421 lines
66 KiB
XML
421 lines
66 KiB
XML
<document id="CAC15D647F7A7DDFE7FA60D6B84DB565" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFAC84578CFBFDE7DFCBF91D" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes viviparus Kluge, 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="52" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="50" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="A547350BAB064FB1FAA28A268523E78B">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:name id="08FC3AEF8858B7A67765DAB337D252F3" type="personal">
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<mods:roleTerm id="5BD3BD279887D0C229D698B140E50DBD">Author</mods:roleTerm>
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<mods:namePart id="0BBD43EF8AE572239B5855CD2BE6D532">Kluge, Nikita J.</mods:namePart>
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<mods:typeOfResource id="F98268C4167A1BD0426335C2143FE59B">text</mods:typeOfResource>
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<mods:title id="E16F7A4B060672044C1251839AB2F696">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E5A9CF8B521926E6D24C93BCAC17299C">
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<mods:date id="2A1CB982102BB923B748D7159B62BAE6">2020</mods:date>
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<mods:number id="B97439DBAE6E54688BA12A3D3CDB8FA7">2020-03-26</mods:number>
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<mods:number id="59866636D12E48790E8687EA0985CBCD">4756</mods:number>
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<mods:classification id="AD75832926EEB571D0F0031036E48115">journal article</mods:classification>
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<mods:identifier id="B333F1D0D00213D88FAAED3303CDB274" type="DOI">10.11646/zootaxa.4756.1.1</mods:identifier>
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<mods:identifier id="AD9E3D0826FF0797C35BBD57387A2829" type="GBIF-Dataset">4d48d52e-6505-4f62-909f-5da5c2cfb489</mods:identifier>
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<mods:identifier id="626666F08CC7E6E2EFB448C2B4F99665" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="5C7C126CD4406E8034E3545E1E0B610B" type="Zenodo-Dep">3736895</mods:identifier>
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</mods:mods>
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<treatment id="039787A6FFAC84578CFBFDE7DFCBF91D" ID-DOI="http://doi.org/10.5281/zenodo.3811769" ID-GBIF-Taxon="163420789" ID-Zenodo-Dep="3811769" LSID="urn:lsid:plazi:treatment:039787A6FFAC84578CFBFDE7DFCBF91D" httpUri="http://treatment.plazi.org/id/039787A6FFAC84578CFBFDE7DFCBF91D" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50">
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<subSubSection id="C324653BFFAC84558CFBFDE7DAC1FDB9" box="[151,511,513,540]" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="8B8136B0FFAC84558CFBFDE7DAC1FDB9" blockId="49.[151,511,513,575]" box="[151,511,513,540]" pageId="49" pageNumber="50">
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<heading id="D0C981DCFFAC84558CFBFDE7DAC1FDB9" bold="true" box="[151,511,513,540]" fontSize="10" level="2" pageId="49" pageNumber="50" reason="2">
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||
<emphasis id="B94AEAA2FFAC84558CFBFDE7DAC1FDB9" bold="true" box="[151,511,513,540]" pageId="49" pageNumber="50">
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5.
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<taxonomicName id="4C3E4D33FFAC84558CD9FDE7DA8BFDB9" box="[181,437,513,540]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
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<emphasis id="B94AEAA2FFAC84558CD9FDE7DA8BFDB9" bold="true" box="[181,437,513,540]" italics="true" pageId="49" pageNumber="50">Thraulodes viviparus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27957D9FFAC84558DD1FDE7DAC1FDB9" box="[445,511,513,539]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C324653BFFAC84558CFBFDC3DA7DFD9D" box="[151,323,548,575]" pageId="49" pageNumber="50" type="description">
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<paragraph id="8B8136B0FFAC84558CFBFDC3DA7DFD9D" blockId="49.[151,511,513,575]" box="[151,323,548,575]" pageId="49" pageNumber="50">
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||
(
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<figureCitation id="13052A35FFAC84558CF3FDC3DA05FD9D" box="[159,315,548,575]" captionStart-0="FIGURES 151–155" captionStart-1="FIGURES 156–168" captionStart-2="FIGURES 169–173" captionStartId-0="50.[151,264,1474,1499]" captionStartId-1="52.[151,264,1899,1924]" captionStartId-2="53.[151,264,1539,1564]" captionTargetBox-0="[156,1436,166,1455]" captionTargetBox-1="[151,1436,166,1879]" captionTargetBox-2="[164,1425,173,1511]" captionTargetId-0="figure@50.[151,1436,166,1455]" captionTargetId-1="figure@52.[151,1436,166,1879]" captionTargetId-2="figure@53.[151,1436,166,1520]" captionTargetPageId-0="50" captionTargetPageId-1="52" captionTargetPageId-2="53" captionText-0="FIGURES 151–155. Larva presumably attributed to Thraulodes viviparus sp. n. 151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings)." captionText-1="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." captionText-2="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi-0="http://doi.org/10.5281/zenodo.3736941" figureDoi-1="http://doi.org/10.5281/zenodo.3736943" figureDoi-2="http://doi.org/10.5281/zenodo.3736945" httpUri-0="https://zenodo.org/record/3736941/files/figure.png" httpUri-1="https://zenodo.org/record/3736943/files/figure.png" httpUri-2="https://zenodo.org/record/3736945/files/figure.png" pageId="49" pageNumber="50">Figs 151–173</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C324653BFFAC84558CFBFD80D822FD22" box="[151,796,614,641]" pageId="49" pageNumber="50" type="etymology">
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<paragraph id="8B8136B0FFAC84558CFBFD80D822FD22" blockId="49.[151,1437,614,2044]" box="[151,796,614,641]" pageId="49" pageNumber="50">
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<emphasis id="B94AEAA2FFAC84558CFBFD80DA20FD22" bold="true" box="[151,286,614,640]" pageId="49" pageNumber="50">Etymology.</emphasis>
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This species is assumed to be ovoviviparous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C324653BFFAC84558CABFD6FDE4DFD4F" pageId="49" pageNumber="50" type="materials_examined">
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<paragraph id="8B8136B0FFAC84558CABFD6FDE4DFD4F" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
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<emphasis id="B94AEAA2FFAC84558CABFD6FDA91FD06" bold="true" box="[199,431,649,676]" pageId="49" pageNumber="50">Material examined.</emphasis>
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<materialsCitation id="3B563CEDFFAC84558DD8FD6CDEA3FD6A" ID-GBIF-Occurrence="2594509921" collectingDate="2018-01-29" collectionCode="XIV" collectorName="N. Kluge & L. Sheyko." country="Panama" county="Bosque" latitude="8.793612" location="Altos del Valle" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Provincia Bocas del Toro" typeStatus="holotype">
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<emphasis id="B94AEAA2FFAC84558DD8FD6CD914FD06" bold="true" box="[436,554,650,676]" pageId="49" pageNumber="50">
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<typeStatus id="54858812FFAC84558DD8FD6CD91BFD06" box="[436,549,650,676]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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S-I
