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<document id="6A153E900C59775F53F71DE5C2442B1E" ID-CLB-Dataset="21377" ID-DOI="10.5852/ejt.2020.692" ID-GBIF-Dataset="ed3d3142-6449-4d72-bea7-51186ee48f37" ID-Zenodo-Dep="3973317" ID-ZooBank="EF16E197-0A18-45A9-8BA7-5C9154CDF4C3" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1596643342424" checkinUser="valdenar" docAuthor="Czerwiński, Tomasz, Szawaryn, Karol &amp; Tomaszewska, Wioletta" docDate="2020" docId="03E5883D462E304EFDF635E7010DFEC4" docLanguage="en" docName="ejt-692_czerwinski_szawaryn_tomaszewska.pdf" docOrigin="European Journal of Taxonomy 692" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhyzobius albinos Czerwiński &amp; Szawaryn &amp; Tomaszewska 2020, sp. nov." docType="treatment" docUuid="BA648A58-DE41-49CE-B3DA-F7D06F2E868C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="7" masterDocId="FFDCF045462F3048FF8C33030348FF97" masterDocTitle="Three new species of the genus Rhyzobius Stephens, 1829 from New Guinea (Coleoptera: Coccinellidae: Coccidulini)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="2" updateTime="1698844045825" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B3D159002F66F61B6EA7E3E5FE332B71">Three new species of the genus Rhyzobius Stephens, 1829 from New Guinea (Coleoptera: Coccinellidae: Coccidulini)</mods:title>
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<mods:name id="C696DE1C8538C8B964192C796F4A2278" type="personal">
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<mods:namePart id="E28173821275ADD41E229C258F714E61">Czerwiński, Tomasz</mods:namePart>
<mods:nameIdentifier id="623613CA858B8BC83E1900876718D967" type="LSID">B7AF2729-3590-48B3-A065-7825623EF8B3</mods:nameIdentifier>
<mods:affiliation id="A2E23530F639CFEBC938FB2D5DCC64E3">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland.</mods:affiliation>
<mods:nameIdentifier id="88751CB02C39F52984D861107025DDC8" type="email">tczerwinski@miiz.waw.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="DE305819D88075BDA6241C316263E8DD" type="personal">
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<mods:namePart id="938B9899F89896CA043309BA4A2AFB7D">Szawaryn, Karol</mods:namePart>
<mods:nameIdentifier id="7ADCD117A041437CD27D1A3760F58FD3" type="LSID">D741C759-6CDD-4B61-BE93-3ECC2918A73F</mods:nameIdentifier>
<mods:affiliation id="F53B1FC24105122972BCBD33AF5509B7">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland.</mods:affiliation>
<mods:nameIdentifier id="4AD63E77A4A8451398422BA842F05C2B" type="email">k.szawaryn@gmail.com</mods:nameIdentifier>
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<mods:namePart id="15525B8933608FF1E79CD2B1338C190D">Tomaszewska, Wioletta</mods:namePart>
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<mods:affiliation id="D94EFB6A24A4A0A08BF713E6362379D5">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland.</mods:affiliation>
<mods:nameIdentifier id="BB7DEED414C5CAC95DF357BD2C4EBC56" type="email">wiolkat@miiz.waw.pl</mods:nameIdentifier>
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<mods:title id="001BA564EECA98B666D9E4B1E7CE1E2F">European Journal of Taxonomy</mods:title>
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<mods:date id="F9D7E9E638D412BA150FFB5199796A2B">2020</mods:date>
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<mods:number id="2C97223233826051B04CB7F6566439AF">2020-07-28</mods:number>
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<mods:number id="C68A699C1CBF6B768F5A22A64B794590">692</mods:number>
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<mods:classification id="0FC98879E7DBEE9F6B76AA4520EA9695">journal article</mods:classification>
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<treatment id="03E5883D462E304EFDF635E7010DFEC4" ID-DOI="http://doi.org/10.5281/zenodo.4328099" ID-GBIF-Taxon="166095762" ID-Zenodo-Dep="4328099" LSID="urn:lsid:zoobank.org:act:BA648A58-DE41-49CE-B3DA-F7D06F2E868C" httpUri="http://treatment.plazi.org/id/03E5883D462E304EFDF635E7010DFEC4" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2">
<subSubSection id="C3566AA0462E3049FDF635E70786F8B4" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BF3392B462E3049FDF635E700F1F968" blockId="1.[357,1230,1764,1862]" box="[634,953,1764,1791]" pageId="1" pageNumber="2">
<heading id="D0BB8E47462E3049FDF635E700F1F968" box="[634,953,1764,1791]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
<taxonomicName id="4C4C42A8462E3049FDF635E7001FF968" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[634,855,1764,1791]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="albinos" status="sp. nov.">
<emphasis id="B938E539462E3049FDF635E7001FF968" bold="true" box="[634,855,1764,1791]" italics="true" pageId="1" pageNumber="2">Rhyzobius albinos</emphasis>
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<taxonomicNameLabel id="A20B5842462E3049FCD335E600F1F968" box="[863,953,1765,1791]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3566AA0462E3049FDD23428009DF8D1" box="[606,981,1835,1862]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BF3392B462E3049FDD23428009DF8D1" blockId="1.[357,1230,1764,1862]" box="[606,981,1835,1862]" pageId="1" pageNumber="2">
<figureCitation id="137725AE462E3049FDD2342801EAF8D2" box="[606,674,1835,1861]" captionStart="Fig" captionStartId="3.[189,232,1923,1949]" captionTargetBox="[247,1322,301,1838]" captionTargetId="figure@3.[244,1344,265,1883]" captionTargetPageId="3" captionText="Fig. 1. Habitus of new species of Rhyzobius spp. (holotypes). A–D. R. albinos sp. nov., ♂ (MNHN). E–H. R. atramentarius sp. nov., ♂ (MNHN). I–L. R. serratus sp. nov., ♂ (MNHN). A, E, I. Dorsal view. B, F, J. Ventral view. C, G, K. Lateral view. D, H, L. Frontal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973320" httpUri="https://zenodo.org/record/3973320/files/figure.png" pageId="1" pageNumber="2">Figs 1</figureCitation>
A–D, 2A–I, 4A–C, 5A–D
</paragraph>
</subSubSection>
<subSubSection id="C3566AA0462E304AFF31346F001AFE42" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BF3392B462E3049FF31346F027EF811" blockId="1.[189,310,1900,1926]" box="[189,310,1900,1926]" pageId="1" pageNumber="2">
<heading id="D0BB8E47462E3049FF31346F027EF811" bold="true" box="[189,310,1900,1926]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
<emphasis id="B938E539462E3049FF31346F027EF811" bold="true" box="[189,310,1900,1926]" pageId="1" pageNumber="2">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3392B462E304AFF313498001AFE42" blockId="1.[189,1399,1947,2009]" lastBlockId="2.[189,1399,267,470]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
This species is most similar to
<taxonomicName id="4C4C42A8462E3049FDCC349F00F7F822" authority="Tomaszewska, 2010" authorityName="Tomaszewska" authorityYear="2010" box="[576,959,1947,1974]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="poorani">
<emphasis id="B938E539462E3049FDCC349F011AF822" box="[576,594,1948,1973]" italics="true" pageId="1" pageNumber="2">R</emphasis>
.
