-JVe277, in Eocene Baltic amber. A, dorsal habitus; arrows point to the spider web.B, thorax in dorsal view, showing the distribution of macrosetae; the box labelled C corresponds to panel C. C, detail of microdenticles of the cuticle.D, head and thorax in ventral view." figureDoi="http://doi.org/10.5281/zenodo.11241070" httpUri="https://zenodo.org/record/11241070/files/figure.png" pageId="13" pageNumber="149">10B</figureCitation>
);
<emphasis id="B94AEAF02D1FFFB3EABEFEE7FD4FF849" box="[664,695,327,350]" italics="true" pageId="13" pageNumber="149">ma</emphasis>
and
<emphasis id="B94AEAF02D1FFFB3EAD1FEE7FCF1F848" box="[759,777,327,351]" italics="true" pageId="13" pageNumber="149">la</emphasis>
macrosetae short and thin;
<emphasis id="B94AEAF02D1FFFB3E9BCFEC6FE55F869" box="[410,429,358,382]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
macrosetae longer and robust; all macrosetae with very short barbs on one side, giving a serrate appearance; marginal setae longer than clothing setae and robust, with barbs as for macrosetae. Legs slightly elongated, with metathoracic leg reaching abdominal segment VI; length of metathoracic leg segments: femur
<quantity id="4CC69B072D1FFFB3EA23FDA3FD9CFB0D" box="[517,612,515,539]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="149" unit="mm" value="0.17">0.17 mm</quantity>
, tibia
<quantity id="4CC69B072D1FFFB3EA80FDA2FCFEFB0D" box="[678,774,514,538]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="149" unit="mm" value="0.16">0.16 mm</quantity>
, tarsus including pretarsus
<quantity id="4CC69B072D1FFFB3E9A8FD82FE16FB2E" box="[398,494,546,570]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="149" unit="mm" value="0.15">0.15 mm</quantity>
; femur with one long, thin, smooth ventral macroseta; tibia with one long, thin, smooth ventral macroseta and two regular calcars distally (
<figureCitation id="13052A672D1FFFB3EABCFDC1FCFCFB6F" box="[666,772,609,633]" captionStart="Figure 8" captionStartId="10.[113,178,1836,1860]" captionTargetBox="[169,1403,147,1805]" captionTargetId="figure-8@10.[166,1406,144,1808]" captionTargetPageId="10" captionText="Figure 8. Camera lucida drawings of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of thorax, showing the distribution of macrosetae.B, detail of basal antennomeres.C, dorsal habitus.D, detail of stylus.E, detail of pretarsus and tibia of leg II; asterisk indicates the ventral macroseta on the tibia. F, detail of pretarsus and tibia of leg III; asterisk indicates the ventral macroseta on the tibia. Abbreviation:c, calcar." figureDoi="http://doi.org/10.5281/zenodo.11241066" httpUri="https://zenodo.org/record/11241066/files/figure.png" pageId="13" pageNumber="149">Figs 8E, F</figureCitation>
,
<figureCitation id="13052A672D1FFFB3E8A6FD20FF46FB8F" box="[128,190,640,664]" captionStart="Figure 9" captionStartId="11.[129,194,1794,1818]" captionTargetBox="[132,1470,148,1763]" captionTargetId="figure-9@11.[129,1473,144,1766]" captionTargetPageId="11" captionText="Figure 9. Photomicrographs of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of left leg II; asterisk indicates the ventral macroseta on the tibia. B, detail of pretarsus in A. C, detail of left leg III; asterisk indicates the ventral macroseta on the tibia. D, detail of pretarsus in C. E, distribution of macrosetae on pronotum and mesonotum. F, distribution of macrosetae on metanotum. Abbreviation: c, calcar." figureDoi="http://doi.org/10.5281/zenodo.11241068" httpUri="https://zenodo.org/record/11241068/files/figure.png" pageId="13" pageNumber="149">9A–D</figureCitation>
); femur and tibia without dorsal macrosetae; tarsus with typical setae along ventral side; pretarsus with subequal elbowlike claws with small and smooth lateral crests; pretarsal lateral processes setiform and smooth.
