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<document id="ACB9E83D7F99F890EE1CE2199FBD8C6D" ID-DOI="10.11646/phytotaxa.311.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13700563" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725574901492" checkinUser="felipe" docAuthor="Ranjbar, Massoud &amp; Ranjbar, Zahra" docDate="2017" docId="039787EDFF8B9C65DEBFD553FCDDA0CE" docLanguage="en" docName="phytotaxa.311.2.1.pdf" docOrigin="Phytotaxa 311 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.311.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Cephalaria golestanica Ranjbar &amp; Z. Ranjbar 2017, sp. nov." docType="treatment" docVersion="1" lastPageNumber="110" masterDocId="FFAEFF95FF8C9C6CDE37D27DFF83A467" masterDocTitle="A taxonomic revision of Cephalaria subgen. Cephalaria sect. Atrocephalae (Caprifoliaceae) from Iran" masterLastPageNumber="138" masterPageNumber="101" pageNumber="108" updateTime="1725584670209" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="569705220729FAE72C91B840181F2B82" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="89299FCB2946446E1874801236675267">A taxonomic revision of Cephalaria subgen. Cephalaria sect. Atrocephalae (Caprifoliaceae) from Iran</mods:title>
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<mods:name id="A2DF40C4F2AB0AE381C868B71AF1EDCF" type="personal">
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<mods:namePart id="BEF900987F1C802EDA5B66F2785D7C8E">Ranjbar, Massoud</mods:namePart>
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<mods:namePart id="FDC3CB50B0EE634FAE393F716CB91487">Ranjbar, Zahra</mods:namePart>
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<mods:title id="13830F2E005F005FA502F4679F33C96B">Phytotaxa</mods:title>
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<mods:part id="DD09B1890184A9DF70088F295D580846">
<mods:date id="75BE3B50D362E6843988885C5921B81F">2017</mods:date>
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<mods:number id="B5DDF7FAC7100888DEF032CB4B8AA7AA">2017-06-27</mods:number>
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<treatment id="039787EDFF8B9C65DEBFD553FCDDA0CE" LSID="urn:lsid:plazi:treatment:039787EDFF8B9C65DEBFD553FCDDA0CE" httpUri="http://treatment.plazi.org/id/039787EDFF8B9C65DEBFD553FCDDA0CE" lastPageId="9" lastPageNumber="110" pageId="7" pageNumber="108">
<subSubSection id="C3246570FF8B9C6BDEBFD553FC93A320" box="[136,784,1836,1863]" pageId="7" pageNumber="108" type="nomenclature">
<paragraph id="8B8136FBFF8B9C6BDEBFD553FC93A320" blockId="7.[136,1451,1836,1934]" box="[136,784,1836,1863]" pageId="7" pageNumber="108">
<heading id="D0C98197FF8B9C6BDEBFD553FC93A320" box="[136,784,1836,1863]" fontSize="11" level="2" pageId="7" pageNumber="108" reason="4">
<emphasis id="B94AEAE9FF8B9C6BDEBFD553FF1EA320" bold="true" box="[136,157,1838,1863]" pageId="7" pageNumber="108">4.</emphasis>
<taxonomicName id="4C3E4D78FF8B9C6BDE93D551FD2FA320" ID-CoL="69J72" authority="Ranjbar &amp; Z. Ranjbar" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[164,684,1836,1863]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="7" pageNumber="108" phylum="Tracheophyta" rank="species" species="golestanica" status="sp. nov.">
<emphasis id="B94AEAE9FF8B9C6BDE93D551FE2CA320" bold="true" box="[164,431,1836,1863]" italics="true" pageId="7" pageNumber="108">Cephalaria golestanica</emphasis>
Ranjbar &amp; Z. Ranjbar
</taxonomicName>
,
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<taxonomicNameLabel id="A2795792FF8B9C6BDC80D553FC93A320" box="[695,784,1838,1863]" pageId="7" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3246570FF8B9C65DEBFD528FB7BA6B2" lastPageId="9" lastPageNumber="110" pageId="7" pageNumber="108" type="materials_examined">
<paragraph id="8B8136FBFF8B9C6BDEBFD528FC98A3E9" blockId="7.[136,1451,1836,1934]" pageId="7" pageNumber="108">
Type:—
<collectingCountry id="F329766BFF8B9C6BDEE0D528FE97A30D" box="[215,276,1877,1898]" name="Iran" pageId="7" pageNumber="108">IRAN</collectingCountry>
. Prov.