<bibRefCitation id="EFDD44DA462E3049FDE8349F00F7F822" author="Tomaszewska W." box="[612,959,1947,1973]" firstAuthor="poorani Tomaszewska" pageId="1" pageNumber="2" refId="ref8317" refString="Tomaszewska W. 2010. Rhyzobius Stephens 1829 (Coleoptera: Coccinellidae), a Revision of the World Species. Fauna Mundi 2, Natura optima dux Foundation, Warsaw." type="book" year="2010">
<emphasis id="B938E539462E3049FDE8349F0189F822" box="[612,705,1948,1973]" italics="true" pageId="1" pageNumber="2">poorani</emphasis>
Tomaszewska, 2010
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C4C42A8462E3049FB8C349F061CF822" authority="Tomaszewska, 2010" authorityName="Tomaszewska" authorityYear="2010" box="[1024,1364,1947,1974]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="jaya">
<emphasis id="B938E539462E3049FB8C349F075AF822" box="[1024,1042,1948,1973]" italics="true" pageId="1" pageNumber="2">R</emphasis>
.
<emphasis id="B938E539462E3049FBA9349F0710F822" box="[1061,1112,1948,1973]" italics="true" pageId="1" pageNumber="2">jaya</emphasis>
<bibRefCitation id="EFDD44DA462E3049FBEF3498061CF822" author="Tomaszewska W." box="[1123,1364,1947,1973]" firstAuthor="Tomaszewska" journalOrPublisher="Fauna Mundi 2, Natura optima dux Foundation, Warsaw" pageId="1" pageNumber="2" refId="ref8317" refString="Tomaszewska W. 2010. Rhyzobius Stephens 1829 (Coleoptera: Coccinellidae), a Revision of the World Species. Fauna Mundi 2, Natura optima dux Foundation, Warsaw." title="Rhyzobius Stephens 1829 (Coleoptera: Coccinellidae), a Revision of the World Species" type="book" year="2010">
Tomaszewska,
<quantity id="4CB494CE462E3049FA953498061CF822" box="[1305,1364,1947,1973]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="1" pageNumber="2" unit="in" value="2010.0">2010</quantity>
</bibRefCitation>
</taxonomicName>
in body colouration. However,
<taxonomicName id="4C4C42A8462E3049FD9C34BC01CFF84F" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[528,647,1982,2009]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="albinos" status="sp. nov.">
<emphasis id="B938E539462E3049FD9C34BC016AF84F" box="[528,546,1983,2008]" italics="true" pageId="1" pageNumber="2">R</emphasis>
.
<emphasis id="B938E539462E3049FDBD34BD01CFF84F" box="[561,647,1982,2008]" italics="true" pageId="1" pageNumber="2">albinos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462E3049FD1C34BC01A4F84E" box="[656,748,1983,2009]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C4C42A8462E3049FC0634BC0741F84F" authorityName="Tomaszewska" authorityYear="2010" box="[906,1033,1983,2009]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="poorani">
<emphasis id="B938E539462E3049FC0634BC00D4F84F" box="[906,924,1983,2008]" italics="true" pageId="1" pageNumber="2">R</emphasis>
.
<emphasis id="B938E539462E3049FC2034BC0741F84F" box="[940,1033,1983,2008]" italics="true" pageId="1" pageNumber="2">poorani</emphasis>
</taxonomicName>
by having simple, not swollen pronotal anterior corners and a prosternal process with complete lateral carinae, joined anteriorly in the form of a triangle.
<taxonomicName id="4C4C42A8462D304AFE16322C0105FEDE" authorityName="Tomaszewska" authorityYear="2010" box="[410,589,303,329]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="jaya">
<emphasis id="B938E539462D304AFE16322C0105FEDE" box="[410,589,303,329]" italics="true" pageId="2" pageNumber="3">Rhyzobius jaya</emphasis>
</taxonomicName>
has a smooth prosternal process without carinae, and straight ventral antennal grooves reaching beyond the posterior margin of the eyes, whereas
<taxonomicName id="4C4C42A8462D304AFBEF32500795FEFB" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[1123,1245,338,364]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="albinos" status="sp. nov.">
<emphasis id="B938E539462D304AFBEF3250073DFEFB" box="[1123,1141,339,364]" italics="true" pageId="2" pageNumber="3">R</emphasis>
.
<emphasis id="B938E539462D304AFB0B32510795FEFB" box="[1159,1245,338,364]" italics="true" pageId="2" pageNumber="3">albinos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462D304AFB643251060EFEFB" box="[1256,1350,338,364]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
has ventral antennal grooves distinctly circularly bent towards the outer margin of the eyes. From all other New Guinean species of
<taxonomicName id="4C4C42A8462D304AFE6D329B0112FE25" authorityName="Stephens" authorityYear="1829" box="[481,602,408,434]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B938E539462D304AFE6D329B0112FE25" box="[481,602,408,434]" italics="true" pageId="2" pageNumber="3">Rhyzobius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C42A8462D304AFDE4329A0197FE25" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[616,735,408,435]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="albinos" status="sp. nov.">
<emphasis id="B938E539462D304AFDE4329A0132FE25" box="[616,634,409,434]" italics="true" pageId="2" pageNumber="3">R</emphasis>
.
<emphasis id="B938E539462D304AFD05329B0197FE25" box="[649,735,408,434]" italics="true" pageId="2" pageNumber="3">albinos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462D304AFD6A329A0008FE24" box="[742,832,409,435]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished by the bright, almost white ventral body surface, excluding meso- and metaventrite.