</paragraph>
<paragraph id="8B8136E22D1FFFB3E8A6FCB7FF41FC95" blockId="13.[128,778,791,1410]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3E8A6FCB7FF1DFA38" box="[128,229,791,815]" italics="true" pageId="13" pageNumber="149">Abdomen:</emphasis>
Length
<quantity id="4CC69B072D1FFFB3E91DFCB7FE60FA38" box="[315,408,791,815]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="149" unit="mm" value="1.3">1.30 mm</quantity>
, 0.55 × length of body. Distribution of macrosetae on tergites: VI–VII with 1 + 1
<emphasis id="B94AEAF02D1FFFB3EA7AFC96FD96FA59" box="[604,622,822,846]" italics="true" pageId="13" pageNumber="149">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3EA9DFC96FD36FA59" box="[699,718,822,846]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; VIII with 3 + 3
<emphasis id="B94AEAF02D1FFFB3E8DEFCF6FEF3FA79" box="[248,267,854,878]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; IX at least with 3 + 3
<emphasis id="B94AEAF02D1FFFB3EA2AFCF6FDE7FA79" box="[524,543,854,878]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
; macrosetae long and thick, with very short barbs on one side along distal half or third, giving a serrate appearance. Urosternite details mostly obscured (stylus II and eversible vesicles not visible). Pairs of styli visible from sternites III to VII; stylus (
<figureCitation id="13052A672D1FFFB3E9FCFC73FDDFFAFC" box="[474,551,979,1003]" captionStart="Figure 8" captionStartId="10.[113,178,1836,1860]" captionTargetBox="[169,1403,147,1805]" captionTargetId="figure-8@10.[166,1406,144,1808]" captionTargetPageId="10" captionText="Figure 8. Camera lucida drawings of holotype of Litocampa eobaltica, AMNH Ba-JVe66, in Eocene Baltic amber. A, detail of thorax, showing the distribution of macrosetae.B, detail of basal antennomeres.C, dorsal habitus.D, detail of stylus.E, detail of pretarsus and tibia of leg II; asterisk indicates the ventral macroseta on the tibia. F, detail of pretarsus and tibia of leg III; asterisk indicates the ventral macroseta on the tibia. Abbreviation:c, calcar." figureDoi="http://doi.org/10.5281/zenodo.11241066" httpUri="https://zenodo.org/record/11241066/files/figure.png" pageId="13" pageNumber="149">Fig. 8D</figureCitation>
) with smooth apical, subapical and ventromedial setae; subapical and ventromedial setae apparently longer than apical setae. Both cerci complete (non-regenerated), length
<quantity id="4CC69B072D1FFFB3E9A9FB91FE15FD5E" box="[399,493,1073,1097]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="149" unit="mm" value="2.54">2.54 mm</quantity>
, 1.08 × length of body; divided into a basal article plus five primary articles; basal article length
<quantity id="4CC69B072D1FFFB3E8E0FBD0FEDAFD90" box="[198,290,1136,1160]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="149" unit="mm" value="0.41">0.41 mm</quantity>
, subdivided into five secondary articles; primary articles increasing in length towards the apex, with four to five constrictions (first primary article
<quantity id="4CC69B072D1FFFB3E9CEFB0FFDB0FDD1" box="[488,584,1199,1223]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="149" unit="mm" value="0.27">0.27 mm</quantity>
; second
<quantity id="4CC69B072D1FFFB3EA82FB0FFCFDFDD1" box="[676,773,1199,1223]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="149" unit="mm" value="0.32">0.32 mm</quantity>
; third
<quantity id="4CC69B072D1FFFB3E89CFB6EFEE1FDF2" box="[186,281,1230,1254]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="149" unit="mm" value="0.45">0.45 mm</quantity>
; fourth
<quantity id="4CC69B072D1FFFB3E94BFB6EFE34FDF2" box="[365,460,1230,1254]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="13" pageNumber="149" unit="mm" value="0.52">0.52 mm</quantity>
; fifth
<quantity id="4CC69B072D1FFFB3EA2FFB6EFD92FDF1" box="[521,618,1230,1254]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="13" pageNumber="149" unit="mm" value="0.57">0.57 mm</quantity>
); cercal articles with four to six whorls of smooth macrosetae accompanied by whorls of long, thin, smooth setae; the number of whorls increasing from first to fifth primary articles; each constriction between articles preceded by a whorl of short, thin, smooth setae.