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<specimenCount id="9D38FD39FFAC84558E3FFD6CD958FD06" box="[595,614,650,676]" pageId="49" pageNumber="50" type="male">♂</specimenCount>
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||
{specimen [
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<collectionCode id="ED2FAE75FFAC84558E82FD6FD819FD06" box="[750,807,649,676]" pageId="49" pageNumber="50">XIV</collectionCode>
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](B)2013}:
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||
<collectingCountry id="F3297620FFAC84558FCFFD6CDF20FD06" box="[931,1054,650,676]" name="Panama" pageId="49" pageNumber="50">PANAMA</collectingCountry>
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||
:
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||
<collectingRegion id="49FAF852FFAC8455884BFD6CDE01FD07" box="[1063,1343,650,677]" country="Panama" name="Bocas del Toro" pageId="49" pageNumber="50">Provincia Bocas del Toro</collectingRegion>
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||
,
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<collectingCounty id="62E04E3CFFAC84558924FD6DDEA2FD07" box="[1352,1436,651,677]" pageId="49" pageNumber="50">Bosque</collectingCounty>
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protector
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<collectingMunicipality id="6BE5ACCAFFAC84558D6FFD48DA4BFD6A" box="[259,373,686,712]" pageId="49" pageNumber="50">Palo Seco</collectingMunicipality>
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||
,
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<location id="8EE1606BFFAC84558D13FD48D919FD6A" LSID="urn:lsid:plazi:treatment:039787A6FFAC84578CFBFDE7DFCBF91D:8EE1606BFFAC84558D13FD48D919FD6A" box="[383,551,686,712]" country="Panama" county="Bosque" latitude="8.793612" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" name="Altos del Valle" pageId="49" pageNumber="50" stateProvince="Provincia Bocas del Toro">Altos del Valle</location>
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||
,
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||
<location id="8EE1606BFFAC84558E5FFD48D9A1FD6A" LSID="urn:lsid:plazi:treatment:039787A6FFAC84578CFBFDE7DFCBF91D:8EE1606BFFAC84558E5FFD48D9A1FD6A" box="[563,671,686,712]" country="Panama" county="Bosque" latitude="8.793612" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" name="Rio Buris" pageId="49" pageNumber="50" stateProvince="Provincia Bocas del Toro">Rio Buris</location>
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||
(
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<geoCoordinate id="EE0A5077FFAC84558EC3FD48D80EFD6A" box="[687,816,686,712]" degrees="8" direction="north" minutes="47" orientation="latitude" pageId="49" pageNumber="50" precision="15" seconds="37" value="8.793612">8°47’37’’N</geoCoordinate>
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,
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<geoCoordinate id="EE0A5077FFAC84558F51FD4BD8EFFD6A" box="[829,977,685,712]" degrees="82" direction="west" minutes="11" orientation="longitude" pageId="49" pageNumber="50" precision="15" seconds="35" value="-82.193054">82°11’35’’W</geoCoordinate>
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),
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||
<date id="FF801070FFAC84558F8AFD48DF6BFD6A" box="[998,1109,686,712]" pageId="49" pageNumber="50" value="2018-01-29">
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<collectingDate id="EFC4E998FFAC84558F8AFD48DF6BFD6A" box="[998,1109,686,712]" pageId="49" pageNumber="50" value="2018-01-29">29.I.2018</collectingDate>
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</date>
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, coll.
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<collectorName id="26CB5366FFAC845588F4FD49DFC0FD6A" box="[1176,1278,686,713]" pageId="49" pageNumber="50">N. Kluge</collectorName>
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&
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<collectorName id="26CB5366FFAC8455894EFD49DEA3FD6A" box="[1314,1437,686,713]" pageId="49" pageNumber="50">L. Sheyko.</collectorName>
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</materialsCitation>
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||
<materialsCitation id="3B563CEDFFAC84558CFBFD34DE4DFD4F" ID-GBIF-Occurrence="2594509900" box="[151,1395,722,749]" collectingDate="2018-01-24" collectingDateMax="2018-01-28" collectingDateMin="2018-01-24" collectionCode="XIV" collectorName="N. Kluge & L. Sheyko." country="Panama" county="Bosque" latitude="8.793612" location="Altos del Valle" longLatPrecision="21" longitude="-82.193054" municipality="Palo Seco" pageId="49" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-larva="1" specimenCount-male="1" stateProvince="Provincia Bocas del Toro" typeStatus="paratype">
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<emphasis id="B94AEAA2FFAC84558CFBFD34DA26FD4E" bold="true" box="[151,280,722,748]" pageId="49" pageNumber="50">
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<typeStatus id="54858812FFAC84558CFBFD34DA2DFD4E" box="[151,275,722,748]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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the same locality and collectors,
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<date id="FF801070FFAC84558EE2FD34D9C3FD4E" box="[654,765,722,748]" pageId="49" pageNumber="50" value="2018-01-26">
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<collectingDate id="EFC4E998FFAC84558EE2FD34D9C3FD4E" box="[654,765,722,748]" pageId="49" pageNumber="50" value="2018-01-26">26.I.2018</collectingDate>
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</date>
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: 1 S-I
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<specimenCount id="9D38FD39FFAC84558F2BFD34D869FD4E" box="[839,855,722,748]" pageId="49" pageNumber="50" type="female">♀</specimenCount>
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;
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<date id="FF801070FFAC84558F0FFD34D8C3FD4E" box="[867,1021,722,748]" pageId="49" pageNumber="50" value="2018-01-24" valueMax="2018-01-28" valueMin="2018-01-24">
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||
<collectingDate id="EFC4E998FFAC84558F0FFD34D8C3FD4E" box="[867,1021,722,748]" pageId="49" pageNumber="50" value="2018-01-24" valueMax="2018-01-28" valueMin="2018-01-24">24–28.I.2018</collectingDate>
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</date>
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||
:
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<specimenCount id="9D38FD39FFAC84558864FD34DF6BFD4E" box="[1032,1109,722,748]" pageId="49" pageNumber="50" type="larva">1 larva</specimenCount>
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(presumably associated).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C324653BFFAC84578CABFD13D8CBFBB1" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" type="description">
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<paragraph id="8B8136B0FFAC84558CABFD13D9A5FCDE" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
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||
<emphasis id="B94AEAA2FFAC84558CABFD13DA8EFCB3" bold="true" box="[199,432,757,785]" pageId="49" pageNumber="50">
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||
Descriptions.