<collectingRegion id="49FAF819FF8B9C6BDF62D528FE2DA30D" box="[341,430,1877,1898]" country="Iran" name="Golestan" pageId="7" pageNumber="108">Golestan</collectingRegion>
: Almeh, near
<collectingRegion id="49FAF819FF8B9C6BDC03D528FD0AA30D" box="[564,649,1877,1898]" country="Iran" name="Golestan" pageId="7" pageNumber="108">Golestan</collectingRegion>
National Park in
<collectingRegion id="49FAF819FF8B9C6BDD03D528FC67A30D" box="[820,996,1877,1898]" country="Iran" name="Golestan" pageId="7" pageNumber="108">Golestan Province</collectingRegion>
NE Iran,
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37º 23
<emphasis id="B94AEAE9FF8B9C6BDAB5D528FB04A30D" bold="true" box="[1154,1159,1877,1898]" italics="true" pageId="7" pageNumber="108">ʹ</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE0A503CFF8B9C6BDA9CD528FA85A30D" box="[1195,1286,1877,1898]" degrees="55" direction="east" minutes="38" orientation="longitude" pageId="7" pageNumber="108" precision="925" value="55.633335">
55º 38
<emphasis id="B94AEAE9FF8B9C6BDADED528FB6DA30D" bold="true" box="[1257,1262,1877,1898]" italics="true" pageId="7" pageNumber="108">ʹ</emphasis>
E
</geoCoordinate>
,
<quantity id="4CC69B1EFF8B9C6BDB27D528FAD8A30D" box="[1296,1371,1877,1898]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="108" unit="m" value="1700.0">1700 m</quantity>
,
<date id="FF80103BFF8B9C6BDB52D528FF6CA3E9" pageId="7" pageNumber="108" value="1984-07-15">15 July 1984</date>
,
<emphasis id="B94AEAE9FF8B9C6BDECED504FD96A3E9" box="[249,533,1913,1934]" italics="true" pageId="7" pageNumber="108">Ayatollahi &amp; Zangooei 11616</emphasis>
(
<collectionCode id="ED2FAE3EFF8B9C6BDC15D504FDE8A3E9" box="[546,619,1913,1934]" country="Iran" lsid="urn:lsid:biocol.org:col:15384" name="Ferdowsi University" pageId="7" pageNumber="108" type="Herbarium">FUMH</collectionCode>
,
<typeStatus id="54858859FF8B9C6BDC42D504FD4EA3E9" box="[629,717,1913,1934]" pageId="7" pageNumber="108" type="holotype">holotype</typeStatus>
).
<figureCitation id="13052A7EFF8B9C6BDCEAD504FC94A3E9" box="[733,791,1913,1934]" captionStart="FIGURE 4" captionStartId="8.[136,229,1745,1766]" captionTargetBox="[165,1422,193,1701]" captionTargetId="figure-16@8.[151,1435,187,1717]" captionTargetPageId="8" captionText="FIGURE 4. A. Cephalaria golestanica Ranjbar &amp; Z. Ranjbar (Ayatollahi &amp; Zangooei 11616, FUMH). B. Capitula. C. &amp; D. Corolla. E. Calyx. F. Involucel. G. Involucral bracts. H. Receptacular bracts." figureDoi="http://doi.org/10.5281/zenodo.13700573" httpUri="https://zenodo.org/record/13700573/files/figure.png" pageId="7" pageNumber="108">Fig. 4</figureCitation>
.