</paragraph>
<paragraph id="8B8136E22D1FFFB3E8A6FA04FE9AFFA1" blockId="13.[128,778,1444,1718]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3E8A6FA04FEBBFCAC" box="[128,323,1444,1467]" italics="true" pageId="13" pageNumber="149">Descriptive notes on</emphasis>
<materialsCitation id="3B563CBF2D1FFFB3E96DFA04FDEDFCCC" collectionCode="AMNH" pageId="13" pageNumber="149" specimenCount="1" typeStatus="paratype">
<emphasis id="B94AEAF02D1FFFB3E96DFA04FD88FCAB" box="[331,624,1444,1468]" italics="true" pageId="13" pageNumber="149">
<typeStatus id="548588402D1FFFB3E96DFA04FE59FCAC" box="[331,417,1444,1467]" pageId="13" pageNumber="149" type="paratype">paratype</typeStatus>
<collectionCode id="ED2FAE272D1FFFB3E98EFA04FE0FFCAC" box="[424,503,1444,1467]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="149">AMNH</collectionCode>
Ba-JVe277:
</emphasis>
Sex unknown, adult. General description is as for the
</materialsCitation>
<typeStatus id="548588402D1FFFB3EA3BFA63FD83FCCC" box="[541,635,1475,1499]" pageId="13" pageNumber="149" type="holotype">holotype</typeStatus>
, with the following measurements: body length
<quantity id="4CC69B072D1FFFB3E9D4FA42FDA9FCED" box="[498,593,1506,1530]" metricMagnitude="-3" metricUnit="m" metricValue="1.91" pageId="13" pageNumber="149" unit="mm" value="1.91">1.91 mm</quantity>
; head length
<quantity id="4CC69B072D1FFFB3EAFBFA42FF53FF0E" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="149" unit="mm" value="0.29">0.29 mm</quantity>
, 0.15 × length of body, showing the occipital Y-suture; antenna length
<quantity id="4CC69B072D1FFFB3E92AF981FE94FF2F" box="[268,364,1569,1593]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="13" pageNumber="149" unit="mm" value="1.09">1.09 mm</quantity>
, 0.57 × length of body, with the distal-most antennomere more robust than in the
<typeStatus id="548588402D1FFFB3EA13F9E0FD69FF4F" box="[565,657,1600,1624]" pageId="13" pageNumber="149" type="holotype">holotype</typeStatus>
(
<figureCitation id="13052A672D1FFFB3EA86F9E0FD0EFF4F" box="[672,758,1600,1624]" captionStart="Figure 10" captionStartId="12.[113,178,1836,1860]" captionTargetBox="[141,1431,147,1805]" captionTargetId="figure-7@12.[138,1434,144,1808]" captionTargetPageId="12" captionText="Figure 10. Photomicrograph of paratype of Litocampa eobaltica, AMNH Ba-JVe277, in Eocene Baltic amber. A, dorsal habitus; arrows point to the spider web.B, thorax in dorsal view, showing the distribution of macrosetae; the box labelled C corresponds to panel C. C, detail of microdenticles of the cuticle.D, head and thorax in ventral view." figureDoi="http://doi.org/10.5281/zenodo.11241070" httpUri="https://zenodo.org/record/11241070/files/figure.png" pageId="13" pageNumber="149">Fig. 10D</figureCitation>
); thorax length
<quantity id="4CC69B072D1FFFB3E934F9C0FE9CFF60" box="[274,356,1632,1656]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="149" unit="mm" value="0.7">0.7 mm</quantity>
, 0.37 × length of body; abdomen length
<quantity id="4CC69B072D1FFFB3E8A6F9DFFF1BFF81" box="[128,227,1663,1687]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="13" pageNumber="149" unit="mm" value="0.92">0.92 mm</quantity>
, 0.48 × length of body; and cerci length
<quantity id="4CC69B072D1FFFB3EA85F9DFFCFDFF81" box="[675,773,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="3.91" pageId="13" pageNumber="149" unit="mm" value="3.91">3.91 mm</quantity>
, 2.05 × length of body.
