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<mods:title id="A09C63F909DF1453A8FBE00FDFA3F1D6">Ovilyra, a new genus of leucosiid crab (Crustacea: Decapoda: Brachyura) from the West Pacific</mods:title>
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<emphasis id="B930EAA8FF9CFFBEFF2DFE2681414A49" bold="true" box="[151,370,510,537]" italics="true" pageId="2" pageNumber="371">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF9CFFBEFE3BFE2682964A48" author="Targioni-Tozzetti, A." box="[385,677,510,536]" pageId="2" pageNumber="371" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
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<taxonomicNameLabel id="A20357D3FF9CFFBEFD0EFE27832C4A49" box="[692,799,511,537]" pageId="2" pageNumber="371" rank="species">n. comb.</taxonomicNameLabel>
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(
<figureCitation id="137F2A3FFF9CFFBEFF25FDFB813A4A6D" box="[159,265,547,573]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1921,1945]" captionStartId-1="4.[151,250,1934,1958]" captionStartId-2="5.[151,250,1254,1278]" captionStartId-3="6.[151,250,1167,1191]" captionStartId-4="7.[151,250,1928,1952]" captionStartId-5="8.[151,250,1906,1930]" captionStartId-6="9.[151,250,1754,1778]" captionTargetBox-0="[169,1418,181,1897]" captionTargetBox-1="[164,1422,181,1910]" captionTargetBox-2="[151,1436,181,1230]" captionTargetBox-3="[151,1436,417,1143]" captionTargetBox-4="[215,1387,688,1892]" captionTargetBox-5="[193,1352,191,1868]" captionTargetBox-6="[151,1436,181,1730]" captionTargetId-0="figure-15@3.[169,1418,181,1897]" captionTargetId-1="figure-19@4.[164,1422,181,1910]" captionTargetId-2="figure-15@5.[151,1436,181,1230]" captionTargetId-3="figure-141@6.[151,1436,417,1143]" captionTargetId-4="figure-278@7.[151,1436,669,1905]" captionTargetId-5="figure-19@8.[178,1409,181,1882]" captionTargetId-6="figure-15@9.[151,1436,181,1730]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." captionText-1="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." captionText-2="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." captionText-3="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." captionText-4="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." captionText-5="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).AD, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; EH, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; IL, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EL, = 0.5 mm." captionText-6="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi-0="http://doi.org/10.5281/zenodo.4674239" figureDoi-1="http://doi.org/10.5281/zenodo.4674241" figureDoi-2="http://doi.org/10.5281/zenodo.4674243" figureDoi-3="http://doi.org/10.5281/zenodo.4674245" figureDoi-4="http://doi.org/10.5281/zenodo.4674247" figureDoi-5="http://doi.org/10.5281/zenodo.4674249" figureDoi-6="http://doi.org/10.5281/zenodo.4674251" httpUri-0="https://zenodo.org/record/4674239/files/figure.png" httpUri-1="https://zenodo.org/record/4674241/files/figure.png" httpUri-2="https://zenodo.org/record/4674243/files/figure.png" httpUri-3="https://zenodo.org/record/4674245/files/figure.png" httpUri-4="https://zenodo.org/record/4674247/files/figure.png" httpUri-5="https://zenodo.org/record/4674249/files/figure.png" httpUri-6="https://zenodo.org/record/4674251/files/figure.png" pageId="2" pageNumber="371">Figs. 17</figureCitation>
)
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<taxonomicName id="4C444D39FF9CFFBEFF2DFDBF80D04A2F" authorityName="Leach" authorityYear="1817" box="[151,227,615,639]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9CFFBEFF2DFDBF80D04A2F" box="[151,227,615,639]" italics="true" pageId="2" pageNumber="371">Philyra</emphasis>
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sp. Targioni-Tozzetti 1872: 396.
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<taxonomicName id="4C444D39FF9CFFBEFF2DFD5E824B4ACE" ID-CoL="856BT" authority="Targioni-Tozzetti, 1877: 201" authorityName="Targioni-Tozzetti" authorityPageNumber="201" authorityYear="1877" box="[151,632,646,670]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9CFFBEFF2DFD5E817C4ACE" box="[151,335,646,670]" italics="true" pageId="2" pageNumber="371">Philyra fuliginosa</emphasis>
<bibRefCitation id="EFD54B4BFF9CFFBEFEEFFD5E824B4ACE" author="Targioni-Tozzetti, A." box="[341,632,646,670]" pageId="2" pageNumber="371" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877: 201</bibRefCitation>
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, pl. 12 fig. 3ag;
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<bibRefCitation id="EFD54B4BFF9CFFBEFC90FD5E83E14ACE" author="Serene, R." box="[810,978,646,670]" pageId="2" pageNumber="371" pagination="33 - 120" refId="ref7333" refString="Serene, R. (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, No. 1, 33 - 120." type="book chapter" year="1968">Serène 1968: 46</bibRefCitation>
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;
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<bibRefCitation id="EFD54B4BFF9CFFBEFC66FD5E84A44ACE" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[988,1175,646,670]" pageId="2" pageNumber="371" pagination="1 - 286" refId="ref6834" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B930EAA8FF9CFFBEFBB8FD5F84064ACE" box="[1026,1077,646,670]" italics="true" pageId="2" pageNumber="371">et al.</emphasis>
2008: 93
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;
<treatmentCitation id="0AE510ABFF9CFFBEFB1BFD5E85704ACE" author="Galil, B. S." box="[1185,1347,646,670]" page="281" pageId="2" pageNumber="371" year="2009">
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.
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<emphasis id="B930EAA8FF9CFFBEFF2DFD7C81734AEC" box="[151,320,676,700]" italics="true" pageId="2" pageNumber="371">Philyra olivacea</emphasis>
<bibRefCitation id="EFD54B4BFF9CFFBEFEFCFD7C82214AED" author="Rathbun, M. J." box="[326,530,676,701]" pageId="2" pageNumber="371" pagination="107 - 114" refId="ref7110" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909: 108</bibRefCitation>
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;
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, fig. 4, p1. 11, fig. 17;
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;
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<bibRefCitation id="EFD54B4BFF9CFFBEFA84FD7C81114A8B" author="Dai, A. - Y. &amp; Yang, S. - L. &amp; Song, Y. - Z. &amp; Chen, G. - X." pageId="2" pageNumber="371" refId="ref6365" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. &amp; Chen, G. - X. (1986) Crabs of the China Seas. China Ocean Press, Beijing. 11 + 642 pp." type="book" year="1986">
Dai
<emphasis id="B930EAA8FF9CFFBEFAD3FD7D85AE4AEC" box="[1385,1437,676,700]" italics="true" pageId="2" pageNumber="371">et al.</emphasis>
1986: 81
</bibRefCitation>
</treatmentCitation>
, fig. 42, pl. 10(3);
<treatmentCitation id="0AE510ABFF9CFFBEFE65FD1B828B4A8B" author="Dai, A. - Y. &amp; Yang, S. - L." box="[479,696,707,732]" page="91" pageId="2" pageNumber="371" year="1991">
<bibRefCitation id="EFD54B4BFF9CFFBEFE65FD1B828B4A8B" author="Dai, A. - Y. &amp; Yang, S. - L." box="[479,696,707,732]" pageId="2" pageNumber="371" pagination="1 - 295" refId="ref6313" refString="Dai, A. - Y. &amp; Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, pp. 21 + 608, figs. 1 - 295, pls. 1 - 74." type="journal article" year="1991">Dai &amp; Yang 1991: 91</bibRefCitation>
</treatmentCitation>
, fig. 42, pl. 10(3); Tan 1995: 469, figs. 3b, d;
<treatmentCitation id="0AE510ABFF9CFFBEFB3EFD1B855C4A8B" author="Chen, H. &amp; Sun, H." box="[1156,1391,707,731]" page="401" pageId="2" pageNumber="371" year="2002">
<bibRefCitation id="EFD54B4BFF9CFFBEFB3EFD1B855C4A8B" author="Chen, H. &amp; Sun, H." box="[1156,1391,707,731]" pageId="2" pageNumber="371" pagination="1 - 16" refId="ref6263" refString="Chen, H. &amp; Sun, H. (2002) Arthropoda Crustacea. Brachyura. Marine primitive crabs. Fauna Sinica. Invertebrata, 30, Science Press, Beijing, 597 pp., colour pls. 1 - 4, pls. 1 - 16." type="book chapter" year="2002">Chen &amp; Sun 2002: 401</bibRefCitation>
</treatmentCitation>
, fig. 181;
<treatmentCitation id="0AE510ABFF9CFFBEFF41FD3A81864AAA" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[251,437,738,762]" page="93" pageId="2" pageNumber="371" year="2008">
<bibRefCitation id="EFD54B4BFF9CFFBEFF41FD3A81864AAA" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[251,437,738,762]" pageId="2" pageNumber="371" pagination="1 - 286" refId="ref6834" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B930EAA8FF9CFFBEFE9AFD3B81604AAA" box="[288,339,738,762]" italics="true" pageId="2" pageNumber="371">et al.</emphasis>
2008: 93
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE510ABFF9CFFBEFE7AFD3A82514AAA" author="Galil, B. S." box="[448,610,738,762]" page="281" pageId="2" pageNumber="371" year="2009">
<bibRefCitation id="EFD54B4BFF9CFFBEFE7AFD3A82514AAA" author="Galil, B. S." box="[448,610,738,762]" pageId="2" pageNumber="371" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009: 281</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFB36BAFF9CFFBEFF2DFCD8822D4B48" blockId="2.[151,1437,615,823]" box="[151,542,768,793]" pageId="2" pageNumber="371">
<treatmentCitationGroup id="AB541194FF9CFFBEFF2DFCD8822D4B48" box="[151,542,768,793]" pageId="2" pageNumber="371">
<taxonomicName id="4C444D39FF9CFFBEFF2DFCD881444B48" ID-CoL="856BT" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[151,375,768,792]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9CFFBEFF2DFCD880D04B48" box="[151,227,768,792]" italics="true" pageId="2" pageNumber="371">Philyra</emphasis>
aff.
<emphasis id="B930EAA8FF9CFFBEFEA8FCD881444B48" box="[274,375,768,792]" italics="true" pageId="2" pageNumber="371">fuliginosa</emphasis>
</taxonomicName>
<treatmentCitation id="0AE510ABFF9CFFBEFE2AFCD982294B48" author="Yang, C. M." box="[400,538,768,793]" page="5" pageId="2" pageNumber="371" year="1979">
<bibRefCitation id="EFD54B4BFF9CFFBEFE2AFCD982294B48" author="Yang, C. M." box="[400,538,768,793]" pageId="2" pageNumber="371" refId="ref7562" refString="Yang, C. M. (1979) A list of Brachyura in the Zoological Reference Collection of the Department of Zoology. No. 14. Department of Zoology, National University of Singapore, 60 pp." type="book" year="1979">Yang 1979: 5</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFB36BAFF9CFFBEFF2DFCC7831E4B67" blockId="2.[151,1437,615,823]" box="[151,813,799,823]" pageId="2" pageNumber="371">
<treatmentCitationGroup id="AB541194FF9CFFBEFF2DFCC7831E4B67" box="[151,813,799,823]" pageId="2" pageNumber="371">
<taxonomicName id="4C444D39FF9CFFBEFF2DFCC781B44B67" authorityName="Rathbun" authorityYear="1909" box="[151,391,799,823]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF9CFFBEFF2DFCC781B44B67" box="[151,391,799,823]" italics="true" pageId="2" pageNumber="371">Pseudophilyra olivacea</emphasis>
</taxonomicName>
<treatmentCitation id="0AE510ABFF9CFFBEFE1BFCC782494B67" author="Shen, C. J. &amp; Dai A. Y." box="[417,634,799,823]" page="29" pageId="2" pageNumber="371" year="1964">
<bibRefCitation id="EFD54B4BFF9CFFBEFE1BFCC782494B67" author="Shen, C. J. &amp; Dai A. Y." box="[417,634,799,823]" pageId="2" pageNumber="371" refId="ref7383" refString="Shen, C. J. &amp; Dai A. Y. (1964) Illustrations of animals in China (Crustacea part II). (unnumbered). Beijing. pp. 1 - 142, 277 figs." type="journal volume" year="1964">Shen &amp; Dai 1964: 29</bibRefCitation>
</treatmentCitation>
, unnumbered fig.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF9CFFBEFF2DFCBD80E44CB7" pageId="2" pageNumber="371" type="materials_examined">
<paragraph id="8BFB36BAFF9CFFBEFF2DFCBD80E44CB7" blockId="2.[151,1437,869,1976]" pageId="2" pageNumber="371">
<emphasis id="B930EAA8FF9CFFBEFF2DFCBD81484B2F" bold="true" box="[151,379,869,895]" pageId="2" pageNumber="371">Material examined</emphasis>
.
