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<document id="9AA677A3431DCE5D56E7E375DDF88A0C" ID-DOI="10.1080/00222933.2018.1515381" ID-ISSN="1464-5262" ID-Zenodo-Dep="5177265" ID-ZooBank="C01D7920-61AD-4651-8861-9D76C3CD86F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628614838824" checkinUser="felipe" docAuthor="Gong, Lu &amp; Chen, Hongwei" docDate="2018" docId="039787CDFFDF06093F84FDD6FB4BFDCF" docLanguage="en" docName="JNatHist.52.31-32.2097-2119.pdf" docOrigin="Journal of Natural History 52 (31 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2018.1515381" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Pseudostegana latiparma" docType="key" docVersion="3" lastPageNumber="2102" masterDocId="FFAEFFB5FFDA060F3E59FFE9FFC8FFFC" masterDocTitle="Seven new species of the genus Pseudostegana (Diptera: Drosophilidae) from South-west China, with DNA barcoding information" masterLastPageNumber="2119" masterPageNumber="2097" pageNumber="2101" updateTime="1699047551353" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="ED28CC8331DDBB09E6CEC59D4CB468C1">Seven new species of the genus Pseudostegana (Diptera: Drosophilidae) from South-west China, with DNA barcoding information</mods:title>
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The
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species group
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<emphasis id="B94AEAC9FFDF060A3EF9FD9BFE7FFD77" box="[160,439,626,651]" italics="true" pageId="5" pageNumber="2101">Pseudostegana latiparma</emphasis>
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species group:
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.
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<emphasis id="B94AEAC9FFDF060A3EF9FD51FEDCFD2D" bold="true" box="[160,276,696,721]" italics="true" pageId="5" pageNumber="2101">Diagnosis</emphasis>
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Wing basally with distinct, cross band (
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(c, i)).
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Key to species of the
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<emphasis id="B94AEAC9FFDF060A3FF1FCDBFDD3FCB7" bold="true" box="[424,539,818,843]" italics="true" pageId="5" pageNumber="2101">latiparma</emphasis>
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group from
<collectingCountry id="F329764BFFDF060A3CEEFCDAFD34FCB7" box="[695,764,819,843]" name="China" pageId="5" pageNumber="2101">China</collectingCountry>
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<key id="FFAF61EAFFDF06093EF9FC8EFB4BFDCF" lastPageId="6" lastPageNumber="2102" pageId="5" pageNumber="2101">
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<keyLead id="30CF95EEFFDF060A3EF9FC8EFB4BFC63" pageId="5" pageNumber="2101">
<keyStep id="30CA2E7EFFDF060A3EF9FC8EFB4BFC63" pageId="5" pageNumber="2101">1. Palpus expanded, medially one-third to half as wide as long......................................... 2 Palpus slender, rod shaped.............................................................................................................. 4</keyStep>
</keyLead>
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<keyLead id="30CF95EEFFDF060A3EF9FC56FB4BFBA3" pageId="5" pageNumber="2101">
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2. Paramere apically shallowly bifurcated, subapically lacking small projection, dorsomedially slightly roundly protruding in lateral view (
<bibRefCitation id="EFAF4B2AFFDF060A3D65FC0BFC33FC06" author="Chen HW &amp; Toda MJ &amp; Wang BC" box="[828,1019,994,1018]" pageId="5" pageNumber="2101" pagination="407 - 442" refId="ref8858" refString="Chen HW, Toda MJ, Wang BC. 2005. A taxonomic revision of the genus Pseudostegana Okada, 1978 (Diptera: Drosophilidae). Insect Syst Evol. 36: 407 - 442." type="journal article" year="2005">Chen et al. 2005</bibRefCitation>
, figs 95, 96) .......................................................................
<taxonomicName id="4C3E4D58FFDF060A3C30FBEDFC5BFBE1" ID-CoL="4PC23" authority="Chen and Wang" authorityName="Chen and Wang" authorityYear="2005" box="[617,915,1028,1053]" class="Insecta" family="Drosophilidae" genus="Pseudostegana" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="2101" phylum="Arthropoda" rank="species" species="bifasciata">
<emphasis id="B94AEAC9FFDF060A3C30FBEDFD1AFBE1" box="[617,722,1028,1053]" italics="true" pageId="5" pageNumber="2101">bifasciata</emphasis>
Chen and Wang
</taxonomicName>
in
<bibRefCitation id="EFAF4B2AFFDF060A3DE3FBECFB4BFBE1" author="Chen HW &amp; Toda MJ &amp; Wang BC" box="[954,1155,1029,1053]" pageId="5" pageNumber="2101" pagination="407 - 442" refId="ref8858" refString="Chen HW, Toda MJ, Wang BC. 2005. A taxonomic revision of the genus Pseudostegana Okada, 1978 (Diptera: Drosophilidae). Insect Syst Evol. 36: 407 - 442." type="journal article" year="2005">Chen et al. (2005)</bibRefCitation>
Paramere apically not bifurcated, subapically with one small, acute projection, dorsomedially triangularly protruding in lateral view.......................................................... 3
</keyStep>
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<keyLead id="30CF95EEFFDF060A3EF9FB96FB4CFA98" pageId="5" pageNumber="2101">
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3. Ocellar triangle black to brown, broadly elongated to anterior margin of frons (
<bibRefCitation id="EFAF4B2AFFDF060A3A29FB96FE9FFB46" author="Li T &amp; Gao JJ &amp; Chen HW" pageId="5" pageNumber="2101" pagination="1401 - 1418" refId="ref9015" refString="Li T, Gao JJ, Chen HW. 2010. Six new species of the genus Pseudostegana (Diptera: Drosophilidae) from the Oriental Region. J Nat Hist. 44: 1401 - 1418." type="journal article" year="2010">Li et al. 2010</bibRefCitation>
, fig. 1A); scutum yellow on anterior two-thirds, brownish black on posterior one-third; aedeagus gradually expanded from median to base (
<bibRefCitation id="EFAF4B2AFFDF060A3A73FB2CFEC3FB03" author="Li T &amp; Gao JJ &amp; Chen HW" pageId="5" pageNumber="2101" pagination="1401 - 1418" refId="ref9015" refString="Li T, Gao JJ, Chen HW. 2010. Six new species of the genus Pseudostegana (Diptera: Drosophilidae) from the Oriental Region. J Nat Hist. 44: 1401 - 1418." type="journal article" year="2010">Li et al. 2010</bibRefCitation>
, fig. 3D)..................................................................
<taxonomicName id="4C3E4D58FFDF060A3CB8FB0FFEE8FAE3" ID-CoL="9HK6F" authority="Li, Gao and Chen, 2010 Ocellar" authorityName="Li, Gao and Chen" authorityYear="2010" class="Insecta" family="Drosophilidae" genus="Pseudostegana" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="2101" phylum="Arthropoda" rank="species" species="acutifoliolata">
<emphasis id="B94AEAC9FFDF060A3CB8FB0FFCB9FB03" box="[737,881,1254,1279]" italics="true" pageId="5" pageNumber="2101">acutifoliolata</emphasis>
<bibRefCitation id="EFAF4B2AFFDF060A3D23FB0EFB4CFB03" author="Li T &amp; Gao JJ &amp; Chen HW" box="[890,1156,1255,1279]" pageId="5" pageNumber="2101" pagination="1401 - 1418" refId="ref9015" refString="Li T, Gao JJ, Chen HW. 2010. Six new species of the genus Pseudostegana (Diptera: Drosophilidae) from the Oriental Region. J Nat Hist. 44: 1401 - 1418." type="journal article" year="2010">Li, Gao and Chen, 2010</bibRefCitation>
Ocellar
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triangle brownish yellow, narrowly elongated to anterior margin of frons (
<figureCitation id="13052A5EFFDF060A3E81FAC3FEF2FABE" box="[216,314,1322,1346]" captionStart="Figure 3" captionStartId="8.[160,225,121,144]" captionText="Figure 3. Head, thorax, wing and abdomen of male: (af) Pseudostegana latiapicula sp. nov.; (gl) P. nannanae sp. nov." pageId="5" pageNumber="2101">Figure 3</figureCitation>
(j)); scutum nearly yellow (
<figureCitation id="13052A5EFFDF060A3C35FAC3FD06FABE" box="[620,718,1322,1346]" captionStart="Figure 3" captionStartId="8.[160,225,121,144]" captionText="Figure 3. Head, thorax, wing and abdomen of male: (af) Pseudostegana latiapicula sp. nov.; (gl) P. nannanae sp. nov." pageId="5" pageNumber="2101">Figure 3</figureCitation>
(h,k)); aedeagus medially expanded in ventral view (
<figureCitation id="13052A5EFFDF060A3F32FAA5FE00FA98" box="[363,456,1356,1380]" captionStart="Figure 7" captionStartId="12.[160,225,121,144]" captionText="Figure 7. Pseudostegana nannanae sp. nov., male terminalia. (a) epandrium, cercus and surstylus; (b, c) hypandrium, paramere, gonopods, aedeagus and aedeagal apodeme (ventral and lateral views). Scale bars: 0.1 mm." pageId="5" pageNumber="2101">Figure 7</figureCitation>
(b))....................................... .......................................
<taxonomicName id="4C3E4D58FFDF060A3DEDFAA2FBEBFA98" authority="Gong &amp; Chen, 2018" authorityName="Gong &amp; Chen" authorityYear="2018" box="[948,1059,1355,1380]" class="Insecta" family="Drosophilidae" genus="Pseudostegana" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="2101" phylum="Arthropoda" rank="species" species="nannanae" status="sp. nov.">
<emphasis id="B94AEAC9FFDF060A3DEDFAA2FBEBFA98" box="[948,1059,1355,1380]" italics="true" pageId="5" pageNumber="2101">nannanae</emphasis>
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<taxonomicNameLabel id="A27957B2FFDF060A3A72FAA5FB4CFA98" box="[1067,1156,1356,1380]" pageId="5" pageNumber="2101" rank="species">sp. nov.</taxonomicNameLabel>
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4. R-m crossvein clear; medial, dark-coloured band neither broadened nor with protruding part............................................................................................................................................. 5 R-m crossvein clouded; medial, dark-coloured band much broadened and with one distinct, protruding part submedially (
<bibRefCitation id="EFAF4B2AFFDF060A3CDAFA03FCD0F9FE" author="Li T &amp; Gao JJ &amp; Chen HW" box="[643,792,1514,1538]" pageId="5" pageNumber="2101" pagination="1401 - 1418" refId="ref9015" refString="Li T, Gao JJ, Chen HW. 2010. Six new species of the genus Pseudostegana (Diptera: Drosophilidae) from the Oriental Region. J Nat Hist. 44: 1401 - 1418." type="journal article" year="2010">Li et al. 2010</bibRefCitation>
, fig. 2B)................................................. .......................................................................................................
<taxonomicName id="4C3E4D58FFDF060A3D4FF9E2FB4CF9D8" ID-CoL="9HK6K" authority="Li, Gao and Chen, 2010" authorityName="Li, Gao and Chen" authorityYear="2010" box="[790,1156,1547,1572]" class="Insecta" family="Drosophilidae" genus="Pseudostegana" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="2101" phylum="Arthropoda" rank="species" species="bilobata">
<emphasis id="B94AEAC9FFDF060A3D4FF9E2FCB8F9D8" box="[790,880,1547,1572]" italics="true" pageId="5" pageNumber="2101">bilobata</emphasis>
<bibRefCitation id="EFAF4B2AFFDF060A3D23F9E5FB4CF9D8" author="Li T &amp; Gao JJ &amp; Chen HW" box="[890,1156,1548,1572]" pageId="5" pageNumber="2101" pagination="1401 - 1418" refId="ref9015" refString="Li T, Gao JJ, Chen HW. 2010. Six new species of the genus Pseudostegana (Diptera: Drosophilidae) from the Oriental Region. J Nat Hist. 44: 1401 - 1418." type="journal article" year="2010">Li, Gao and Chen, 2010</bibRefCitation>
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<paragraph id="8B8136DBFFDF060A3EF9F9ADFB4BF963" blockId="5.[160,1156,1604,1695]" pageId="5" pageNumber="2101">
<keyLead id="30CF95EEFFDF060A3EF9F9ADFB4BF963" pageId="5" pageNumber="2101">
<keyStep id="30CA2E7EFFDF060A3EF9F9ADFB4BF963" pageId="5" pageNumber="2101">5. Paramere apically with expanded lobe-like process and strongly sclerotized, acute projection................................................................................................................................................. 6 Paramere apically pointed, lacking lobe-like process........................................................... 7</keyStep>
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<paragraph id="8B8136DBFFDC06093EF9FF67FB4BFEAD" blockId="6.[160,1156,142,337]" pageId="6" pageNumber="2102">
<keyLead id="30CF95EEFFDC06093EF9FF67FB4BFEAD" pageId="6" pageNumber="2102">
<keyStep id="30CA2E7EFFDC06093EF9FF67FB4BFEAD" pageId="6" pageNumber="2102">
6. Paramere subapically triangularly expanded in ventral view (
<figureCitation id="13052A5EFFDC06093DDDFF67FC2CFF5A" box="[900,996,142,166]" captionStart="Figure 6" captionStartId="11.[160,225,121,144]" captionText="Figure 6. Pseudostegana latiapicula sp. nov., male terminalia. (a) epandrium (epan), cercus (cerc) and surstylus (sur) (lateral view); (b, c) hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed) and aedeagal apodeme (aed a) (ventral and lateral views). Scale bars: 0.1 mm." pageId="6" pageNumber="2102">Figure 6</figureCitation>
(b)); aedeagus with slender basal processes, distally roundly expanded in lateral view (
<figureCitation id="13052A5EFFDC06093A5BFF58FBA8FF35" box="[1026,1120,177,201]" captionStart="Figure 6" captionStartId="11.[160,225,121,144]" captionText="Figure 6. Pseudostegana latiapicula sp. nov., male terminalia. (a) epandrium (epan), cercus (cerc) and surstylus (sur) (lateral view); (b, c) hypandrium (hypd), paramere (pm), gonopods (gon), aedeagus (aed) and aedeagal apodeme (aed a) (ventral and lateral views). Scale bars: 0.1 mm." pageId="6" pageNumber="2102">Figure 6</figureCitation>
(c)) ...................................................................................................................................
<taxonomicName id="4C3E4D58FFDC06093DE8FF3AFBEBFF10" authority="Gong &amp; Chen, 2018" authorityName="Gong &amp; Chen" authorityYear="2018" box="[945,1059,211,236]" class="Insecta" family="Drosophilidae" genus="Pseudostegana" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="2102" phylum="Arthropoda" rank="species" species="latiapicula" status="sp. nov.">
<emphasis id="B94AEAC9FFDC06093DE8FF3AFBEBFF10" box="[945,1059,211,236]" italics="true" pageId="6" pageNumber="2102">latiapicula</emphasis>
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Paramere subapically roundly expanded in ventral view (
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, fig. 6D); aedeagus with lobe-like basal processes, distally narrowed, apically hammer-like in lateral view (
<bibRefCitation id="EFAF4B2AFFDC06093F3DFED0FE31FEAD" author="Li T &amp; Gao JJ &amp; Chen HW" box="[356,505,313,337]" pageId="6" pageNumber="2102" pagination="1401 - 1418" refId="ref9015" refString="Li T, Gao JJ, Chen HW. 2010. Six new species of the genus Pseudostegana (Diptera: Drosophilidae) from the Oriental Region. J Nat Hist. 44: 1401 - 1418." type="journal article" year="2010">Li et al. 2010</bibRefCitation>
, fig. 6E).......... ..........
<taxonomicName id="4C3E4D58FFDC06093C8FFED1FB4BFEAD" ID-CoL="9HK6Q" authority="Li, Gao and Chen, 2010" authorityName="Li, Gao and Chen" authorityYear="2010" box="[726,1155,312,337]" class="Insecta" family="Drosophilidae" genus="Pseudostegana" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="2102" phylum="Arthropoda" rank="species" species="minutipalpata">
<emphasis id="B94AEAC9FFDC06093C8FFED1FCB9FEAD" box="[726,881,312,337]" italics="true" pageId="6" pageNumber="2102">minutipalpata</emphasis>
<bibRefCitation id="EFAF4B2AFFDC06093D23FED0FB4BFEAD" author="Li T &amp; Gao JJ &amp; Chen HW" box="[890,1155,313,337]" pageId="6" pageNumber="2102" pagination="1401 - 1418" refId="ref9015" refString="Li T, Gao JJ, Chen HW. 2010. Six new species of the genus Pseudostegana (Diptera: Drosophilidae) from the Oriental Region. J Nat Hist. 44: 1401 - 1418." type="journal article" year="2010">Li, Gao and Chen, 2010</bibRefCitation>
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7. Paramere subapically triangularly expanded in ventral view (
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, fig. 92); basal process of aedeagus membranous, lacking finger-like processes (
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, fig. 93)................... ..................
<taxonomicName id="4C3E4D58FFDC06093C61FE5CFC5FFE32" ID-CoL="4PBZZ" authority="Chen and Wang" authorityName="Chen and Wang" authorityYear="2005" box="[568,919,437,462]" class="Insecta" family="Drosophilidae" genus="Pseudostegana" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="2102" phylum="Arthropoda" rank="species" species="angustifasciata">
<emphasis id="B94AEAC9FFDC06093C61FE5CFD12FE32" box="[568,730,437,462]" italics="true" pageId="6" pageNumber="2102">angustifasciata</emphasis>
Chen and Wang
</taxonomicName>
in
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Paramere subapically roundly expanded in lateral view (
<bibRefCitation id="EFAF4B2AFFDC06093D0CFE3FFBDDFE12" author="Chen HW &amp; Toda MJ &amp; Wang BC" box="[853,1045,470,494]" pageId="6" pageNumber="2102" pagination="407 - 442" refId="ref8858" refString="Chen HW, Toda MJ, Wang BC. 2005. A taxonomic revision of the genus Pseudostegana Okada, 1978 (Diptera: Drosophilidae). Insect Syst Evol. 36: 407 - 442." type="journal article" year="2005">Chen et al. 2005</bibRefCitation>
, fig. 106); basal process of aedeagus with finger-like processes (
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, fig. 105)....... ............................................................
<emphasis id="B94AEAC9FFDC06093C70FDF3FD1AFDCF" box="[553,722,538,563]" italics="true" pageId="6" pageNumber="2102">pallidimaculata</emphasis>
Chen and Wang in
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