</paragraph>
</subSubSection>
<subSubSection id="C3566AA0462D304AFF3132FE0763FDFE" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BF3392B462D304AFF3132FE020CFD80" blockId="2.[189,324,509,535]" box="[189,324,509,535]" pageId="2" pageNumber="3">
<heading id="D0BB8E47462D304AFF3132FE020CFD80" bold="true" box="[189,324,509,535]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B938E539462D304AFF3132FE020CFD80" bold="true" box="[189,324,509,535]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3392B462D304AFF3131280763FDFE" blockId="2.[189,1399,555,617]" pageId="2" pageNumber="3">
The specific epithet is a noun in apposition, it means ‘albino’ in Polish and refers to the bright colouration of most of the ventral surfaces, not observed in other species of
<taxonomicName id="4C4C42A8462D304AFC20314C076DFDFE" authorityName="Stephens" authorityYear="1829" box="[940,1061,591,617]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B938E539462D304AFC20314C076DFDFE" box="[940,1061,591,617]" italics="true" pageId="2" pageNumber="3">Rhyzobius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3566AA0462D304AFF3131930088FA2A" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BF3392B462D304AFF31319302E3FD3D" blockId="2.[189,427,656,682]" box="[189,427,656,682]" pageId="2" pageNumber="3">
<heading id="D0BB8E47462D304AFF31319302E3FD3D" bold="true" box="[189,427,656,682]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B938E539462D304AFF31319302E3FD3D" bold="true" box="[189,427,656,682]" pageId="2" pageNumber="3">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B243376462D304AFF6731BC023FFCD4" ID-GBIF-Occurrence="2824181308" collectingDate="2012-11-09" collectingDateMax="2012-11-11" collectingDateMin="2012-11-09" collectionCode="MNHN" country="NEW GUINEA" county="Bananumbo" elevation="1700" latitude="-5.759278" location="Madang Province" longLatPrecision="1" longitude="145.23561" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8BF3392B462D304AFF6731BC0215FD4E" blockId="2.[189,1399,703,835]" box="[235,349,703,729]" pageId="2" pageNumber="3">
<emphasis id="B938E539462D304AFF6731BC0215FD4E" bold="true" box="[235,349,703,729]" pageId="2" pageNumber="3">
<typeStatus id="54F78789462D304AFF6731BC0215FD4E" box="[235,349,703,729]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF3392B462D304AFF3131E1023FFCD4" blockId="2.[189,1399,703,835]" pageId="2" pageNumber="3">
<collectingCountry id="F35B79BB462D304AFF3131E1023DFD6B" box="[189,373,738,764]" pageId="2" pageNumber="3">NEW GUINEA</collectingCountry>
•
<specimenCount id="9D4AF2A2462D304AFE0031E302EBFD6B" box="[396,419,736,764]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
;
<location id="8E936FF0462D304AFE3C31E101C9FD6B" LSID="urn:lsid:plazi:treatment:03E5883D462E304EFDF635E7010DFEC4:8E936FF0462D304AFE3C31E101C9FD6B" box="[432,641,738,764]" country="NEW GUINEA" county="Bananumbo" latitude="-5.759278" longLatPrecision="1" longitude="145.23561" municipality="Mt Wilhelm" name="Madang Province" pageId="2" pageNumber="3">Madang Province</location>
,
<collectingMunicipality id="6B97A351462D304AFD0031E10055FD6B" box="[652,797,738,764]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
,
<collectingCounty id="629241A7462D304AFCAB31E100F1FD6B" box="[807,953,738,764]" pageId="2" pageNumber="3">Bananumbo</collectingCounty>
;
<geoCoordinate id="EE785FEC462D304AFC4831E3071CFD6B" box="[964,1108,736,764]" degrees="5" direction="south" minutes="45" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="33.4" value="-5.759278">5º45ʹ33.4ʺ S</geoCoordinate>
,
<geoCoordinate id="EE785FEC462D304AFBEE31E3065CFD6B" box="[1122,1300,736,764]" degrees="145" direction="east" minutes="14" orientation="longitude" pageId="2" pageNumber="3" precision="1" seconds="08.2" value="145.23561">145º14ʹ08.2ʺ E</geoCoordinate>
;
<elevation id="0061DE18462D304AFA9331E103BEFC88" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="3" unit="m" value="1700.0">
<quantity id="4CB494CE462D304AFA9331E1063FFD6B" box="[1311,1399,738,764]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="3" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFF21FEB462D304AFE883006028CFC88" box="[260,452,773,800]" pageId="2" pageNumber="3" value="2012-11-09" valueMax="2012-11-11" valueMin="2012-11-09">
<collectingDate id="EFB6E603462D304AFE883006028CFC88" box="[260,452,773,800]" pageId="2" pageNumber="3" value="2012-11-09" valueMax="2012-11-11" valueMin="2012-11-09">9–11 Nov. 2012</collectingDate>
</date>
; “06686-
<location id="8E936FF0462D304AFDA230060077FC88" LSID="urn:lsid:plazi:treatment:03E5883D462E304EFDF635E7010DFEC4:8E936FF0462D304AFDA230060077FC88" box="[558,831,773,799]" country="NEW GUINEA" county="Bananumbo" latitude="-5.759278" longLatPrecision="1" longitude="145.23561" municipality="Mt Wilhelm" name="CoCocc Ibisca Niugini" pageId="2" pageNumber="3">CoCocc Ibisca Niugini</location>
2012”; FIT-MW1700-O-8/8-d16; plot 15, order 17260;
<collectionCode id="ED5DA1EE462D304AFE9A302A0238FCD4" box="[278,368,809,835]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BF3392B462D304AFF6730690220FC13" blockId="2.[189,1399,874,1358]" box="[235,360,874,900]" pageId="2" pageNumber="3">
<emphasis id="B938E539462D304AFF6730690220FC13" bold="true" box="[235,360,874,900]" pageId="2" pageNumber="3">
<typeStatus id="54F78789462D304AFF6730690220FC13" box="[235,360,874,900]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF3392B462D304AFF31308E021BFAD9" blockId="2.[189,1399,874,1358]" pageId="2" pageNumber="3">
<materialsCitation id="3B243376462D304AFF31308E0653FC30" ID-GBIF-Occurrence="2824181314" box="[189,1307,907,935]" collectingDate="2012-11-09" collectingDateMax="2012-11-11" collectingDateMin="2012-11-09" collectionCode="MNHN" country="NEW GUINEA" county="Bananumbo" elevation="1700" latitude="-5.759278" location="Madang Province" longLatPrecision="1" longitude="145.23561" municipality="Madang Province" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F35B79BB462D304AFF31308E0233FC30" box="[189,379,909,935]" pageId="2" pageNumber="3">NEW GUINEA</collectingCountry>
–
<emphasis id="B938E539462D304AFE2E308E01CDFC30" bold="true" box="[418,645,909,935]" pageId="2" pageNumber="3">
<collectingMunicipality id="6B97A351462D304AFE2E308E01CDFC30" box="[418,645,909,935]" pageId="2" pageNumber="3">Madang Province</collectingMunicipality>
</emphasis>
•
<specimenCount id="9D4AF2A2462D304AFD24308E0192FC30" box="[680,730,907,935]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED5DA1EE462D304AFB4D308E0653FC30" box="[1217,1307,909,935]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFAB2308E0115FC79" ID-GBIF-Occurrence="2824181304" collectingDate="2012-10-18" collectingDateMax="2012-10-20" collectingDateMin="2012-10-18" collectionCode="MNHN" elevation="2200" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFAB2308E0638FC30" box="[1342,1392,907,935]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFF3130B20218FC5D" box="[189,336,944,971]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFED130B302FFFC5C" box="[349,439,944,971]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">
<elevation id="0061DE18462D304AFED130B302FFFC5C" box="[349,439,944,971]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">2200 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFD8A30B30198FC5D" box="[518,720,944,970]" pageId="2" pageNumber="3" value="2012-10-18" valueMax="2012-10-20" valueMin="2012-10-18">
<collectingDate id="EFB6E603462D304AFD8A30B30198FC5D" box="[518,720,944,970]" pageId="2" pageNumber="3" value="2012-10-18" valueMax="2012-10-20" valueMin="2012-10-18">18–20 Oct. 2012</collectingDate>
</date>
; “04064-CoCocc Ibisca Niugini 2012”; FIT-MW2200-C- 2/8-d03; plot 3, order 1603;
<collectionCode id="ED5DA1EE462D304AFD8F30D70115FC79" box="[515,605,980,1006]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFDFF30D0077DFB86" ID-GBIF-Occurrence="2824181311" collectingDate="2012-10-19" collectingDateMax="2012-10-21" collectingDateMin="2012-10-19" collectionCode="MNHN" elevation="2200" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFDFF30D001D6FC79" box="[627,670,978,1006]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFD2730D70074FC7A" box="[683,828,979,1006]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFCCB30D000D6FC79" box="[839,926,979,1006]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">
