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<document id="96BB6E0714E4267461E9A5E225628891" ID-CLB-Dataset="20639" ID-DOI="10.11646/zootaxa.5099.4.5" ID-GBIF-Dataset="f0a97603-c958-4814-a1a2-44a2622b286f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6092195" ID-ZooBank="C2244780-3616-4188-A23E-0291BD463F03" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644912521413" checkinUser="plazi" docAuthor="Archibald, S. Bruce, Cannings, Robert A. &amp; Greenwalt, Dale E." docDate="2022" docId="03ED5075EA21FFF23CCC79E4C6F7FDFD" docLanguage="en" docName="zootaxa.5099.4.5.pdf" docOrigin="Zootaxa 5099 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kishenehna prima Archibald &amp; Cannings &amp; Greenwalt 2022, new species" docType="treatment" docVersion="7" lastPageNumber="499" masterDocId="FFD4280DEA23FFF13C5B7F09C701FFC0" masterDocTitle="Kishenehna prima, a new genus and species of darner dragonfly (Odonata, Aeshnidae, Gomphaeschninae) from the early middle Eocene Kishenehn Formation of Montana, USA" masterLastPageNumber="500" masterPageNumber="496" pageNumber="498" updateTime="1699312088466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F06EB5CB83C495FED469D563EDECC60B">Kishenehna prima, a new genus and species of darner dragonfly (Odonata, Aeshnidae, Gomphaeschninae) from the early middle Eocene Kishenehn Formation of Montana, USA</mods:title>
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<mods:namePart id="26B2926D8DF135B926F0FC8E8451AC1F">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="CD4BCB6EF1BCCDC8FD6D733E11C80FB6">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America &amp; Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada. RCannings @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0001 - 9491 - 8186</mods:affiliation>
<mods:nameIdentifier id="B28E36424C55360D1C6E9506D9A6A5D1" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="75B0B2C700001BD7FF3CD7CAC6E6B78B">Cannings, Robert A.</mods:namePart>
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<mods:affiliation id="88A7C7BFFA9DC38F114B07BFF361C1E7">Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada. RCannings @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0001 - 9491 - 8186</mods:affiliation>
<mods:nameIdentifier id="2C2C9C5D91E79638EDE6A13877894B8B" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="D871DD770D09536960A94685B03F38CC">Greenwalt, Dale E.</mods:namePart>
<mods:nameIdentifier id="B9C557D3B62CE6EF6B9589D73321BE7A" type="ORCID">0000-0002-9811-6356</mods:nameIdentifier>
<mods:affiliation id="E97AC603323DAE4A4F39AD6E6C3DE15B">Department of Paleobiology, NMNH, Smithsonian Institution, P. O. Box 37012 MRC 121, Washington, DC, 20013 - 7012, USA. greenwaltde @ gmail. com; https: // orcid. org / 0000 - 0002 - 9811 - 6356</mods:affiliation>
<mods:nameIdentifier id="3E172AB0382175063AF72AD1A998219B" type="email">greenwaltde@gmail.com</mods:nameIdentifier>
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<mods:date id="9EF4A6F67BC647E3DE510E6B6A0E7E17">2022</mods:date>
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<subSubSection id="C35EB2E8EA21FFF33CCC79E4C430F8C7" box="[151,817,1773,1800]" pageId="2" pageNumber="498" type="nomenclature">
<paragraph id="8BFBE163EA21FFF33CCC79E4C430F8C7" blockId="2.[151,817,1773,1800]" box="[151,817,1773,1800]" pageId="2" pageNumber="498">
<heading id="D0B3560FEA21FFF33CCC79E4C430F8C7" bold="true" box="[151,817,1773,1800]" fontSize="11" level="1" pageId="2" pageNumber="498" reason="1">
<emphasis id="B9303D71EA21FFF33CCC79E4C430F8C7" bold="true" box="[151,817,1773,1800]" pageId="2" pageNumber="498">
<taxonomicName id="4C449AE0EA21FFF33CCC79E4C679F8C7" authority="Archibald &amp; Cannings &amp; Greenwalt, 2022" authorityName="Archibald &amp; Cannings &amp; Greenwalt" authorityYear="2022" box="[151,376,1773,1800]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="prima" status="sp. nov.">
<emphasis id="B9303D71EA21FFF33CCC79E4C679F8C7" bold="true" box="[151,376,1773,1800]" italics="true" pageId="2" pageNumber="498">Kishenehna prima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A203800AEA21FFF33D2479E7C50FF8C7" box="[383,526,1773,1800]" pageId="2" pageNumber="498" rank="species">new species</taxonomicNameLabel>
Archibald &amp; Cannings
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35EB2E8EA21FFF33CCC783CC6ACF890" box="[151,429,1845,1872]" pageId="2" pageNumber="498" type="diagnosis">
<paragraph id="8BFBE163EA21FFF33CCC783CC6ACF890" blockId="2.[151,1437,1845,2016]" box="[151,429,1845,1872]" pageId="2" pageNumber="498">
<emphasis id="B9303D71EA21FFF33CCC783CC613F88F" bold="true" box="[151,274,1845,1871]" pageId="2" pageNumber="498">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="C35EB2E8EA21FFF33C9C7850C6D1F820" pageId="2" pageNumber="498" type="materials_examined">
<paragraph id="8BFBE163EA21FFF33C9C7850C6D1F820" blockId="2.[151,1437,1845,2016]" pageId="2" pageNumber="498">
<emphasis id="B9303D71EA21FFF33C9C7850C637F8B3" bold="true" box="[199,310,1881,1907]" pageId="2" pageNumber="498">Material.</emphasis>
<collectionCode id="ED5579A6EA21FFF33D667853C68FF8B4" box="[317,398,1882,1908]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="498" type="Museum">USNM</collectionCode>
621304, a finely preserved female hind wing, complete except broken across its basal region and other small portions missing, collected at the Park site of the Coal Creek Member of the Kishenehn Formation by DEG July, 2015. Housed in the collections of the Paleobiology Department of the National Museum of Natural History in
<collectingRegion id="49802F81EA21FFF33D5578CCC697F81F" box="[270,406,1989,2015]" country="United States of America" name="Washington" pageId="2" pageNumber="498">Washington</collectingRegion>
, DC.