</paragraph>
<caption id="DF416673FF849C64DEBFD4ACFD78A36D" ID-DOI="http://doi.org/10.5281/zenodo.13700573" ID-Zenodo-Dep="13700573" httpUri="https://zenodo.org/record/13700573/files/figure.png" pageId="8" pageNumber="109" startId="8.[136,229,1745,1766]" targetBox="[165,1422,193,1701]" targetPageId="8" targetType="figure">
<paragraph id="8B8136FBFF849C64DEBFD4ACFD78A36D" blockId="8.[136,1452,1745,1802]" pageId="8" pageNumber="109">
<emphasis id="B94AEAE9FF849C64DEBFD4ACFE9FA281" bold="true" box="[136,284,1745,1766]" pageId="8" pageNumber="109">FIGURE 4. A.</emphasis>
<taxonomicName id="4C3E4D78FF849C64DF15D4ACFD5DA281" authority="Ranjbar &amp; Z. Ranjbar" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[290,734,1745,1766]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="golestanica">
<emphasis id="B94AEAE9FF849C64DF15D4ACFD86A281" box="[290,517,1745,1766]" italics="true" pageId="8" pageNumber="109">Cephalaria golestanica</emphasis>
Ranjbar &amp; Z. Ranjbar
</taxonomicName>
(
<emphasis id="B94AEAE9FF849C64DCDAD4ACFB88A281" box="[749,1035,1745,1766]" italics="true" pageId="8" pageNumber="109">Ayatollahi &amp; Zangooei 11616</emphasis>
, FUMH).
<emphasis id="B94AEAE9FF849C64DA46D4ACFB0BA281" bold="true" box="[1137,1160,1745,1766]" pageId="8" pageNumber="109">B.</emphasis>
Capitula.
<emphasis id="B94AEAE9FF849C64DADBD4ACFA80A281" bold="true" box="[1260,1283,1745,1766]" pageId="8" pageNumber="109">C.</emphasis>
&amp;
<emphasis id="B94AEAE9FF849C64DB14D4ACFAB8A281" bold="true" box="[1315,1339,1745,1766]" pageId="8" pageNumber="109">D.</emphasis>
Corolla.
<emphasis id="B94AEAE9FF849C64DBA2D4ACFA2FA281" bold="true" box="[1429,1452,1745,1766]" pageId="8" pageNumber="109">E.</emphasis>
Calyx.
<emphasis id="B94AEAE9FF849C64DEFAD488FF63A36D" bold="true" box="[205,224,1781,1802]" pageId="8" pageNumber="109">F.</emphasis>
Involucel.
<emphasis id="B94AEAE9FF849C64DF7BD488FEE6A36D" bold="true" box="[332,357,1781,1802]" pageId="8" pageNumber="109">G.</emphasis>
Involucral bracts.
<emphasis id="B94AEAE9FF849C64DC2ED488FDB1A36D" bold="true" box="[537,562,1781,1802]" pageId="8" pageNumber="109">H.</emphasis>
Receptacular bracts.