</paragraph>
<paragraph id="8B8136E22D1FFFB3E8A6F978FBBFF8D6" blockId="13.[128,778,1752,1964]" lastBlockId="13.[825,1474,144,449]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3E8A6F978FF26FFE7" box="[128,222,1752,1776]" italics="true" pageId="13" pageNumber="149">Remarks:</emphasis>
The new
<taxonomicName id="4C3E4D612D1FFFB3E96FF978FDFBFFE7" box="[329,515,1752,1776]" pageId="13" pageNumber="149">
<emphasis id="B94AEAF02D1FFFB3E96FF978FE4AFFF8" box="[329,434,1752,1775]" italics="true" pageId="13" pageNumber="149">Litocampa</emphasis>
species
</taxonomicName>
fits within the subfamily
<taxonomicName id="4C3E4D612D1FFFB3E8A6F957FEE8FE18" authorityName="Conde" authorityYear="1956" box="[128,272,1783,1807]" class="Diplura" family="Campodeidae" kingdom="Animalia" pageId="13" pageNumber="149" phylum="Arthropoda" rank="subFamily" subFamily="Campodeinae">Campodeinae</taxonomicName>
owing to the presence of a maximum of 3 + 3 macrosetae on the pronotum (
<bibRefCitation id="EFAF4B132D1FFFB3E9E1F8B6FDB2FE39" author="Conde B" box="[455,586,1814,1838]" pageId="13" pageNumber="149" pagination="1 - 202" refId="ref10405" refString="Conde B. Materiaux pour une Monographie des Diploures Campodeides. Memoires du Museum National d'Histoire naturelle, Serie A. Zoologie 1956; 12: 1 - 202." type="journal article" year="1956">Condé 1956</bibRefCitation>
, Paclt 1957). The presence of claws with lateral crests, in combination with the presence of
<emphasis id="B94AEAF02D1FFFB3E8D9F8F5FEEAFE7A" box="[255,274,1877,1901]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
macrosetae on the pro-, meso- and metanotum is characteristic of the genus
<taxonomicName id="4C3E4D612D1FFFB3E99CF8D5FCFCFE9B" authority="(Sendra et al. 2016)" baseAuthorityName="Sendra" baseAuthorityYear="2016" box="[442,772,1908,1932]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAF02D1FFFB3E99CF8D5FDDBFE9B" box="[442,547,1909,1932]" italics="true" pageId="13" pageNumber="149">Litocampa</emphasis>
(
<bibRefCitation id="EFAF4B132D1FFFB3EA11F8D4FD00FE9B" author="Sendra A &amp; Palacios J &amp; Garcia A" box="[567,760,1908,1932]" pageId="13" pageNumber="149" pagination="540 - 58" refId="ref11386" refString="Sendra A, Palacios J, Garcia A et al. New species of Campodeidae (Diplura) from Mexican caves. Zootaxa 2016; 4072: 540 - 58." type="journal article" year="2016">
Sendra
<emphasis id="B94AEAF02D1FFFB3EAAEF8D5FD44FE9B" box="[648,700,1908,1932]" italics="true" pageId="13" pageNumber="149">et al.</emphasis>
2016
</bibRefCitation>
)
</taxonomicName>
. The genus
<taxonomicName id="4C3E4D612D1FFFB3E8C8F834FEAFFEBC" box="[238,343,1940,1963]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAF02D1FFFB3E8C8F834FEAFFEBC" box="[238,343,1940,1963]" italics="true" pageId="13" pageNumber="149">Litocampa</emphasis>
</taxonomicName>