<specimenCount id="9D42FD33FF9CFFBEFE30FCBD81E44B2F" box="[394,471,869,895]" pageId="2" pageNumber="371" type="male">1 male</specimenCount>
(10.4 ×
<quantity id="4CBC9B5FFF9CFFBEFD81FCBD82964BD0" box="[571,677,869,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="371" unit="mm" value="12.2">12.2 mm</quantity>
),
<materialsCitation id="3B2C3CE7FF9CFFBEFD03FCBD83FB4B97" ID-GBIF-Occurrence="3073881303" collectingDate="2012-10-16" collectionCode="ZRC" collectorName="J. J. Eeo" country="Malaysia" county="West" latitude="3.2241666" location="Jeram Polychaete Reef" longLatPrecision="21" longitude="101.30361" municipality="Kuala" pageId="2" pageNumber="371" specimenCode="ZRC 2020.0345" specimenCount="1" specimenCount-female="1" stateProvince="Selangor">
<specimenCount id="9D42FD33FF9CFFBEFD03FCBD83284B2F" box="[697,795,869,895]" pageId="2" pageNumber="371" type="female">1 female</specimenCount>
(8.5 ×
<quantity id="4CBC9B5FFF9CFFBEFCCBFCBD83EB4BD0" box="[881,984,869,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="2" pageNumber="371" unit="mm" value="10.1">10.1 mm</quantity>
) (
<specimenCode id="DBE29EC1FF9CFFBEFC4AFCBD849F4BD0" box="[1008,1196,869,896]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="371">ZRC 2020.0345</specimenCode>
), station BLB,
<location id="8E9B6061FF9CFFBEFAE3FCBE81604BF3" LSID="urn:lsid:plazi:treatment:03ED87ACFF9CFFB6FF2DFE2683844E01:8E9B6061FF9CFFBEFAE3FCBE81604BF3" country="Malaysia" county="West" latitude="3.2241666" longLatPrecision="21" longitude="101.30361" municipality="Kuala" name="Jeram Polychaete Reef" pageId="2" pageNumber="371" stateProvince="Selangor">Jeram Polychaete Reef</location>
, intertidal muddy shore and intermittent sabellariid reefs, middle shore,
<location id="8E9B6061FF9CFFBEFB29FC5284EA4BF4" LSID="urn:lsid:plazi:treatment:03ED87ACFF9CFFB6FF2DFE2683844E01:8E9B6061FF9CFFBEFB29FC5284EA4BF4" box="[1171,1241,906,932]" country="Malaysia" county="West" latitude="3.2241666" longLatPrecision="21" longitude="101.30361" municipality="Kuala" name="Jeram" pageId="2" pageNumber="371" stateProvince="Selangor">Jeram</location>
,
<collectingMunicipality id="6B9FACC0FF9CFFBEFB5DFC5185194BF3" box="[1255,1322,905,931]" pageId="2" pageNumber="371">Kuala</collectingMunicipality>
<collectingRegion id="4980F858FF9CFFBEFA89FC5185AB4BF3" box="[1331,1432,905,931]" country="Malaysia" name="Selangor" pageId="2" pageNumber="371">Selangor</collectingRegion>
,
<collectingCounty id="629A4E36FF9CFFBEFF2DFC7680FD4B98" box="[151,206,942,968]" pageId="2" pageNumber="371">West</collectingCounty>
<collectingCountry id="F353762AFF9CFFBEFF6FFC75810D4B97" box="[213,318,941,967]" name="Malaysia" pageId="2" pageNumber="371">Malaysia</collectingCountry>
,
<geoCoordinate id="EE70507DFF9CFFBEFEF2FC7581F04B97" box="[328,451,941,967]" degrees="3" direction="north" minutes="13" orientation="latitude" pageId="2" pageNumber="371" precision="15" seconds="27" value="3.2241666">3°1327”N</geoCoordinate>
<geoCoordinate id="EE70507DFF9CFFBEFE73FC7582534B97" box="[457,608,941,967]" degrees="101" direction="east" minutes="18" orientation="longitude" pageId="2" pageNumber="371" precision="15" seconds="13" value="101.30361">101°1813”E</geoCoordinate>
, coll.
<collectorName id="26B1536CFF9CFFBEFD18FC7683334B98" box="[674,768,942,968]" pageId="2" pageNumber="371">J. J. Eeo</collectorName>
,
<date id="FFFA107AFF9CFFBEFCB0FC7583FB4B97" box="[778,968,941,967]" pageId="2" pageNumber="371" value="2012-10-16">
<collectingDate id="EFBEE992FF9CFFBEFCB0FC7583FB4B97" box="[778,968,941,967]" pageId="2" pageNumber="371" value="2012-10-16">16 October 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2C3CE7FF9CFFBEFC68FC7583804BBB" ID-GBIF-Occurrence="3073881305" collectingDate="1938-10" collectionCode="ZRC" collectorName="M. W. F. Tweedie" country="Malaysia" location="Muar" municipality="Peninsular" pageId="2" pageNumber="371" specimenCount="1" specimenCount-male="1" stateProvince="Johor">
<specimenCount id="9D42FD33FF9CFFBEFC68FC75842F4B97" box="[978,1052,941,967]" pageId="2" pageNumber="371" type="male">1 male</specimenCount>
(6.9 ×
<quantity id="4CBC9B5FFF9CFFBEFBD1FC7584F24B98" box="[1131,1217,941,968]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="371" unit="mm" value="8.7">8.7 mm</quantity>
) (
<collectionCode id="ED55AE7FFF9CFFBEFB6DFC75853C4B97" box="[1239,1295,941,967]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="371">ZRC</collectionCode>
1970.8.8.7),
<location id="8E9B6061FF9CFFBEFF2DFC0A80E64BBC" LSID="urn:lsid:plazi:treatment:03ED87ACFF9CFFB6FF2DFE2683844E01:8E9B6061FF9CFFBEFF2DFC0A80E64BBC" box="[151,213,978,1004]" country="Malaysia" municipality="Peninsular" name="Muar" pageId="2" pageNumber="371" stateProvince="Johor">Muar</location>
,
<collectingRegion id="4980F858FF9CFFBEFF65FC09812C4BBB" box="[223,287,977,1003]" country="Malaysia" name="Johor" pageId="2" pageNumber="371">Johor</collectingRegion>
,
<collectingMunicipality id="6B9FACC0FF9CFFBEFE92FC0981AD4BBB" box="[296,414,977,1003]" pageId="2" pageNumber="371">Peninsular</collectingMunicipality>
<collectingCountry id="F353762AFF9CFFBEFE1EFC09823E4BBB" box="[420,525,977,1003]" name="Malaysia" pageId="2" pageNumber="371">Malaysia</collectingCountry>
, coll.
<collectorName id="26B1536CFF9CFFBEFDF7FC0A833C4BBB" box="[589,783,977,1004]" pageId="2" pageNumber="371">M. W. F. Tweedie</collectorName>
,
<date id="FFFA107AFF9CFFBEFCA3FC0983804BBB" box="[793,947,977,1003]" pageId="2" pageNumber="371" value="1938-10">
<collectingDate id="EFBEE992FF9CFFBEFCA3FC0983804BBB" box="[793,947,977,1003]" pageId="2" pageNumber="371" value="1938-10">October 1938</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2C3CE7FF9CFFBEFC07FC0984434C5F" ID-GBIF-Occurrence="3073881307" collectingDate="2011-11-13" collectionCode="ZRC" collectorName="Peninsular" country="Malaysia" location="Sungei Buloh" municipality="Kuala" pageId="2" pageNumber="371" specimenCode="ZRC 2020.0363" specimenCount="1" specimenCount-male="1" stateProvince="Selangor">
<specimenCount id="9D42FD33FF9CFFBEFC07FC0984344BBB" box="[957,1031,977,1003]" pageId="2" pageNumber="371" type="male">1 male</specimenCount>
(11.0 ×
<quantity id="4CBC9B5FFF9CFFBEFBDAFC0984F04BBC" box="[1120,1219,977,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="2" pageNumber="371" unit="mm" value="13.1">13.1 mm</quantity>
) (
<specimenCode id="DBE29EC1FF9CFFBEFB6DFC0985A24BBC" box="[1239,1425,977,1004]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="371">ZRC 2020.0363</specimenCode>
),
<location id="8E9B6061FF9CFFBEFF2DFC2D81004C5F" LSID="urn:lsid:plazi:treatment:03ED87ACFF9CFFB6FF2DFE2683844E01:8E9B6061FF9CFFBEFF2DFC2D81004C5F" box="[151,307,1013,1039]" country="Malaysia" municipality="Kuala" name="Sungei Buloh" pageId="2" pageNumber="371" stateProvince="Selangor">Sungei Buloh</location>
,
<collectingMunicipality id="6B9FACC0FF9CFFBEFE85FC2D81B14C5F" box="[319,386,1013,1039]" pageId="2" pageNumber="371">Kuala</collectingMunicipality>
<collectingRegion id="4980F858FF9CFFBEFE33FC2D81DD4C5F" box="[393,494,1013,1039]" country="Malaysia" name="Selangor" pageId="2" pageNumber="371">Selangor</collectingRegion>
,
<collectingRegion id="4980F858FF9CFFBEFE40FC2D826C4C5F" box="[506,607,1013,1039]" country="Malaysia" name="Selangor" pageId="2" pageNumber="371">Selangor</collectingRegion>
,
<collectorName id="26B1536CFF9CFFBEFDD1FC2D82D24C5F" box="[619,737,1013,1039]" pageId="2" pageNumber="371">Peninsular</collectorName>
<collectingCountry id="F353762AFF9CFFBEFD52FC2D83624C5F" box="[744,849,1013,1039]" name="Malaysia" pageId="2" pageNumber="371">Malaysia</collectingCountry>
, coll.
<date id="FFFA107AFF9CFFBEFC2FFC2D84434C5F" box="[917,1136,1013,1039]" pageId="2" pageNumber="371" value="2011-11-13">
<collectingDate id="EFBEE992FF9CFFBEFC2FFC2D84434C5F" box="[917,1136,1013,1039]" pageId="2" pageNumber="371" value="2011-11-13">13 November 2011</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2C3CE7FF9CFFBEFBC6FC2D847F4C07" ID-GBIF-Occurrence="3073881302" collectingDate="1996-04-17" collectionCode="ZRC" collectorName="Changi &amp; P. K. L. Ng" country="Singapore" location="Singapore" pageId="2" pageNumber="371" specimenCode="ZRC 2009.0217" specimenCount="4" specimenCount-male="4">
<specimenCount id="9D42FD33FF9CFFBEFBC6FC2D84E14C5F" box="[1148,1234,1013,1039]" pageId="2" pageNumber="371" type="male">4 males</specimenCount>
(11.2 × 14.0 mm, 12.3 ×
<quantity id="4CBC9B5FFF9CFFBEFF52FBC181634C64" box="[232,336,1049,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="371" unit="mm" value="14.8">14.8 mm</quantity>
, 12.7 ×
<quantity id="4CBC9B5FFF9CFFBEFE14FBC182264C64" box="[430,533,1049,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="2" pageNumber="371" unit="mm" value="15.8">15.8 mm</quantity>
, 13.3 ×
<quantity id="4CBC9B5FFF9CFFBEFDC9FBC182EE4C64" box="[627,733,1049,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="2" pageNumber="371" unit="mm" value="16.2">16.2 mm</quantity>
), 3 ovigerous females (9.3 ×
<quantity id="4CBC9B5FFF9CFFBEFB90FBC184A34C64" box="[1066,1168,1049,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="2" pageNumber="371" unit="mm" value="11.6">11.6 mm</quantity>
, 9.9 ×
<quantity id="4CBC9B5FFF9CFFBEFB5AFBC185754C64" box="[1248,1350,1049,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="371" unit="mm" value="11.8">11.8 mm</quantity>
, 10.4 ×
<quantity id="4CBC9B5FFF9CFFBEFF2DFBE580CF4C08" box="[151,252,1085,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="371" unit="mm" value="12.2">12.2 mm</quantity>
) (
<specimenCode id="DBE29EC1FF9CFFBEFEA9FBE581FC4C07" box="[275,463,1085,1111]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="371">ZRC 2009.0217</specimenCode>
), off
<collectorName id="26B1536CFF9CFFBEFDB3FBE5826E4C07" box="[521,605,1085,1111]" pageId="2" pageNumber="371">Changi</collectorName>
,
<collectingCountry id="F353762AFF9CFFBEFDD2FBE582EF4C07" box="[616,732,1085,1111]" name="Singapore" pageId="2" pageNumber="371">Singapore</collectingCountry>
, coll.