</paragraph>
</caption>
<paragraph id="8B8136FBFF849C65DEBFD546FD65A60E" blockId="8.[136,1452,1851,2092]" lastBlockId="9.[136,1452,160,1193]" lastPageId="9" lastPageNumber="110" pageId="8" pageNumber="109">
Perennial plants, slightly stout, erect, with greyish indumentum. Stem up to
<quantity id="4CC69B1EFF849C64DA35D546FBC9A333" box="[1026,1098,1851,1876]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="109" unit="cm" value="35.0">35 cm</quantity>
tall, simple, slightly branched (consisting of 23 racemes), rather densely pilose. Leaves coriaceous, verticillate (with 6 leaves in each node), blades symmetrical, leaves greyish-green when dried, pilose on both surfaces, with distinct vagina. Lower cauline leaves simple, oblanceolate in outline, 67 ×
<quantity id="4CC69B1EFF849C64DC1DD5DAFD2BA3A7" box="[554,680,1959,1984]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="8" pageNumber="109" unit="cm" value="0.8" valueMax="0.9" valueMin="0.7">0.70.9 cm</quantity>
, acute to subacute at apex, serrate, 69 teeth (apical side straight, basal side flexuous), angle of teeth with reference to the midvein 20º45º, teeth
<quantity id="4CC69B1EFF849C64DDF4D5B6FBCAA383" box="[963,1097,1995,2020]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="8" pageNumber="109" unit="mm" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.30.6 mm</quantity>
long, midvein impressed to flat adaxially and sharply keeled abaxially, tertiary veins weakly percurrent, widely spaced, oblique. Middle cauline leaves lyrate oblanceolate-obovate to spathulate in outline, 2.56.5 ×
<quantity id="4CC69B1EFF849C64DD7ADA6EFC3AAC4B" box="[845,953,2067,2092]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="8" pageNumber="109" unit="cm" value="1.25" valueMax="2.0" valueMin="0.5">0.52 cm</quantity>
, with 46 lateral segments, lateral segments linear-lanceolate, 0.51.5 ×
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, entire, acute at apex, terminal segment larger than lateral ones, obovate to elliptic, 25.5 ×
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, obtuse and round at apex, serrate, 34 teeth (apical side flexuous, basal side concave), angle of teeth with reference to the midvein 45º80º, teeth
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long, midvein flat adaxially and sharply keeled abaxially, tertiary veins reticulate, oblique. Upper cauline leaves simple, lanceolate to linear in outline, 0.41 ×
<quantity id="4CC69B1EFF859C65DB6FD371FF2EA52E" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="9" pageNumber="110" unit="cm" value="0.2" valueMax="0.3" valueMin="0.1">0.10.3 cm</quantity>
, entire, acute at apex, midvein flat adaxially and rounded abaxially, tertiary veins close together, oblique. Peduncle
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long, pubescent. Capitula globose or ovoid,
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in diameter in flower, subradiant. Involucral bracts in two series, ovate, ca. 3.54 ×
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, straw-greenish and very slightly blackish at apex, tomentose (hairs up to
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), obtuse at apex. Receptacular bracts in five series, oblong to narrowly ovate to lanceolate, 57 ×
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, spineless, straw-coloured dorsally at base, blackish and attenuate at apex, pilose or sericeous, ciliate at margins. Calyx cupuliform, 11.5 ×
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, dark green, pubescent, with irregular teeth
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long. Corolla
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long, yellow, densely covered with appressed hairs outside, stamens slightly longer than corolla, anthers deeply yellow to light brown. Involucel 22.5 ×
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in flower, 4-angled, hairy (hairs up to
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), 4 long and 4 short teeth at apex; long teeth ca.
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long, short teeth ca.
<quantity id="4CC69B1EFF859C65DC67D02DFD25A60E" box="[592,678,592,617]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="110" unit="mm" value="0.5">0.5 mm</quantity>
long.
</paragraph>
<paragraph id="8B8136FBFF859C65DE8AD009FB7BA6B2" blockId="9.[136,1452,160,1193]" pageId="9" pageNumber="110">
<materialsCitation id="3B563CA6FF859C65DE8AD009FE70A6D6" collectionCode="TARI" collectorName="Roshan" country="Iran" elevation="1720" location="National Park" municipality="Almeh" pageId="9" pageNumber="110" specimenCount="2" stateProvince="Golestan">
<emphasis id="B94AEAE9FF859C65DE8AD009FD52A6EA" bold="true" box="[189,721,628,653]" pageId="9" pageNumber="110">
<typeStatus id="54858859FF859C65DE8AD009FDBEA6EA" box="[189,573,628,653]" isOtherTypeStatus="true" pageId="9" pageNumber="110">Additional specimens examined</typeStatus>
(
<typeStatus id="54858859FF859C65DC78D009FD4AA6EA" box="[591,713,628,653]" pageId="9" pageNumber="110" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—
<collectingCountry id="F329766BFF859C65DCC5D009FCBFA6EA" box="[754,828,628,653]" name="Iran" pageId="9" pageNumber="110">IRAN</collectingCountry>
. Prov.
