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<document id="A82729ADDF1912D833333A531E840EEF" ID-CLB-Dataset="22081" ID-DOI="10.5852/ejt.2018.490" ID-GBIF-Dataset="52356089-4633-4477-99c9-45f18606f5b9" ID-Zenodo-Dep="3830845" ID-ZooBank="BE023137-CC5E-4DC5-94F6-B549BB140361" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589637568005" checkinUser="valdenar" docAuthor="Koromyslova, Anna V., Taylor, Paul D., Martha, Silviu O. &amp; Riley, Matthew" docDate="2018" docId="03D587D79653FF86F326F906FD0DF91C" docLanguage="en" docName="ejt-490_koromyslova_taylor_martha_riley.pdf.imf" docOrigin="European Journal of Taxonomy 490" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhagasostoma tchvanovi" docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="FFECFFAF9676FFAEF12BFFD3FFE7FF87" masterDocTitle="Rhagasostoma (Bryozoa) from the Late Cretaceous of Eurasia: taxonomic revision, stratigraphy and palaeobiogeography" masterLastPageNumber="66" masterPageNumber="1" pageNumber="38" updateTime="1698840019227" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="508F798C74789E353B2F25B2BB086EDF">Rhagasostoma (Bryozoa) from the Late Cretaceous of Eurasia: taxonomic revision, stratigraphy and palaeobiogeography</mods:title>
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<mods:affiliation id="7F85E7F78912A479108E67C4BA006E74">Borissiak Paleontological Institute of the Russian Academy of Science, Profsoyuznaya st. 123, Moscow, 117997, Russian Federation. Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Senckenberg Forschungsinstitute und Naturmuseun, Sektion Marine Evertebraten III (Bryozoologie), Senckenberganlage 25, 60325 Frankfurt am Main, Germany. The Sedgwick Museum of Earth Sciences, Department of Earth Sciences, University of Cambridge, UK.</mods:affiliation>
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<subSubSection id="C366654A9653FF8BF326F906FBC1F977" box="[525,1062,1749,1776]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BC336C19653FF8BF326F906FBC1F977" blockId="37.[525,1062,1749,1812]" box="[525,1062,1749,1776]" pageId="37" pageNumber="38">
<heading id="D08B81AD9653FF8BF326F906FBC1F977" box="[525,1062,1749,1776]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="4C7C4D429653FF8BF326F906FBC1F977" authority="(Favorskaya, 1992)" baseAuthorityName="Favorskaya" baseAuthorityYear="1992" box="[525,1062,1749,1776]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="37" pageNumber="38" phylum="Bryozoa" rank="species" species="tchvanovi">
<emphasis id="B908EAD39653FF8BF326F906FCDCF977" bold="true" box="[525,827,1749,1776]" italics="true" pageId="37" pageNumber="38">Rhagasostoma tchvanovi</emphasis>
(
<bibRefCitation id="EFED4B309653FF8BF261F906FBF9F977" author="Favorskaya T. A." box="[842,1054,1749,1776]" pageId="37" pageNumber="38" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. &amp; Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya, 1992</bibRefCitation>
)
</taxonomicName>
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</paragraph>
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<subSubSection id="C366654A9653FF86F395F92AFAB1FDEE" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38" type="description">
<subSubSection id="C366654A9653FF8BF395F92AFC91F894" box="[702,886,1785,1812]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BC336C19653FF8BF395F92AFC91F894" blockId="37.[525,1062,1749,1812]" box="[702,886,1785,1812]" pageId="37" pageNumber="38">
<figureCitation id="13472A449653FF8BF395F92AFCF5F893" box="[702,786,1785,1812]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="37" pageNumber="38">Fig. 15</figureCitation>
,
<tableCitation id="C6FE037A9653FF8BF234F92AFC91F894" box="[799,886,1785,1811]" captionStart="Table 9" captionStartId="39.[189,259,268,294]" captionTargetBox="[510,1080,433,1876]" captionTargetId="graphics@39.[482,1105,620,1800]" captionTargetPageId="39" captionText="Table 9. Summary of measurements of Rhagasostoma tchvanovi (Favorskaya, 1992). For each parameter the range is given with the number of measurements in brackets. The arithmetic mean is given ± standard deviation. All measurements in µm." httpUri="http://table.plazi.org/id/DF0366499651FF89F196FEDFFCD5FEEB" pageId="37" pageNumber="38" tableUuid="DF0366499651FF89F196FEDFFCD5FEEB">Table 9</tableCitation>
</paragraph>
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<paragraph id="8BC336C19653FF8BF196F8E9FC7AF8D3" blockId="37.[189,1133,1850,1912]" box="[189,925,1850,1876]" pageId="37" pageNumber="38">
<taxonomicName id="4C7C4D429653FF8BF196F8E9FD3DF8D3" ID-CoL="86NWL" authority="Favorskaja, 1992: 126" authorityName="Favorskaja" authorityPageNumber="126" authorityYear="1992" box="[189,730,1850,1876]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="37" pageNumber="38" phylum="Bryozoa" rank="species" species="tchvanovi">
<emphasis id="B908EAD39653FF8BF196F8E9FE2CF8D3" box="[189,459,1850,1876]" italics="true" pageId="37" pageNumber="38">Onychocella tchvanovi</emphasis>
Favorskaja, 1992: 126
</taxonomicName>
, pl. 68, figs 68.
</paragraph>
<paragraph id="8BC336C19653FF8BF196F88DFB8AF8FF" blockId="37.[189,1133,1850,1912]" box="[189,1133,1885,1912]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="AB6C11EF9653FF8BF196F88DFB8AF8FF" box="[189,1133,1885,1912]" pageId="37" pageNumber="38">
?