<collectorName id="26B1536CFF9CFFBEFC9AFBE683914C08" box="[800,930,1086,1112]" pageId="2" pageNumber="371">P. K. L. Ng</collectorName>
,
<date id="FFFA107AFF9CFFBEFC17FBE5847F4C07" box="[941,1100,1085,1111]" pageId="2" pageNumber="371" value="1996-04-17">
<collectingDate id="EFBEE992FF9CFFBEFC17FBE5847F4C07" box="[941,1100,1085,1111]" pageId="2" pageNumber="371" value="1996-04-17">17 April 1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2C3CE7FF9CFFBEFBE2FBE584E74C2B" ID-GBIF-Occurrence="3073881306" collectingDate="1987-11-18" collectionCode="ZRC" collectorName="Reef Ecology Study Team" country="Singapore" location="River" pageId="2" pageNumber="371" specimenCode="ZRC 2014.0176" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D42FD33FF9CFFBEFBE2FBE584974C07" box="[1112,1188,1085,1111]" pageId="2" pageNumber="371" type="male">1 male</specimenCount>
(6.8 × 8.0 mm) (
<specimenCode id="DBE29EC1FF9CFFBEFADEFBE581204C2C" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="371">ZRC 2014.0176</specimenCode>
),
<collectingCountry id="F353762AFF9CFFBEFE9FFBB981A44C2B" box="[293,407,1121,1147]" name="Singapore" pageId="2" pageNumber="371">Singapore</collectingCountry>
<location id="8E9B6061FF9CFFBEFE27FBB981E84C2B" LSID="urn:lsid:plazi:treatment:03ED87ACFF9CFFB6FF2DFE2683844E01:8E9B6061FF9CFFBEFE27FBB981E84C2B" box="[413,475,1121,1147]" country="Singapore" name="River" pageId="2" pageNumber="371">River</location>
mouth, dredge, coll.
<collectorName id="26B1536CFF9CFFBEFD72FBB983DC4C2C" box="[712,1007,1121,1148]" pageId="2" pageNumber="371">Reef Ecology Study Team</collectorName>
,
<date id="FFFA107AFF9CFFBEFC43FBB984E74C2B" box="[1017,1236,1121,1147]" pageId="2" pageNumber="371" value="1987-11-18">
<collectingDate id="EFBEE992FF9CFFBEFC43FBB984E74C2B" box="[1017,1236,1121,1147]" pageId="2" pageNumber="371" value="1987-11-18">18 November 1987</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2C3CE7FF9CFFBEFB65FBB985B14CCF" ID-GBIF-Occurrence="3073881301" collectingDate="1989-02-22" collectionCode="ZRC" collectorName="Reef Ecology Study Team" country="Singapore" location="Kallang River" pageId="2" pageNumber="371" specimenCode="ZRC 1990.8338" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D42FD33FF9CFFBEFB65FBB9851A4C2B" box="[1247,1321,1121,1147]" pageId="2" pageNumber="371" type="male">1 male</specimenCount>
(6.9 ×
<quantity id="4CBC9B5FFF9CFFBEFAC3FBB980F74CF0" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="2" pageNumber="371" unit="mm" value="8.1">8.1 mm</quantity>
) (
<specimenCode id="DBE29EC1FF9CFFBEFF61FB5D81A64CCF" box="[219,405,1157,1183]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="371">ZRC 1990.8338</specimenCode>
),
<location id="8E9B6061FF9CFFBEFE13FB5D82754CCF" LSID="urn:lsid:plazi:treatment:03ED87ACFF9CFFB6FF2DFE2683844E01:8E9B6061FF9CFFBEFE13FB5D82754CCF" box="[425,582,1157,1183]" country="Singapore" name="Kallang River" pageId="2" pageNumber="371">Kallang River</location>
basin, dredge, dredge, coll.
<collectorName id="26B1536CFF9CFFBEFC3EFB5D849E4CF0" box="[900,1197,1157,1184]" pageId="2" pageNumber="371">Reef Ecology Study Team</collectorName>
,
<date id="FFFA107AFF9CFFBEFB02FB5D85B14CCF" box="[1208,1410,1157,1183]" pageId="2" pageNumber="371" value="1989-02-22">
<collectingDate id="EFBEE992FF9CFFBEFB02FB5D85B14CCF" box="[1208,1410,1157,1183]" pageId="2" pageNumber="371" value="1989-02-22">22 February 1989</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2C3CE7FF9CFFBEFA34FB5D80E14CB7" ID-GBIF-Occurrence="3073881304" collectingDate="1989-02-23" collectionCode="ZRC" collectorName="Reef Ecology Study Team" country="Singapore" location="Kallang River" pageId="2" pageNumber="371" specimenCode="ZRC 2001.2242" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D42FD33FF9CFFBEFA34FB5D80D04C93" pageId="2" pageNumber="371" type="female">1 female</specimenCount>
(7.8 ×
<quantity id="4CBC9B5FFF9CFFBEFE88FB7181BB4C94" box="[306,392,1193,1220]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="371" unit="mm" value="9.3">9.3 mm</quantity>
) (
<specimenCode id="DBE29EC1FF9CFFBEFE27FB71826B4C93" box="[413,600,1193,1219]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="371">ZRC 2001.2242</specimenCode>
),
<location id="8E9B6061FF9CFFBEFDD3FB7183354C93" LSID="urn:lsid:plazi:treatment:03ED87ACFF9CFFB6FF2DFE2683844E01:8E9B6061FF9CFFBEFDD3FB7183354C93" box="[617,774,1193,1219]" country="Singapore" name="Kallang River" pageId="2" pageNumber="371">Kallang River</location>
basin, dredge, coll.
<collectorName id="26B1536CFF9CFFBEFC5FFB7185384C94" box="[997,1291,1193,1220]" pageId="2" pageNumber="371">Reef Ecology Study Team</collectorName>
,
<date id="FFFA107AFF9CFFBEFAAFFB7180E14CB7" pageId="2" pageNumber="371" value="1989-02-23">
<collectingDate id="EFBEE992FF9CFFBEFAAFFB7180E14CB7" pageId="2" pageNumber="371" value="1989-02-23">23 February 1989</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF9CFFBEFF7DFB2981EE4D5C" box="[199,477,1265,1292]" pageId="2" pageNumber="371" type="diagnosis">
<paragraph id="8BFB36BAFF9CFFBEFF7DFB2981EE4D5C" blockId="2.[151,1437,869,1976]" box="[199,477,1265,1292]" pageId="2" pageNumber="371">
<emphasis id="B930EAA8FF9CFFBEFF7DFB2981094D5B" bold="true" box="[199,314,1265,1291]" pageId="2" pageNumber="371">Diagnosis</emphasis>
. As for genus.
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF9CFFBAFF7DFACD83624E93" lastPageId="6" lastPageNumber="375" pageId="2" pageNumber="371" type="description">
<paragraph id="8BFB36BAFF9CFFBEFF7DFACD82E84F53" blockId="2.[151,1437,869,1976]" pageId="2" pageNumber="371">
<emphasis id="B930EAA8FF9CFFBEFF7DFACD81864D7F" bold="true" box="[199,437,1301,1327]" pageId="2" pageNumber="371">Description of male</emphasis>
. Carapace longitudinally ovate, distinctly longer than wide (length to width ratio 1.17 1.26); most regions indistinct, branchio-cardiac grooves just visible, cardiac region distinct, swollen; dorsal surface glabrous, covered with minute to small granules and small depressions; branchial region usually with oblique row of granules, sometimes undiscernible (
<figureCitation id="137F2A3FFF9CFFBEFDFEFA5982F54DCC" box="[580,710,1409,1436]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1A, B</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFD6BFA5983264DCC" box="[721,789,1409,1436]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2A, B</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFC9AFA5983544DCB" box="[800,871,1409,1435]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="2" pageNumber="371">3AC</figureCitation>
). Frontal region not produced anteriorly, with shallow median longitudinal depression; frontal margin granulated, with low median triangular tooth (
<figureCitation id="137F2A3FFF9CFFBEFB63FA7D850A4DEF" box="[1241,1337,1445,1471]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1B</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFAFFFA7D855B4DEF" box="[1349,1384,1445,1471]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2B</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFAC9FA7D809F4DB4" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="2" pageNumber="371">5D, E</figureCitation>
). Antennules folding transversely. Antennae short, longitudinally inserted between antennular fossa and base of ocular peduncle. Orbits small, rounded, upper orbital margin entire. Eyes short, retractable (
<figureCitation id="137F2A3FFF9CFFBEFB2DFA3584C94E57" box="[1175,1274,1517,1543]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1D</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFABCFA35851F4E57" box="[1286,1324,1517,1543]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2D</figureCitation>
). Anterior margin of efferent branchial channel strongly concave, not projecting beyond frontal margin, separated from crenulate efferent margin on subhepatic region by longitudinal groove. Hepatic facet well defined by rows of granules and cristae, elongate; distal part with low granules, almost reaching frontal margin; distal part of lower margin prominently angular, with broad tooth on distal one-third (
<figureCitation id="137F2A3FFF9CFFBEFC9AF9A583904EC8" box="[800,931,1661,1688]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1B, D</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFC14F9A583C04EC8" box="[942,1011,1661,1688]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2B, D</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFC44F9A584174EC7" box="[1022,1060,1661,1687]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="2" pageNumber="371">4A</figureCitation>
). Lateral and posterior margins of carapace irregularly granulated; posterolateral margin convex, usually demarcated from posterior margin by sharp lobiform tooth; posterior margin gently convex (
<figureCitation id="137F2A3FFF9CFFBEFD08F91D83064EB0" box="[690,821,1733,1760]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1A, B</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFCFBF91D83B64EB0" box="[833,901,1733,1760]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2A, B</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFC2AF91D83E54E8F" box="[912,982,1733,1760]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="2" pageNumber="371">3AC</figureCitation>
). Pterygostomial region granulated; subbranchial region almost smooth (
<figureCitation id="137F2A3FFF9CFFBEFDB2F93182584F53" box="[520,619,1769,1795]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1D</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFDC2F93182AF4F53" box="[632,668,1769,1795]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2D</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFD12F93182FD4F53" box="[680,718,1769,1795]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="2" pageNumber="371">4A</figureCitation>
).
</paragraph>
<paragraph id="8BFB36BAFF9CFFBEFF7DF8D585154FE8" blockId="2.[151,1437,869,1976]" pageId="2" pageNumber="371">
Third maxilliped exopod wider than basal part of endopod, forming petaliform structure with strongly convex outer margin, with submarginal row of low to very low granules along inner margin (
<figureCitation id="137F2A3FFF9CFFBEFBE9F8E984874F1B" box="[1107,1204,1841,1867]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1C</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFB7BF8E984D74F1B" box="[1217,1252,1841,1867]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2C</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFB4AF8E985224F1B" box="[1264,1297,1841,1867]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="2" pageNumber="371">3E</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFAA7F8E985564F1C" box="[1309,1381,1841,1868]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="2" pageNumber="371">5AC</figureCitation>
); endopod with completely fused basis-ischium, distinctly longer than merus, with shallow submarginal sulcus near inner margin; merus acutely triangular with pointed tip, margins unevenly granulated; palp on inner surface, dactylus longest (
<figureCitation id="137F2A3FFF9CFFBEFF4DF845816A4FE7" box="[247,345,1949,1975]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1C</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFEDFF84581BB4FE7" box="[357,392,1949,1975]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2C</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFE2EF84581864FE7" box="[404,437,1949,1975]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="2" pageNumber="371">3E</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFE7BF845823A4FE8" box="[449,521,1949,1976]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="2" pageNumber="371">5AC</figureCitation>
). Coxa prominent, forming curved plate (
<figureCitation id="137F2A3FFF9CFFBEFC63F84584094FE7" box="[985,1082,1949,1975]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="2" pageNumber="371">Figs. 1C</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFBFDF845845A4FE7" box="[1095,1129,1949,1975]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="2" pageNumber="371">2C</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFBCFF84584A44FE7" box="[1141,1175,1949,1975]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="2" pageNumber="371">3E</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFB18F84584F44FE7" box="[1186,1223,1949,1975]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="2" pageNumber="371">4A</figureCitation>
,
<figureCitation id="137F2A3FFF9CFFBEFB69F84585294FE8" box="[1235,1306,1949,1976]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="2" pageNumber="371">5AC</figureCitation>
).