</paragraph>
</subSubSection>
<subSubSection id="C35EB2E8EA20FFF23C9C7F9FC444FE6D" pageId="3" pageNumber="499" type="description">
<paragraph id="8BFBE163EA20FFF23C9C7F9FC444FE6D" blockId="3.[151,1437,150,573]" pageId="3" pageNumber="499">
<emphasis id="B9303D71EA20FFF23C9C7F9FC658FF70" bold="true" box="[199,345,150,176]" pageId="3" pageNumber="499">Description.</emphasis>
Female hind wing with character states of diagnosis and: recognisable humeral, axillary plates at base (
<emphasis id="B9303D71EA20FFF23C817FB2C7EFFF15" box="[218,238,187,213]" italics="true" pageId="3" pageNumber="499">cf</emphasis>
.
<bibRefCitation id="EFD59C92EA20FFF23CA67FB2C50CFF15" author="Grimaldi, D. &amp; Engel, M. S." box="[253,525,186,213]" pageId="3" pageNumber="499" refId="ref3132" refString="Grimaldi, D. &amp; Engel, M. S. (2005) Evolution of the Insects. Cambridge University Press, New York, 755 pp." type="book" year="2005">Grimaldi &amp; Engel 2005</bibRefCitation>
, Fig. 4.6);
<quantity id="4CBC4C86EA20FFF23ED27FB2C5D8FF15" box="[649,729,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="499" unit="mm" value="36.0">36 mm</quantity>
long, maximum width
<quantity id="4CBC4C86EA20FFF23FB87FB2C334FF15" box="[995,1077,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="499" unit="mm" value="11.0">11 mm</quantity>
; membrane hyaline throughout; pterostigma length four times width, subtends 1.5 cells; 11 antenodal crossveins; no crossveins stouter, identifiable as Ax1 or Ax2 as preserved; 9 postnodal, 9 postsubnodal crossveins, not aligned; pterostigmal brace vein present; oblique vein O present, first crossvein in RP2IRP2 space distal to origin of RP2; RP2, IRP2 subparallel distal to bend of RP2 to margin; triangle with distal side (MAb) kinked; radial, medial planates rather straight, curving toward posterior margin only in their last cell; CuAA space apparently maximum five cells wide; only one vein (CuP) crosses the cubito-anal space,
<emphasis id="B9303D71EA20FFF23E687E9AC54EFE6C" box="[563,591,403,428]" italics="true" pageId="3" pageNumber="499">i.e</emphasis>
., basal to subtriangle.
</paragraph>
</subSubSection>
<subSubSection id="C35EB2E8EA20FFF23C9C7EBEC6F7FDFD" pageId="3" pageNumber="499" type="etymology">
<paragraph id="8BFBE163EA20FFF23C9C7EBEC5D5FE35" blockId="3.[151,1437,150,573]" pageId="3" pageNumber="499">
<emphasis id="B9303D71EA20FFF23C9C7EBEC64FFE11" bold="true" box="[199,334,439,465]" pageId="3" pageNumber="499">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C449AE0EA20FFF23E677EBEC57EFE10" authorityName="Archibald &amp; Cannings &amp; Greenwalt" authorityYear="2022" box="[572,639,439,464]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="B9303D71EA20FFF23E677EBEC57EFE10" box="[572,639,439,464]" italics="true" pageId="3" pageNumber="499">prima</emphasis>
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is from the Latin
<emphasis id="B9303D71EA20FFF23F167EBEC49AFE10" box="[845,923,439,464]" italics="true" pageId="3" pageNumber="499">primus</emphasis>
, “the first”, as this is the first dragonfly (
<taxonomicName id="4C449AE0EA20FFF239347EBEC7F3FE36" authorityName="Selys" authorityYear="1854" class="Magnoliopsida" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="496" phylum="Tracheophyta" rank="subOrder" subOrder="Anisoptera">Anisoptera</taxonomicName>
) described from the Kishenehn Formation.
</paragraph>
<paragraph id="8BFBE163EA20FFF23C9C7EF6C6F7FDFD" blockId="3.[151,1437,150,573]" pageId="3" pageNumber="499">
<emphasis id="B9303D71EA20FFF23C9C7EF6C67FFDD9" bold="true" box="[199,382,511,537]" pageId="3" pageNumber="499">Range and age.</emphasis>
The early-middle Lutetian Coal Creek Member of the Kishenehn Formation, northwest
<collectingRegion id="49802F81EA20FFF239057EF6C7C9FDFD" country="United States of America" name="Montana" pageId="3" pageNumber="499">Montana</collectingRegion>
,
<collectingCountry id="F353A1F3EA20FFF23C8F7D2AC6F3FDFD" box="[212,498,547,573]" name="United States of America" pageId="3" pageNumber="499">United States of America</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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