<treatmentCitation id="0ADD10D09653FF8BF1E1F88EFC59F8FF" author="Brydone R. M." box="[202,958,1885,1912]" page="48" pageId="37" pageNumber="38" year="1930">
<taxonomicName id="4C7C4D429653FF8BF1E1F88EFC59F8FF" authority="Brydone, 1930: 48" authorityName="Brydone" authorityPageNumber="48" authorityYear="1930" box="[202,958,1885,1912]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="37" pageNumber="38" phylum="Bryozoa" rank="variety" species="gibbosum" variety="weybournensis">
<emphasis id="B908EAD39653FF8BF1E1F88EFE17F8F0" box="[202,496,1885,1911]" italics="true" pageId="37" pageNumber="38">Rhagasostoma gibbosum</emphasis>
var.
<emphasis id="B908EAD39653FF8BF301F88EFD3FF8F0" box="[554,728,1885,1911]" italics="true" pageId="37" pageNumber="38">weybournensis</emphasis>
<bibRefCitation id="EFED4B309653FF8BF3CBF88EFC59F8FF" author="Brydone R. M." box="[736,958,1885,1912]" pageId="37" pageNumber="38" refId="ref29975" refString="Brydone R. M. 1930. Further Notes on New or Imperfectly Known Chalk Polyzoa. Part II. (Vincularia, Onychocella, Rhagasostoma, Porina, etc.). Dulau and Co., London." type="book" year="1930">Brydone, 1930: 48</bibRefCitation>
</taxonomicName>
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, pl. 26, fig. 12.
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</paragraph>
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non
<taxonomicName id="4C7C4D429653FF8BF1DBF84CFD19F83E" authorityName="Brydone" authorityYear="1930" box="[240,766,1951,1977]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="37" pageNumber="38" phylum="Bryozoa" rank="variety" species="gibbosum" variety="weybournensis">
<emphasis id="B908EAD39653FF8BF1DBF84CFDF1F83E" box="[240,534,1951,1977]" italics="true" pageId="37" pageNumber="38">Rhagasostoma gibbosum</emphasis>
var.
<emphasis id="B908EAD39653FF8BF37BF84CFD19F83E" box="[592,766,1951,1977]" italics="true" pageId="37" pageNumber="38">weybournensis</emphasis>
</taxonomicName>
<treatmentCitation id="0ADD10D09653FF8BF237F84CFC14F83E" author="Brydone R. M." box="[796,1011,1951,1977]" page="48" pageId="37" pageNumber="38" year="1930">
<bibRefCitation id="EFED4B309653FF8BF237F84CFC14F83E" author="Brydone R. M." box="[796,1011,1951,1977]" pageId="37" pageNumber="38" refId="ref29975" refString="Brydone R. M. 1930. Further Notes on New or Imperfectly Known Chalk Polyzoa. Part II. (Vincularia, Onychocella, Rhagasostoma, Porina, etc.). Dulau and Co., London." type="book" year="1930">Brydone 1930: 48</bibRefCitation>
</treatmentCitation>
, pl. 26, figs 1314.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF0366499650FF88F196F91DFB11F867" ID-DOI="http://doi.org/10.5281/zenodo.3830877" ID-Zenodo-Dep="3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="38" pageNumber="39" startId="38.[189,232,1742,1768]" targetBox="[189,1398,265,1690]" targetPageId="38">
<paragraph id="8BC336C19650FF88F196F91DFB11F867" blockId="38.[189,1399,1742,2016]" pageId="38" pageNumber="39">
<emphasis id="B908EAD39650FF88F196F91DFEFDF96E" bold="true" box="[189,282,1742,1769]" pageId="38" pageNumber="39">Fig. 15.</emphasis>
<taxonomicName id="4C7C4D429650FF88F008F91CFCDCF96E" authority="(Favorskaya, 1992)" baseAuthorityName="Favorskaya" baseAuthorityYear="1992" box="[291,827,1743,1769]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="38" pageNumber="39" phylum="Bryozoa" rank="species" species="tchvanovi">
<emphasis id="B908EAD39650FF88F008F91CFDADF96E" box="[291,586,1743,1769]" italics="true" pageId="38" pageNumber="39">Rhagasostoma tchvanovi</emphasis>
(
<bibRefCitation id="EFED4B309650FF88F371F91CFCD5F96E" author="Favorskaya T. A." box="[602,818,1743,1769]" pageId="38" pageNumber="39" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. &amp; Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya, 1992</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B908EAD39650FF88F26CF91DFC99F96F" bold="true" box="[839,894,1742,1768]" pageId="38" pageNumber="39">AE</emphasis>
. Late Campanian (
<taxonomicName id="4C7C4D429650FF88F548F91CFF1AF88B" authority="Zone" authorityName="Zone" class="Globothalamea" family="Anomalinidae" genus="Cibicidoides" kingdom="Chromista" order="Rotaliida" pageId="38" pageNumber="39" phylum="Foraminifera" rank="species" species="voltzianus">
<emphasis id="B908EAD39650FF88F548F91CFA90F96E" box="[1123,1399,1743,1769]" italics="true" pageId="38" pageNumber="39">Cibicidoides voltzianus</emphasis>
Zone
</taxonomicName>
; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan.
<emphasis id="B908EAD39650FF88F59EF921FB09F88B" bold="true" box="[1205,1262,1778,1804]" pageId="38" pageNumber="39">AC</emphasis>
. Holotype, TsNIGR Museum 33/12582.
<emphasis id="B908EAD39650FF88F30DF8C6FDDCF8A8" bold="true" box="[550,571,1813,1839]" pageId="38" pageNumber="39">A</emphasis>
. Overview of erect bifoliate colony.