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||
<emphasis id="B94AEAA2FFAC84558D00FD11DA8EFCB3" bold="true" box="[364,432,759,785]" italics="true" pageId="49" pageNumber="50">Larva</emphasis>
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||
</emphasis>
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||
(presumably associated). CUTICULAR COLORATION (
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<figureCitation id="13052A35FFAC84558829FD10DF94FCB2" box="[1093,1194,757,784]" captionStart="FIGURES 151–155" captionStartId="50.[151,264,1474,1499]" captionTargetBox="[156,1436,166,1455]" captionTargetId="figure@50.[151,1436,166,1455]" captionTargetPageId="50" captionText="FIGURES 151–155. Larva presumably attributed to Thraulodes viviparus sp. n. 151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings)." figureDoi="http://doi.org/10.5281/zenodo.3736941" httpUri="https://zenodo.org/record/3736941/files/figure.png" pageId="49" pageNumber="50">Fig. 154</figureCitation>
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): Dorsal side of head dark brown. Dorsal side of thorax mostly dark brown with small contrasting blanks. Abdominal terga brown, with blanks near tergalii bases, terga IV, V, VIII and IX with contrasting unpaired blanks medially. Femora brown with blanks; tibiae and tarsi uniformly light brown.
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</paragraph>
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<paragraph id="8B8136B0FFAC84558CABFC60D8BCFC4A" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
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HYPODERMAL COLORATION.Abdominal terga III–VI, besides cuticular coloration (see above) with pair of transverse blackish spots (corresponding to midway gray spots of imago) and blackish spiracular spots (
|
||
<figureCitation id="13052A35FFAC84558941FC4CDEAEFC66" box="[1325,1424,937,964]" captionStart="FIGURES 151–155" captionStartId="50.[151,264,1474,1499]" captionTargetBox="[156,1436,166,1455]" captionTargetId="figure@50.[151,1436,166,1455]" captionTargetPageId="50" captionText="FIGURES 151–155. Larva presumably attributed to Thraulodes viviparus sp. n. 151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings)." figureDoi="http://doi.org/10.5281/zenodo.3736941" httpUri="https://zenodo.org/record/3736941/files/figure.png" pageId="49" pageNumber="50">Fig. 155</figureCitation>
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||
). Tergalii brownish with costal areas gray, tracheae black (
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||
<figureCitation id="13052A35FFAC84558F7FFC28D848FC4A" box="[787,886,973,1000]" captionStart="FIGURES 151–155" captionStartId="50.[151,264,1474,1499]" captionTargetBox="[156,1436,166,1455]" captionTargetId="figure@50.[151,1436,166,1455]" captionTargetPageId="50" captionText="FIGURES 151–155. Larva presumably attributed to Thraulodes viviparus sp. n. 151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings)." figureDoi="http://doi.org/10.5281/zenodo.3736941" httpUri="https://zenodo.org/record/3736941/files/figure.png" pageId="49" pageNumber="50">Fig. 153</figureCitation>
|
||
).