comprises ~30 species, mostly distributed in the Holartic region in soil and cave habitats. The absence of dorsal macrosetae on femora III and the macrosetae formula 1 + 1
<emphasis id="B94AEAF02D1FFFB3EB50FF6EFC70F9F1" box="[886,904,206,230]" italics="true" pageId="13" pageNumber="149">la</emphasis>
, 1 + 1
<emphasis id="B94AEAF02D1FFFB3EBF6FF6EFC1BF9F1" box="[976,995,206,230]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
, without
<emphasis id="B94AEAF02D1FFFB3EC60FF6FFB9DF9F1" box="[1094,1125,207,230]" italics="true" pageId="13" pageNumber="149">ma</emphasis>
, on abdominal tergites VI–VII excludes most of the
<taxonomicName id="4C3E4D612D1FFFB3EBDBFF4EFB43F811" box="[1021,1211,238,262]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94AEAF02D1FFFB3EBDBFF4EFB9EF812" box="[1021,1126,238,261]" italics="true" pageId="13" pageNumber="149">Litocampa</emphasis>
species.
</taxonomicName>
In this sense,
<taxonomicName id="4C3E4D612D1FFFB3ED6DFF4EFA3AF811" box="[1355,1474,238,262]" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="species" species="eobaltica">
<emphasis id="B94AEAF02D1FFFB3ED6DFF4EFA3AF811" box="[1355,1474,238,262]" italics="true" pageId="13" pageNumber="149">Li. eobaltica</emphasis>
</taxonomicName>
resembles a few species, such as the extant species
<taxonomicName id="4C3E4D612D1FFFB3ED7FFEAEFB1CF853" authority="Sendra and Garcia, 2016" authorityName="Sendra and Garcia" authorityYear="2016" class="Diplura" family="Campodeidae" genus="Litocampa" higherTaxonomySource="CoL" kingdom="Animalia" order="Diplura" pageId="13" pageNumber="149" phylum="Arthropoda" rank="species" species="enriqueriojai">
<emphasis id="B94AEAF02D1FFFB3ED7FFEAEFC4DF853" italics="true" pageId="13" pageNumber="149">Litocampa enriqueriojai</emphasis>
<bibRefCitation id="EFAF4B132D1FFFB3EBE3FE8CFB1CF853" author="Sendra A &amp; Palacios J &amp; Garcia A" box="[965,1252,300,324]" pageId="13" pageNumber="149" pagination="540 - 58" refId="ref11386" refString="Sendra A, Palacios J, Garcia A et al. New species of Campodeidae (Diplura) from Mexican caves. Zootaxa 2016; 4072: 540 - 58." type="journal article" year="2016">Sendra and García, 2016</bibRefCitation>
</taxonomicName>
, but differs in the microdenticles of the cuticle, the number of antennomeres, the robust
<emphasis id="B94AEAF02D1FFFB3EBA4FECBFC6DF894" box="[898,917,363,387]" italics="true" pageId="13" pageNumber="149">lp</emphasis>
compared with
<emphasis id="B94AEAF02D1FFFB3EC18FECCFBA5F894" box="[1086,1117,364,387]" italics="true" pageId="13" pageNumber="149">ma</emphasis>
and
<emphasis id="B94AEAF02D1FFFB3ECB4FECBFB5CF894" box="[1170,1188,363,387]" italics="true" pageId="13" pageNumber="149">la</emphasis>
macrosetae, and the robust marginal setae. The abundance of clothing setae indicates that both specimens are adults.
</paragraph>
</subSubSection>
</treatment>
</document>
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