<emphasis id="B908EAD39650FF88F53AF8C6FBC2F8A8" bold="true" box="[1041,1061,1813,1839]" pageId="38" pageNumber="39">B</emphasis>
. Ovicellate autozooids and avicularia.
<emphasis id="B908EAD39650FF88F015F8EBFEB4F8D5" bold="true" box="[318,339,1848,1874]" pageId="38" pageNumber="39">C</emphasis>
. Non-ovicellate autozooids and avicularia.
<emphasis id="B908EAD39650FF88F260F8EBFC65F8D5" bold="true" box="[843,898,1848,1874]" pageId="38" pageNumber="39">DE</emphasis>
. TsNIGR Museum 34/12582.
<emphasis id="B908EAD39650FF88F5CBF8EBFB12F8D5" bold="true" box="[1248,1269,1848,1874]" pageId="38" pageNumber="39">D</emphasis>
. Overview erect bifoliate of colony.
<emphasis id="B908EAD39650FF88F0C2F88FFE1AF8F1" bold="true" box="[489,509,1884,1910]" pageId="38" pageNumber="39">E</emphasis>
. Non-ovicellate autozooids and avicularia.
<emphasis id="B908EAD39650FF88F528F88FFBF2F8F1" bold="true" box="[1027,1045,1884,1910]" pageId="38" pageNumber="39">F</emphasis>
. TsNIGR Museum 32/12582, middle Campanian (
<taxonomicName id="4C7C4D429650FF88F090F8ACFCA6F81E" authority="Zone" authorityName="Zone" box="[443,833,1919,1945]" class="Globothalamea" family="Cancrisidae" genus="Brotzenella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Rotaliida" pageId="38" pageNumber="39" phylum="Foraminifera" rank="species" species="monterelensis">
<emphasis id="B908EAD39650FF88F090F8ACFD15F81E" box="[443,754,1919,1945]" italics="true" pageId="38" pageNumber="39">Brotzenella monterelensis</emphasis>
Zone
</taxonomicName>
; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm.
</paragraph>
</caption>
<caption id="DF0366499651FF89F196FEDFFCD5FEEB" ID-Table-UUID="DF0366499651FF89F196FEDFFCD5FEEB" httpUri="http://table.plazi.org/id/DF0366499651FF89F196FEDFFCD5FEEB" pageId="39" pageNumber="40" targetBox="[510,1080,433,1876]" targetIsTable="true" targetPageId="39">
<paragraph id="8BC336C19651FF89F196FEDFFCD5FEEB" blockId="39.[189,1399,267,364]" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F196FEDFFECFFEA2" bold="true" box="[189,296,267,294]" pageId="39" pageNumber="40">Table 9.</emphasis>
Summary of measurements of
<taxonomicName id="4C7C4D429651FF89F3E8FED8FB12FEA2" authority="(Favorskaya, 1992)" baseAuthorityName="Favorskaya" baseAuthorityYear="1992" box="[707,1269,267,293]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="39" pageNumber="40" phylum="Bryozoa" rank="species" species="tchvanovi">
<emphasis id="B908EAD39651FF89F3E8FED8FC14FEA2" box="[707,1011,267,293]" italics="true" pageId="39" pageNumber="40">Rhagasostoma tchvanovi</emphasis>
(
<bibRefCitation id="EFED4B309651FF89F520FED8FB0BFEA2" author="Favorskaya T. A." box="[1035,1260,267,293]" pageId="39" pageNumber="40" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. &amp; Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya, 1992</bibRefCitation>
)
</taxonomicName>
. For each parameter the range is given with the number of measurements in brackets. The arithmetic mean is given ± standard deviation. All measurements in µm.
</paragraph>
</caption>
<paragraph id="8BC336C19651FF89F310FE62FC32F8D4" pageId="39" pageNumber="40">
<table id="F97CC46196510051F0D5FE62FBDFF8D3" box="[510,1080,433,1876]" gridcols="3" gridrows="11" pageId="39" pageNumber="40">
<tr id="354C348396510051F0D5FE62FBDFFDDF" box="[510,1080,433,600]" gridrow="0" pageId="39" pageNumber="40">
<th id="769D5DFF96510051F0D5FE62FD42FDDF" box="[510,677,433,600]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F310FE62FD9CFE4E" bold="true" box="[571,635,433,457]" pageId="39" pageNumber="40">Basin</emphasis>
<emphasis id="B908EAD39651FF89F339FE24FD42FD88" bold="true" box="[530,677,503,527]" pageId="39" pageNumber="40">Stratigraphy</emphasis>
<emphasis id="B908EAD39651FF89F306FD93FD6DFDDF" bold="true" box="[557,650,576,600]" pageId="39" pageNumber="40">Locality</emphasis>
</th>
<th id="769D5DFF96510051F3C5FE62FBDFFDDF" box="[750,1080,433,600]" gridcol="2" gridrow="0" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F22CFE61FBF9FE4F" bold="true" box="[775,1054,434,456]" pageId="39" pageNumber="40">North Ust-YurtAral Basin</emphasis>
<emphasis id="B908EAD39651FF89F3C5FE2BFBDFFD89" bold="true" box="[750,1080,504,526]" pageId="39" pageNumber="40">Early to middle late Campanian</emphasis>
<emphasis id="B908EAD39651FF89F225FD93FBFFFDD1" bold="true" box="[782,1048,576,598]" pageId="39" pageNumber="40">Southern Aral Sea Region</emphasis>
</th>
</tr>
<tr id="354C348396510051F0D5FD55FBDFFD3E" box="[510,1080,646,697]" gridrow="1" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348FD55FD42FD3E" box="[611,677,646,697]" gridcol="1" gridrow="1" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F341FD40FD7CFD2C" bold="true" box="[618,667,659,683]" pageId="39" pageNumber="40">AzL</emphasis>
</td>
<td id="769D5DFF96510051F3C5FD55FBDFFD3E" box="[750,1080,646,697]" gridcol="2" gridrow="1" pageId="39" pageNumber="40">340550 (18) 418.89 ± 50.98</td>
</tr>
<tr id="354C348396510051F0D5FD32FBDFFC94" box="[510,1080,737,787]" gridrow="2" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348FD32FD42FC94" box="[611,677,737,787]" gridcol="1" gridrow="2" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F34EFD3DFD78FC81" bold="true" box="[613,671,750,774]" pageId="39" pageNumber="40">AzW</emphasis>
</td>
<td id="769D5DFF96510051F3C5FD32FBDFFC94" box="[750,1080,737,787]" gridcol="2" gridrow="2" pageId="39" pageNumber="40">230360 (18) 264.44 ± 22.29</td>
</tr>
<tr id="354C348396510051F0D5FCEFFBDFFBA3" box="[510,1080,828,1060]" gridrow="3" pageId="39" pageNumber="40">
<th id="769D5DFF96510051F0D5FCEFFDFAFBA3" box="[510,541,828,1060]" gridcol="0" gridrow="3" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F0D5FC8DFDFAFC70" bold="true" box="[510,541,862,1015]" pageId="39" pageNumber="40">Autozooids</emphasis>
</th>
<td id="769D5DFF96510051F348FCEFFD42FBA3" box="[611,677,828,1060]" gridcol="1" gridrow="3" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F342FC9AFD7CFCE6" bold="true" box="[617,667,841,865]" pageId="39" pageNumber="40">CvL</emphasis>
<emphasis id="B908EAD39651FF89F343FC70FD7AFC3C" bold="true" box="[616,669,931,955]" pageId="39" pageNumber="40">OpL</emphasis>
<emphasis id="B908EAD39651FF89F348FC2DFD46FB91" bold="true" box="[611,673,1022,1046]" pageId="39" pageNumber="40">OpW</emphasis>
</td>
<td id="769D5DFF96510051F3C5FCEFFBDFFBA3" box="[750,1080,828,1060]" gridcol="2" gridrow="3" pageId="39" pageNumber="40">70100 (4) 80 ± 14.14 110140 (14) 127.86 ± 9.75 110140 (14) 120.00 ± 2.96</td>
</tr>
<tr id="354C348396510051F0D5FB98FBDFFBF9" box="[510,1080,1099,1150]" gridrow="4" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348FB98FD42FBF9" box="[611,677,1099,1150]" gridcol="1" gridrow="4" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F343FB8AFD7BFBF6" bold="true" box="[616,668,1113,1137]" pageId="39" pageNumber="40">OoL</emphasis>
</td>
<td id="769D5DFF96510051F3C5FB98FBDFFBF9" box="[750,1080,1099,1150]" gridcol="2" gridrow="4" pageId="39" pageNumber="40">140230 (5) 198.00 ± 35.64</td>
</tr>
<tr id="354C348396510051F0D5FB75FBDFFB5F" box="[510,1080,1190,1240]" gridrow="5" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348FB75FD42FB5F" box="[611,677,1190,1240]" gridcol="1" gridrow="5" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F348FB67FD47FB4B" bold="true" box="[611,672,1204,1228]" pageId="39" pageNumber="40">OoW</emphasis>
</td>
<td id="769D5DFF96510051F3C5FB75FBDFFB5F" box="[750,1080,1190,1240]" gridcol="2" gridrow="5" pageId="39" pageNumber="40">180230 (5) 210.00 ± 18.71</td>
</tr>
<tr id="354C348396510051F0D5FAD2FBDFFAB3" box="[510,1080,1281,1332]" gridrow="6" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348FAD2FD42FAB3" box="[611,677,1281,1332]" gridcol="1" gridrow="6" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F341FADDFD7FFAA1" bold="true" box="[618,664,1294,1318]" pageId="39" pageNumber="40">AvL</emphasis>
</td>
<td id="769D5DFF96510051F3C5FAD2FBDFFAB3" box="[750,1080,1281,1332]" gridcol="2" gridrow="6" pageId="39" pageNumber="40">290530 (11) 381.82 ± 71.25</td>
</tr>
<tr id="354C348396510051F0D5FA88FBDFFA09" box="[510,1080,1371,1422]" gridrow="7" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348FA88FD42FA09" box="[611,677,1371,1422]" gridcol="1" gridrow="7" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F35BFABAFD72FA06" bold="true" box="[624,661,1385,1409]" pageId="39" pageNumber="40">RL</emphasis>
</td>
<td id="769D5DFF96510051F3C5FA88FBDFFA09" box="[750,1080,1371,1422]" gridcol="2" gridrow="7" pageId="39" pageNumber="40">160350 (11) 254.55 ± 58.20</td>
</tr>
<tr id="354C348396510051F0D5FA65FBDFF919" box="[510,1080,1462,1694]" gridrow="8" pageId="39" pageNumber="40">
<th id="769D5DFF96510051F0D5FA65FDFAF919" box="[510,541,1462,1694]" gridcol="0" gridrow="8" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F0D5FA0CFDFAF9EA" bold="true" box="[510,541,1503,1645]" pageId="39" pageNumber="40">Avicularia</emphasis>
</th>
<td id="769D5DFF96510051F348FA65FD42F919" box="[611,677,1462,1694]" gridcol="1" gridrow="8" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F340FA17FD70FA5B" bold="true" box="[619,663,1476,1500]" pageId="39" pageNumber="40">RW</emphasis>
<emphasis id="B908EAD39651FF89F340F9CCFD7EF9B0" bold="true" box="[619,665,1567,1591]" pageId="39" pageNumber="40">PrL</emphasis>
<emphasis id="B908EAD39651FF89F34DF9AAFD7AF916" bold="true" box="[614,669,1657,1681]" pageId="39" pageNumber="40">PrW</emphasis>
</td>