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||
</paragraph>
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||
<paragraph id="8B8136B0FFAC84558CABFC14DA57FBDB" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
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||
SHAPE AND SETATION. Clypeus widened distally; labrum 1.4 times wider than clypeus. Labrum widest at 1/3 of length from base; initial fore margin (turned ventrally) without median emargination, with all 5 denticles wide (as in
|
||
<figureCitation id="13052A35FFAC84558D75FBDCDA55FBF6" box="[281,363,1082,1108]" captionStart="FIGURES 86–93" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="49" pageNumber="50">Fig. 88</figureCitation>
|
||
); anterior transverse setal row regular, wider than 3 denticles. Maxilla with 18 pectinate setae in apical-ventral row.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABFB65DA95FB46" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
|
||
Femora: Stout setae on anterior surface parallel-sided or narrowed distally, occasionally widened distally, apically rounded or truncate (
|
||
<figureCitation id="13052A35FFAC84558DA5FB40D907FB62" box="[457,569,1189,1216]" captionStart="FIGURES 151–155" captionStartId="50.[151,264,1474,1499]" captionTargetBox="[156,1436,166,1455]" captionTargetId="figure@50.[151,1436,166,1455]" captionTargetPageId="50" captionText="FIGURES 151–155. Larva presumably attributed to Thraulodes viviparus sp. n. 151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings)." figureDoi="http://doi.org/10.5281/zenodo.3736941" httpUri="https://zenodo.org/record/3736941/files/figure.png" pageId="49" pageNumber="50">Figs. 152</figureCitation>
|
||
). Irregular row of hairs near inner margin absent on fore femur, present on middle and hind femora.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABFB09D8B2FAF3" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">Fore tibia: outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inner-anterior row of stout setae absent; inner field of stout pointed setae moderately dense (setae lengths subequal to distances between them), its setae subequal in lengths, forming mostly one row.</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABFABCD82EFA42" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
|
||
Hind tibia (
|
||
<figureCitation id="13052A35FFAC84558D24FABCDA95FAD6" box="[328,427,1369,1396]" captionStart="FIGURES 151–155" captionStartId="50.[151,264,1474,1499]" captionTargetBox="[156,1436,166,1455]" captionTargetId="figure@50.[151,1436,166,1455]" captionTargetPageId="50" captionText="FIGURES 151–155. Larva presumably attributed to Thraulodes viviparus sp. n. 151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings)." figureDoi="http://doi.org/10.5281/zenodo.3736941" httpUri="https://zenodo.org/record/3736941/files/figure.png" pageId="49" pageNumber="50">Fig. 151</figureCitation>
|
||
): both outer-anterior and outer-posterior rows of stout setae consist of various stout setae: short rounded, long spoon-like and intermediate ones; posterior hairs located between these rows, form more than one row (besides row of hairs posteriad of outer-posterior row of stout setae); stout setae of inner-anterior row moderately long, mostly narrowed distally, apically rounded.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABFA0CDFB2F9A6" blockId="49.[151,1437,614,2044]" box="[199,1164,1514,1541]" pageId="49" pageNumber="50">Claws with 6 denticles on rigid portion, with pointed denticles on articulatory portion.</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABF9E8DF8FF98A" blockId="49.[151,1437,614,2044]" box="[199,1201,1549,1576]" pageId="49" pageNumber="50">
|
||
Tergalii (
|
||
<figureCitation id="13052A35FFAC84558D41F9E8DAB1F98A" box="[301,399,1549,1576]" captionStart="FIGURES 151–155" captionStartId="50.[151,264,1474,1499]" captionTargetBox="[156,1436,166,1455]" captionTargetId="figure@50.[151,1436,166,1455]" captionTargetPageId="50" captionText="FIGURES 151–155. Larva presumably attributed to Thraulodes viviparus sp. n. 151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings)." figureDoi="http://doi.org/10.5281/zenodo.3736941" httpUri="https://zenodo.org/record/3736941/files/figure.png" pageId="49" pageNumber="50">Fig. 153</figureCitation>
|
||
): each lamella moderately wide, tracheae with few small side branches.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABF9D4DA5FF936" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
|
||
Male genitalia in last larval instar (
|
||
<figureCitation id="13052A35FFAC84558E3FF9D4D989F9EE" box="[595,695,1586,1612]" captionStart="FIGURES 169–173" captionStartId="53.[151,264,1539,1564]" captionTargetBox="[164,1425,173,1511]" captionTargetId="figure@53.[151,1436,166,1520]" captionTargetPageId="53" captionText="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736945" httpUri="https://zenodo.org/record/3736945/files/figure.png" pageId="49" pageNumber="50">Fig. 172</figureCitation>
|
||
): protogonostyli short and separated one from another by shallow emargination. Each protopenis lobe with small, sharply projected gonopore-bearing process; gonopores opened caudally-laterally.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABF97BDF01F832" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
|
||
<emphasis id="B94AEAA2FFAC84558CABF97BDA02F91A" bold="true" box="[199,316,1693,1720]" italics="true" pageId="49" pageNumber="50">Subimago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFAC84558D26F978DA91F91A" box="[330,431,1693,1720]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="49" pageNumber="50">Figs 165</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFAC84558DD7F978DAD7F91A" box="[443,489,1694,1720]" captionStart="FIGURES 169–173" captionStartId="53.[151,264,1539,1564]" captionTargetBox="[164,1425,173,1511]" captionTargetId="figure@53.[151,1436,166,1520]" captionTargetPageId="53" captionText="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736945" httpUri="https://zenodo.org/record/3736945/files/figure.png" pageId="49" pageNumber="50">170</figureCitation>
|
||
). CUTICULAR COLORATION. Cuticle mostly colorless, with brown areas. Flagellum of antenna brown. Pronotum colorless. Mesonotum mostly colorless, with contrasting brown antelateroparapsidal and lateroparapsidal sutures, anteronotal transverse impression and chromozone of medioscutum; chromozone of submedioscutum nearly colorless (
|
||
<figureCitation id="13052A35FFAC84558E51F8ECD99EF886" box="[573,672,1801,1828]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="49" pageNumber="50">Fig. 165</figureCitation>