<elevation id="0061DE18462D304AFCCB30D000D6FC79" box="[839,926,979,1006]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">2200 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFC6530D007E6FC7A" box="[1001,1198,979,1005]" pageId="2" pageNumber="3" value="2012-10-19" valueMax="2012-10-21" valueMin="2012-10-19">
<collectingDate id="EFB6E603462D304AFC6530D007E6FC7A" box="[1001,1198,979,1005]" pageId="2" pageNumber="3" value="2012-10-19" valueMax="2012-10-21" valueMin="2012-10-19">19–21 Oct. 2012</collectingDate>
</date>
; “09588-CoCocc Ibisca Niugini 2012”; FIT-MW2200-C-2/8-d04; plot 11, order 6111;
<collectionCode id="ED5DA1EE462D304AFC5730F4077DFB86" box="[987,1077,1015,1041]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFBC730F40228FBC0" ID-GBIF-Occurrence="2824181309" collectingDate="2012-10-29" collectingDateMax="2012-10-31" collectingDateMin="2012-10-29" collectionCode="MNHN" elevation="1200" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFBC730F4073FFB86" box="[1099,1143,1013,1041]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFB0830F4065DFB87" box="[1156,1301,1014,1041]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFA9330F4063FFB86" box="[1311,1399,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="1200.0">
<elevation id="0061DE18462D304AFA9330F4063FFB86" box="[1311,1399,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="1200.0">1200 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFE8837190285FBA3" box="[260,461,1050,1076]" pageId="2" pageNumber="3" value="2012-10-29" valueMax="2012-10-31" valueMin="2012-10-29">
<collectingDate id="EFB6E603462D304AFE8837190285FBA3" box="[260,461,1050,1076]" pageId="2" pageNumber="3" value="2012-10-29" valueMax="2012-10-31" valueMin="2012-10-29">29–31 Oct. 2012</collectingDate>
</date>
; “16881-CoCocc Ibisca Niugini 2012”; FIT-MW1200-J-3/8-d05; plot 10, order 6033;
<collectionCode id="ED5DA1EE462D304AFE8A373E0228FBC0" box="[262,352,1085,1111]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFEF9373E0001FBEC" ID-GBIF-Occurrence="2824181313" collectingDate="2012-10-27" collectingDateMax="2012-10-29" collectingDateMin="2012-10-27" collectionCode="MNHN" elevation="1700" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFEF9373E02E8FBC0" box="[373,416,1083,1111]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFE21373E0175FBC0" box="[429,573,1085,1111]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFDCB373E01D6FBC0" box="[583,670,1085,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="3" unit="m" value="1700.0">
<elevation id="0061DE18462D304AFDCB373E01D6FBC0" box="[583,670,1085,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="3" unit="m" value="1700.0">1700 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFD64373E00E4FBC0" box="[744,940,1085,1111]" pageId="2" pageNumber="3" value="2012-10-27" valueMax="2012-10-29" valueMin="2012-10-27">
<collectingDate id="EFB6E603462D304AFD64373E00E4FBC0" box="[744,940,1085,1111]" pageId="2" pageNumber="3" value="2012-10-27" valueMax="2012-10-29" valueMin="2012-10-27">27–29 Oct. 2012</collectingDate>
</date>
; “P1927-CoCocc Ibisca Niugini 2012”; FIT-MW1700-G-2/8-d03; plot 7, order 18341;
<collectionCode id="ED5DA1EE462D304AFD6337620001FBEC" box="[751,841,1121,1147]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFCE93763062BFB09" ID-GBIF-Occurrence="2824181315" collectingDate="2012-10-20" collectingDateMax="2012-10-22" collectingDateMin="2012-10-20" collectionCode="MNHN" elevation="2200" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFCE9376300DCFBED" box="[869,916,1118,1146]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFC2937620771FBED" box="[933,1081,1120,1147]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFBCB376307E9FBEC" box="[1095,1185,1120,1147]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">
<elevation id="0061DE18462D304AFBCB376307E9FBEC" box="[1095,1185,1120,1147]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">2200 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFB7F376303B4FB0A" pageId="2" pageNumber="3" value="2012-10-20" valueMax="2012-10-22" valueMin="2012-10-20">
<collectingDate id="EFB6E603462D304AFB7F376303B4FB0A" pageId="2" pageNumber="3" value="2012-10-20" valueMax="2012-10-22" valueMin="2012-10-20">20–22 Oct. 2012</collectingDate>
</date>
; “P2334-CoCocc Ibisca Niugini 2012”; FIT-MW
<date id="FFF21FEB462D304AFCC6378000F8FB09" box="[842,944,1155,1182]" pageId="2" pageNumber="3" value="2200-01-03">2200-I-3</date>
/8-d05; plot 9, order 18623;
<collectionCode id="ED5DA1EE462D304AFA853787062BFB09" box="[1289,1379,1156,1182]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFF3237A40128FB73" ID-GBIF-Occurrence="2824181312" collectingDate="2012-11-09" collectingDateMax="2012-11-11" collectingDateMin="2012-11-09" elevation="1700" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFF3237A403A2FB56" box="[190,234,1189,1217]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFF7437A402C1FB57" box="[248,393,1190,1217]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFE1837A402A5FB56" box="[404,493,1191,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="3" unit="m" value="1700.0">
<elevation id="0061DE18462D304AFE1837A402A5FB56" box="[404,493,1191,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="3" unit="m" value="1700.0">1700 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFDB537A401BDFB56" box="[569,757,1191,1217]" pageId="2" pageNumber="3" value="2012-11-09" valueMax="2012-11-11" valueMin="2012-11-09">
<collectingDate id="EFB6E603462D304AFDB537A401BDFB56" box="[569,757,1191,1217]" pageId="2" pageNumber="3" value="2012-11-09" valueMax="2012-11-11" valueMin="2012-11-09">9–11 Nov. 2012</collectingDate>
</date>
; “06696-CoCocc Ibisca Niugini 2012”; FIT-MW1700- S-8/8-d16; plot 19, order 5202; MIZ
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFDFA37C9076FFA90" ID-GBIF-Occurrence="2824181302" collectingDate="2012-10-27" collectingDateMax="2012-10-29" collectingDateMin="2012-10-27" elevation="2200" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFDFA37C901E9FB73" box="[630,673,1224,1252]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFD2237C90076FB73" box="[686,830,1226,1252]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFCC437C900E8FB73" box="[840,928,1226,1252]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">
<elevation id="0061DE18462D304AFCC437C900E8FB73" box="[840,928,1226,1252]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">2200 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFC6637C907E6FB73" box="[1002,1198,1226,1252]" pageId="2" pageNumber="3" value="2012-10-27" valueMax="2012-10-29" valueMin="2012-10-27">
<collectingDate id="EFB6E603462D304AFC6637C907E6FB73" box="[1002,1198,1226,1252]" pageId="2" pageNumber="3" value="2012-10-27" valueMax="2012-10-29" valueMin="2012-10-27">27–29 Oct. 2012</collectingDate>
</date>
; “02308-CoCocc Ibisca Niugini 2012”; FIT-MW2200-N-6/8-d12; plot 14, order 1630; MIZ
</materialsCitation>
•
<materialsCitation id="3B243376462D304AFBCD37EE021BFAD9" ID-GBIF-Occurrence="2824181306" collectingDate="2012-11-20" collectingDateMax="2012-11-22" collectingDateMin="2012-11-20" elevation="2200" location="Mt Wilhelm" municipality="Mt Wilhelm" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4AF2A2462D304AFBCD37EE0727FA90" box="[1089,1135,1259,1287]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6B97A351462D304AFBF237EE0659FA90" box="[1150,1297,1261,1287]" pageId="2" pageNumber="3">Mt Wilhelm</collectingMunicipality>
;
<quantity id="4CB494CE462D304AFA9137EE063FFA90" box="[1309,1399,1261,1287]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">
<elevation id="0061DE18462D304AFA9137EE063FFA90" box="[1309,1399,1261,1287]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">2200 m</elevation>
</quantity>
a.s.l.;
<date id="FFF21FEB462D304AFE883613029BFABD" box="[260,467,1296,1323]" pageId="2" pageNumber="3" value="2012-11-20" valueMax="2012-11-22" valueMin="2012-11-20">
<collectingDate id="EFB6E603462D304AFE883613029BFABD" box="[260,467,1296,1323]" pageId="2" pageNumber="3" value="2012-11-20" valueMax="2012-11-22" valueMin="2012-11-20">20–22 Nov. 2012</collectingDate>
</date>
; “P2294-CoCocc Ibisca Niugini 2012”; FIT-MW2200-D-3/8-d05; plot 4, order 18478; MIZ.