<td id="769D5DFF96510051F3C5FA65FBDFF919" box="[750,1080,1462,1694]" gridcol="2" gridrow="8" pageId="39" pageNumber="40">80120 (11) 93.64 ± 13.62 110190 (10) 137.00 ± 25.84 70120 (11) 94.55 ± 17.53</td>
</tr>
<tr id="354C348396510051F0D5F914FBDFF97E" box="[510,1080,1735,1785]" gridrow="9" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348F914FD42F97E" box="[611,677,1735,1785]" gridcol="1" gridrow="9" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F343F907FD7AF96B" bold="true" box="[616,669,1748,1772]" pageId="39" pageNumber="40">OpL</emphasis>
</td>
<td id="769D5DFF96510051F3C5F914FBDFF97E" box="[750,1080,1735,1785]" gridcol="2" gridrow="9" pageId="39" pageNumber="40">70100 (8) 77.50 ± 10.35</td>
</tr>
<tr id="354C348396510051F0D5F8F2FBDFF8D3" box="[510,1080,1825,1876]" gridrow="10" pageId="39" pageNumber="40" rowspan-0="1">
<td id="769D5DFF96510051F348F8F2FD42F8D3" box="[611,677,1825,1876]" gridcol="1" gridrow="10" pageId="39" pageNumber="40">
<emphasis id="B908EAD39651FF89F348F8FCFD46F8C0" bold="true" box="[611,673,1839,1863]" pageId="39" pageNumber="40">OpW</emphasis>
</td>
<subSubSection id="C366654A9651FF89F24FF8F2FC32F8D4" pageId="39" pageNumber="40" type="description">
<td id="769D5DFF96510051F3C5F8F2FBDFF8D3" box="[750,1080,1825,1876]" gridcol="2" gridrow="10" pageId="39" pageNumber="40">3070 (8) 53.75 ± 11.88</td>
</subSubSection>
</tr>
</table>
</paragraph>
<subSubSection id="C366654A965EFF86F196FED8FAB1FDEE" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8BC336C1965EFF86F196FED8FE6BFEA2" blockId="40.[189,396,267,293]" box="[189,396,267,293]" pageId="40" pageNumber="41">
<heading id="D08B81AD965EFF86F196FED8FE6BFEA2" bold="true" box="[189,396,267,293]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B908EAD3965EFF86F196FED8FE6BFEA2" bold="true" box="[189,396,267,293]" pageId="40" pageNumber="41">Material studied</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B143C9C965EFF86F1C1FEE9FEBCFE39" ID-GBIF-Occurrence="2620322310" collectionCode="LS, TsNIGR" country="Uzbekistan" county="southern Aral Sea Region" location="Republic of Karakalpakstan" municipality="Campanian" pageId="40" pageNumber="41" specimenCount="1" typeStatus="holotype">
<paragraph id="8BC336C1965EFF86F1C1FEE9FEBBFED3" blockId="40.[189,1399,314,446]" box="[234,348,314,340]" pageId="40" pageNumber="41">
<emphasis id="B908EAD3965EFF86F1C1FEE9FEBBFED3" bold="true" box="[234,348,314,340]" pageId="40" pageNumber="41">
<typeStatus id="54C78863965EFF86F1C1FEE9FEBBFED3" box="[234,348,314,340]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC336C1965EFF86F196FE8EFEBCFE39" blockId="40.[189,1399,314,446]" pageId="40" pageNumber="41">
<collectingCountry id="F36B7651965EFF86F196FE8EFE94FEF0" box="[189,371,349,375]" name="Uzbekistan" pageId="40" pageNumber="41">UZBEKISTAN</collectingCountry>
<location id="8EA3601A965EFF86F0A0FE8EFD32FEF0" LSID="urn:lsid:plazi:treatment:03D587D79653FF86F326F906FD0DF91C:8EA3601A965EFF86F0A0FE8EFD32FEF0" box="[395,725,349,375]" country="Uzbekistan" county="southern Aral Sea Region" municipality="Campanian" name="Republic of Karakalpakstan" pageId="40" pageNumber="41">Republic of Karakalpakstan</location>
; late
<collectingMunicipality id="6BA7ACBB965EFF86F239FE8EFC7FFEF0" box="[786,920,349,375]" pageId="40" pageNumber="41">Campanian</collectingMunicipality>
(
<taxonomicName id="4C7C4D42965EFF86F282FE8EFAE5FEF0" authority="Zone" authorityName="Zone" box="[937,1282,349,375]" class="Globothalamea" family="Anomalinidae" genus="Cibicidoides" kingdom="Chromista" order="Rotaliida" pageId="40" pageNumber="41" phylum="Foraminifera" rank="species" species="voltzianus">
<emphasis id="B908EAD3965EFF86F282FE8EFB5CFEF0" box="[937,1211,349,375]" italics="true" pageId="40" pageNumber="41">Cibicidoides voltzianus</emphasis>
Zone
</taxonomicName>
;
<collectionCode id="ED6DAE04965EFF86F425FE8FFAD4FEF0" box="[1294,1331,348,375]" collectionName="United Kingdom, London, Linnean Society" pageId="40" pageNumber="41">LS</collectionCode>
15) of the
<collectingCounty id="62A24E4D965EFF86F1C2FE53FDF9FE1D" box="[233,542,384,410]" pageId="40" pageNumber="41">southern Aral Sea Region</collectingCounty>
;
<collectionCode id="ED6DAE04965EFF86F301FE53FD6AFE1D" box="[554,653,384,410]" pageId="40" pageNumber="41">TsNIGR</collectionCode>
Museum 33/12582 (
<figureCitation id="13472A44965EFF86F2AEFE53FC3FFE1C" box="[901,984,384,411]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="40" pageNumber="41">Fig. 15</figureCitation>
AC; figured by
<bibRefCitation id="EFED4B30965EFF86F58FFE53FA94FE1D" author="Favorskaya T. A." box="[1188,1395,384,410]" pageId="40" pageNumber="41" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. &amp; Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya 1992</bibRefCitation>
: pl. 68, fig. 6).