|
||
). Thoracic pleura and sterna colorless. On each leg femur colorless, in distal part with inner and outer margins shed with brownish; tibia and tarsus light brown. Cuticle of wings colorless, microtrichia light brownish. Abdominal terga mostly colorless; terga II–IX with diffusive brown median stripe; tergum VI with wider median area diffusively light brownish. Caudalii light brown.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAC84558CABF87DDA1CF85E" blockId="49.[151,1437,614,2044]" pageId="49" pageNumber="50">
|
||
TEXTURE. On tarsi of all legs, 1st tarsomere with microtrichia (as tibia), 2nd–5th tarsomeres coved by blunt microlepides; pointed microlepides present near apical margins of 2nd–4th tarsomeres of middle and hind legs (as in
|
||
<figureCitation id="13052A35FFAC84558CD8F804DA2BF85E" box="[180,277,2018,2044]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="49" pageNumber="50">Fig. 473</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF416638FFAF84568CFBFA24D983F981" ID-DOI="http://doi.org/10.5281/zenodo.3736941" ID-Zenodo-Dep="3736941" httpUri="https://zenodo.org/record/3736941/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,264,1474,1499]" targetBox="[156,1436,166,1455]" targetPageId="50">
|
||
<paragraph id="8B8136B0FFAF84568CFBFA24D983F981" blockId="50.[151,1437,1474,1571]" pageId="50" pageNumber="51">
|
||
<emphasis id="B94AEAA2FFAF84568CFBFA24DA50FA79" bold="true" box="[151,366,1474,1499]" pageId="50" pageNumber="51">FIGURES 151–155.</emphasis>
|
||
Larva presumably attributed to
|
||
<taxonomicName id="4C3E4D33FFAF84568EDAFA25D8B1FA79" box="[694,911,1475,1499]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAF84568EDAFA25D8B1FA79" box="[694,911,1475,1499]" italics="true" pageId="50" pageNumber="51">Thraulodes viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAF84568FFAFA25D8EEFA79" bold="true" box="[918,976,1475,1499]" pageId="50" pageNumber="51">
|
||
<taxonomicNameLabel id="A27957D9FFAF84568FFAFA25D8EEFA79" box="[918,976,1475,1499]" pageId="50" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
151, anterior side of hind tibia; 152, anterior side of hind femur; 153, tergalius VII; 154, general view; 155, abdominal terga spread on slide (with brown and whitish cuticular coloration and blackish paired hypodermal markings).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8136B0FFAF84568CABF993D86BF916" blockId="50.[151,1437,1653,2040]" pageId="50" pageNumber="51">
|
||
<emphasis id="B94AEAA2FFAF84568CABF993DA6DF932" bold="true" box="[199,339,1653,1680]" italics="true" pageId="50" pageNumber="51">Male imago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFAF84568D0EF993D94CF932" box="[354,626,1653,1680]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="50" pageNumber="51">Figs 156–161, 165–168</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFAF84568E13F993D9DBF932" box="[639,741,1653,1680]" captionStart="FIGURES 169–173" captionStartId="53.[151,264,1539,1564]" captionTargetBox="[164,1425,173,1511]" captionTargetId="figure@53.[151,1436,166,1520]" captionTargetPageId="53" captionText="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736945" httpUri="https://zenodo.org/record/3736945/files/figure.png" pageId="50" pageNumber="51">169, 171</figureCitation>
|
||
). Head brown. Antenna with scapus ocher, pedicellus brown, flagellum ocher. Dorsal eyes contiguous medially, dull orange.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAF84568CABF95BD823F882" blockId="50.[151,1437,1653,2040]" pageId="50" pageNumber="51">
|
||
Prothorax light ocher with pair of dark brown spots above coxa. Pterothorax brown with ocher; metathoracic scutellum apically brown (
|
||
<figureCitation id="13052A35FFAF84568DAFF907D918F95E" box="[451,550,1761,1788]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="50" pageNumber="51">Fig. 159</figureCitation>
|
||
); prosternum anteriorly dark, posteriorly ocher; furcasternal protuberances and furcasternal median impression equally brown (
|
||
<figureCitation id="13052A35FFAF84568EC2F8E3D82EF882" box="[686,784,1797,1824]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="50" pageNumber="51">Fig. 158</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAF84578CABF8CFDAEEFF59" blockId="50.[151,1437,1653,2040]" lastBlockId="51.[151,1437,153,1727]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
|
||
Cuticle of legs mostly colorless; on fore leg apex of femur with cuticle light brownish. Legs with following hypodermal pigmentation on whitish background (
|
||
<figureCitation id="13052A35FFAF84568EB4F8ABD847F8CA" box="[728,889,1869,1896]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="50" pageNumber="51">Figs 160–161</figureCitation>
|
||
): Fore femur with triangular orange spot on anterior side at 1/4 from base and with multicolored apical band occupying distal 0.4 of femur; apical band orange, proximally and apically narrowly bordered by blackish-brown, on inner and outer sides with blackish-brown longitudinal stripes. Fore tibia with dark brown apex. Middle and hind femora with multicolored pre-apical band within apical 0.4 of femur; pre-apical band mainly orange-brown, proximally bordered by blackish-brown, on inner side with blackish-brown longitudinal stripe; apex of femur non-colored. In
|
||
<typeStatus id="54858812FFAE84578FDDFF7FDF28FF11" box="[945,1046,153,179]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
|
||
, relation of fore femur to fore wing length 82:355; proportions femur/tibia/tarsomeres on fore leg 82:110:3:29:22:15:10; on middle leg 77:85:3:5:3:3:9; on hind leg 95:90:3:5:3:3:9.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFEE3DA7FFE71" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">
|
||
Fore wing with dark brown macula just distad of costal brace and light brown band crossing wing posteriad of costal brace; costal brace lighter orange; very base of fore wing non-colored. Hind wing with dark brown macula on costal brace and bases of Sc and R. On all wings longitudinal and cross veins colorless. Costal cross veins proximad of bulla few, colorless and very thin, so that visible only under high magnification (as in
|
||
<figureCitation id="13052A35FFAE845788E8FE97DFD8FE29" box="[1156,1254,369,395]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="51" pageNumber="52">Fig. 142</figureCitation>
|
||
). Pterostigmatic cross veins moderately dense, oblique, arched, non-branched (
|
||
<figureCitation id="13052A35FFAE84578F3BFE73D885FE0D" box="[855,955,404,431]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="51" pageNumber="52">Fig. 156</figureCitation>