</materialsCitation>
</paragraph>
<paragraph id="8BF3392B462D304AFF3236760215FA18" blockId="2.[190,349,1397,1423]" box="[190,349,1397,1423]" pageId="2" pageNumber="3">
<heading id="D0BB8E47462D304AFF3236760215FA18" bold="true" box="[190,349,1397,1423]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B938E539462D304AFF3236760215FA18" bold="true" box="[190,349,1397,1423]" pageId="2" pageNumber="3">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3392B462D304AFF3236A70088FA2A" blockId="2.[190,960,1443,1470]" box="[190,960,1443,1470]" pageId="2" pageNumber="3">
<collectingCountry id="F35B79BB462D304AFF3236A702EBFA2A" box="[190,419,1443,1470]" name="Papua New Guinea" pageId="2" pageNumber="3">Papua New Guinea</collectingCountry>
,
<collectingRegion id="4988F7C9462D304AFE2336A001C9FA2A" box="[431,641,1443,1469]" country="Papua New Guinea" name="Madang" pageId="2" pageNumber="3">Madang Province</collectingRegion>
, Mt Wilhelm, Bananumbo.
</paragraph>
</subSubSection>
<subSubSection id="C3566AA0462D304CFF3236E602E5F836" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="8BF3392B462D304AFF3236E60207FA68" blockId="2.[190,335,1509,1535]" box="[190,335,1509,1535]" pageId="2" pageNumber="3">
<heading id="D0BB8E47462D304AFF3236E60207FA68" bold="true" box="[190,335,1509,1535]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B938E539462D304AFF3236E60207FA68" bold="true" box="[190,335,1509,1535]" pageId="2" pageNumber="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3392B462D304AFF313517063FF9BA" blockId="2.[189,1399,1555,1582]" box="[189,1399,1555,1582]" pageId="2" pageNumber="3">
MEASUREMENTS. Length 2.00–
<quantity id="4CB494CE462D304AFD94351001CBF9B9" box="[536,643,1555,1582]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="2" pageNumber="3" unit="mm" value="2.25">2.25 mm</quantity>
; TL/EW = 1.42–1.50; PL/PW = 0.52–0.56; EL/EW = 1.06–1.07.
</paragraph>
<paragraph id="8BF3392B462D304AFF3135560651F888" blockId="2.[189,1399,1621,1823]" pageId="2" pageNumber="3">
BODY (
<figureCitation id="137725AE462D304AFE9F35560214F9F8" box="[275,348,1621,1647]" captionStart="Fig" captionStartId="3.[189,232,1923,1949]" captionTargetBox="[247,1322,301,1838]" captionTargetId="figure@3.[244,1344,265,1883]" captionTargetPageId="3" captionText="Fig. 1. Habitus of new species of Rhyzobius spp. (holotypes). A–D. R. albinos sp. nov., ♂ (MNHN). E–H. R. atramentarius sp. nov., ♂ (MNHN). I–L. R. serratus sp. nov., ♂ (MNHN). A, E, I. Dorsal view. B, F, J. Ventral view. C, G, K. Lateral view. D, H, L. Frontal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973320" httpUri="https://zenodo.org/record/3973320/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
A–C, 2A). Moderately oval and convex dorsally, winged; with antennae, mouthparts, hypomera, apex of elytra, most of abdominal ventrites and legs except coxae whitish; frons, anterior and lateral margins of pronotum, preapical parts of elytra, prosternum, epipleura, coxae and partially abdominal ventrite 1 pale brown; vertex, posterior and central parts of pronotum, most of elytra, meso- and metaventrite blackish or dark brown. Elytra with weak bluish, metallic sheen. Dorsum with moderately long and uniform pubescence, not forming pattern on elytra. Elytral bristles absent.
</paragraph>
<paragraph id="8BF3392B462D304CFF3134440786FD12" blockId="2.[189,1399,1862,1994]" lastBlockId="4.[189,1399,267,646]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
HEAD (
<figureCitation id="137725AE462D304AFE8234450222F8F7" box="[270,362,1862,1888]" captionStart="Fig" captionStartId="3.[189,232,1923,1949]" captionTargetBox="[247,1322,301,1838]" captionTargetId="figure@3.[244,1344,265,1883]" captionTargetPageId="3" captionText="Fig. 1. Habitus of new species of Rhyzobius spp. (holotypes). A–D. R. albinos sp. nov., ♂ (MNHN). E–H. R. atramentarius sp. nov., ♂ (MNHN). I–L. R. serratus sp. nov., ♂ (MNHN). A, E, I. Dorsal view. B, F, J. Ventral view. C, G, K. Lateral view. D, H, L. Frontal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973320" httpUri="https://zenodo.org/record/3973320/files/figure.png" pageId="2" pageNumber="3">Fig. 1D</figureCitation>
). Dorsally exposed with eyes almost entirely visible; ventral antennal grooves distinctly circularly bent towards outer margin of eye; corpotentorium absent. Eyes dorsally with inner orbits arcuate, closest at middle; ocular canthus distinct; interocular distance 0.50–0.52 times as wide as head across eyes; interfacetal setae distinct. Antenna (
<figureCitation id="137725AE462D304AFD6334B30000F85D" box="[751,840,1968,1994]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="2" pageNumber="3">Fig. 2B</figureCitation>
) 0.77–0.80 times as long as head capsule width, composed of 11 antennomeres; scape 1.65–1.70 times as long as pedicel; pedicel narrower than scape, barrel-shaped, 1.25–1.28 times as long as wide; antennomere 3 about 2.5 times longer than wide, and 2.15–2.30 times as long as antennomere 4; antennomere 4 as long as 5; antennomeres 6 and 7 quadrate. Antennal club consisting of 3 antennomeres, with two subterminal segments asymmetrical; penultimate antennomere as long as terminal one; terminal antennomere rounded apically. Anterior clypeal margin (
<figureCitation id="137725AE462B304CFF4A32B8026CFE42" box="[198,292,443,469]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2E</figureCitation>
) distinctly emarginate with rounded lateral lobes and median area membranous. Labrum truncate at apex. Maxillary cardo transverse with outer angle reaching slightly outside of mouth cavity; terminal palpomere 1.30–1.35 times as long as wide, expanded apically. Mentum weakly transverse, 1.5 times as broad as long; anterior margin weakly arcuate; ventral surface with horseshoe-like impression; prementum transverse; ligula parallel-sided; labial palps separated by distance about equal to width of palpiger; apical palpomere as long and as broad as penultimate one; submentum distinct.