</paragraph>
</materialsCitation>
<paragraph id="8BC336C1965EFF86F1C1FE37FDA5FE79" blockId="40.[189,1398,484,617]" box="[234,578,484,510]" pageId="40" pageNumber="41">
<emphasis id="B908EAD3965EFF86F1C1FE37FDA5FE79" bold="true" box="[234,578,484,510]" pageId="40" pageNumber="41">Additional figured material</emphasis>
</paragraph>
<paragraph id="8BC336C1965EFF86F196FDDBFAB1FDEE" blockId="40.[189,1398,484,617]" pageId="40" pageNumber="41">
<materialsCitation id="3B143C9C965EFF86F196FDDBFDB0FDC2" ID-GBIF-Occurrence="2620322332" collectionCode="TsNIGR" country="Uzbekistan" county="southern Aral Sea Region" location="Same" municipality="Campanian" pageId="40" pageNumber="41" specimenCount="1">
<collectingCountry id="F36B7651965EFF86F196FDDBFE94FDA5" box="[189,371,520,546]" name="Uzbekistan" pageId="40" pageNumber="41">UZBEKISTAN</collectingCountry>
<location id="8EA3601A965EFF86F0BDFDDBFE30FDA5" LSID="urn:lsid:plazi:treatment:03D587D79653FF86F326F906FD0DF91C:8EA3601A965EFF86F0BDFDDBFE30FDA5" box="[406,471,520,546]" country="Uzbekistan" name="Same" pageId="40" pageNumber="41">Same</location>
data as for holotype;
<collectionCode id="ED6DAE04965EFF86F3C4FDDBFCB5FDA5" box="[751,850,520,546]" pageId="40" pageNumber="41">TsNIGR</collectionCode>
Museum 34/12582 (
<figureCitation id="13472A44965EFF86F57DFDDBFB4AFDA5" box="[1110,1197,519,546]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="40" pageNumber="41">Fig. 15</figureCitation>
DE; figured by
<bibRefCitation id="EFED4B30965EFF86F196FDF8FE74FDC2" author="Favorskaya T. A." box="[189,403,555,581]" pageId="40" pageNumber="41" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. &amp; Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya 1992</bibRefCitation>
: pl. 68, fig. 7)
</materialsCitation>
<materialsCitation id="3B143C9C965EFF86F3ABFDF8FAB5FDEE" ID-GBIF-Occurrence="2620322318" collectionCode="TsNIGR" country="Uzbekistan" county="southern Aral Sea Region" location="Republic of Karakalpakstan" municipality="Campanian" pageId="40" pageNumber="41" specimenCount="1">
<location id="8EA3601A965EFF86F3ABFDF8FC3BFDC2" LSID="urn:lsid:plazi:treatment:03D587D79653FF86F326F906FD0DF91C:8EA3601A965EFF86F3ABFDF8FC3BFDC2" box="[640,988,555,581]" country="Uzbekistan" county="southern Aral Sea Region" municipality="Campanian" name="Republic of Karakalpakstan" pageId="40" pageNumber="41">Republic of Karakalpakstan</location>
; middle
<collectingMunicipality id="6BA7ACBB965EFF86F57AFDF8FB30FDC2" box="[1105,1239,555,581]" pageId="40" pageNumber="41">Campanian</collectingMunicipality>
(
<taxonomicName id="4C7C4D42965EFF86F5DBFDF8FE41FDEE" authority="Zone" authorityName="Zone" class="Globothalamea" family="Cancrisidae" genus="Brotzenella" higherTaxonomySource="GBIF" kingdom="Chromista" order="Rotaliida" pageId="40" pageNumber="41" phylum="Foraminifera" rank="species" species="monterelensis">
<emphasis id="B908EAD3965EFF86F5DBFDF8FEB8FDEF" italics="true" pageId="40" pageNumber="41">Brotzenella monterelensis</emphasis>
Zone
</taxonomicName>
; LS14) of the
<collectingCounty id="62A24E4D965EFF86F365FD9DFC99FDEF" box="[590,894,590,616]" pageId="40" pageNumber="41">southern Aral Sea Region</collectingCounty>
;
<collectionCode id="ED6DAE04965EFF86F2A2FD9DFC0BFDEF" box="[905,1004,590,616]" pageId="40" pageNumber="41">TsNIGR</collectionCode>
Museum 32/12582 (
<figureCitation id="13472A44965EFF86F5CBFD9DFAAFFDEE" box="[1248,1352,590,617]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="40" pageNumber="41">Fig. 15F</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</subSubSection>
<subSubSection id="C366654A965EFF86F196FD5CFCF1FB6F" pageId="40" pageNumber="41" type="description">
<paragraph id="8BC336C1965EFF86F196FD5CFEA9FD2E" blockId="40.[189,334,655,681]" box="[189,334,655,681]" pageId="40" pageNumber="41">
<heading id="D08B81AD965EFF86F196FD5CFEA9FD2E" bold="true" box="[189,334,655,681]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B908EAD3965EFF86F196FD5CFEA9FD2E" bold="true" box="[189,334,655,681]" pageId="40" pageNumber="41">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC336C1965EFF86F196FD6DFCF1FB6F" blockId="40.[189,1399,702,1257]" pageId="40" pageNumber="41">
Colony erect with flattened bifoliate branches (
<quantity id="4C849B24965EFF86F22DFD6DFC89FD5E" box="[774,878,702,729]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="40" pageNumber="41" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
wide). Ancestrula and early astogeny not observed. Autozooids variable in shape, often broad and subrectangular with rounded distal ends; zooidal boundaries raised. Gymnocyst lacking. Cryptocyst extensive, finely pustulose, slightly depressed centrally, sometimes with proximolateral peripheral caverns (
<figureCitation id="13472A44965EFF86F2A7FCFBFC14FCC5" box="[908,1011,807,834]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="40" pageNumber="41">Fig. 15F</figureCitation>
). Opesia terminal, semielliptical, formed by two walls, a thin inner wall forms the distolateral part of the opesia, and a projecting thickened outer wall forms the proximolateral part (
<figureCitation id="13472A44965EFF86F3E8FCBDFCBDFC0E" box="[707,858,878,905]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="40" pageNumber="41">Fig. 15C, E</figureCitation>
). Proximal edge of opesia straight, smooth, with outgrowths at the two proximolateral corners which delineate small opesiules; outgrowths often destroyed giving the opesiules the appearance of opesiular indentations. Septula not observed. Ovicells endozooidal, ooecium is formed by the distal zooid, well-recognizable, with cryptocyst-like surface and arch-like proximal edge with elongated proximolateral processes extending along the cryptocyst of the maternal zooid (
<figureCitation id="13472A44965EFF86F086FBCDFDE5FBBE" box="[429,514,1054,1081]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="40" pageNumber="41">Fig. 15</figureCitation>
AB). Avicularia interzooidal, smaller than autozooids, elongate. Rostrum channeled with elevated wing-like walls and pointed tip, narrow, almost straight, asymmetrical, dextral or sinistral. Top of rostrum reaches opesia of distal autozooid. Wedge-like structures not observed. Proximal part rounded, shorter and wider than rostrum, cryptocyst pustulose, concave. Opesia round with short slit proximally, opesiules lacking (
<figureCitation id="13472A44965EFF86F3E9FB78FCCCFB41" box="[706,811,1195,1222]" captionStart="Fig" captionStartId="38.[189,232,1742,1768]" captionTargetBox="[189,1398,265,1690]" captionTargetId="figure@38.[189,1399,265,1692]" captionTargetPageId="38" captionText="Fig. 15. Rhagasostoma tchvanovi (Favorskaya, 1992). AE. Late Campanian (Cibicidoides voltzianus Zone; LS15), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan. AC. Holotype, TsNIGR Museum 33/12582. A. Overview of erect bifoliate colony. B. Ovicellate autozooids and avicularia. C. Non-ovicellate autozooids and avicularia. DE. TsNIGR Museum 34/12582. D. Overview erect bifoliate of colony. E. Non-ovicellate autozooids and avicularia. F. TsNIGR Museum 32/12582, middle Campanian (Brotzenella monterelensis Zone; LS14), southern Aral Sea Region, Republic of Karakalpakstan, Uzbekistan; overview of erect bifoliate colony, cryptocyst of autozooids with proximolateral caverns. Scale bars: A = 500 µm; B, E = 200 µm; С = 250 µm; D, F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3830877" httpUri="https://zenodo.org/record/3830877/files/figure.png" pageId="40" pageNumber="41">Fig. 15C</figureCitation>
). Closure plates, kenozooids, intramural reparative budding of autozooids and avicularia not observed.
</paragraph>
</subSubSection>
<subSubSection id="C366654A965EFF86F196FAC3FA91F98E" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BC336C1965EFF86F196FAC3FEC8FAAD" blockId="40.[189,303,1296,1322]" box="[189,303,1296,1322]" pageId="40" pageNumber="41">
<heading id="D08B81AD965EFF86F196FAC3FEC8FAAD" bold="true" box="[189,303,1296,1322]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B908EAD3965EFF86F196FAC3FEC8FAAD" bold="true" box="[189,303,1296,1322]" pageId="40" pageNumber="41">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC336C1965EFF86F196FAEDFA91F98E" blockId="40.[189,1398,1342,1545]" pageId="40" pageNumber="41">
<taxonomicName id="4C7C4D42965EFF86F196FAEDFE05FADF" baseAuthorityName="Favorskaya" baseAuthorityYear="1992" box="[189,482,1342,1368]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="species" species="tchvanovi">
<emphasis id="B908EAD3965EFF86F196FAEDFE05FADF" box="[189,482,1342,1368]" italics="true" pageId="40" pageNumber="41">Rhagasostoma tchvanovi</emphasis>
</taxonomicName>
differs from the closely related species
<taxonomicName id="4C7C4D42965EFF86F299FAECFBA4FADF" baseAuthorityName="Marsson" baseAuthorityYear="1887" box="[946,1091,1342,1368]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="species" species="gibbosum">
<emphasis id="B908EAD3965EFF86F299FAECFBA4FADF" box="[946,1091,1342,1368]" italics="true" pageId="40" pageNumber="41">R. gibbosum</emphasis>
</taxonomicName>
in the avicularian rostrum being long and almost straight instead of short and tilted to the right or left. The specimen described by
<bibRefCitation id="EFED4B30965EFF86F1C8FA57FE7DFA18" author="Brydone R. M." box="[227,410,1412,1439]" pageId="40" pageNumber="41" refId="ref29975" refString="Brydone R. M. 1930. Further Notes on New or Imperfectly Known Chalk Polyzoa. Part II. (Vincularia, Onychocella, Rhagasostoma, Porina, etc.). Dulau and Co., London." type="book" year="1930">Brydone (1930</bibRefCitation>
: pl. 26, fig. 12) as
<taxonomicName id="4C7C4D42965EFF86F357FA57FBA5FA19" box="[636,1090,1412,1438]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="subSpecies" species="gibbosum" subSpecies="weybournensis">
<emphasis id="B908EAD3965EFF86F357FA57FBA5FA19" box="[636,1090,1412,1438]" italics="true" pageId="40" pageNumber="41">Onychocella gibbosum weybournensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C7C4D42965EFF86F5C9FA56FE98FA45" authority="(Marsson, 1887)" baseAuthorityName="Marsson" baseAuthorityYear="1887" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="species" species="gibbosum">
<emphasis id="B908EAD3965EFF86F5C9FA56FA91FA19" box="[1250,1398,1412,1438]" italics="true" pageId="40" pageNumber="41">R. gibbosum</emphasis>
(
<bibRefCitation id="EFED4B30965EFF86F1EEFA7BFE91FA45" author="Marsson T." box="[197,374,1448,1474]" pageId="40" pageNumber="41" pagination="1 - 112" refId="ref31259" refString="Marsson T. 1887. Die Bryozoen der weissen Schreibkreide der Insel Rugen. Palaeontologische Abhandlungen 4: 1 - 112." type="journal article" year="1887">Marsson, 1887</bibRefCitation>
)
</taxonomicName>
in having a long and straight avicularian rostrum and is conspecific with
<taxonomicName id="4C7C4D42965EFF86F5F7FA7BFA88FA45" baseAuthorityName="Favorskaya" baseAuthorityYear="1992" box="[1244,1391,1448,1474]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="species" species="tchvanovi">
<emphasis id="B908EAD3965EFF86F5F7FA7BFA88FA45" box="[1244,1391,1448,1474]" italics="true" pageId="40" pageNumber="41">R. tchvanovi</emphasis>
</taxonomicName>
. Specimen 32/
<quantity id="4C849B24965EFF86F077FA18FE26FA62" box="[348,449,1483,1510]" metricMagnitude="2" metricUnit="m" metricValue="3.195828" pageId="40" pageNumber="41" unit="in" value="12582.0">12582 in</quantity>
the collections of the TsNIGR Museum and labelled as “
<taxonomicName id="4C7C4D42965EFF86F562FA18FF1DF98F" authority="Brydone, 1930" authorityName="Brydone" authorityYear="1930" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="species" species="subgibbosum">
<emphasis id="B908EAD3965EFF86F562FA18FAE2FA62" box="[1097,1285,1483,1509]" italics="true" pageId="40" pageNumber="41">O. subgibbosum</emphasis>
<bibRefCitation id="EFED4B30965EFF86F421FA18FF1DF98F" author="Brydone R. M." pageId="40" pageNumber="41" refId="ref29975" refString="Brydone R. M. 1930. Further Notes on New or Imperfectly Known Chalk Polyzoa. Part II. (Vincularia, Onychocella, Rhagasostoma, Porina, etc.). Dulau and Co., London." type="book" year="1930">Brydone, 1930</bibRefCitation>
</taxonomicName>
” is not the one that was depicted by
<bibRefCitation id="EFED4B30965EFF86F383FA3DFC67F98F" author="Favorskaya T. A." box="[680,896,1518,1544]" pageId="40" pageNumber="41" pagination="115 - 136" refId="ref30441" refString="Favorskaya T. A. 1992. Мsaнki kaмpaнa i мaaсtрikta Юga СССР [Campanian and Maastrichtian bryozoans of the southern USSR]. In: Okuneva T. M., Titova M. V., Favorskaya T. A., Zonova T. D. &amp; Rostovtsev K. O. (eds) Atlas rukovodyashchikh grupp fauny mesozoya Yuga i Vostoka SSSR. Trudy Vserossiyskogo Nauchno-issledovatelskogo Geologicheskogo Instituta, new series 350: 115 - 136. Nedra, St Petersburg." type="book chapter" year="1992">Favorskaya (1992</bibRefCitation>
: pl. 67, fig. 3) and belongs to
<taxonomicName id="4C7C4D42965EFF86F5F7FA3CFA88F98F" baseAuthorityName="Favorskaya" baseAuthorityYear="1992" box="[1244,1391,1518,1544]" class="Gymnolaemata" family="Onychocellidae" genus="Rhagasostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="species" species="tchvanovi">
<emphasis id="B908EAD3965EFF86F5F7FA3CFA88F98F" box="[1244,1391,1518,1544]" italics="true" pageId="40" pageNumber="41">R. tchvanovi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C366654A965EFF86F196F9FCFD0DF91C" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8BC336C1965EFF86F196F9FCFEB1F9CE" blockId="40.[189,342,1583,1609]" box="[189,342,1583,1609]" pageId="40" pageNumber="41">
<heading id="D08B81AD965EFF86F196F9FCFEB1F9CE" bold="true" box="[189,342,1583,1609]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B908EAD3965EFF86F196F9FCFEB1F9CE" bold="true" box="[189,342,1583,1609]" pageId="40" pageNumber="41">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC336C1965EFF86F196F98DFD0DF91C" blockId="40.[189,1399,1630,1691]" pageId="40" pageNumber="41">
Middle to late Campanian Republic of Karakalpakstan,
<collectingCountry id="F36B7651965EFF86F274F98DFC0FF9FF" box="[863,1000,1630,1656]" name="Uzbekistan" pageId="40" pageNumber="41">Uzbekistan</collectingCountry>
: six localities between Chimboy/ Shımbay (Чимбой/Шымбай) and the Aral Sea.
</paragraph>
</subSubSection>
</treatment>
</document>