|
||
). Hind wing with prominent costal projection (
|
||
<figureCitation id="13052A35FFAE84578CBEFE5FDA0AFE71" box="[210,308,441,467]" captionStart="FIGURES 169–173" captionStartId="53.[151,264,1539,1564]" captionTargetBox="[164,1425,173,1511]" captionTargetId="figure@53.[151,1436,166,1520]" captionTargetPageId="53" captionText="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736945" httpUri="https://zenodo.org/record/3736945/files/figure.png" pageId="51" pageNumber="52">Fig. 173</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFE3BD957FD09" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">Abdominal hypodermal coloration: Tergum I brown. Terga II–VI ocher, translucent, with light brown band on posterior margin; terga III–VI with pair of small gray midway spots; terga IV–VI with small diffusive orange stripe medially; pleura with blackish stigmatic dots. Tergum VII mainly orange, tergum VIII anteriorly-medially orange and posteriorly-laterally ocher, terga XI–X mainly orange. Sterna I–VI ocher, translucent; sterna VII–IX ocher with lateral sides colored with brownish and orange. Caudalii with segments alternated as following: black, white, white with base black, white; all joinings black.</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFD53DA4FFC99" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">
|
||
Genitalia (
|
||
<figureCitation id="13052A35FFAE84578D50FD53DAE7FD6D" box="[316,473,693,719]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="51" pageNumber="52">Figs 167–168</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFAE84578D89FD53D972FD72" box="[485,588,693,720]" captionStart="FIGURES 169–173" captionStartId="53.[151,264,1539,1564]" captionTargetBox="[164,1425,173,1511]" captionTargetId="figure@53.[151,1436,166,1520]" captionTargetPageId="53" captionText="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736945" httpUri="https://zenodo.org/record/3736945/files/figure.png" pageId="51" pageNumber="52">169–171</figureCitation>
|
||
): Styliger and gonostyli light ocher. Dorsal extension of styliger narrow and prominent. Penis lobes not long, markedly widened apically, with narrow lateral pouches. Telopenes in form of spear-like rolls, attached on dorsal side and directed medially-dorsally, nearly straight, with groove opened dorsallylaterally (
|
||
<figureCitation id="13052A35FFAE84578D6EFCC7DA5AFC99" box="[258,356,801,827]" captionStart="FIGURES 169–173" captionStartId="53.[151,264,1539,1564]" captionTargetBox="[164,1425,173,1511]" captionTargetId="figure@53.[151,1436,166,1520]" captionTargetPageId="53" captionText="FIGURES 169–173. Thraulodes viviparus sp. n. 169, genitalia of male imago in ventral view; 170, their subimaginal exuviae at the same magnification; 171, apex of penis in dorsal view; 172, protogonostyli and protopenis of presumably associated larva; 173, hind wing (169–171, 173, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736945" httpUri="https://zenodo.org/record/3736945/files/figure.png" pageId="51" pageNumber="52">Fig. 171</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFCA2D89FFC21" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">
|
||
<emphasis id="B94AEAA2FFAE84578CABFCA2DA53FCFD" bold="true" box="[199,365,836,863]" italics="true" pageId="51" pageNumber="52">Female imago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFAE84578D10FCA3D925FCC2" box="[380,539,837,864]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="51" pageNumber="52">Figs 162–164</figureCitation>
|
||
). Pronotum medially brown with ocher, laterally whitish; abdominal terga II–VI brown with ocher; in other respects coloration similar to that of male.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFC6BDBF2FC4D" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">
|
||
<emphasis id="B94AEAA2FFAE84578CABFC6BDA36FC05" bold="true" box="[199,264,909,935]" italics="true" pageId="51" pageNumber="52">Eggs.</emphasis>
|
||
Chorion absent. In the single examined non-fecundated female imago, abdomen is entirely filled with polygonal eggs with smooth surfaces closely pressed one to another; no one egg has developed or developing chorion.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFC1ED8CBFBB1" blockId="51.[151,1437,153,1727]" box="[199,1013,1016,1044]" pageId="51" pageNumber="52">
|
||
<emphasis id="B94AEAA2FFAE84578CABFC1EDA70FBB1" bold="true" box="[199,334,1016,1043]" pageId="51" pageNumber="52">Dimension.</emphasis>
|
||
Fore wing length (and approximate body length)
|
||
<quantity id="4CC69B55FFAE84578F10FC1ED8CFFBB1" box="[892,1009,1016,1043]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="51" pageNumber="52" unit="mm" value="8.75" valueMax="9.0" valueMin="8.5">8.5–9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324653BFFAE84578CABFBFADFCBF91D" pageId="51" pageNumber="52" type="discussion">
|
||
<paragraph id="8B8136B0FFAE84578CABFBFADEB9FB49" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">
|
||
<emphasis id="B94AEAA2FFAE84578CABFBFADA71FB95" bold="true" box="[199,335,1052,1079]" pageId="51" pageNumber="52">Comments.</emphasis>
|
||
Male and female imagines, reared from subimagines, were associated by similar coloration of thorax, wings, legs and caudalii of imago and coloration of subimaginal exuviae; particularly, presence of brown band posteriad of costal brace (
|
||
<figureCitation id="13052A35FFAE84578DDBFB83D9FDFB22" box="[439,707,1124,1152]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="51" pageNumber="52">Figs 157–159, 162–164</figureCitation>
|
||
) allowed to separate them from the similar sympatric species
|
||
<taxonomicName id="4C3E4D33FFAE84578914FB83DA25FB01" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578914FB83DA25FB01" italics="true" pageId="51" pageNumber="52">Th. panamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578D4FFB6FDA5DFB01" bold="true" box="[291,355,1161,1187]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578D4FFB6FDA5DFB01" box="[291,355,1161,1187]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C3E4D33FFAE84578D1DFB6FD93EFB01" authorityName="Chacon, Segnini & Dominguez" authorityYear="1999" box="[369,512,1161,1187]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="marreroi">
|
||
<emphasis id="B94AEAA2FFAE84578D1DFB6FD93EFB01" box="[369,512,1161,1187]" italics="true" pageId="51" pageNumber="52">Th. marreroi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E4D33FFAE84578E62FB6FD9A4FB01" authorityName="Mariano & Flowers (in Mariano, Flowers & Froehlich" authorityYear="2011" box="[526,666,1161,1187]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="sinuosus">
|
||
<emphasis id="B94AEAA2FFAE84578E62FB6FD9A4FB01" box="[526,666,1161,1187]" italics="true" pageId="51" pageNumber="52">Th. sinuosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3E4D33FFAE84578EBEFB6FD85EFB01" box="[722,864,1161,1187]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578EBEFB6FD85EFB01" box="[722,864,1161,1187]" italics="true" pageId="51" pageNumber="52">Th. ludmilae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578F0BFB6FD899FB01" bold="true" box="[871,935,1161,1187]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578F0BFB6FD899FB01" box="[871,935,1161,1187]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; in addition to this, brown color of metathoracic scutellum (
|
||
<figureCitation id="13052A35FFAE84578D3FFB4BDACFFB6A" box="[339,497,1196,1224]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="51" pageNumber="52">Figs 159, 164</figureCitation>
|
||
) distinguishes
|
||
<taxonomicName id="4C3E4D33FFAE84578EF0FB4BD80FFB65" box="[668,817,1197,1223]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus">
|
||
<emphasis id="B94AEAA2FFAE84578EF0FB4BD80FFB65" box="[668,817,1197,1223]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C3E4D33FFAE84578F14FB4BDF17FB65" box="[888,1065,1197,1223]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578F14FB4BDF17FB65" box="[888,1065,1197,1223]" italics="true" pageId="51" pageNumber="52">Th. panamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE8457885DFB4BDF4CFB65" bold="true" box="[1073,1138,1197,1223]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE8457885DFB4BDF4CFB65" box="[1073,1138,1197,1223]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; brown medioscutal chromozone on subimaginal exuviae (
|
||
<figureCitation id="13052A35FFAE84578E7CFB37D951FB49" box="[528,623,1232,1259]" captionStart="FIGURES 156–168" captionStartId="52.[151,264,1899,1924]" captionTargetBox="[151,1436,166,1879]" captionTargetId="figure@52.[151,1436,166,1879]" captionTargetPageId="52" captionText="FIGURES 156–168. Thraulodes viviparus sp. n. 156, pterostigma; 157–159, head and thorax of male imago in lateral in ventral and dorsal view; 160–161, fore and hind legs of male imago; 162–164, female imago in ventral in dorsal and lateral view; 165, left half of subimaginal exuviae of mesonotum; 166, abdominal terga and sterna; 167–168, penis with focus on dorsal and ventral side (156–161, 165–168, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736943" httpUri="https://zenodo.org/record/3736943/files/figure.png" pageId="51" pageNumber="52">Fig. 165</figureCitation>
|
||
) distinguishes
|
||
<taxonomicName id="4C3E4D33FFAE84578F74FB37D892FB49" box="[792,940,1233,1259]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus">
|
||
<emphasis id="B94AEAA2FFAE84578F74FB37D892FB49" box="[792,940,1233,1259]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C3E4D33FFAE84578F9CFB37DF42FB49" authorityName="Mariano & Flowers (in Mariano, Flowers & Froehlich" authorityYear="2011" box="[1008,1148,1233,1259]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="sinuosus">
|
||
<emphasis id="B94AEAA2FFAE84578F9CFB37DF42FB49" box="[1008,1148,1233,1259]" italics="true" pageId="51" pageNumber="52">Th. sinuosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3E4D33FFAE845788DFFB37DE7EFB49" box="[1203,1344,1233,1259]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="ludmilae" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE845788DFFB37DE7EFB49" box="[1203,1344,1233,1259]" italics="true" pageId="51" pageNumber="52">Th. ludmilae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE8457892BFB37DEB9FB49" bold="true" box="[1351,1415,1233,1259]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE8457892BFB37DEB9FB49" box="[1351,1415,1233,1259]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFB13D8D2FA61" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">
|
||
A single male larva of last instar has no developing imaginal details, and venation of its protoptera is not preserved. Its association with
|
||
<taxonomicName id="4C3E4D33FFAE84578DBEFAFFD959FA91" box="[466,615,1305,1331]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578DBEFAFFD959FA91" box="[466,615,1305,1331]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578E1CFAFFD98EFA91" bold="true" box="[624,688,1305,1331]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578E1CFAFFD98EFA91" box="[624,688,1305,1331]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is based only on blackish spots corresponding to the midway and spiracular spots of imago, the same size as in imago and the same locality. Among species of
|
||
<taxonomicName id="4C3E4D33FFAE845788A4FADBDE76FAF5" authorityName="Ulmer" authorityYear="1920" box="[1224,1352,1341,1367]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94AEAA2FFAE845788A4FADBDE76FAF5" box="[1224,1352,1341,1367]" italics="true" pageId="51" pageNumber="52">Thraulodes</emphasis>
|
||
</taxonomicName>
|
||
known from
|
||
<collectingCountry id="F3297620FFAE84578CB9FA87DA0FFAD9" box="[213,305,1377,1403]" name="Panama" pageId="51" pageNumber="52">Panama</collectingCountry>
|
||
, midway spots on abdominal terga III–VI and spiracular spots are present also in
|
||
<taxonomicName id="4C3E4D33FFAE845788A8FA87DE4DFAD9" box="[1220,1395,1377,1403]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE845788A8FA87DE4DFAD9" box="[1220,1395,1377,1403]" italics="true" pageId="51" pageNumber="52">Th. panamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578917FA87DB90FA3D" bold="true" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578917FA87DB90FA3D" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C3E4D33FFAE84578CD1FA63DA74FA3D" authorityName="Mariano & Flowers (in Mariano, Flowers & Froehlich" authorityYear="2011" box="[189,330,1413,1439]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="sinuosus">
|
||
<emphasis id="B94AEAA2FFAE84578CD1FA63DA74FA3D" box="[189,330,1413,1439]" italics="true" pageId="51" pageNumber="52">Th. sinuosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3E4D33FFAE84578DE8FA63D92AFA3D" authorityName="Chacon, Segnini & Dominguez" authorityYear="1999" box="[388,532,1413,1439]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="marreroi">
|
||
<emphasis id="B94AEAA2FFAE84578DE8FA63D92AFA3D" box="[388,532,1413,1439]" italics="true" pageId="51" pageNumber="52">Th. marreroi</emphasis>
|
||
</taxonomicName>
|
||
; these species have smaller size; larvae of these three species are associated by rearing and differ from the larva presumably ascribed to
|
||