</paragraph>
<caption id="DF3B6632FF9DFFBFFF2DF85982824FB1" ID-DOI="http://doi.org/10.5281/zenodo.4674239" ID-Zenodo-Dep="4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="3" pageNumber="372" startId="3.[151,250,1921,1945]" targetBox="[169,1418,181,1897]" targetPageId="3">
<paragraph id="8BFB36BAFF9DFFBFFF2DF85982824FB1" blockId="3.[151,1437,1921,2017]" pageId="3" pageNumber="372">
<emphasis id="B930EAA8FF9DFFBFFF2DF859813F4FC9" bold="true" box="[151,268,1921,1945]" pageId="3" pageNumber="372">FIGURE 1</emphasis>
.
<taxonomicName id="4C444D39FF9DFFBFFEA2F85982E44FC9" authority="(Targioni-Tozzetti, 1877)" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[280,727,1921,1945]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9DFFBFFEA2F85981E34FC9" box="[280,464,1921,1945]" italics="true" pageId="3" pageNumber="372">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF9DFFBFFE67F85982FC4FC9" author="Targioni-Tozzetti, A." box="[477,719,1921,1945]" pageId="3" pageNumber="372" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
)
</taxonomicName>
, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg.
</paragraph>
</caption>
<caption id="DF3B6632FF9AFFB8FF2DF85683534FBE" ID-DOI="http://doi.org/10.5281/zenodo.4674241" ID-Zenodo-Dep="4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="4" pageNumber="373" startId="4.[151,250,1934,1958]" targetBox="[164,1422,181,1910]" targetPageId="4">
<paragraph id="8BFB36BAFF9AFFB8FF2DF85683534FBE" blockId="4.[151,1437,1934,2030]" pageId="4" pageNumber="373">
<emphasis id="B930EAA8FF9AFFB8FF2DF856813E4FF6" bold="true" box="[151,269,1934,1958]" pageId="4" pageNumber="373">FIGURE 2</emphasis>
.
<taxonomicName id="4C444D39FF9AFFB8FEA3F85682E94FF6" authority="(Targioni-Tozzetti, 1877)" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[281,730,1934,1958]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9AFFB8FEA3F85681E14FF6" box="[281,466,1934,1958]" italics="true" pageId="4" pageNumber="373">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF9AFFB8FE65F85682E14FF6" author="Targioni-Tozzetti, A." box="[479,722,1934,1958]" pageId="4" pageNumber="373" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
)
</taxonomicName>
, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg.
</paragraph>
</caption>
<caption id="DF3B6632FF9BFFB9FF2DFB3E84F64D16" ID-DOI="http://doi.org/10.5281/zenodo.4674243" ID-Zenodo-Dep="4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="5" pageNumber="374" startId="5.[151,250,1254,1278]" targetBox="[151,1436,181,1230]" targetPageId="5">
<paragraph id="8BFB36BAFF9BFFB9FF2DFB3E84F64D16" blockId="5.[151,1436,1254,1350]" pageId="5" pageNumber="374">
<emphasis id="B930EAA8FF9BFFB9FF2DFB3E813C4CAE" bold="true" box="[151,271,1254,1278]" pageId="5" pageNumber="374">FIGURE 3</emphasis>
.
<taxonomicName id="4C444D39FF9BFFB9FEA4FB3E82D44CAE" authority="(Targioni-Tozzetti, 1877)" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[286,743,1254,1278]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9BFFB9FEA4FB3E81EA4CAE" box="[286,473,1254,1278]" italics="true" pageId="5" pageNumber="374">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF9BFFB9FE53FB3E82D34CAE" author="Targioni-Tozzetti, A." box="[489,736,1254,1278]" pageId="5" pageNumber="374" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
)
</taxonomicName>
. A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia. AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped.
</paragraph>
</caption>
<paragraph id="8BFB36BAFF9BFFB9FF7DFAAC81A04E12" blockId="5.[151,1437,1396,2035]" pageId="5" pageNumber="374">
Chelipeds subequal, longer, more robust in adults; merus subcylindrical, symmetrical along entire length, surface minutely granulate, especially along margins; carpus smooth, unarmed (
<figureCitation id="137F2A3FFF9BFFB9FBB4FA4084464DE2" box="[1038,1141,1432,1458]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1A</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FB39FA40849B4DE2" box="[1155,1192,1432,1458]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2A</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FB0CFA4084CD4DE2" box="[1206,1278,1432,1458]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="5" pageNumber="374">3AC</figureCitation>
). Chela stout, surfaces smooth; fingers as long as or subequal to length of palm, pollex bent at angle of 4560º along longitudinal axis; cutting edge of pollex with broad, large subproximal tooth lined with denticles, rest of edge with small denticles; cutting edge of dactylus with subproximal concavity lined with denticles, rest of edge with small denticles (
<figureCitation id="137F2A3FFF9BFFB9FF25F9F081304E12" box="[159,259,1576,1602]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1A</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FEB5F9F081004E12" box="[271,307,1576,1602]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2A</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FE85F9F081B44E12" box="[319,391,1576,1602]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="5" pageNumber="374">3AC</figureCitation>
).
</paragraph>
<paragraph id="8BFB36BAFF9BFFB9FF7DF99484474EFF" blockId="5.[151,1437,1396,2035]" pageId="5" pageNumber="374">
First to fifth ambulatory legs slender, short; second and third legs longest, fourth leg shortest; merus distinctly longer than carpus and propodus, margins lined with small granules (more prominent on ventral margin); dactylus longer than propodus, lanceolate, terminating in cornute tips (
<figureCitation id="137F2A3FFF9BFFB9FCF0F94C83F94EFF" box="[842,970,1684,1711]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1A, F</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FC6FF94C84264EFF" box="[981,1045,1684,1711]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2A, F</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FB9BF94C84554EFF" box="[1057,1126,1684,1711]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="5" pageNumber="374">3A, B</figureCitation>
).
</paragraph>
<paragraph id="8BFB36BAFF9BFFB9FF7DF960834B4FFA" blockId="5.[151,1437,1396,2035]" pageId="5" pageNumber="374">
Thoracic sternites transversely narrow, surface finely granulated (
<figureCitation id="137F2A3FFF9BFFB9FC0DF96084294E82" box="[951,1050,1720,1746]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FB93F96084794E82" box="[1065,1098,1720,1746]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FBE3F960844E4E82" box="[1113,1149,1720,1746]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="5" pageNumber="374">3D</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FB31F96084814E82" box="[1163,1202,1720,1746]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">4A</figureCitation>
); sternites 13 completely fused without trace of sutures; sternite 3 separated from sternite 4 by shallow groove (
<figureCitation id="137F2A3FFF9BFFB9FB77F90485194EA6" box="[1229,1322,1756,1782]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">Fig. 4A</figureCitation>
); exposed sternites 47 progressively narrower (
<figureCitation id="137F2A3FFF9BFFB9FD86F8D882864F4B" box="[572,693,1792,1819]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">Fig. 4A, B</figureCitation>
); sternite 8 visible when pleon closed, between margins of somites 2 and 3 (
<figureCitation id="137F2A3FFF9BFFB9FF46F8FC81654F6E" box="[252,342,1828,1854]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">Fig. 4B</figureCitation>
). Penis arising under constriction between sternites 7 and 8. Sternopleonal cavity deep, reaching to mid-distance between fused thoracic sternites 13 (
<figureCitation id="137F2A3FFF9BFFB9FD4AF890837C4F32" box="[752,847,1864,1890]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FCE1F890834E4F32" box="[859,893,1864,1890]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FC32F890839D4F32" box="[904,942,1864,1890]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">4A</figureCitation>
); margin lined with granules, those on distal part proportionately larger (
<figureCitation id="137F2A3FFF9BFFB9FE74F8B4821A4FD6" box="[462,553,1900,1926]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">Fig. 4A</figureCitation>
); pleonal locking mechanism formed by small, low projection at distomarginal edge of sternite 5 (adjacent to sternite 4) with shallow depression.
</paragraph>
<paragraph id="8BFB36BAFF9BFFBAFF7DF86C85BB48A9" blockId="5.[151,1437,1396,2035]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374">
Pleon narrow, slender, long (
<figureCitation id="137F2A3FFF9BFFB9FDB0F86C82594F9E" box="[522,618,1972,1998]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FDCDF86C82AB4F9E" box="[631,664,1972,1998]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FD1EF86C82FA4F9E" box="[676,713,1972,1998]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="5" pageNumber="374">3D</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FD6FF86C83254F9E" box="[725,790,1972,1999]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="5" pageNumber="374">5F, G</figureCitation>
); somite 1 longitudinally narrow, wide (
<figureCitation id="137F2A3FFF9BFFB9FB63F86C850A4F9E" box="[1241,1337,1972,1998]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FAFCF86C85544F9E" box="[1350,1383,1972,1998]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FAC9F86C85AB4F9E" box="[1395,1432,1972,1998]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="5" pageNumber="374">3D</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FF2DF80080894FA2" box="[151,186,2008,2034]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">4B</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FF72F800813F4FA2" box="[200,268,2008,2035]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="5" pageNumber="374">5F, G</figureCitation>
); somite 2 yoke-like, reaching coxae of fourth ambulatory leg (
<figureCitation id="137F2A3FFF9BFFB9FC5DF80084794FA2" box="[999,1098,2008,2034]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="5" pageNumber="374">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FBE3F80084494FA2" box="[1113,1146,2008,2034]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="5" pageNumber="374">2E</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FB32F80084984FA2" box="[1160,1195,2008,2034]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="5" pageNumber="374">4B</figureCitation>
,
<figureCitation id="137F2A3FFF9BFFB9FB03F80084CE4FA2" box="[1209,1277,2008,2035]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="5" pageNumber="374">5F, G</figureCitation>
); somites 36 fused, forming elongate trapezoidal plate, very shallow suture just visible between somites 3 and 4, surface smooth (
<figureCitation id="137F2A3FFF98FFBAFF25FF6381304885" box="[159,259,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="6" pageNumber="375">Figs. 3D</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFEB4FF6381024885" box="[270,305,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="6" pageNumber="375">4B</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFE87FF62814D4885" box="[317,382,186,213]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="6" pageNumber="375">5F, G</figureCitation>
); somite 6 longitudinally subrectangular, free, surface unarmed, lateral margins gently sinuous to gently convex (
<figureCitation id="137F2A3FFF98FFBAFEDBFF0781F248A9" box="[353,449,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="6" pageNumber="375">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFE74FF0781DC48A9" box="[462,495,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="6" pageNumber="375">2E</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFE41FF07822C48A9" box="[507,543,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="6" pageNumber="375">3D</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFD91FF07826348A9" box="[555,592,223,249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="6" pageNumber="375">4A</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFDE1FF0682AF48A9" box="[603,668,222,249]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="6" pageNumber="375">5F, G</figureCitation>
); telson triangular, longer than wide (
<figureCitation id="137F2A3FFF98FFBAFBFAFF07849348A9" box="[1088,1184,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="6" pageNumber="375">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFB16FF0784FD48A9" box="[1196,1230,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="6" pageNumber="375">2E</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFB63FF0784CD48A9" box="[1241,1278,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="6" pageNumber="375">3D</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFAB3FF07851D48A9" box="[1289,1326,223,249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="6" pageNumber="375">4A</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFA80FF06854848A9" box="[1338,1403,222,249]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="6" pageNumber="375">5F, G</figureCitation>
).
</paragraph>
<paragraph id="8BFB36BAFF98FFBAFF7DFEDB84D849D9" blockId="6.[151,1437,151,393]" pageId="6" pageNumber="375">
G1 elongate, slender, shaft gradually tapering distally; distal quarter sharply bent backwards 120150º from the longitudinal axis; with curved, tapering subdistal process, outer margin lined with 1 or 2 rows of 58 spines; distal part elongate, curved, hook-shape, reaches to, overlaps or overreaches shaft, opening distal (
<figureCitation id="137F2A3FFF98FFBAFB21FE9385A34935" box="[1179,1424,331,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).AD, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; EH, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; IL, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EL, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="6" pageNumber="375">Fig. 6AC, EG, IK</figureCitation>
). G2 short, ca. one-third length of straight part of G1; apex elongate, subspatuliform (
<figureCitation id="137F2A3FFF98FFBAFBF9FEB784EC49D9" box="[1091,1247,367,393]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).AD, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; EH, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; IL, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EL, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="6" pageNumber="375">Fig. 6D, H, L</figureCitation>
).
</paragraph>
<caption id="DF3B6632FF98FFBAFF2DFB5785B04CBF" ID-DOI="http://doi.org/10.5281/zenodo.4674245" ID-Zenodo-Dep="4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="6" pageNumber="375" startId="6.[151,250,1167,1191]" targetBox="[151,1436,417,1143]" targetPageId="6">
<paragraph id="8BFB36BAFF98FFBAFF2DFB5785B04CBF" blockId="6.[151,1437,1167,1263]" pageId="6" pageNumber="375">
<emphasis id="B930EAA8FF98FFBAFF2DFB57813F4CF7" bold="true" box="[151,268,1167,1191]" pageId="6" pageNumber="375">FIGURE 4</emphasis>
.
<taxonomicName id="4C444D39FF98FFBAFEA2FB5782E44CF7" authority="(Targioni-Tozzetti, 1877)" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[280,727,1167,1191]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF98FFBAFEA2FB5781E34CF7" box="[280,464,1167,1191]" italics="true" pageId="6" pageNumber="375">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF98FFBAFE67FB5782FC4CF7" author="Targioni-Tozzetti, A." box="[477,719,1167,1191]" pageId="6" pageNumber="375" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
)
</taxonomicName>
, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson.
</paragraph>
</caption>
<paragraph id="8BFB36BAFF98FFBAFF7DFAC583424D98" blockId="6.[151,1437,1309,2020]" pageId="6" pageNumber="375">
<emphasis id="B930EAA8FF98FFBAFF7DFAC5811A4D67" bold="true" box="[199,297,1309,1335]" pageId="6" pageNumber="375">Females</emphasis>
. The carapaces of the female specimens are similar to the males in all aspects, but the chelipeds are proportionately shorter (
<figureCitation id="137F2A3FFF98FFBAFE10FA99821A4D0C" box="[426,553,1345,1372]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="6" pageNumber="375">Fig. 7A, D</figureCitation>
). The female pleon is longitudinally ovate, with somites 1 and 2 free, somites 36 completely fused to form a domed plate that completely covers the thoracic sternum (
<figureCitation id="137F2A3FFF98FFBAFB2DFABD85224DD0" box="[1175,1297,1381,1408]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="6" pageNumber="375">Fig. 7B, E</figureCitation>
). The telson is triangular and mobile (
<figureCitation id="137F2A3FFF98FFBAFE0DFA5182024DF4" box="[439,561,1417,1444]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="6" pageNumber="375">Fig. 7B, E</figureCitation>
). The vulvae are large, obliquely ovate and positioned on the anterior part of sternite 6, without any sign of a sternal vulvar cover (
<figureCitation id="137F2A3FFF98FFBAFD54FA7583574D98" box="[750,868,1453,1480]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="6" pageNumber="375">Fig. 7C, F</figureCitation>
).
</paragraph>
<paragraph id="8BFB36BAFF98FFBAFF7DFA0983624E93" blockId="6.[151,1437,1309,2020]" pageId="6" pageNumber="375">
<emphasis id="B930EAA8FF98FFBAFF7DFA0981284DBB" bold="true" box="[199,283,1489,1515]" pageId="6" pageNumber="375">Colour</emphasis>
.
<bibRefCitation id="EFD54B4BFF98FFBAFE91FA0982484DBB" author="Targioni-Tozzetti, A." box="[299,635,1489,1515]" pageId="6" pageNumber="375" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti (1877: 203)</bibRefCitation>
described the fresh colours: “Non ho che un solo maschio perfetto, e una femmina in istato di muta assai guasta; il primo è colorato in giallastro e coperto come di una patina bruno fuliginosa, che lascia scoperte le zampe rosso-brune, e i tarsi giallo-crocei.” [I have only one perfect male, and one female in which is a poor state due to imperfect moulting; the first is yellowish colored and covered as with a sooty brown patina which covers the reddish-brown and reticulated yellow legs.] This “sooty appearance” as described by
<bibRefCitation id="EFD54B4BFF98FFBAFF2DF95D819A4ECF" author="Targioni-Tozzetti, A." box="[151,425,1669,1695]" pageId="6" pageNumber="375" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti (1877)</bibRefCitation>
is very obvious in several of the specimens from
<collectingCountry id="F353762AFF98FFBAFC75F95D84054ECF" box="[975,1078,1669,1695]" name="Malaysia" pageId="6" pageNumber="375">Malaysia</collectingCountry>
and
<collectingCountry id="F353762AFF98FFBAFBD0F95D84EF4ECF" box="[1130,1244,1669,1695]" name="Singapore" pageId="6" pageNumber="375">Singapore</collectingCountry>
(see
<figureCitation id="137F2A3FFF98FFBAFAAFF95D85A44EF0" box="[1301,1431,1669,1696]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="6" pageNumber="375">Figs. 3A, B</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFF2DF972808E4E94" box="[151,189,1706,1732]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="6" pageNumber="375">7D</figureCitation>
). In life, the specimens from
<collectingCountry id="F353762AFF98FFBAFDBFF97182444E93" box="[517,631,1705,1731]" name="Singapore" pageId="6" pageNumber="375">Singapore</collectingCountry>
are greyish-brown.
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF98FFB6FF7DF91585BC4D05" lastPageId="10" lastPageNumber="379" pageId="6" pageNumber="375" type="discussion">
<paragraph id="8BFB36BAFF98FFBBFF7DF915849649D9" blockId="6.[151,1437,1309,2020]" lastBlockId="7.[151,1437,151,646]" lastPageId="7" lastPageNumber="376" pageId="6" pageNumber="375">
<emphasis id="B930EAA8FF98FFBAFF7DF91581074EB7" bold="true" box="[199,308,1741,1767]" pageId="6" pageNumber="375">Remarks</emphasis>
.
<taxonomicName id="4C444D39FF98FFBAFEFFF91582204EB7" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[325,531,1741,1767]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF98FFBAFEFFF91582204EB7" box="[325,531,1741,1767]" italics="true" pageId="6" pageNumber="375">Philyra fuliginosa</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFD54B4BFF98FFBAFD56F91584314EB7" author="Targioni-Tozzetti, A." box="[748,1026,1741,1767]" pageId="6" pageNumber="375" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti (1877)</bibRefCitation>
from a male (6.5 × 8.0 mm) and a female (7.0 ×
<quantity id="4CBC9B5FFF98FFBAFE8FF92981BF4F5C" box="[309,396,1777,1804]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="375" unit="mm" value="8.5">8.5 mm</quantity>
) from Java. The present specimens agree with the
<typeStatus id="54FF8818FF98FFBAFC7EF92A83C74F5C" box="[964,1012,1778,1804]" pageId="6" pageNumber="375">type</typeStatus>
description and figures well, the only apparent difference being in the proportions of the exopod of the third maxilliped. That figured by
<bibRefCitation id="EFD54B4BFF98FFBAFB61F8CD80EE4F03" author="Targioni-Tozzetti, A." pageId="6" pageNumber="375" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti (1877</bibRefCitation>
: fig.
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) shows a relatively more slender exopod but this is wider in the present specimens (
<figureCitation id="137F2A3FFF98FFBAFB43F8E1856F4F03" box="[1273,1372,1849,1875]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="6" pageNumber="375">Figs. 1C</figureCitation>
,
<figureCitation id="137F2A3FFF98FFBAFAD0F8E185A34F04" box="[1386,1424,1849,1876]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="6" pageNumber="375">5A</figureCitation>
). This may just be because they were depicted in situ; in life the exopod fits onto the sloping sides of the pterygostomial region and appear more slender than they actually are.
<bibRefCitation id="EFD54B4BFF98FFBAFCFDF85984004FCB" author="Rathbun, M. J." box="[839,1075,1921,1947]" pageId="6" pageNumber="375" pagination="107 - 114" refId="ref7110" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun (1909: 108)</bibRefCitation>
described
<taxonomicName id="4C444D39FF98FFBAFB0AF85A851F4FCB" authorityName="Rathbun" authorityYear="1909" box="[1200,1324,1921,1947]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF98FFBAFB0AF85A851F4FCB" box="[1200,1324,1921,1947]" italics="true" pageId="6" pageNumber="375">P. olivacea</emphasis>
</taxonomicName>
from one male (9.0 ×
<quantity id="4CBC9B5FFF98FFBAFEA1F87D81434F90" box="[283,368,1957,1984]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="375" unit="mm" value="7.8">7.8 mm</quantity>
) collected by a seine from the coast of Lem Ngob in the Gulf of
<collectingCountry id="F353762AFF98FFBAFB88F87D84AA4FEF" box="[1074,1177,1957,1983]" name="Thailand" pageId="6" pageNumber="375">Thailand</collectingCountry>
. She later described the species in more detail (
<bibRefCitation id="EFD54B4BFF98FFBAFE27F811827B4FB3" author="Rathbun, M. J." box="[413,584,1993,2019]" pageId="6" pageNumber="375" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">Rathbun, 1910</bibRefCitation>
), providing a photograph of the specimen and figures of the carapace, third maxillipeds and pleon.
<bibRefCitation id="EFD54B4BFF99FFBBFE1AFF4F82BF48E1" author="Rathbun, M. J." box="[416,652,151,177]" pageId="7" pageNumber="376" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">Rathbun (1910: 313)</bibRefCitation>
commented that
<taxonomicName id="4C444D39FF99FFBBFCE8FF4F83E748E1" authorityName="Rathbun" authorityYear="1909" box="[850,980,151,177]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF99FFBBFCE8FF4F83E748E1" box="[850,980,151,177]" italics="true" pageId="7" pageNumber="376">O. olivacea</emphasis>
</taxonomicName>
was close to
<taxonomicName id="4C444D39FF99FFBBFBCAFF4F853748E1" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[1136,1284,151,177]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF99FFBBFBCAFF4F853748E1" box="[1136,1284,151,177]" italics="true" pageId="7" pageNumber="376">O. fuliginosa</emphasis>
</taxonomicName>
but the latter differed in “in wanting a postero-lateral angulation, and in the form of the
<specimenCount id="9D42FD33FF99FFBBFC6CFF6383D84885" box="[982,1003,187,213]" pageId="7" pageNumber="376" type="male"></specimenCount>
abdomen and chela.” Comparing their published figures, the differences would be that in
<taxonomicName id="4C444D39FF99FFBBFD68FF07835648A9" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[722,869,223,249]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF99FFBBFD68FF07835648A9" box="[722,869,223,249]" italics="true" pageId="7" pageNumber="376">O. fuliginosa</emphasis>
</taxonomicName>
, the posterior carapace margin has two low lateral lobes (
<bibRefCitation id="EFD54B4BFF99FFBBFF65FEDB81D7494D" author="Targioni-Tozzetti, A." box="[223,484,259,285]" pageId="7" pageNumber="376" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti 1877</bibRefCitation>
: pl. 12 fig. 3a) (versus with 2 prominent lobiform teeth in
<taxonomicName id="4C444D39FF99FFBBFBD9FEDB84D0494D" authorityName="Rathbun" authorityYear="1909" box="[1123,1251,259,285]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF99FFBBFBD9FEDB84D0494D" box="[1123,1251,259,285]" italics="true" pageId="7" pageNumber="376">O. olivacea</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFD54B4BFF99FFBBFB4BFEDB85AB494D" author="Rathbun, M. J." box="[1265,1432,259,285]" pageId="7" pageNumber="376" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">Rathbun, 1910</bibRefCitation>
: p1. 11, fig. 17); the lateral margin of male pleonal somite 6 is sinuous (
<bibRefCitation id="EFD54B4BFF99FFBBFC18FEFF84944911" author="Targioni-Tozzetti, A." box="[930,1191,295,321]" pageId="7" pageNumber="376" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti 1877</bibRefCitation>
: pl. 12 fig. 3c) (versus gently convex in
<taxonomicName id="4C444D39FF99FFBBFEEDFE9381EB4935" authorityName="Rathbun" authorityYear="1909" box="[343,472,331,357]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF99FFBBFEEDFE9381EB4935" box="[343,472,331,357]" italics="true" pageId="7" pageNumber="376">O. olivacea</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFD54B4BFF99FFBBFE5CFE9382BD4935" author="Rathbun, M. J." box="[486,654,331,357]" pageId="7" pageNumber="376" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">Rathbun, 1910</bibRefCitation>
: fig. 4c); and the fingers of the chela are more strongly bent (Targioni-Tozzetti 1877: pl. 12 fig.
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) (versus less bent in
<taxonomicName id="4C444D39FF99FFBBFD06FEB7830E49D9" authorityName="Rathbun" authorityYear="1909" box="[700,829,367,393]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF99FFBBFD06FEB7830E49D9" box="[700,829,367,393]" italics="true" pageId="7" pageNumber="376">O. olivacea</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFD54B4BFF99FFBBFCF6FEB783DD49D9" author="Rathbun, M. J." box="[844,1006,367,393]" pageId="7" pageNumber="376" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">Rathbun 1910</bibRefCitation>
: p1. 11, fig. 17).
</paragraph>
<paragraph id="8BFB36BAFF99FFBBFF7DFE4B84D54AD5" blockId="7.[151,1437,151,646]" pageId="7" pageNumber="376">
In the ZRC are two lots which have originally been identified as
<taxonomicName id="4C444D39FF99FFBBFC1AFE4B841F49FD" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[928,1068,403,429]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF99FFBBFC1AFE4B841F49FD" box="[928,1068,403,429]" italics="true" pageId="7" pageNumber="376">P. fuliginosa</emphasis>
</taxonomicName>
, both from Peninsular
<collectingCountry id="F353762AFF99FFBBFA95FE4B85AB49FD" box="[1327,1432,403,429]" name="Malaysia" pageId="7" pageNumber="376">Malaysia</collectingCountry>
, one from Muar in
<collectingRegion id="4980F858FF99FFBBFED7FE6F81984981" box="[365,427,439,465]" country="Malaysia" name="Johor" pageId="7" pageNumber="376">Johor</collectingRegion>
(ZRC 1970.8.8.7) (
<bibRefCitation id="EFD54B4BFF99FFBBFD35FE6F83274981" author="Yang, C. M." box="[655,788,439,465]" pageId="7" pageNumber="376" refId="ref7562" refString="Yang, C. M. (1979) A list of Brachyura in the Zoological Reference Collection of the Department of Zoology. No. 14. Department of Zoology, National University of Singapore, 60 pp." type="book" year="1979">Yang, 1979</bibRefCitation>
) and another from Jeram in
<collectingRegion id="4980F858FF99FFBBFBE5FE6F84F04981" box="[1119,1219,439,465]" country="Malaysia" name="Selangor" pageId="7" pageNumber="376">Selangor</collectingRegion>
(ZRC 2020.0345). There are also several lots from Peninsular
<collectingCountry id="F353762AFF99FFBBFDC1FE0382D149A5" box="[635,738,475,501]" name="Malaysia" pageId="7" pageNumber="376">Malaysia</collectingCountry>
and
<collectingCountry id="F353762AFF99FFBBFCADFE0383BA49A5" box="[791,905,475,501]" name="Singapore" pageId="7" pageNumber="376">Singapore</collectingCountry>
which were identified as
<taxonomicName id="4C444D39FF99FFBBFB12FE03851149A5" authorityName="Rathbun" authorityYear="1909" box="[1192,1314,475,501]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF99FFBBFB12FE03851149A5" box="[1192,1314,475,501]" italics="true" pageId="7" pageNumber="376">P. olivacea</emphasis>
</taxonomicName>
by the late H.-L. Chen. A re-examination of this material showed that the specimens from Jeram (ZRC 2020.0345) agreed very well with the
<typeStatus id="54FF8818FF99FFBBFE8BFDFB81524A6D" box="[305,353,547,573]" pageId="7" pageNumber="376">type</typeStatus>
description and figures of
<taxonomicName id="4C444D39FF99FFBBFD35FDFB83284A6D" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[655,795,547,573]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF99FFBBFD35FDFB83284A6D" box="[655,795,547,573]" italics="true" pageId="7" pageNumber="376">P. fuliginosa</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFD54B4BFF99FFBBFCFFFDFB84694A6D" author="Targioni-Tozzetti, A." box="[837,1114,547,573]" pageId="7" pageNumber="376" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti (1877)</bibRefCitation>
, with the rest of the material (including the Muar specimen, ZRC 1970.8.8.7) matching that for
<taxonomicName id="4C444D39FF99FFBBFC38FD9F83CF4A31" authorityName="Rathbun" authorityYear="1909" box="[898,1020,583,609]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF99FFBBFC38FD9F83CF4A31" box="[898,1020,583,609]" italics="true" pageId="7" pageNumber="376">P. olivacea</emphasis>
</taxonomicName>
as described and figured by
<bibRefCitation id="EFD54B4BFF99FFBBFA87FD9F80E84AD5" author="Rathbun, M. J." pageId="7" pageNumber="376" pagination="107 - 114" refId="ref7110" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun (1909</bibRefCitation>
,
<bibRefCitation id="EFD54B4BFF99FFBBFF5DFDB381104AD5" author="Rathbun, M. J." box="[231,291,619,645]" pageId="7" pageNumber="376" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">1910</bibRefCitation>
),
<bibRefCitation id="EFD54B4BFF99FFBBFE8CFDB381C24AD6" author="Dai, A. - Y. &amp; Yang, S. - L. &amp; Song, Y. - Z. &amp; Chen, G. - X." box="[310,497,619,646]" pageId="7" pageNumber="376" refId="ref6365" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. &amp; Chen, G. - X. (1986) Crabs of the China Seas. China Ocean Press, Beijing. 11 + 642 pp." type="book" year="1986">
Dai
<emphasis id="B930EAA8FF99FFBBFEDFFDB381AD4AD5" box="[357,414,619,645]" italics="true" pageId="7" pageNumber="376">et al.</emphasis>
(1986)
</bibRefCitation>
,
<bibRefCitation id="EFD54B4BFF99FFBBFE47FDB382E54AD5" author="Dai, A. - Y. &amp; Yang, S. - L." box="[509,726,619,645]" pageId="7" pageNumber="376" pagination="1 - 295" refId="ref6313" refString="Dai, A. - Y. &amp; Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, pp. 21 + 608, figs. 1 - 295, pls. 1 - 74." type="journal article" year="1991">Dai &amp; Yang (1991)</bibRefCitation>
, Tan (1995) and
<bibRefCitation id="EFD54B4BFF99FFBBFC29FDB3845C4AD5" author="Chen, H. &amp; Sun, H." box="[915,1135,619,645]" pageId="7" pageNumber="376" pagination="1 - 16" refId="ref6263" refString="Chen, H. &amp; Sun, H. (2002) Arthropoda Crustacea. Brachyura. Marine primitive crabs. Fauna Sinica. Invertebrata, 30, Science Press, Beijing, 597 pp., colour pls. 1 - 4, pls. 1 - 16." type="book chapter" year="2002">Chen &amp; Sun (2002)</bibRefCitation>
very well.
</paragraph>
<caption id="DF3B6632FF99FFBBFF2DF85085724FB8" ID-DOI="http://doi.org/10.5281/zenodo.4674247" ID-Zenodo-Dep="4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="7" pageNumber="376" startId="7.[151,250,1928,1952]" targetBox="[215,1387,688,1892]" targetPageId="7">
<paragraph id="8BFB36BAFF99FFBBFF2DF85085724FB8" blockId="7.[151,1436,1928,2025]" pageId="7" pageNumber="376">
<emphasis id="B930EAA8FF99FFBBFF2DF850813E4FF1" bold="true" box="[151,269,1928,1953]" pageId="7" pageNumber="376">FIGURE 5</emphasis>
.
<taxonomicName id="4C444D39FF99FFBBFEA3F85082EA4FF1" authority="(Targioni-Tozzetti, 1877)" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[281,729,1928,1953]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF99FFBBFEA3F85081E24FF0" box="[281,465,1928,1952]" italics="true" pageId="7" pageNumber="376">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF99FFBBFE64F85082E14FF1" author="Targioni-Tozzetti, A." box="[478,722,1928,1953]" pageId="7" pageNumber="376" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
)
</taxonomicName>
. A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm.
</paragraph>
</caption>
<caption id="DF3B6632FF96FFB4FF2DF8AA83214FA5" ID-DOI="http://doi.org/10.5281/zenodo.4674249" ID-Zenodo-Dep="4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="8" pageNumber="377" startId="8.[151,250,1906,1930]" targetBox="[193,1352,191,1868]" targetPageId="8">
<paragraph id="8BFB36BAFF96FFB4FF2DF8AA83214FA5" blockId="8.[151,1437,1906,2038]" pageId="8" pageNumber="377">
<emphasis id="B930EAA8FF96FFB4FF2DF8AA813F4FDA" bold="true" box="[151,268,1906,1930]" pageId="8" pageNumber="377">FIGURE 6</emphasis>
.
<taxonomicName id="4C444D39FF96FFB4FEA2F8AA82E44FDA" authority="(Targioni-Tozzetti, 1877)" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[280,727,1906,1930]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF96FFB4FEA2F8AA81E34FDA" box="[280,464,1906,1930]" italics="true" pageId="8" pageNumber="377">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF96FFB4FE66F8AA82E34FDA" author="Targioni-Tozzetti, A." box="[476,720,1906,1930]" pageId="8" pageNumber="377" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
)
</taxonomicName>
. AD, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; EH, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; IL, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EL, = 0.5 mm.
</paragraph>
</caption>
<caption id="DF3B6632FF97FFB5FF2DF90284FD4F46" ID-DOI="http://doi.org/10.5281/zenodo.4674251" ID-Zenodo-Dep="4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="9" pageNumber="378" startId="9.[151,250,1754,1778]" targetBox="[151,1436,181,1730]" targetPageId="9">
<paragraph id="8BFB36BAFF97FFB5FF2DF90284FD4F46" blockId="9.[151,1436,1754,1814]" pageId="9" pageNumber="378">
<emphasis id="B930EAA8FF97FFB5FF2DF90281234EA2" bold="true" box="[151,272,1754,1778]" pageId="9" pageNumber="378">FIGURE 7</emphasis>
.
<taxonomicName id="4C444D39FF97FFB5FEA5F90282DA4EA2" authority="(Targioni-Tozzetti, 1877)" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[287,745,1754,1778]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF97FFB5FEA5F90281E94EA2" box="[287,474,1754,1778]" italics="true" pageId="9" pageNumber="378">Ovilyra fuliginosa</emphasis>
(
<bibRefCitation id="EFD54B4BFF97FFB5FE51F90282D14EA2" author="Targioni-Tozzetti, A." box="[491,738,1754,1778]" pageId="9" pageNumber="378" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
)
</taxonomicName>
. AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae.
</paragraph>
</caption>
<paragraph id="8BFB36BAFF97FFB5FF7DF89C85514F9A" blockId="9.[151,1437,1860,2031]" pageId="9" pageNumber="378">
Several characters appear to differentiate the specimens from Jeram (ZRC 2020.0345) from the rest of the material, notably in the proportionately shorter cheliped merus and ambulatory merus (
<figureCitation id="137F2A3FFF97FFB5FB97F8B084964FD3" box="[1069,1189,1896,1923]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="9" pageNumber="378">Fig. 1A, F</figureCitation>
); the lateral margin of male pleonal somite 6 is slightly sinuous (
<figureCitation id="137F2A3FFF97FFB5FDC9F85482E04FF6" box="[627,723,1932,1958]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="9" pageNumber="378">Figs. 1E</figureCitation>
,
<figureCitation id="137F2A3FFF97FFB5FD5AF85483364FF6" box="[736,773,1932,1958]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 36 and telson; B, posterior thoracic sternum and pleonal somites 14. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1so6 = pleonal somites 16; s1s8 = sternites 18; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="9" pageNumber="378">4A</figureCitation>
,
<figureCitation id="137F2A3FFF97FFB5FCABF85483014FF7" box="[785,818,1932,1959]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="9" pageNumber="378">5F</figureCitation>
); and the distal projection of the G1 prominently overreaches the main shaft of the (
<figureCitation id="137F2A3FFF97FFB5FE52F86882504F9B" box="[488,611,1968,1995]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).AD, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; EH, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; IL, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EL, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="9" pageNumber="378">Fig. 6AC</figureCitation>
). These differences, however, are not significant at the species level.
</paragraph>
<paragraph id="8BFB36BAFF97FFB6FF7DF80C83444981" blockId="9.[151,1437,1860,2031]" lastBlockId="10.[151,1437,151,1617]" lastPageId="10" lastPageNumber="379" pageId="9" pageNumber="378">
The male cheliped merus of the Jeram
<taxonomicName id="4C444D39FF97FFB5FDC3F80D83364FBE" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[633,773,2004,2030]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF97FFB5FDC3F80D83364FBE" box="[633,773,2004,2030]" italics="true" pageId="9" pageNumber="378">P. fuliginosa</emphasis>
</taxonomicName>
is proportionately shorter (
<figureCitation id="137F2A3FFF97FFB5FB8CF80C84BF4FBE" box="[1078,1164,2004,2030]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="9" pageNumber="378">Fig. 1A</figureCitation>
) as figured by Targioni-Tozzetti 1877: pl. 12 fig.
<quantity id="4CBC9B5FFF94FFB6FE14FF4F81FF48E1" box="[430,460,151,177]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="10" pageNumber="379" unit="g" value="3.0">3g</quantity>
), and this structure is distinctly longer in most of the specimens of
<taxonomicName id="4C444D39FF94FFB6FB0AFF4F851A48E1" authorityName="Rathbun" authorityYear="1909" box="[1200,1321,151,177]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FB0AFF4F851A48E1" box="[1200,1321,151,177]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
examined (
<figureCitation id="137F2A3FFF94FFB6FF25FF6381374885" box="[159,260,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="10" pageNumber="379">Figs. 2A</figureCitation>
,
<figureCitation id="137F2A3FFF94FFB6FEAAFF6381654885" box="[272,342,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">3A, B</figureCitation>
), even when specimens of similar sizes are compared. In a few smaller specimens of
<taxonomicName id="4C444D39FF94FFB6FAA0FF6385A64885" authorityName="Rathbun" authorityYear="1909" box="[1306,1429,187,213]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FAA0FF6385A64885" box="[1306,1429,187,213]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
, however, the merus is somewhat shorter (
<figureCitation id="137F2A3FFF94FFB6FDCCFF0782E248A9" box="[630,721,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">Fig. 3C</figureCitation>
). The same pattern is observed for female specimens (
<figureCitation id="137F2A3FFF94FFB6FAFFFF0785AF48A9" box="[1349,1436,223,249]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="10" pageNumber="379">Fig. 7A</figureCitation>
versus
<figureCitation id="137F2A3FFF94FFB6FF5DFEDB8171494D" box="[231,322,259,285]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="10" pageNumber="379">Fig. 7D</figureCitation>
). Slight differences in proportions of the cheliped merus, however, can be explained by variation. The ambulatory merus of the Jeram male is proportionately shorter (
<figureCitation id="137F2A3FFF94FFB6FC20FEFF83DF4911" box="[922,1004,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="10" pageNumber="379">Fig. 1F</figureCitation>
) and looks identical to that figured by
<bibRefCitation id="EFD54B4BFF94FFB6FF2DFE9381954935" author="Targioni-Tozzetti, A." box="[151,422,331,357]" pageId="10" pageNumber="379" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti (1877</bibRefCitation>
: pl. 12 fig. 3b). This is also true for the female specimen from Jeram (
<figureCitation id="137F2A3FFF94FFB6FB78FE93852B4935" box="[1218,1304,331,357]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="10" pageNumber="379">Fig. 7A</figureCitation>
versus
<figureCitation id="137F2A3FFF94FFB6FACAFE93808E49D9" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). AC, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. DF, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="10" pageNumber="379">Fig. 7D</figureCitation>
). Most of the specimens of
<taxonomicName id="4C444D39FF94FFB6FE4BFEB7825F49D9" authorityName="Rathbun" authorityYear="1909" box="[497,620,367,393]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FE4BFEB7825F49D9" box="[497,620,367,393]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
have the ambulatory merus slightly longer (
<figureCitation id="137F2A3FFF94FFB6FBE0FEB7849C49D9" box="[1114,1199,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="10" pageNumber="379">Fig. 2F</figureCitation>
), but we observe that larger specimens tend to have more slender legs, although not always the case. The differences, observed, in any case, are minor and are unlikely to be reliable at the species level.
</paragraph>
<paragraph id="8BFB36BAFF94FFB6FF7DFE0381274B69" blockId="10.[151,1437,151,1617]" pageId="10" pageNumber="379">
The differences in the male pleonal somite 6 and G1 appear more significant. Most of the specimens of
<taxonomicName id="4C444D39FF94FFB6FAE3FE0380EB4A48" authorityName="Rathbun" authorityYear="1909" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FAE3FE0380EB4A48" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
have the lateral margins of male pleonal somite 6 gently convex (
<figureCitation id="137F2A3FFF94FFB6FC04FE27842C4A49" box="[958,1055,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="10" pageNumber="379">Figs. 2E</figureCitation>
,
<figureCitation id="137F2A3FFF94FFB6FB91FE27847C4A49" box="[1067,1103,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">3D</figureCitation>
,
<figureCitation id="137F2A3FFF94FFB6FBE1FE26844D4A49" box="[1115,1150,510,537]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="10" pageNumber="379">5G</figureCitation>
) and the distal projection of the G1 reaches or overlaps the main shaft but does not overreach it (
<figureCitation id="137F2A3FFF94FFB6FC71FDFB84B14A6D" box="[971,1154,547,573]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).AD, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; EH, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; IL, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EL, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="10" pageNumber="379">Fig. 6EG, IK</figureCitation>
). A male (6.8 × 8.0 mm, ZRC 2014.0176) from
<collectingCountry id="F353762AFF94FFB6FE2DFD9F823A4A31" box="[407,521,583,609]" name="Singapore" pageId="10" pageNumber="379">Singapore</collectingCountry>
is a typical
<taxonomicName id="4C444D39FF94FFB6FD37FD9F83344A31" authorityName="Rathbun" authorityYear="1909" box="[653,775,583,609]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FD37FD9F83344A31" box="[653,775,583,609]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
with regards to the frontal lobe, proportions of the cheliped merus, ambulatory merus and third maxilliped exopod, with the lateral margins of pleonal somite 6 clearly convex, but its G1 distal process overlaps the main shaft by a short distance, albeit not to the degree of the Jeram
<taxonomicName id="4C444D39FF94FFB6FA86FD5780FF4A9C" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FA86FD5780FF4A9C" italics="true" pageId="10" pageNumber="379">P. fuliginosa</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A3FFF94FFB6FF67FD6B816A4A9E" box="[221,345,691,718]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).AD, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; EH, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; IL, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EL, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="10" pageNumber="379">Fig. 6AC</figureCitation>
). Another male (11.2 × 14.0 mm, ZRC 2009.0217) of
<taxonomicName id="4C444D39FF94FFB6FC7EFD6B84734A9D" authorityName="Rathbun" authorityYear="1909" box="[964,1088,691,717]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FC7EFD6B84734A9D" box="[964,1088,691,717]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
has the proportionately longer cheliped merus, ambulatory merus and narrower third maxilliped exopod like the other specimens in the lot except that the lateral margins of its pleonal somite 6 are sinuous and the G1 has the distal process slightly overlapping the main shaft.
</paragraph>
<paragraph id="8BFB36BAFF94FFB6FF7DFC9B80FB4C41" blockId="10.[151,1437,151,1617]" pageId="10" pageNumber="379">
In addition, the frontal median lobe is relatively lower in
<taxonomicName id="4C444D39FF94FFB6FCFEFC9B83FC4B0D" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[836,975,835,861]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FCFEFC9B83FC4B0D" box="[836,975,835,861]" italics="true" pageId="10" pageNumber="379">P. fuliginosa</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A3FFF94FFB6FC67FC9B84074B0D" box="[989,1076,834,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="10" pageNumber="379">Fig. 5D</figureCitation>
) while in most of the specimens of
<taxonomicName id="4C444D39FF94FFB6FF0CFCBF81034BD1" authorityName="Rathbun" authorityYear="1909" box="[182,304,871,897]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FF0CFCBF81034BD1" box="[182,304,871,897]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
, the frontal median lobe is usually more developed (
<figureCitation id="137F2A3FFF94FFB6FCC4FCBF83E44BD1" box="[894,983,870,897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="10" pageNumber="379">Fig. 5D</figureCitation>
), but this appears to be partly associated with size as smaller specimens tend to have somewhat lower frontal lobes. The exopod of the third maxilliped is relatively wider in the specimens of
<taxonomicName id="4C444D39FF94FFB6FD89FC77828C4B99" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[563,703,943,969]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FD89FC77828C4B99" box="[563,703,943,969]" italics="true" pageId="10" pageNumber="379">P. fuliginosa</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A3FFF94FFB6FD75FC7783144B99" box="[719,807,942,969]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="10" pageNumber="379">Fig. 5A</figureCitation>
) and while most of the specimens of
<taxonomicName id="4C444D39FF94FFB6FB6AFC77857F4B99" authorityName="Rathbun" authorityYear="1909" box="[1232,1356,943,969]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FB6AFC77857F4B99" box="[1232,1356,943,969]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
have a slightly narrower exopod (
<figureCitation id="137F2A3FFF94FFB6FE7FFC0B82144BBD" box="[453,551,979,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">Figs. 3E</figureCitation>
,
<figureCitation id="137F2A3FFF94FFB6FD8EFC0A826B4BBD" box="[564,600,978,1005]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="10" pageNumber="379">5B</figureCitation>
), this is not always the case with a few also possessing a wider one (
<figureCitation id="137F2A3FFF94FFB6FAD5FC0B808F4C41" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.AC, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, EG = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="10" pageNumber="379">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8BFB36BAFF94FFB6FF7DFBC385B74CF1" blockId="10.[151,1437,151,1617]" pageId="10" pageNumber="379">
As for the three characters used by
<bibRefCitation id="EFD54B4BFF94FFB6FDE9FBC382CE4C65" author="Rathbun, M. J." box="[595,765,1051,1077]" pageId="10" pageNumber="379" pagination="107 - 114" refId="ref7110" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun (1909</bibRefCitation>
,
<bibRefCitation id="EFD54B4BFF94FFB6FCB3FBC383714C65" author="Rathbun, M. J." box="[777,834,1051,1077]" pageId="10" pageNumber="379" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">1910</bibRefCitation>
) to separate
<taxonomicName id="4C444D39FF94FFB6FC6BFBC384784C65" authorityName="Rathbun" authorityYear="1909" box="[977,1099,1051,1077]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FC6BFBC384784C65" box="[977,1099,1051,1077]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
</taxonomicName>
from
<taxonomicName id="4C444D39FF94FFB6FB35FBC385284C65" authorityName="Ng" authorityYear="2021" baseAuthorityName="Targioni-Tozzetti" baseAuthorityYear="1877" box="[1167,1307,1051,1077]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FB35FBC385284C65" box="[1167,1307,1051,1077]" italics="true" pageId="10" pageNumber="379">P. fuliginosa</emphasis>
</taxonomicName>
, it was discussed earlier that the form of the male pleon varies.As to the strength of the tooth at the angles of the posterior carapace margin, this character varies substantially, from low (
<figureCitation id="137F2A3FFF94FFB6FCA6FBBB834F4C2D" box="[796,892,1123,1149]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="10" pageNumber="379">Figs. 1B</figureCitation>
,
<figureCitation id="137F2A3FFF94FFB6FC3DFBBB839B4C2D" box="[903,936,1123,1149]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">3C</figureCitation>
) to medium size (
<figureCitation id="137F2A3FFF94FFB6FBD7FBBB84F74C2D" box="[1133,1220,1123,1149]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 36 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="10" pageNumber="379">Fig. 2B</figureCitation>
,
<figureCitation id="137F2A3FFF94FFB6FB74FBBB84C14C2D" box="[1230,1266,1123,1149]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">3A</figureCitation>
) to strong (
<figureCitation id="137F2A3FFF94FFB6FAD5FBBB80884CF1" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">Fig. 3B</figureCitation>
). The same applies to the slightly different degree of bending of the cheliped fingers varies (e.g.,
<figureCitation id="137F2A3FFF94FFB6FB47FB5F854B4CF1" box="[1277,1400,1159,1185]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; CE, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.AC, overall dorsal view; D, anterior thoracic sternum, pleonal somites 36 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="10" pageNumber="379">Fig. 3AC</figureCitation>
).
</paragraph>
<paragraph id="8BFB36BAFF94FFB6FF7DFB7385BC4D05" blockId="10.[151,1437,151,1617]" pageId="10" pageNumber="379">
In summary, there are no clear differences between
<taxonomicName id="4C444D39FF94FFB6FCB2FB7384904C95" authority="Targioni-Tozzetti, 1877" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[776,1187,1195,1221]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FCB2FB7383A04C95" box="[776,915,1195,1221]" italics="true" pageId="10" pageNumber="379">P. fuliginosa</emphasis>
<bibRefCitation id="EFD54B4BFF94FFB6FC23FB7384904C95" author="Targioni-Tozzetti, A." box="[921,1187,1195,1221]" pageId="10" pageNumber="379" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C444D39FF94FFB6FB67FB7381384CB9" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF94FFB6FB67FB7385644C95" box="[1245,1367,1195,1221]" italics="true" pageId="10" pageNumber="379">P. olivacea</emphasis>
<bibRefCitation id="EFD54B4BFF94FFB6FAE4FB7381384CB9" author="Rathbun, M. J." pageId="10" pageNumber="379" pagination="107 - 114" refId="ref7110" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
</taxonomicName>
, and the author therefore synonymises the two taxa. The types of
<taxonomicName id="4C444D39FF94FFB6FC54FB1784494CB9" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[1006,1146,1231,1257]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FC54FB1784494CB9" box="[1006,1146,1231,1257]" italics="true" pageId="10" pageNumber="379">P. fuliginosa</emphasis>
</taxonomicName>
are not extant and almost certainly lost.
<bibRefCitation id="EFD54B4BFF94FFB6FE8FFB2B82324D5D" author="Lucas, J. S." box="[309,513,1267,1293]" pageId="10" pageNumber="379" pagination="148 - 247" refId="ref6766" refString="Lucas, J. S. (1981) Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: Systematics and biology. Records of the Australian Museum, 1980, 33 (4), 148 - 247, figs. 1 - 10. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1981">Lucas (1981: 200)</bibRefCitation>
searched for them for his study on hymenosomatids and commented they were lost in the Second World War, and a fresh search has confirmed this (see also
<bibRefCitation id="EFD54B4BFF94FFB6FC05FACF84D24D61" author="Schubart, C. D. &amp; Ng, P. K. L." box="[959,1249,1302,1329]" pageId="10" pageNumber="379" pagination="891 - 994" refId="ref7258" refString="Schubart, C. D. &amp; Ng, P. K. L. (2020) Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serene &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species. Raffles Bulletin of Zoology, 68, 891 - 994." type="journal article" year="2020">Schubart &amp; Ng 2020: 956</bibRefCitation>
). There is no immediate need to select a
<typeStatus id="54FF8818FF94FFB6FE10FAE382304D05" box="[426,515,1339,1365]" pageId="10" pageNumber="379" type="neotype">neotype</typeStatus>
as the specimens on hand from
<collectingCountry id="F353762AFF94FFB6FCD0FAE383E24D05" box="[874,977,1339,1365]" name="Malaysia" pageId="10" pageNumber="379">Malaysia</collectingCountry>
and
<collectingCountry id="F353762AFF94FFB6FBBDFAE3844A4D05" box="[1031,1145,1339,1365]" name="Singapore" pageId="10" pageNumber="379">Singapore</collectingCountry>
are clearly
<taxonomicName id="4C444D39FF94FFB6FB46FAE385BB4D05" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[1276,1416,1339,1365]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FB46FAE385BB4D05" box="[1276,1416,1339,1365]" italics="true" pageId="10" pageNumber="379">P. fuliginosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF94FFB6FF7DFA87854C4DB5" pageId="10" pageNumber="379" type="biology_ecology">
<paragraph id="8BFB36BAFF94FFB6FF7DFA87854C4DB5" blockId="10.[151,1437,151,1617]" pageId="10" pageNumber="379">
<emphasis id="B930EAA8FF94FFB6FF7DFA8781154D29" bold="true" box="[199,294,1375,1401]" pageId="10" pageNumber="379">Ecology</emphasis>
. The
<collectingCountry id="F353762AFF94FFB6FED1FA8781EE4D29" box="[363,477,1375,1401]" name="Singapore" pageId="10" pageNumber="379">Singapore</collectingCountry>
specimens were all collected from the southern part of the island, in sandy-muddy habitats in shallow water less than
<quantity id="4CBC9B5FFF94FFB6FDA0FA5B82604DCD" box="[538,595,1411,1437]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="379" unit="m" value="20.0">20 m</quantity>
deep. Two specimens of
<taxonomicName id="4C444D39FF94FFB6FCD1FA5B83CE4DCD" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[875,1021,1411,1437]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF94FFB6FCD1FA5B83CE4DCD" box="[875,1021,1411,1437]" italics="true" pageId="10" pageNumber="379">O. fuliginosa</emphasis>
</taxonomicName>
were obtained from an unusual habitat, a polychaete reef in Jeram,
<collectingCountry id="F353762AFF94FFB6FE40FA7F82524D91" box="[506,609,1447,1473]" name="Malaysia" pageId="10" pageNumber="379">Malaysia</collectingCountry>
(ZRC 2020.0345), where a number of new decapods were also collected (see
<bibRefCitation id="EFD54B4BFF94FFB6FF71FA1381A54DB5" author="Polgar, G. &amp; Nishi, E. &amp; Idris, I. &amp; Glasby, C. J." box="[203,406,1482,1509]" pageId="10" pageNumber="379" pagination="401 - 417" refId="ref7056" refString="Polgar, G., Nishi, E., Idris, I. &amp; Glasby, C. J. (2015) Tropical polychaete community and reef dynamics: insights from a Malayan Sabellaria (Annelida: Sabellariidae) reef. Raffles Bulletin of Zoology, 63, 401 - 417." type="journal article" year="2015">
Polgar
<emphasis id="B930EAA8FF94FFB6FEA1FA1381674DB5" box="[283,340,1483,1509]" italics="true" pageId="10" pageNumber="379">et al.</emphasis>
2015
</bibRefCitation>
), including two rare species of porcellanid crabs (Osawa
<emphasis id="B930EAA8FF94FFB6FBA2FA1384634DB5" box="[1048,1104,1483,1509]" italics="true" pageId="10" pageNumber="379">et al.</emphasis>
2018;
<bibRefCitation id="EFD54B4BFF94FFB6FB27FA1385474DB5" author="Osawa, M. &amp; Ng, P. K. L." box="[1181,1396,1483,1509]" pageId="10" pageNumber="379" pagination="111 - 126" refId="ref6968" refString="Osawa, M. &amp; Ng, P. K. L. (2018) A new species of the genus Raphidopus Stimpson, 1858 (Crustacea: Decapoda: Anomura: Porcellanidae) from Peninsular Malaysia, with additional records of R. johnsoni Ng &amp; Nakasone, 1994 from the Southeast Asia, and a key to species in the genus. Zootaxa, 4433 (1), 111 - 126. https: // doi. org / 10.11646 / zootaxa. 4433.1.6" type="journal article" year="2018">Osawa &amp; Ng 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF94FFB6FF7DFA3683844E01" pageId="10" pageNumber="379" type="distribution">
<paragraph id="8BFB36BAFF94FFB6FF7DFA3683844E01" blockId="10.[151,1437,151,1617]" pageId="10" pageNumber="379">
<emphasis id="B930EAA8FF94FFB6FF7DFA36816A4E58" bold="true" box="[199,345,1518,1544]" pageId="10" pageNumber="379">Distribution</emphasis>
. The species has been reported from Java, Peninsular
<collectingCountry id="F353762AFF94FFB6FC79FA37841F4E59" box="[963,1068,1519,1545]" name="Malaysia" pageId="10" pageNumber="379">Malaysia</collectingCountry>
,
<collectingCountry id="F353762AFF94FFB6FB83FA37849E4E59" box="[1081,1197,1519,1545]" name="Singapore" pageId="10" pageNumber="379">Singapore</collectingCountry>
, Gulf of
<collectingCountry id="F353762AFF94FFB6FAAEFA3785494E59" box="[1300,1402,1519,1545]" name="Thailand" pageId="10" pageNumber="379">Thailand</collectingCountry>
, to
<collectingRegion id="4980F858FF94FFB6FF2DF9CB80CD4E7D" box="[151,254,1555,1581]" country="China" name="Zhejiang" pageId="10" pageNumber="379">Zhejiang</collectingRegion>
,
<collectingRegion id="4980F858FF94FFB6FEB1F9CB81604E7D" box="[267,339,1555,1581]" country="China" name="Fujian" pageId="10" pageNumber="379">Fujian</collectingRegion>
and
<collectingRegion id="4980F858FF94FFB6FE36F9CB81EE4E7D" box="[396,477,1555,1581]" country="China" name="Hainan" pageId="10" pageNumber="379">Hainan</collectingRegion>
Island in southern
<collectingCountry id="F353762AFF94FFB6FD01F9CB82CD4E7D" box="[699,766,1555,1581]" name="China" pageId="10" pageNumber="379">China</collectingCountry>
(
<bibRefCitation id="EFD54B4BFF94FFB6FCB4F9CB84264E7D" author="Targioni-Tozzetti, A." box="[782,1045,1555,1581]" pageId="10" pageNumber="379" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti 1877</bibRefCitation>
;
<bibRefCitation id="EFD54B4BFF94FFB6FB99F9CB84F54E7D" author="Rathbun, M. J." box="[1059,1222,1555,1581]" pageId="10" pageNumber="379" pagination="107 - 114" refId="ref7110" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun 1909</bibRefCitation>
,
<bibRefCitation id="EFD54B4BFF94FFB6FB69F9CB853D4E7D" author="Rathbun, M. J." box="[1235,1294,1555,1581]" pageId="10" pageNumber="379" pagination="301 - 367" refId="ref7141" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Konelige Danske Videnskabernes Selskat, Naturvidenskabelig og Matematisk Afdeling, 5 (4), 301 - 367, figs. 1 - 44, pls. 1, 2." type="journal article" year="1910">1910</bibRefCitation>
,
<bibRefCitation id="EFD54B4BFF94FFB6FAA1F9CB85654E7D" author="Rathbun, M. J." box="[1307,1366,1555,1581]" pageId="10" pageNumber="379" pagination="75 - 104" refId="ref7194" refString="Rathbun, M. J. (1931) New and rare Chinese crabs. Lingnan Science Journal, 1929, 8, 75 - 104, pls. 5 - 15." type="journal article" year="1931">1931</bibRefCitation>
;
<bibRefCitation id="EFD54B4BFF94FFB6FAD8F9CB80E14E01" author="Yang, C. M." pageId="10" pageNumber="379" refId="ref7562" refString="Yang, C. M. (1979) A list of Brachyura in the Zoological Reference Collection of the Department of Zoology. No. 14. Department of Zoology, National University of Singapore, 60 pp." type="book" year="1979">Yang 1979</bibRefCitation>
;
<bibRefCitation id="EFD54B4BFF94FFB6FF64F9EF81B44E01" author="Dai, A. - Y. &amp; Yang, S. - L. &amp; Song, Y. - Z. &amp; Chen, G. - X." box="[222,391,1591,1617]" pageId="10" pageNumber="379" refId="ref6365" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. &amp; Chen, G. - X. (1986) Crabs of the China Seas. China Ocean Press, Beijing. 11 + 642 pp." type="book" year="1986">
Dai
<emphasis id="B930EAA8FF94FFB6FEB7F9EF81754E01" box="[269,326,1591,1617]" italics="true" pageId="10" pageNumber="379">et al.</emphasis>
1986
</bibRefCitation>
;
<bibRefCitation id="EFD54B4BFF94FFB6FE29F9EF82694E01" author="Dai, A. - Y. &amp; Yang, S. - L." box="[403,602,1591,1617]" pageId="10" pageNumber="379" pagination="1 - 295" refId="ref6313" refString="Dai, A. - Y. &amp; Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, pp. 21 + 608, figs. 1 - 295, pls. 1 - 74." type="journal article" year="1991">Dai &amp; Yang 1991</bibRefCitation>
; Tan 1995;
<bibRefCitation id="EFD54B4BFF94FFB6FD67F9EF83984E01" author="Chen, H. &amp; Sun, H." box="[733,939,1591,1617]" pageId="10" pageNumber="379" pagination="1 - 16" refId="ref6263" refString="Chen, H. &amp; Sun, H. (2002) Arthropoda Crustacea. Brachyura. Marine primitive crabs. Fauna Sinica. Invertebrata, 30, Science Press, Beijing, 597 pp., colour pls. 1 - 4, pls. 1 - 16." type="book chapter" year="2002">Chen &amp; Sun 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>