<collectingRegion id="49FAF819FF859C65DDB9D009FC75A6EA" box="[910,1014,628,653]" country="Iran" name="Golestan" pageId="9" pageNumber="110">Golestan</collectingRegion>
:
<collectingMunicipality id="6BE5AC81FF859C65DA35D009FBD2A6EA" box="[1026,1105,628,653]" pageId="9" pageNumber="110">Almeh</collectingMunicipality>
, near
<collectingRegion id="49FAF819FF859C65DAAED009FB7EA6EA" box="[1177,1277,628,653]" country="Iran" name="Golestan" pageId="9" pageNumber="110">Golestan</collectingRegion>
<location id="8EE16020FF859C65DB30D009FA24A6EA" LSID="urn:lsid:plazi:treatment:039787EDFF8B9C65DEBFD553FCDDA0CE:8EE16020FF859C65DB30D009FA24A6EA" box="[1287,1447,628,653]" country="Iran" municipality="Almeh" name="National Park" pageId="9" pageNumber="110" stateProvince="Golestan">National Park</location>
,
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<elevation id="0013D1C8FF859C65DEBFD0E5FF60A6D6" box="[136,227,664,689]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="9" pageNumber="110" unit="m" value="1720.0">1720 m</elevation>
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,
<emphasis id="B94AEAE9FF859C65DEC4D0E5FE09A6D6" box="[243,394,664,689]" italics="true" pageId="9" pageNumber="110">
<collectorName id="26CB532DFF859C65DEC4D0E5FEC4A6D6" box="[243,327,664,689]" pageId="9" pageNumber="110">Roshan</collectorName>
6978
</emphasis>
(
<collectionCode id="ED2FAE3EFF859C65DFAAD0E5FE64A6D6" box="[413,487,664,689]" country="Taiwan" name="Taiwan Agricultural Research Institute" pageId="9" pageNumber="110">TARI</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="3B563CA6FF859C65DC34D0E5FF5DA6B2" collectionCode="TARI" country="Iran" location="Sharlogh" pageId="9" pageNumber="110" specimenCount="1" stateProvince="Golestan">
<collectingRegion id="49FAF819FF859C65DC34D0E5FDE8A6D6" box="[515,619,664,689]" country="Iran" name="Golestan" pageId="9" pageNumber="110">Golestan</collectingRegion>
:
<collectingRegion id="49FAF819FF859C65DC4ED0E5FD5EA6D6" box="[633,733,664,689]" country="Iran" name="Golestan" pageId="9" pageNumber="110">Golestan</collectingRegion>
forest, between
<location id="8EE16020FF859C65DD95D0E5FB84A6D6" LSID="urn:lsid:plazi:treatment:039787EDFF8B9C65DEBFD553FCDDA0CE:8EE16020FF859C65DD95D0E5FB84A6D6" box="[930,1031,664,689]" country="Iran" name="Sharlogh" pageId="9" pageNumber="110" stateProvince="Golestan">Sharlogh</location>
and Cheshme Khan,
<emphasis id="B94AEAE9FF859C65DB3FD0E5FA28A6D6" box="[1288,1451,664,689]" italics="true" pageId="9" pageNumber="110">Zahzad 82261</emphasis>
(
<collectionCode id="ED2FAE3EFF859C65DEA7D0C1FF55A6B2" box="[144,214,700,725]" country="Taiwan" name="Taiwan Agricultural Research Institute" pageId="9" pageNumber="110">TARI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B563CA6FF859C65DEDED0C1FB77A6B2" box="[233,1268,700,725]" collectionCode="FUMH" collectorName="W Bojnurd &amp; Rafeie &amp; Zangooie" country="Iran" elevation="1700" location="Almeh" pageId="9" pageNumber="110" specimenCount="1" stateProvince="Khorasan-e Shomali">
Prov.
<collectingRegion id="49FAF819FF859C65DF1DD0C1FD97A6B2" box="[298,532,700,725]" country="Iran" name="Khorasan-e Shomali" pageId="9" pageNumber="110">Khorasan-e Shomali</collectingRegion>
:
<collectorName id="26CB532DFF859C65DC2AD0C1FD1EA6B2" box="[541,669,700,725]" pageId="9" pageNumber="110">W Bojnurd</collectorName>
,
<location id="8EE16020FF859C65DC91D0C1FD76A6B2" LSID="urn:lsid:plazi:treatment:039787EDFF8B9C65DEBFD553FCDDA0CE:8EE16020FF859C65DC91D0C1FD76A6B2" box="[678,757,700,725]" country="Iran" name="Almeh" pageId="9" pageNumber="110" stateProvince="Khorasan-e Shomali">Almeh</location>
,
<quantity id="4CC69B1EFF859C65DD36D0C1FCDBA6B2" box="[769,856,700,725]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="110" unit="m" value="1700.0">
<elevation id="0013D1C8FF859C65DD36D0C1FCDBA6B2" box="[769,856,700,725]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="110" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<emphasis id="B94AEAE9FF859C65DD54D0C1FB05A6B2" box="[867,1158,700,725]" italics="true" pageId="9" pageNumber="110">
<collectorName id="26CB532DFF859C65DD54D0C1FC28A6B2" box="[867,939,700,725]" pageId="9" pageNumber="110">Rafeie</collectorName>
&amp;
<collectorName id="26CB532DFF859C65DDF8D0C1FBBAA6B2" box="[975,1081,700,725]" pageId="9" pageNumber="110">Zangooie</collectorName>
29607
</emphasis>
(
<collectionCode id="ED2FAE3EFF859C65DAA3D0C1FB68A6B2" box="[1172,1259,700,725]" country="Iran" lsid="urn:lsid:biocol.org:col:15384" name="Ferdowsi University" pageId="9" pageNumber="110" type="Herbarium">FUMH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3246570FF859C65DE8AD09DFCDDA0CE" pageId="9" pageNumber="110" type="distribution">
<paragraph id="8B8136FBFF859C65DE8AD09DFEA9A7CA" blockId="9.[136,1452,160,1193]" pageId="9" pageNumber="110">
<emphasis id="B94AEAE9FF859C65DE8AD09DFE6DA69E" bold="true" box="[189,494,736,761]" pageId="9" pageNumber="110">Distribution and habitat:</emphasis>
<taxonomicName id="4C3E4D78FF859C65DC3DD09DFC97A69E" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[522,788,736,761]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="golestanica">
<emphasis id="B94AEAE9FF859C65DC3DD09DFC97A69E" box="[522,788,736,761]" italics="true" pageId="9" pageNumber="110">Cephalaria golestanica</emphasis>
</taxonomicName>
is a rare endemic to NE
<collectingCountry id="F329766BFF859C65DA0ED09DFBEAA69E" box="[1081,1129,736,761]" name="Iran" pageId="9" pageNumber="110">Iran</collectingCountry>
, an Irano-Turanian element, and is considered threatened. It is known only from mountain meadows with grasses, thorn-cushions and
<taxonomicName id="4C3E4D78FF859C65DB79D179FE9AA726" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="macranthera">
<emphasis id="B94AEAE9FF859C65DB79D179FE9AA726" italics="true" pageId="9" pageNumber="110">Quercus macranthera</emphasis>
</taxonomicName>
forest around Almeh,
<collectingRegion id="49FAF819FF859C65DC29D155FD01A726" box="[542,642,808,833]" country="Iran" name="Golestan" pageId="9" pageNumber="110">Golestan</collectingRegion>
National Park in
<collectingRegion id="49FAF819FF859C65DD67D155FBA1A726" box="[848,1058,808,833]" country="Iran" name="Golestan" pageId="9" pageNumber="110">Golestan province</collectingRegion>
, within the boundary of
<collectingRegion id="49FAF819FF859C65DB77D155FA28A726" box="[1344,1451,808,833]" country="Iran" name="Khorasan-e Shomali" pageId="9" pageNumber="110">Khorasan</collectingRegion>
and
<collectingRegion id="49FAF819FF859C65DE8CD131FE9CA702" box="[187,287,844,869]" country="Iran" name="Golestan" pageId="9" pageNumber="110">Golestan</collectingRegion>
provinces. This area is a transitional zone between two completely different climates, i.e. temperate sub-humid and cold-arid (
<bibRefCitation id="EFAF4B0AFF859C65DF9CD10DFDC0A7EE" author="Akhani, H." box="[427,579,880,905]" pageId="9" pageNumber="110" pagination="1 - 412" refId="ref21041" refString="Akhani, H. (1998) Plant biodiversity of Golestan National Park, Iran. Stapfia 53: 1 - 412." type="journal article" year="1998">Akhani 1998</bibRefCitation>
).
<taxonomicName id="4C3E4D78FF859C65DC60D10DFCDCA7EE" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[599,863,880,905]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="golestanica">
<emphasis id="B94AEAE9FF859C65DC60D10DFCDCA7EE" box="[599,863,880,905]" italics="true" pageId="9" pageNumber="110">Cephalaria golestanica</emphasis>
</taxonomicName>
grows on agricultural land as weed, at elevations of
<quantity id="4CC69B1EFF859C65DEBFD1E9FEA5A7CA" box="[136,294,916,941]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="9" pageNumber="110" unit="m" value="1950.0" valueMax="2200.0" valueMin="1700.0">17002200 m</quantity>
.
</paragraph>
<paragraph id="8B8136FBFF859C65DE8AD1C5FCDDA0CE" blockId="9.[136,1452,160,1193]" pageId="9" pageNumber="110">
<emphasis id="B94AEAE9FF859C65DE8AD1C5FE13A7B6" bold="true" box="[189,400,952,977]" pageId="9" pageNumber="110">Taxonomic notes:</emphasis>
—The species is similar to
<taxonomicName id="4C3E4D78FF859C65DCF0D1C5FC64A7B6" authorityName="Boissier" authorityYear="1856" box="[711,999,952,977]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="microcephala">
<emphasis id="B94AEAE9FF859C65DCF0D1C5FC64A7B6" box="[711,999,952,977]" italics="true" pageId="9" pageNumber="110">Cephalaria microcephala</emphasis>
</taxonomicName>
in the shape of lower leaf teeth, middle leaf teeth angle with reference to midvein, upper leaf margin, involucral bracts shape and shape and size of involucel, but it differs by its height (35 vs.
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), leaves pilose to hairy on both surfaces (vs. glabrous to pubescent), peduncle length (up to 12 vs. up to
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long) and receptacular bracts in 5 series (vs. 6), straw-coloured dorsally at base, blackish and attenuate at apex (vs. straw), pilose or sericeous (vs. velvety-hairy). Furthermore,
<taxonomicName id="4C3E4D78FF859C65DA8ED635FAD8A006" authorityName="Ranjbar &amp; Z. Ranjbar" authorityYear="2017" box="[1209,1371,1096,1121]" class="Magnoliopsida" family="Caprifoliaceae" genus="Cephalaria" kingdom="Plantae" order="Dipsacales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="golestanica">
<emphasis id="B94AEAE9FF859C65DA8ED635FB4FA006" box="[1209,1228,1096,1121]" italics="true" pageId="9" pageNumber="110">C</emphasis>
.
<emphasis id="B94AEAE9FF859C65DAEDD635FAD8A006" box="[1242,1371,1096,1121]" italics="true" pageId="9" pageNumber="110">golestanica</emphasis>
</taxonomicName>
differs from other related species by greyish indumentum, leaves coriaceous, verticillate (with 6 leaves in each node), greyish-green, pilose on both surfaces, with distinct vagina (see
<tableCitation id="C6BC0340FF859C65DCCCD6EDFCD2A0CE" box="[763,849,1168,1193]" captionStart="TABLE 4" captionStartId="33.[136,224,334,357]" captionText="TABLE 4. Morphological comparison of Cephalaria axillaris, C. golestanica, C. integerrima, C. microcephala, C. subindivisa, C. sublanata and C. torbatejamensis." pageId="9" pageNumber="110">Table 4</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>