</paragraph>
<caption id="DF3369A3462C304BFF313480079BF873" ID-DOI="http://doi.org/10.5281/zenodo.3973320" ID-Zenodo-Dep="3973320" httpUri="https://zenodo.org/record/3973320/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1923,1949]" targetBox="[247,1322,301,1838]" targetPageId="3">
<paragraph id="8BF3392B462C304BFF313480079BF873" blockId="3.[189,1399,1922,2021]" pageId="3" pageNumber="4">
<emphasis id="B938E539462C304BFF3134800245F80A" bold="true" box="[189,269,1923,1949]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
Habitus of new species of
<taxonomicName id="4C4C42A8462C304BFDDB34870198F809" authorityName="Stephens" authorityYear="1829" box="[599,720,1924,1950]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B938E539462C304BFDDB34870198F809" box="[599,720,1924,1950]" italics="true" pageId="3" pageNumber="4">Rhyzobius</emphasis>
</taxonomicName>
spp. (holotypes).
<emphasis id="B938E539462C304BFC25348000AAF80A" bold="true" box="[937,994,1923,1949]" pageId="3" pageNumber="4">A–D</emphasis>
.
<taxonomicName id="4C4C42A8462C304BFC7F34870723F809" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[1011,1131,1924,1950]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="albinos" status="sp. nov.">
<emphasis id="B938E539462C304BFC7F3487074DF80A" box="[1011,1029,1924,1949]" italics="true" pageId="3" pageNumber="4">R</emphasis>
.
<emphasis id="B938E539462C304BFB9934870723F809" box="[1045,1131,1924,1950]" italics="true" pageId="3" pageNumber="4">albinos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462C304BFBF93487079DF809" box="[1141,1237,1924,1950]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (MNHN).
<emphasis id="B938E539462C304BFF3134A403BEF856" bold="true" box="[189,246,1959,1985]" pageId="3" pageNumber="4">E–H</emphasis>
.
<taxonomicName id="4C4C42A8462C304BFE8F34AB0280F856" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[259,456,1959,1985]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="atramentarius" status="sp. nov.">
<emphasis id="B938E539462C304BFE8F34AB025DF856" box="[259,277,1960,1985]" italics="true" pageId="3" pageNumber="4">R</emphasis>
.
<emphasis id="B938E539462C304BFEAE34AB0280F856" box="[290,456,1960,1985]" italics="true" pageId="3" pageNumber="4">atramentarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462C304BFE4334A40162F856" box="[463,554,1959,1985]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (MNHN).
<emphasis id="B938E539462C304BFD4734A401B1F856" bold="true" box="[715,761,1959,1985]" pageId="3" pageNumber="4">I–L</emphasis>
.
<taxonomicName id="4C4C42A8462C304BFC8A34AB00CDF856" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[774,901,1959,1985]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="serratus" status="sp. nov.">
<emphasis id="B938E539462C304BFC8A34AB0050F856" box="[774,792,1960,1985]" italics="true" pageId="3" pageNumber="4">R</emphasis>
.
<emphasis id="B938E539462C304BFCA934AB00CDF856" box="[805,901,1960,1985]" italics="true" pageId="3" pageNumber="4">serratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462C304BFC0734A400AFF856" box="[907,999,1959,1985]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (MNHN).
<emphasis id="B938E539462C304BFB0434A407D5F856" bold="true" box="[1160,1181,1959,1985]" pageId="3" pageNumber="4">A</emphasis>
,
<emphasis id="B938E539462C304BFB2634A407F6F856" bold="true" box="[1194,1214,1959,1985]" pageId="3" pageNumber="4">E</emphasis>
,
<emphasis id="B938E539462C304BFB4734A4079EF856" bold="true" box="[1227,1238,1959,1985]" pageId="3" pageNumber="4">I</emphasis>
. Dorsal view.
<emphasis id="B938E539462C304BFF3134C90399F873" bold="true" box="[189,209,1994,2020]" pageId="3" pageNumber="4">B</emphasis>
,
<emphasis id="B938E539462C304BFF5334C903B9F873" bold="true" box="[223,241,1994,2020]" pageId="3" pageNumber="4">F</emphasis>
,
<emphasis id="B938E539462C304BFE8C34C90247F873" bold="true" box="[256,271,1994,2020]" pageId="3" pageNumber="4">J</emphasis>
. Ventral view.
<emphasis id="B938E539462C304BFE3234C9029BF873" bold="true" box="[446,467,1994,2020]" pageId="3" pageNumber="4">C</emphasis>
,
<emphasis id="B938E539462C304BFE6E34C902B1F873" bold="true" box="[482,505,1994,2020]" pageId="3" pageNumber="4">G</emphasis>
,
<emphasis id="B938E539462C304BFD8434C90157F873" bold="true" box="[520,543,1994,2020]" pageId="3" pageNumber="4">K</emphasis>
. Lateral view.
<emphasis id="B938E539462C304BFD4134C901AAF873" bold="true" box="[717,738,1994,2020]" pageId="3" pageNumber="4">D</emphasis>
,
<emphasis id="B938E539462C304BFD7D34C90040F873" bold="true" box="[753,776,1994,2020]" pageId="3" pageNumber="4">H</emphasis>
,
<emphasis id="B938E539462C304BFC9B34C90063F873" bold="true" box="[791,811,1994,2020]" pageId="3" pageNumber="4">L</emphasis>
. Frontal view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BF3392B462B304CFF3131AE0022FC2A" blockId="4.[189,1399,685,958]" pageId="4" pageNumber="5">
PRONOTUM (
<figureCitation id="137725AE462B304CFECB31AE02E8FD50" box="[327,416,685,711]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2F</figureCitation>
). With anterolateral angles rounded, scarcely produced anteriorly, not swollen with regular border; anterior and hind margin without border; lateral margin with entire border. Prothoracic hypomeron with weak, broad, concave area along anterior half of prothoracic lateral margin; notosternal suture weakly visible, simple; prosternal process (
<figureCitation id="137725AE462B304CFD7230150010FCA7" box="[766,856,790,816]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2G</figureCitation>
) 0.55 times as broad as longest coxal diameter, its surface with lateral carinae complete, convergent and joined anteriorly forming a triangle; prosternum in front of coxa about 0.45 times as long as coxal longitudinal diameter at the same position; anterior margin continuing as weakly arcuate line, much more posterior than anterior pronotal margin; procoxal cavity distinctly transverse, without visible bordering line.
</paragraph>
<paragraph id="8BF3392B462B304CFF3130E6010CFAAC" blockId="4.[189,1399,997,1339]" pageId="4" pageNumber="5">
PTEROTHORAX. Anterior margin of mesoventrite with complete raised border (
<figureCitation id="137725AE462B304CFBE630E60785FC68" box="[1130,1229,997,1023]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2G</figureCitation>
); mesoventral process at median length of coxa 1.25–1.30 times as broad as corresponding coxal diameter; mesometaventrite articulation with suture visible; junction angulate posteriorly. Scutellar shield transverse, triangular; surface punctate and setose. Elytra with sides subparallel; with lateral margins very narrow but entirely visible from above; surface with single size punctures (
<figureCitation id="137725AE462B304CFC1A377200ACFB1B" box="[918,996,1137,1164]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2I</figureCitation>
), elytral epipleuron comparatively narrow, 1.5 times as wide as corresponding metaepisternum, incomplete apically (
<figureCitation id="137725AE462B304CFBFD37960786FB38" box="[1137,1230,1173,1199]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
), inner margin with border area narrow throughout and border line fading before base of elytron. Metaventrite with discrimen long but incomplete anteriorly; metaventral postcoxal lines distinctly separated at middle, laterally complete and recurved; metaepisternum with external process interlocking with fovea on elytron; metaepimeron indistinct.
</paragraph>
<paragraph id="8BF3392B462B304CFF313660065EFA37" blockId="4.[189,1398,1379,1440]" pageId="4" pageNumber="5">
LEGS. With trochanters angulately produced (
<figureCitation id="137725AE462B304CFD443660006CFAEA" box="[712,804,1379,1405]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
, G–H); tibiae without visible apical spurs; fore and mid tarsal claws appendiculate (
<figureCitation id="137725AE462B304CFDBA368501DBFA37" box="[566,659,1414,1440]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2C</figureCitation>
); metatarsal claws with quadrate basal tooth (
<figureCitation id="137725AE462B304CFB2636850641FA37" box="[1194,1289,1414,1440]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8BF3392B462B304CFF3136CB063EF9DC" blockId="4.[189,1399,1479,1612]" pageId="4" pageNumber="5">
ABDOMEN (
<figureCitation id="137725AE462B304CFEB336C402EAFA75" box="[319,418,1479,1506]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[248,1340,266,1821]" captionTargetId="figure@5.[248,1340,266,1821]" captionTargetPageId="5" captionText="Fig. 2. SEM images of Rhyzobius spp. (holotypes). A–I. R. albinos sp. nov., ♂ (MNHN). J. R. atramentarius sp. nov., ♂ (MNHN). A. Habitus, ventral. B. Antenna. C. Protarsal claws. D. Hind tarsal claws. E. Head, ventral. F. Pronotum. G. Pro-, meso- and metathorax, ventral. H. Abdomen, ventral. I–J. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="4" pageNumber="5">Figs 2H</figureCitation>
,
<figureCitation id="137725AE462B304CFE2336CB029BFA75" box="[431,467,1480,1506]" captionStart="Fig" captionStartId="9.[189,232,1923,1949]" captionTargetBox="[195,1381,399,1764]" captionTargetId="figure@9.[189,1398,370,1775]" captionTargetPageId="9" captionText="Fig. 4. Abdomens and male terminalia of Rhyzobius spp. A–C. R. albinos sp. nov. D–F. R. atramentarius sp. nov. G–I. R. serratus sp. nov. A, D, G. Abdomen. B, E, H. Abdominal segment VIII, ventral. C, F, I. Male genital segment, dorsal. Scale bars: A, D, G = 0.5 mm; B–C, E–F, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3973326" httpUri="https://zenodo.org/record/3973326/files/figure.png" pageId="4" pageNumber="5">4A</figureCitation>
). With five ventrites, ventrite 6 partially visible; ventrite 1- 1.6 times as long as ventrite 2; abdominal postcoxal lines separate medially, recurved and complete anteriorly, deep, posteriorly almost reaches ventrite 2; hind margin of ventrite 5 covered with long setae and deeply emarginate; hind margin of ventrite 6 weakly emarginate, tergite VIII widely truncate apically (
<figureCitation id="137725AE462B304CFA8135320621F9DC" box="[1293,1385,1585,1611]" captionStart="Fig" captionStartId="9.[189,232,1923,1949]" captionTargetBox="[195,1381,399,1764]" captionTargetId="figure@9.[189,1398,370,1775]" captionTargetPageId="9" captionText="Fig. 4. Abdomens and male terminalia of Rhyzobius spp. A–C. R. albinos sp. nov. D–F. R. atramentarius sp. nov. G–I. R. serratus sp. nov. A, D, G. Abdomen. B, E, H. Abdominal segment VIII, ventral. C, F, I. Male genital segment, dorsal. Scale bars: A, D, G = 0.5 mm; B–C, E–F, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3973326" httpUri="https://zenodo.org/record/3973326/files/figure.png" pageId="4" pageNumber="5">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="8BF3392B462B304CFF3135700105F8F7" blockId="4.[189,1399,1650,1888]" pageId="4" pageNumber="5">
MALE TERMINALIA AND GENITALIA (
<figureCitation id="137725AE462B304CFDCE357101E9F91A" box="[578,673,1650,1677]" captionStart="Fig" captionStartId="9.[189,232,1923,1949]" captionTargetBox="[195,1381,399,1764]" captionTargetId="figure@9.[189,1398,370,1775]" captionTargetPageId="9" captionText="Fig. 4. Abdomens and male terminalia of Rhyzobius spp. A–C. R. albinos sp. nov. D–F. R. atramentarius sp. nov. G–I. R. serratus sp. nov. A, D, G. Abdomen. B, E, H. Abdominal segment VIII, ventral. C, F, I. Male genital segment, dorsal. Scale bars: A, D, G = 0.5 mm; B–C, E–F, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3973326" httpUri="https://zenodo.org/record/3973326/files/figure.png" pageId="4" pageNumber="5">Figs 4C</figureCitation>
,
<figureCitation id="137725AE462B304CFD22357101F5F91A" box="[686,701,1650,1677]" captionStart="Fig" captionStartId="12.[189,232,1926,1952]" captionTargetBox="[266,1337,288,1867]" captionTargetId="figure@12.[242,1344,266,1887]" captionTargetPageId="12" captionText="Fig. 5. Male and female genitalia of Rhyzobius spp. A–D. R. albinos sp. nov. E–I. R. atramentarius sp. nov. J–M. R. serratus sp. nov. A, E, J. Penis, lateral. B, F, K. Apex of penis. C, G, L. Tegmen, inner view. D, H, M. Tegmen, lateral. I. Female genitalia. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3973332" httpUri="https://zenodo.org/record/3973332/files/figure.png" pageId="4" pageNumber="5">5</figureCitation>
A–D). Sternite IX with central part membranous (
<figureCitation id="137725AE462B304CFA8735710621F91A" box="[1291,1385,1650,1677]" captionStart="Fig" captionStartId="9.[189,232,1923,1949]" captionTargetBox="[195,1381,399,1764]" captionTargetId="figure@9.[189,1398,370,1775]" captionTargetPageId="9" captionText="Fig. 4. Abdomens and male terminalia of Rhyzobius spp. A–C. R. albinos sp. nov. D–F. R. atramentarius sp. nov. G–I. R. serratus sp. nov. A, D, G. Abdomen. B, E, H. Abdominal segment VIII, ventral. C, F, I. Male genital segment, dorsal. Scale bars: A, D, G = 0.5 mm; B–C, E–F, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3973326" httpUri="https://zenodo.org/record/3973326/files/figure.png" pageId="4" pageNumber="5">Fig. 4C</figureCitation>
); apodeme of male sternum IX with its apical half rod-like, base of spiculum broadly widened. Penis base with outer arm reduced, inner well developed; penis apex as in
<figureCitation id="137725AE462B304CFC5035BA0768F943" box="[988,1056,1721,1748]" captionStart="Fig" captionStartId="12.[189,232,1926,1952]" captionTargetBox="[266,1337,288,1867]" captionTargetId="figure@12.[242,1344,266,1887]" captionTargetPageId="12" captionText="Fig. 5. Male and female genitalia of Rhyzobius spp. A–D. R. albinos sp. nov. E–I. R. atramentarius sp. nov. J–M. R. serratus sp. nov. A, E, J. Penis, lateral. B, F, K. Apex of penis. C, G, L. Tegmen, inner view. D, H, M. Tegmen, lateral. I. Female genitalia. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3973332" httpUri="https://zenodo.org/record/3973332/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
A–B. Parameres (
<figureCitation id="137725AE462B304CFB7C35BA067CF943" box="[1264,1332,1721,1748]" captionStart="Fig" captionStartId="12.[189,232,1926,1952]" captionTargetBox="[266,1337,288,1867]" captionTargetId="figure@12.[242,1344,266,1887]" captionTargetPageId="12" captionText="Fig. 5. Male and female genitalia of Rhyzobius spp. A–D. R. albinos sp. nov. E–I. R. atramentarius sp. nov. J–M. R. serratus sp. nov. A, E, J. Penis, lateral. B, F, K. Apex of penis. C, G, L. Tegmen, inner view. D, H, M. Tegmen, lateral. I. Female genitalia. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3973332" httpUri="https://zenodo.org/record/3973332/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
C–D) articulated with phallobase, well developed, simple and separated, slightly longer than penis guide, with apices covered with long setae; penis guide simple, subtriangular in inner view, without additional processes, with acute apex and lateral sides symmetrical throughout and pointed apex; tegminal strut simple, slightly expanded at apex.
</paragraph>
<paragraph id="8BF3392B462B304CFF31348402E5F836" blockId="4.[189,429,1927,1954]" box="[189,429,1927,1954]" pageId="4" pageNumber="5">Female. Not known.</paragraph>
</subSubSection>
<caption id="DF3369A3462A304DFF313441013CF87E" ID-DOI="http://doi.org/10.5281/zenodo.3973322" ID-Zenodo-Dep="3973322" httpUri="https://zenodo.org/record/3973322/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1858,1884]" targetBox="[248,1340,266,1821]" targetPageId="5">
<paragraph id="8BF3392B462A304DFF313441013CF87E" blockId="5.[189,1399,1856,2025]" pageId="5" pageNumber="6">
<emphasis id="B938E539462A304DFF313441025EF8CB" bold="true" box="[189,278,1858,1884]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
SEM images of
<taxonomicName id="4C4C42A8462A304DFD86344101CBF8CB" authorityName="Stephens" authorityYear="1829" box="[522,643,1858,1884]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938E539462A304DFD86344101CBF8CB" box="[522,643,1858,1884]" italics="true" pageId="5" pageNumber="6">Rhyzobius</emphasis>
</taxonomicName>
spp. (holotypes).
<emphasis id="B938E539462A304DFCF5344100E0F8CB" bold="true" box="[889,936,1858,1884]" pageId="5" pageNumber="6">A–I</emphasis>
.
<taxonomicName id="4C4C42A8462A304DFC4F3440070DF8CB" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[963,1093,1858,1884]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="albinos" status="sp. nov.">
<emphasis id="B938E539462A304DFC4F3440009DF8CB" box="[963,981,1859,1884]" italics="true" pageId="5" pageNumber="6">R</emphasis>
.
<emphasis id="B938E539462A304DFC633441070DF8CB" box="[1007,1093,1858,1884]" italics="true" pageId="5" pageNumber="6">albinos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462A304DFBD534410789F8CB" box="[1113,1217,1858,1884]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (MNHN).
<emphasis id="B938E539462A304DFF3134660384F8E8" bold="true" box="[189,204,1893,1919]" pageId="5" pageNumber="6">J</emphasis>
.
<taxonomicName id="4C4C42A8462A304DFF57346502EAF8E8" authority="Czerwiński &amp; Szawaryn &amp; Tomaszewska, 2020" authorityName="Czerwiński &amp; Szawaryn &amp; Tomaszewska" authorityYear="2020" box="[219,418,1894,1920]" class="Insecta" family="Coccinellidae" genus="Rhyzobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="atramentarius" status="sp. nov.">
<emphasis id="B938E539462A304DFF57346503A5F8E8" box="[219,237,1894,1919]" italics="true" pageId="5" pageNumber="6">R</emphasis>
.
<emphasis id="B938E539462A304DFF70346502EAF8E8" box="[252,418,1894,1919]" italics="true" pageId="5" pageNumber="6">atramentarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5842462A304DFE2634650140F817" box="[426,520,1894,1920]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (MNHN).
<emphasis id="B938E539462A304DFD223466018BF8E8" bold="true" box="[686,707,1893,1919]" pageId="5" pageNumber="6">A</emphasis>
. Habitus, ventral.
<emphasis id="B938E539462A304DFC11346600F9F8E8" bold="true" box="[925,945,1893,1919]" pageId="5" pageNumber="6">B</emphasis>
. Antenna.
<emphasis id="B938E539462A304DFBBD3466070EF8E8" bold="true" box="[1073,1094,1893,1919]" pageId="5" pageNumber="6">C</emphasis>
. Protarsal claws.
<emphasis id="B938E539462A304DFA9B34660664F8E8" bold="true" box="[1303,1324,1893,1919]" pageId="5" pageNumber="6">D</emphasis>
. Hind tarsal claws.
<emphasis id="B938E539462A304DFED1348B0239F835" bold="true" box="[349,369,1928,1954]" pageId="5" pageNumber="6">E</emphasis>
. Head, ventral.
<emphasis id="B938E539462A304DFDB4348B0102F835" bold="true" box="[568,586,1928,1954]" pageId="5" pageNumber="6">F</emphasis>
. Pronotum.
<emphasis id="B938E539462A304DFD6E348B01B1F835" bold="true" box="[738,761,1928,1954]" pageId="5" pageNumber="6">G</emphasis>
. Pro-, meso- and metathorax, ventral.
<emphasis id="B938E539462A304DFB58348B07A3F835" bold="true" box="[1236,1259,1928,1954]" pageId="5" pageNumber="6">H</emphasis>
. Abdomen, ventral.
<emphasis id="B938E539462A304DFE9134AF020EF851" bold="true" box="[285,326,1964,1990]" pageId="5" pageNumber="6">I–J</emphasis>
. Details of elytral punctation. Scale bars: A = 1.00 mm; B = 0.2 mm; C–D = 0.1 mm; E–F = 0.3 mm; G = 0.4 mm; H–J = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C3566AA04629304EFF313208010DFEC4" box="[189,581,267,340]" pageId="6" pageNumber="7" type="distribution">
<paragraph id="8BF3392B4629304EFF313208021EFEB2" blockId="6.[189,342,267,293]" box="[189,342,267,293]" pageId="6" pageNumber="7">
<heading id="D0BB8E474629304EFF313208021EFEB2" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B938E5394629304EFF313208021EFEB2" bold="true" box="[189,342,267,293]" pageId="6" pageNumber="7">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3392B4629304EFF313239010DFEC4" blockId="6.[189,581,313,340]" box="[189,581,313,340]" pageId="6" pageNumber="7">
<collectingCountry id="F35B79BB4629304EFF31323902ECFEC4" box="[189,420,313,340]" name="Papua New Guinea" pageId="6" pageNumber="7">Papua New Guinea</collectingCountry>
: Mt Wilhelm.
</paragraph>
</subSubSection>
</treatment>
</document>
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