<taxonomicName id="4C3E4D33FFAE84578F7EFA4FD89BFA61" box="[786,933,1449,1475]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578F7EFA4FD89BFA61" box="[786,933,1449,1475]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578FC1FA4FD8D2FA61" bold="true" box="[941,1004,1449,1475]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578FC1FA4FD8D2FA61" box="[941,1004,1449,1475]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFAE84578CABFA2ADFCBF91D" blockId="51.[151,1437,153,1727]" pageId="51" pageNumber="52">
|
||
<emphasis id="B94AEAA2FFAE84578CABFA2ADA5CFA45" bold="true" box="[199,354,1484,1511]" pageId="51" pageNumber="52">Comparison.</emphasis>
|
||
<taxonomicName id="4C3E4D33FFAE84578D06FA2BDAC0FA45" box="[362,510,1485,1511]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578D06FA2BDAC0FA45" box="[362,510,1485,1511]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578E6AFA2BD978FA45" bold="true" box="[518,582,1485,1511]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578E6AFA2BD978FA45" box="[518,582,1485,1511]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all other species by absence of sculptured chorion on eggs. By penis structure and coloration of legs and abdomen
|
||
<taxonomicName id="4C3E4D33FFAE84578EB8FA17D859F9A9" box="[724,871,1521,1547]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578EB8FA17D859F9A9" box="[724,871,1521,1547]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578F01FA17D892F9A9" bold="true" box="[877,940,1521,1547]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578F01FA17D892F9A9" box="[877,940,1521,1547]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C3E4D33FFAE84578856FA17DA62F98D" authority="Chacon, Segnini & Dominguez 1999" authorityName="Chacon, Segnini & Dominguez" authorityYear="1999" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="guanare">
|
||
<emphasis id="B94AEAA2FFAE84578856FA17DFFFF9A9" box="[1082,1217,1521,1547]" italics="true" pageId="51" pageNumber="52">Th. guanare</emphasis>
|
||
<bibRefCitation id="EFAF4B41FFAE845788A4FA17DA62F98D" author="Chacon, M. & Segnini, S. & Dominguez, E." pageId="51" pageNumber="52" pagination="249 - 257" refId="ref68968" refString="Chacon, M. M, Segnini, S. & Dominguez, E. (1999) Three new species of Thraulodes (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Venezuela. Aquatic Insects, 21, 249 - 257. https: // doi. org / 10.1076 / aqin. 21.4.249.4504" type="journal article" year="1999">Chacón, Segnini & Domínguez 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
, which was described as male and female imagines from
|
||
<collectingCountry id="F3297620FFAE84578FB1F9F3DF6AF98D" box="[989,1108,1557,1583]" name="Venezuela" pageId="51" pageNumber="52">Venezuela</collectingCountry>
|
||
; egg structure of
|
||
<taxonomicName id="4C3E4D33FFAE84578979F9F3DEA2F98D" authorityName="Chacon, Segnini & Dominguez" authorityYear="1999" box="[1301,1436,1557,1583]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="guanare">
|
||
<emphasis id="B94AEAA2FFAE84578979F9F3DEA2F98D" box="[1301,1436,1557,1583]" italics="true" pageId="51" pageNumber="52">Th. guanare</emphasis>
|
||
</taxonomicName>
|
||
was not reported. Imagines of
|
||
<taxonomicName id="4C3E4D33FFAE84578D9DF9DFD9B8F9F1" box="[497,646,1593,1619]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578D9DF9DFD9B8F9F1" box="[497,646,1593,1619]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578EE2F9DFD9F1F9F1" bold="true" box="[654,719,1593,1619]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578EE2F9DFD9F1F9F1" box="[654,719,1593,1619]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differ from
|
||
<taxonomicName id="4C3E4D33FFAE84578F37F9DFD8DDF9F1" authorityName="Chacon, Segnini & Dominguez" authorityYear="1999" box="[859,995,1593,1619]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="guanare">
|
||
<emphasis id="B94AEAA2FFAE84578F37F9DFD8DDF9F1" box="[859,995,1593,1619]" italics="true" pageId="51" pageNumber="52">Th. guanare</emphasis>
|
||
</taxonomicName>
|
||
by wider multicolored macula at wing base (instead of black spot in
|
||
<taxonomicName id="4C3E4D33FFAE84578D9DF9BBD945F9D5" authorityName="Chacon, Segnini & Dominguez" authorityYear="1999" box="[497,635,1629,1655]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="guanare">
|
||
<emphasis id="B94AEAA2FFAE84578D9DF9BBD945F9D5" box="[497,635,1629,1655]" italics="true" pageId="51" pageNumber="52">Th. guanare</emphasis>
|
||
</taxonomicName>
|
||
) and proportion of penis, which in
|
||
<taxonomicName id="4C3E4D33FFAE8457887BF9BBDF93F9D5" box="[1047,1197,1629,1655]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE8457887BF9BBDF93F9D5" box="[1047,1197,1629,1655]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE845788DBF9BBDFC6F9D5" bold="true" box="[1207,1272,1629,1655]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE845788DBF9BBDFC6F9D5" box="[1207,1272,1629,1655]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is shorter and wider than in
|
||
<taxonomicName id="4C3E4D33FFAE84578D5FF967DA84F939" authorityName="Chacon, Segnini & Dominguez" authorityYear="1999" box="[307,442,1665,1691]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="guanare">
|
||
<emphasis id="B94AEAA2FFAE84578D5FF967DA84F939" box="[307,442,1665,1691]" italics="true" pageId="51" pageNumber="52">Th. guanare</emphasis>
|
||
</taxonomicName>
|
||
. From the sympatric species
|
||
<taxonomicName id="4C3E4D33FFAE84578F68F967D88AF939" box="[772,948,1665,1691]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE84578F68F967D88AF939" box="[772,948,1665,1691]" italics="true" pageId="51" pageNumber="52">Th. panamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578FD7F967D8C5F939" bold="true" box="[955,1019,1665,1691]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578FD7F967D8C5F939" box="[955,1019,1665,1691]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, imagines of
|
||
<taxonomicName id="4C3E4D33FFAE845788F8F967DE19F939" box="[1172,1319,1665,1691]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="viviparus" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFAE845788F8F967DE19F939" box="[1172,1319,1665,1691]" italics="true" pageId="51" pageNumber="52">Th. viviparus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFAE84578943F967DE51F939" bold="true" box="[1327,1391,1665,1691]" pageId="51" pageNumber="52">
|
||
<taxonomicNameLabel id="A27957D9FFAE84578943F967DE51F939" box="[1327,1391,1665,1691]" pageId="51" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by absence of white spot on scutellum, coloration of wing base and shape of penis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |