233 lines
33 KiB
XML
233 lines
33 KiB
XML
<document id="7FFE56798D53C3616A013A45092719C2" ID-CLB-Dataset="157773" ID-DOI="10.11646/zootaxa.5194.1.2" ID-GBIF-Dataset="9c76b088-9845-48f6-9561-d4c6e91c4343" ID-ISSN="1175-5326" ID-Zenodo-Dep="7141848" ID-ZooBank="7C18727C-3AF0-4BE6-AFBC-EA1AC2F2B926" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664862422389" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2022" docId="03EE7A4DC90AFD05FF5EFBD4FAC7F836" docLanguage="en" docName="zootaxa.5194.1.2.pdf" docOrigin="Zootaxa 5194 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nodocepheus Hammer 1958" docType="treatment" docVersion="6" lastPageNumber="44" masterDocId="FFD70235C901FD0EFFC9FFD6FF9AFFDF" masterDocTitle="First records of Tumerozetidae and Nodocepheidae from Australia, with descriptions of new taxa and a re-assessment of the Polypterozetoidea (Oribatida, Brachypylina)" masterLastPageNumber="57" masterPageNumber="33" pageNumber="44" updateTime="1699387054636" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="333EDBFAE4424F8BF3063B191D2B104D">
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<mods:title id="1E33881D037C13D76A2FBA245ADBACCF">First records of Tumerozetidae and Nodocepheidae from Australia, with descriptions of new taxa and a re-assessment of the Polypterozetoidea (Oribatida, Brachypylina)</mods:title>
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<mods:namePart id="2F592958DE6AC286D7A1F40B04416AE0">Colloff, Matthew J.</mods:namePart>
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<mods:typeOfResource id="21DBF3DB8EB24C707E670946E72972D9">text</mods:typeOfResource>
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<mods:title id="740A1543787F43624FE0BB308F0C3129">Zootaxa</mods:title>
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<mods:part id="D497693C0FC32ED0D56A878E813E00BE">
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<mods:date id="632238D96B643219F24E580A672F4A58">2022</mods:date>
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<mods:number id="E6294AFA03762ED00F55AF66E7262089">2022-10-04</mods:number>
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<mods:number id="0B2D4090FA55BBEF3371D1FCE7C83FC9">5194</mods:number>
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<mods:identifier id="66C394D99AE5A6FBC55C90CFDBD29177" type="DOI">10.11646/zootaxa.5194.1.2</mods:identifier>
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<treatment id="03EE7A4DC90AFD05FF5EFBD4FAC7F836" ID-DOI="http://doi.org/10.5281/zenodo.7141890" ID-GBIF-Taxon="202134807" ID-Zenodo-Dep="7141890" LSID="urn:lsid:plazi:treatment:03EE7A4DC90AFD05FF5EFBD4FAC7F836" httpUri="http://treatment.plazi.org/id/03EE7A4DC90AFD05FF5EFBD4FAC7F836" lastPageNumber="44" pageId="11" pageNumber="44">
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<subSubSection id="C35D98D0C90AFD05FF5EFBD4FE76FBC2" box="[151,492,1026,1053]" pageId="11" pageNumber="44" type="nomenclature">
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<paragraph id="8BF8CB5BC90AFD05FF5EFBD4FE76FBC2" blockId="11.[151,1437,1026,2025]" box="[151,492,1026,1053]" pageId="11" pageNumber="44">
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<heading id="D0B07C37C90AFD05FF5EFBD4FE76FBC2" bold="true" box="[151,492,1026,1053]" fontSize="11" level="1" pageId="11" pageNumber="44" reason="1">
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<emphasis id="B9331749C90AFD05FF5EFBD4FE76FBC2" bold="true" box="[151,492,1026,1053]" pageId="11" pageNumber="44">
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Redefinition of
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<taxonomicName id="4C47B0D8C90AFD05FE99FBD4FE76FBC2" ID-CoL="64M5" authorityName="Hammer" authorityYear="1958" box="[336,492,1026,1053]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B9331749C90AFD05FE99FBD4FE76FBC2" bold="true" box="[336,492,1026,1053]" italics="true" pageId="11" pageNumber="44">Nodocepheus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D98D0C90AFD05FF0EFBF0FE56FAE2" pageId="11" pageNumber="44" type="description">
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<paragraph id="8BF8CB5BC90AFD05FF0EFBF0FE56FAE2" blockId="11.[151,1437,1026,2025]" pageId="11" pageNumber="44">
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<emphasis id="B9331749C90AFD05FF0EFBF0FEDCFB9F" bold="true" box="[199,326,1062,1088]" pageId="11" pageNumber="44">Definition.</emphasis>
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Polypterozetoid mites with subcapitulum secondarily anarthric (
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<figureCitation id="137CD7DEC90AFD05FBD2FBF1FB13FB9E" box="[1051,1161,1062,1089]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="11" pageNumber="44">Figure 5b</figureCitation>
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) or diarthric, though the labiogenal articulation may be incomplete (
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<figureCitation id="137CD7DEC90AFD05FD4FFB9DFD63FBBA" box="[646,761,1099,1125]" captionStart="FIGURE 7" captionStartId="17.[152,255,1109,1133]" captionTargetBox="[441,1148,184,1079]" captionTargetId="figure-15@17.[435,1152,181,1085]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of the subcapitulum of polypterozetoid taxa included in the character analysis. (a) Polypterozetes cherubin Berlese 1916; (b) Podopterotegaeus altimonticola Piffl, 1972; (c) Nemacepheus dentatusAoki, 1968; (d) Nodocepheus laterodentatus Piffl, 1972; (e) Tumerozetes roughleyi sp. nov.; (f) Sacculella yarra gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7141872" httpUri="https://zenodo.org/record/7141872/files/figure.png" pageId="11" pageNumber="44">Figure 7d</figureCitation>
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); rutellum bifurcate, apex Y-shaped (
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<figureCitation id="137CD7DEC90AFD05FB68FB9DFA8EFBBA" box="[1185,1300,1099,1125]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="11" pageNumber="44">
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Figure
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<quantity id="4CBF66BEC90AFD05FB3CFB9DFA8EFBBA" box="[1269,1300,1099,1125]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="11" pageNumber="44" unit="g" value="4.0">4g</quantity>
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</figureCitation>
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); chelicerae modified: either with digitus fixus pointed, lacking teeth and digitus mobilis comb-like (
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<figureCitation id="137CD7DEC90AFD05FB4DFBB9FB6EFB56" box="[1156,1268,1135,1161]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="11" pageNumber="44">Figure 4e</figureCitation>
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) or both digits with reduced, blunted teeth, offset from each other (
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<figureCitation id="137CD7DEC90AFD05FD27FB45FCC1FB72" box="[750,859,1171,1197]" captionStart="FIGURE 4" captionStartId="10.[152,255,1502,1526]" captionTargetBox="[292,1305,192,1466]" captionTargetId="figure-22@10.[279,1308,181,1478]" captionTargetPageId="10" captionText="FIGURE 4. Sacculella yarra gen. et sp. nov. (a) palp; (b) chelicera; (c) digitus mobilis of chelicera, ventral aspect; (d) tutorium and sub-tutorium. Nodocepheus luxtoni sp. nov. (e) chelicera; (f) palp; (g) apex of rutellum; (h) tutorium and sub-tutorium; (i) rostrum and rostral setae; (j) Polypterozetes cherubin Grandjean, 1959, chelicera; (k) Podopterotegaeus miyamaensis Fujikawa, 2008, chelicera; (l) Nodocepheus dentatus Hammer, 1958, chelicera. Scale bars: a, b, d, e, f, h = 20 μm; i = 10 μm; c, g = 5 μm" figureDoi="http://doi.org/10.5281/zenodo.7141862" httpUri="https://zenodo.org/record/7141862/files/figure.png" pageId="11" pageNumber="44">Figure 4l</figureCitation>
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). With serrated tutorium and cusped sub-tutorium. With short, rounded single-lobed or bilobed humeral processes. With 11 pairs of smooth notogastral setae positioned marginally;
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<emphasis id="B9331749C90AFD05FEE9FB0DFEB6FB2B" box="[288,300,1243,1268]" italics="true" pageId="11" pageNumber="44">c</emphasis>
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<subScript id="17C3C91EC90AFD05FEE4FB3BFEAFFB24" attach="left" box="[301,309,1261,1275]" fontSize="6" pageId="11" pageNumber="44">1</subScript>
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and
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<emphasis id="B9331749C90AFD05FEA6FB0DFEE1FB2B" box="[367,379,1243,1268]" italics="true" pageId="11" pageNumber="44">c</emphasis>
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<subScript id="17C3C91EC90AFD05FEB2FB3BFE19FB24" attach="left" box="[379,387,1261,1275]" fontSize="6" pageId="11" pageNumber="44">2</subScript>
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marginal, on or near humeral process, sometimes on ventral fold. With five pairs of genital setae. Epimeral setal formula 3-1-3-3. Genital setae 6 pairs; aggenital setae 1 pair, anal setae 2 pairs, adanal setae 3 pairs. Legs monodactylous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D98D0C90AFD05FF0EFA91FAC7F836" pageId="11" pageNumber="44" type="discussion">
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<paragraph id="8BF8CB5BC90AFD05FF0EFA91FD9AFA76" blockId="11.[151,1437,1026,2025]" pageId="11" pageNumber="44">
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<emphasis id="B9331749C90AFD05FF0EFA91FEA1FABE" bold="true" box="[199,315,1351,1377]" pageId="11" pageNumber="44">Remarks.</emphasis>
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The above definition is based on those of
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<bibRefCitation id="EFD6B6AAC90AFD05FCD8FA91FC09FABE" author="Piffl, E." box="[785,915,1351,1377]" pageId="11" pageNumber="44" pagination="269 - 314" refId="ref15415" refString="Piffl, E. (1972) Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972)</bibRefCitation>
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and
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<bibRefCitation id="EFD6B6AAC90AFD05FC01FA91FBC0FABE" author="Woas, S." box="[968,1114,1351,1377]" pageId="11" pageNumber="44" pagination="21 - 291" refId="ref15549" refString="Woas, S. (2002) Acari: Oribatida. In: Adis, J. (Ed.) Amazonian Arachnida and Myriapoda. Pensoft Publishers, Sofia, pp. 21 - 291." type="book chapter" year="2002">Woas (2002)</bibRefCitation>
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.
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<bibRefCitation id="EFD6B6AAC90AFD05FBADFA91FA85FABE" author="Mahunka, S." box="[1124,1311,1351,1377]" pageId="11" pageNumber="44" pagination="151 - 180" refId="ref15225" refString="Mahunka, S. (1983) Oribatids from the eastern part of the Ethiopian region. II. Acta Zoologica Academiae Scientiarum Hungaricae, 29, 151 - 180." type="journal article" year="1983">Mahunka (1983)</bibRefCitation>
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provided a key to the six species of
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<taxonomicName id="4C47B0D8C90AFD05FE63FABDFDDBFA5A" authorityName="Hammer" authorityYear="1958" box="[426,577,1387,1413]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B9331749C90AFD05FE63FABDFDDBFA5A" box="[426,577,1387,1413]" italics="true" pageId="11" pageNumber="44">Nodocepheus</emphasis>
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</taxonomicName>
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known at that time, stressing the morphology of the humeral lobe, bothridial seta, tutorium and sub-tutorium.
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</paragraph>
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<paragraph id="8BF8CB5BC90AFD05FF0EFA65FE7FF9E6" blockId="11.[151,1437,1026,2025]" pageId="11" pageNumber="44">
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The presence of Trägårdh’s organ is based on the examination by
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<bibRefCitation id="EFD6B6AAC90AFD05FC4FFA65FB9CFA12" author="Piffl, E." box="[902,1030,1459,1485]" pageId="11" pageNumber="44" pagination="269 - 314" refId="ref15415" refString="Piffl, E. (1972) Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972)</bibRefCitation>
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of a single specimen of
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<taxonomicName id="4C47B0D8C90AFD05FACCFA65FE3AFA2E" authority="Hammer 1958" authorityName="Hammer" authorityYear="1958" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="dentatus">
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<emphasis id="B9331749C90AFD05FACCFA65FF6CFA2E" italics="true" pageId="11" pageNumber="44">Nodocepheus dentatus</emphasis>
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<bibRefCitation id="EFD6B6AAC90AFD05FF37FA01FE3AFA2E" author="Hammer, M." box="[254,416,1494,1521]" pageId="11" pageNumber="44" pagination="1 - 129" refId="ref14898" refString="Hammer, M. (1958) Investigations on the Oribatid fauna of the Andes Mountains. I. The Argentine and Bolivia. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 10 (1), 1 - 129." type="journal article" year="1958">
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Hammer
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</bibRefCitation>
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</taxonomicName>
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in which the chelicerae were damaged. The structure is described as short, broad and thus lacks the characteristic elongated form of this structure. I could not identify a Trägårdh’s organ on the chelicerae
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<taxonomicName id="4C47B0D8C90AFD05FF5EF9C9FE19F9E6" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[151,387,1567,1593]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="luxtoni" status="sp. nov.">
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<emphasis id="B9331749C90AFD05FF5EF9C9FE19F9E6" box="[151,387,1567,1593]" italics="true" pageId="11" pageNumber="44">Nodocepheus luxtoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A200AA32C90AFD05FE43F9C9FE7FF9E6" box="[394,485,1567,1593]" pageId="11" pageNumber="44" rank="species">
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<emphasis id="B9331749C90AFD05FE43F9C9FE44F9E6" bold="true" box="[394,478,1567,1593]" pageId="11" pageNumber="44">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BF8CB5BC90AFD05FF0EF995FAC7F836" blockId="11.[151,1437,1026,2025]" pageId="11" pageNumber="44">
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In
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<taxonomicName id="4C47B0D8C90AFD05FF2FF995FE7EF982" authorityName="Hammer" authorityYear="1958" box="[230,484,1603,1629]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="dentatus">
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<emphasis id="B9331749C90AFD05FF2FF995FE7EF982" box="[230,484,1603,1629]" italics="true" pageId="11" pageNumber="44">Nodocepheus dentatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C47B0D8C90AFD05FE3BF995FCAEF982" authority="Piffl, 1972" authorityName="Piffl" authorityYear="1972" box="[498,820,1603,1629]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="laterodentatus">
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<emphasis id="B9331749C90AFD05FE3BF995FD2FF982" box="[498,693,1603,1629]" italics="true" pageId="11" pageNumber="44">N. laterodentatus</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FD75F995FCAEF982" author="Piffl, E." box="[700,820,1603,1629]" pageId="11" pageNumber="44" pagination="269 - 314" refId="ref15415" refString="Piffl, E. (1972) Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47B0D8C90AFD05FCA5F995FC41F982" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[876,987,1603,1629]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="luxtoni" status="sp. nov.">
|
||
<emphasis id="B9331749C90AFD05FCA5F995FC41F982" box="[876,987,1603,1629]" italics="true" pageId="11" pageNumber="44">N. luxtoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A200AA32C90AFD05FC2AF995FBA7F982" box="[995,1085,1603,1629]" pageId="11" pageNumber="44" rank="species">
|
||
<emphasis id="B9331749C90AFD05FC2AF995FBA2F982" bold="true" box="[995,1080,1603,1629]" pageId="11" pageNumber="44">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, setae
|
||
<emphasis id="B9331749C90AFD05FB44F995FB03F983" box="[1165,1177,1603,1628]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
<subScript id="17C3C91EC90AFD05FB50F983FB04F9BC" attach="left" box="[1177,1182,1621,1635]" fontSize="6" pageId="11" pageNumber="44">l</subScript>
|
||
are shifted laterally to the humeral region, positioned on a fold of the humeral process adjacent to setae
|
||
<emphasis id="B9331749C90AFD05FBF3F9B1FBDCF95F" box="[1082,1094,1639,1664]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
<subScript id="17C3C91EC90AFD05FB8FF9AFFBD4F958" attach="left" box="[1094,1102,1657,1671]" fontSize="6" pageId="11" pageNumber="44">2</subScript>
|
||
(
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FB97F9B1FB4FF95E" author="Piffl, E." box="[1118,1237,1639,1665]" pageId="11" pageNumber="44" pagination="269 - 314" refId="ref15415" refString="Piffl, E. (1972) Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl 1972</bibRefCitation>
|
||
, Plate 13 therein;
|
||
<figureCitation id="137CD7DEC90AFD05FF5EF95DFE9EF97A" box="[151,260,1674,1701]" captionStart="FIGURE 5" captionStartId="12.[152,255,2006,2030]" captionTargetBox="[269,1320,604,1966]" captionTargetId="figure-164@12.[258,1330,592,1982]" captionTargetPageId="12" captionText="FIGURE 5. Nodocepheus luxtoni sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) lateral aspect. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141866" httpUri="https://zenodo.org/record/7141866/files/figure.png" pageId="11" pageNumber="44">Figure 5a</figureCitation>
|
||
herein). It is likely that
|
||
<taxonomicName id="4C47B0D8C90AFD05FDD7F95DFCCCF97A" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[542,854,1675,1701]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis id="B9331749C90AFD05FDD7F95DFD3EF97A" box="[542,676,1675,1701]" italics="true" pageId="11" pageNumber="44">N. barbatus</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FD64F95DFCCCF97A" author="Hammer, M." box="[685,854,1675,1701]" pageId="11" pageNumber="44" pagination="1 - 108" refId="ref14939" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
(originally described as
|
||
<taxonomicName id="4C47B0D8C90AFD05FBBAF95DFA09F97A" authorityName="Hammer" authorityYear="1966" box="[1139,1427,1675,1701]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="variety" species="dentatus" variety="barbatus">
|
||
<emphasis id="B9331749C90AFD05FBBAF95DFB6EF97A" box="[1139,1268,1675,1701]" italics="true" pageId="11" pageNumber="44">N. dentatus</emphasis>
|
||
var.
|
||
<emphasis id="B9331749C90AFD05FAE6F95DFA09F97A" box="[1327,1427,1675,1701]" italics="true" pageId="11" pageNumber="44">barbatus</emphasis>
|
||
</taxonomicName>
|
||
) shares this character state, giving a likely notogastral setal complement of 11 pairs (cf.
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FBB3F979FB74F916" author="Piffl, E." box="[1146,1262,1711,1737]" pageId="11" pageNumber="44" pagination="269 - 314" refId="ref15415" refString="Piffl, E. (1972) Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl 1972</bibRefCitation>
|
||
, p. 292), rather than 9 pairs as illustrated by
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FE2EF905FD0DF932" author="Hammer, M." box="[487,663,1747,1773]" pageId="11" pageNumber="44" pagination="1 - 108" refId="ref14939" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer (1966</bibRefCitation>
|
||
, Figure 85 therein). In
|
||
<taxonomicName id="4C47B0D8C90AFD05FC6DF905FB7FF932" authority="Mahunka, 1985" authorityName="Mahunka" authorityYear="1985" box="[932,1253,1746,1773]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B9331749C90AFD05FC6DF905FBBDF933" box="[932,1063,1747,1772]" italics="true" pageId="11" pageNumber="44">N. minimus</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FBF9F905FB7FF932" author="Mahunka, S." box="[1072,1253,1746,1773]" pageId="11" pageNumber="44" pagination="295 - 339" refId="ref15256" refString="Mahunka, S. (1985) Oribatids from Africa (Acari, Oribatida), I. Acta Zoologica Hungarica, 31, 295 - 339." type="journal article" year="1985">Mahunka, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
, setae
|
||
<emphasis id="B9331749C90AFD05FAFDF905FADAF933" box="[1332,1344,1747,1772]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
<subScript id="17C3C91EC90AFD05FA89F933FAD2F92C" attach="left" box="[1344,1352,1765,1779]" fontSize="6" pageId="11" pageNumber="44">1</subScript>
|
||
appear to be absent and
|
||
<emphasis id="B9331749C90AFD05FE92F921FEFDF8CF" box="[347,359,1783,1808]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
<subScript id="17C3C91EC90AFD05FEAEF8DFFEF5F8C8" attach="left" box="[359,367,1801,1815]" fontSize="6" pageId="11" pageNumber="44">2</subScript>
|
||
are apparently reduced to their alveoli, giving a complement of 10 pairs (including
|
||
<emphasis id="B9331749C90AFD05FAFCF921FADBF8CF" box="[1333,1345,1783,1808]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
<subScript id="17C3C91EC90AFD05FA88F8DFFAD3F8C8" attach="left" box="[1345,1353,1801,1815]" fontSize="6" pageId="11" pageNumber="44">2</subScript>
|
||
). In
|
||
<taxonomicName id="4C47B0D8C90AFD05FA4AF921FE27F8EA" authority="Mahunka, 1980" authorityName="Mahunka" authorityYear="1980" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="cerebralis">
|
||
<emphasis id="B9331749C90AFD05FA4AF921FE92F8EA" italics="true" pageId="11" pageNumber="44">N. cerebralis</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FEC7F8CDFE27F8EA" author="Mahunka, S." box="[270,445,1819,1845]" pageId="11" pageNumber="44" pagination="155 - 181" refId="ref15182" refString="Mahunka, S. (1980) Neue und interessante Milben aus dem Genfer Museum XXXVIII. Oribatids (Acari) from Monte Susana (Tierra del Fuego, Argentina). Revue Suisse de Zoologie, 87, 155 - 181." type="journal article" year="1980">Mahunka, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47B0D8C90AFD05FE3BF8CDFCBFF8EA" authority="Balogh, 1961" authorityName="Balogh" authorityYear="1961" box="[498,805,1819,1845]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="hammerae">
|
||
<emphasis id="B9331749C90AFD05FE3BF8CDFD12F8EA" box="[498,648,1819,1845]" italics="true" pageId="11" pageNumber="44">N. hammerae</emphasis>
|
||
Balogh, 1961
|
||
</taxonomicName>
|
||
neither of the
|
||
<emphasis id="B9331749C90AFD05FC0CF8CDFC4BF8EB" box="[965,977,1819,1844]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
series are illustrated, yet in the figures of the humeral regions of
|
||
<taxonomicName id="4C47B0D8C90AFD05FE50F8E9FD5DF886" authority="Mahunka, 1983" authorityName="Mahunka" authorityYear="1983" box="[409,711,1855,1881]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="baloghi">
|
||
<emphasis id="B9331749C90AFD05FE50F8E9FD95F886" box="[409,527,1855,1881]" italics="true" pageId="11" pageNumber="44">N. baloghi</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FDDCF8E9FD5DF886" author="Mahunka, S." box="[533,711,1855,1881]" pageId="11" pageNumber="44" pagination="151 - 180" refId="ref15225" refString="Mahunka, S. (1983) Oribatids from the eastern part of the Ethiopian region. II. Acta Zoologica Academiae Scientiarum Hungaricae, 29, 151 - 180." type="journal article" year="1983">Mahunka, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C47B0D8C90AFD05FD1BF8E9FCF8F886" authorityName="Mahunka" authorityYear="1980" box="[722,866,1855,1881]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="cerebralis">
|
||
<emphasis id="B9331749C90AFD05FD1BF8E9FCF8F886" box="[722,866,1855,1881]" italics="true" pageId="11" pageNumber="44">N. cerebralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C47B0D8C90AFD05FCB9F8E9FC75F886" authorityName="Hammer" authorityYear="1958" box="[880,1007,1855,1881]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="B9331749C90AFD05FCB9F8E9FC75F886" box="[880,1007,1855,1881]" italics="true" pageId="11" pageNumber="44">N. dentatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47B0D8C90AFD05FBEDF8E9FB21F886" authorityName="Balogh" authorityYear="1961" box="[1060,1211,1855,1881]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="hammerae">
|
||
<emphasis id="B9331749C90AFD05FBEDF8E9FB21F886" box="[1060,1211,1855,1881]" italics="true" pageId="11" pageNumber="44">N. hammerae</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FB2DF8E9FA0DF886" author="Mahunka, S." box="[1252,1431,1855,1881]" pageId="11" pageNumber="44" pagination="151 - 180" refId="ref15225" refString="Mahunka, S. (1983) Oribatids from the eastern part of the Ethiopian region. II. Acta Zoologica Academiae Scientiarum Hungaricae, 29, 151 - 180." type="journal article" year="1983">Mahunka (1983</bibRefCitation>
|
||
, Figures 47-53 therein),
|
||
<emphasis id="B9331749C90AFD05FE69F8B5FE36F8A3" box="[416,428,1891,1916]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
<subScript id="17C3C91EC90AFD05FE64F8A3FE2FF85C" attach="left" box="[429,437,1909,1923]" fontSize="6" pageId="11" pageNumber="44">1</subScript>
|
||
is present on the ventral surface of the median lobe of the humeral process, as illustrated for
|
||
<taxonomicName id="4C47B0D8C90AFD05FF77F851FEA7F87E" authorityName="Hammer" authorityYear="1958" box="[190,317,1927,1953]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="B9331749C90AFD05FF77F851FEA7F87E" box="[190,317,1927,1953]" italics="true" pageId="11" pageNumber="44">N. dentatus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFD6B6AAC90AFD05FEADF851FE7AF87E" author="Piffl, E." box="[356,480,1927,1953]" pageId="11" pageNumber="44" pagination="269 - 314" refId="ref15415" refString="Piffl, E. (1972) Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972</bibRefCitation>
|
||
, Plate 13, Figure Nd
|
||
<subScript id="17C3C91EC90AFD05FD0AF84FFD51F878" attach="left" box="[707,715,1945,1959]" fontSize="6" pageId="11" pageNumber="44">3</subScript>
|
||
therein). It seems likely that
|
||
<emphasis id="B9331749C90AFD05FBC5F851FB82F87F" box="[1036,1048,1927,1952]" italics="true" pageId="11" pageNumber="44">c</emphasis>
|
||
<subScript id="17C3C91EC90AFD05FBD0F84FFBBBF878" attach="left" box="[1049,1057,1945,1959]" fontSize="6" pageId="11" pageNumber="44">2</subScript>
|
||
is also present in
|
||
<taxonomicName id="4C47B0D8C90AFD05FB2CF851FAEFF87E" authorityName="Mahunka" authorityYear="1980" box="[1253,1397,1927,1953]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="cerebralis">
|
||
<emphasis id="B9331749C90AFD05FB2CF851FAEFF87E" box="[1253,1397,1927,1953]" italics="true" pageId="11" pageNumber="44">N. cerebralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C47B0D8C90AFD05FA4AF851FE97F81A" authorityName="Balogh" authorityYear="1961" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="hammerae">
|
||
<emphasis id="B9331749C90AFD05FA4AF851FE97F81A" italics="true" pageId="11" pageNumber="44">N. hammerae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C47B0D8C90AFD05FE85F87DFE56F81B" authorityName="Mahunka" authorityYear="1985" box="[332,460,1963,1988]" class="Arachnida" family="Nodocepheidae" genus="Nodocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="B9331749C90AFD05FE85F87DFE56F81B" box="[332,460,1963,1988]" italics="true" pageId="11" pageNumber="44">N. minimus</emphasis>
|
||
</taxonomicName>
|
||
on the anterioventral surface of the rugose lateral lobe of the humeral process, but has been overlooked when viewed dorsally because of its position below the heavily textured surface of the lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |