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<document id="37090113D35E812698498AB3301D6F08" ID-CLB-Dataset="300858" ID-DOI="10.37828/em.2017.10.4" ID-GBIF-Dataset="a36708a3-238e-491e-8a45-3589eb668f04" ID-ISSN="2336-9744" ID-Zenodo-Dep="13236524" ID-ZooBank="urn:lsid:zoobank.org:pub:30F90165-9296-4852-93EE-0F40796F090A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1720661961264" checkinUser="felipe" docAuthor="Tuzovskij, Petr V." docDate="2017" docId="03D5878DFFC4FFA5FF1EFA3AFCC97A3C" docLanguage="en" docName="EcolMontenegrina.10.22-25.pdf" docOrigin="Ecologica Montenegrina 10" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Hygrobates (Rivobates) primoryensis Tuzovskij, 2017, sp.n." docType="treatment" docUuid="E8A898C7-8F7A-47E7-A7CF-EB423C1336B8" docUuidSource="ZooBank" docVersion="4" lastPageNumber="25" masterDocId="FFECFFF5FFC4FFA6FF89FFCFFFE87F6C" masterDocTitle="A new water mite species of the genus Hygrobates Koch, 1837 (Acari, Hydrachnidia, Hygrobatidae) from Far East of Russia" masterLastPageNumber="25" masterPageNumber="22" pageNumber="22" updateTime="1722948791579" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="4BD8464B794BF54A44D60B073D09B9BF">
<mods:title id="3442B78084D1110E4D3EDC818FD2E48B">A new water mite species of the genus Hygrobates Koch, 1837 (Acari, Hydrachnidia, Hygrobatidae) from Far East of Russia</mods:title>
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<mods:name id="D6A1DE4FE3BE3C7C08B067937EB96606" type="personal">
<mods:role id="63078CAC532B7A82F1401B168871D762">
<mods:roleTerm id="FDAA6A80BB3741F48F048EF224B91D38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C8BA07AE0B0E16952912ECAFEC59E9F9">Tuzovskij, Petr V.</mods:namePart>
<mods:affiliation id="7E02583665F593FCD494F9CB34CE4A3D">Institute for Biology of Inland waters of the Russian Academy of Sciences. Borok, Yaroslavl Province, 152742 Russia.</mods:affiliation>
<mods:nameIdentifier id="37F8B791C83C84A4A8A149A292A9FEC6" type="email">tuz@ibiw.yaroslavl.ru</mods:nameIdentifier>
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<mods:typeOfResource id="AA004BBE852DEC8141D34BEB53D94949">text</mods:typeOfResource>
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<mods:title id="2D47270FE61D72AF41D2F4777049CF82">Ecologica Montenegrina</mods:title>
</mods:titleInfo>
<mods:part id="7132E619224C4C3E6752668DBA1B7DEA">
<mods:date id="AD33D831EAB97B672D815D52EC8D16E7">2017</mods:date>
<mods:detail id="34DAF74201592E9C5014B245A8177994" type="pubDate">
<mods:number id="FE390128C562BF68A597FB2FA717D121">2017-02-10</mods:number>
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<mods:number id="B52E2E24DB134996BBE2F5C1C5976E45">10</mods:number>
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<mods:classification id="54E8E74EA3F145F35D806EF6B25B02AC">journal article</mods:classification>
<mods:identifier id="D74E46AB0F122F28BD3362B6BDF466EF" type="CLB-Dataset">300858</mods:identifier>
<mods:identifier id="A4E32F0C63B7AC7906630403CEAF2AF7" type="DOI">10.37828/em.2017.10.4</mods:identifier>
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<mods:identifier id="68BF185FB891577EA7553D51AB76C437" type="ISSN">2336-9744</mods:identifier>
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</mods:mods>
<treatment id="03D5878DFFC4FFA5FF1EFA3AFCC97A3C" ID-DOI="http://doi.org/10.5281/zenodo.12717722" ID-Zenodo-Dep="12717722" LSID="urn:lsid:zoobank.org:act:E8A898C7-8F7A-47E7-A7CF-EB423C1336B8" httpUri="http://treatment.plazi.org/id/03D5878DFFC4FFA5FF1EFA3AFCC97A3C" lastPageId="3" lastPageNumber="25" pageId="0" pageNumber="22">
<subSubSection id="C3666510FFC4FFA6FF1EFA3AFD707963" box="[151,664,1525,1551]" pageId="0" pageNumber="22" type="nomenclature">
<paragraph id="8BC3369BFFC4FFA6FF1EFA3AFD707963" blockId="0.[151,664,1525,1585]" box="[151,664,1525,1551]" pageId="0" pageNumber="22">
<heading id="D08B81F7FFC4FFA6FF1EFA3AFD707963" box="[151,664,1525,1551]" centered="true" fontSize="11" level="2" pageId="0" pageNumber="22" reason="2">
<emphasis id="B908EA89FFC4FFA6FF1EFA3AFD707963" bold="true" box="[151,664,1525,1551]" pageId="0" pageNumber="22">
<taxonomicName id="4C7C4D18FFC4FFA6FF1EFA3AFDBD7963" authorityName=". &amp; . &amp; P- &amp; I- &amp; - &amp; -" authorityYear="2017" box="[151,597,1525,1551]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="0" pageNumber="22" phylum="Arthropoda" rank="species" species="primoryensis" status="sp. nov." subGenus="Rivobates">
<emphasis id="B908EA89FFC4FFA6FF1EFA3AFECA7963" bold="true" box="[151,290,1525,1551]" italics="true" pageId="0" pageNumber="22">Hygrobates</emphasis>
(
<emphasis id="B908EA89FFC4FFA6FEBAFA3AFE407963" bold="true" box="[307,424,1525,1551]" italics="true" pageId="0" pageNumber="22">Rivobates</emphasis>
)
<emphasis id="B908EA89FFC4FFA6FE30FA3AFDBD7963" bold="true" box="[441,597,1525,1551]" italics="true" pageId="0" pageNumber="22">primoryensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57F2FFC4FFA6FDD4FA3AFD707963" box="[605,664,1525,1551]" pageId="0" pageNumber="22" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3666510FFC4FFA6FF1EF9D9FEF2795D" box="[151,282,1558,1585]" pageId="0" pageNumber="22" type="description">
<paragraph id="8BC3369BFFC4FFA6FF1EF9D9FEF2795D" blockId="0.[151,664,1525,1585]" box="[151,282,1558,1585]" pageId="0" pageNumber="22">
(
<figureCitation id="13472A1EFFC4FFA6FF16F9D9FEF9795D" box="[159,273,1558,1585]" captionStart-0="Figures 1-6" captionStart-1="Figures 7-10" captionStartId-0="1.[151,237,1087,1111]" captionStartId-1="2.[151,237,1138,1162]" captionTargetBox-0="[153,1424,210,1050]" captionTargetBox-1="[198,1414,202,1120]" captionTargetId-0="figure-17@1.[151,1433,189,1050]" captionTargetId-1="figure-8@2.[151,1427,189,1120]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" captionText-1="Figures 7-10. Hygrobates (Rivobates) primoryensis sp. n.: 7 – I-Leg-4-6, 8 – IV-Leg-4-6, 9 – leg claw; 10 – genital field; 7-9 male, 10 – female. Scale bars: 7, 8, 10 = 100 μm, 9 = 50 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13236526" figureDoi-1="http://doi.org/10.5281/zenodo.13236528" httpUri-0="https://zenodo.org/record/13236526/files/figure.png" httpUri-1="https://zenodo.org/record/13236528/files/figure.png" pageId="0" pageNumber="22">Figs 1-10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3666510FFC4FFA6FF1EF995FCD079B5" pageId="0" pageNumber="22" type="materials_examined">
<paragraph id="8BC3369BFFC4FFA6FF1EF995FCD079B5" blockId="0.[151,1437,1625,1922]" pageId="0" pageNumber="22">
<emphasis id="B908EA89FFC4FFA6FF1EF995FEA07918" bold="true" box="[151,328,1626,1652]" pageId="0" pageNumber="22">Type material</emphasis>
:
<materialsCitation id="3B143CC6FFC4FFA6FED1F996FA7179FA" collectingDate="1983-07-17" collectionCode="T" collectorName="S. Vshivkova" country="Russia" county="Komarovka river" location="Institute" municipality="Primory Territory" pageId="0" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C78839FFC4FFA6FED1F996FE227918" box="[344,458,1625,1652]" pageId="0" pageNumber="22" type="holotype">Holotype</typeStatus>
: male, slide 4967, deposited in the collection of the
<location id="8EA36040FFC4FFA6FBB1F996FB707918" LSID="urn:lsid:plazi:treatment:03D5878DFFC4FFA5FF1EFA3AFCC97A3C:8EA36040FFC4FFA6FBB1F996FB707918" box="[1080,1176,1625,1652]" country="Russia" county="Komarovka river" municipality="Primory Territory" name="Institute" pageId="0" pageNumber="22">Institute</location>
for
<location id="8EA36040FFC4FFA6FB43F996FF0279FA" LSID="urn:lsid:plazi:treatment:03D5878DFFC4FFA5FF1EFA3AFCC97A3C:8EA36040FFC4FFA6FB43F996FF0279FA" country="Russia" county="Komarovka river" municipality="Primory Territory" name="Biology of Inland Waters" pageId="0" pageNumber="22">Biology of Inland Waters</location>
(
<location id="8EA36040FFC4FFA6FE89F9B4FEA679FA" LSID="urn:lsid:plazi:treatment:03D5878DFFC4FFA5FF1EFA3AFCC97A3C:8EA36040FFC4FFA6FE89F9B4FEA679FA" box="[256,334,1659,1686]" country="Russia" county="Komarovka river" municipality="Primory Territory" name="Borok" pageId="0" pageNumber="22">Borok</location>
,
<collectingCountry id="F36B760BFFC4FFA6FEE8F9B4FE5C79FA" box="[353,436,1659,1686]" name="Russia" pageId="0" pageNumber="22">Russia</collectingCountry>
), Russia,
<collectingMunicipality id="6BA7ACE1FFC4FFA6FDBBF9B4FCE479FA" box="[562,780,1659,1686]" pageId="0" pageNumber="22">Primory Territory</collectingMunicipality>
,
<collectingCounty id="62A24E17FFC4FFA6FC96F9B4FC1B79FA" box="[799,1011,1659,1686]" pageId="0" pageNumber="22">Komarovka river</collectingCounty>
,
<date id="FFC2105BFFC4FFA6FB8CF9B4FB6379FA" box="[1029,1163,1659,1686]" pageId="0" pageNumber="22" value="1983-07-17">
<collectingDate id="EF86E9B3FFC4FFA6FB8CF9B4FB6379FA" box="[1029,1163,1659,1686]" pageId="0" pageNumber="22" value="1983-07-17">17.7. 1983</collectingDate>
</date>
, leg.
<collectionCode id="ED6DAE5EFFC4FFA6FB5EF9B4FB0679FA" box="[1239,1262,1659,1686]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="22">T</collectionCode>
.
<collectorName id="2689534DFFC4FFA6FB7BF9B4FA7179FA" box="[1266,1433,1659,1686]" pageId="0" pageNumber="22">S. Vshivkova</collectorName>
</materialsCitation>
.
<materialsCitation id="3B143CC6FFC4FFA6FF1EF951FCDD79B5" collectingDate="1983-07-17" collectionCode="T" collectorName="S. Vshivkova" country="Russia" county="Komarovka river" location="Institute" municipality="Primory Territory" pageId="0" pageNumber="22" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="54C78839FFC4FFA6FF1EF951FEE679D4" box="[151,270,1694,1720]" pageId="0" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7AFD12FFC4FFA6FE95F952FE4079D4" box="[284,424,1693,1720]" count="3" pageId="0" pageNumber="22" type="male">three males</specimenCount>
,
<specimenCount id="9D7AFD12FFC4FFA6FE3EF951FDD479D4" box="[439,572,1693,1720]" count="1" pageId="0" pageNumber="22" type="female">one female</specimenCount>
collected from the same locality and data as holotype, and
<specimenCount id="9D7AFD12FFC4FFA6FA9EF951FA7379D4" box="[1303,1435,1693,1720]" count="1" pageId="0" pageNumber="22" type="female">one female</specimenCount>
from the same locality,
<date id="FFC2105BFFC4FFA6FE24F971FDD479B5" box="[429,572,1726,1753]" pageId="0" pageNumber="22" value="1983-10-28">
<collectingDate id="EF86E9B3FFC4FFA6FE24F971FDD479B5" box="[429,572,1726,1753]" pageId="0" pageNumber="22" value="1983-10-28">28.10. 1983</collectingDate>
</date>
, leg.
<collectionCode id="ED6DAE5EFFC4FFA6FDF3F971FD7979B5" box="[634,657,1726,1753]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="22">T</collectionCode>
.
<collectorName id="2689534DFFC4FFA6FD1DF971FCDD79B5" box="[660,821,1726,1753]" pageId="0" pageNumber="22">S. Vshivkova</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3666510FFC4FFA6FF7FF92FFC9678EE" pageId="0" pageNumber="22" type="diagnosis">
<paragraph id="8BC3369BFFC4FFA6FF7FF92FFC9678EE" blockId="0.[151,1437,1625,1922]" pageId="0" pageNumber="22">
<emphasis id="B908EA89FFC4FFA6FF7FF92FFE9E7996" bold="true" box="[246,374,1760,1786]" pageId="0" pageNumber="22">Diagnosis.</emphasis>
Integument smooth; anterior coxal plates with short posterolateral apodemes, posteromedial margin convex in shape; coxal plate IV trapezoidal, medial margin forming an obtuse angle and with two medially-directed apodemes; genital field with 6-8 pairs of acetabula and 12-18 pairs setae, posterior margin of genital plate of male deeply indented; P-2 with short rounded ventrodistal protrusion; P-4 ventral setae located on the same level near middle of segment.
</paragraph>
</subSubSection>
<caption id="DF036613FFC5FFA7FF1EFBF0FC547B1A" ID-DOI="http://doi.org/10.5281/zenodo.13236526" ID-Zenodo-Dep="13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="1" pageNumber="23" startId="1.[151,237,1087,1111]" targetBox="[153,1424,210,1050]" targetPageId="1" targetType="figure">
<paragraph id="8BC3369BFFC5FFA7FF1EFBF0FC547B1A" blockId="1.[151,1436,1086,1142]" pageId="1" pageNumber="23">
<emphasis id="B908EA89FFC5FFA7FF1EFBF0FEF37B3B" bold="true" box="[151,283,1087,1111]" pageId="1" pageNumber="23">Figures 1-6</emphasis>
.
<taxonomicName id="4C7C4D18FFC5FFA7FEA4FBF1FD2C7B3A" authorityName=". &amp; . &amp; P- &amp; I- &amp; - &amp; -" authorityYear="2017" box="[301,708,1086,1110]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="23" phylum="Arthropoda" rank="species" species="primoryensis" status="sp. nov." subGenus="Rivobates">
<emphasis id="B908EA89FFC5FFA7FEA4FBF1FE417B3A" box="[301,425,1086,1110]" italics="true" pageId="1" pageNumber="23">Hygrobates</emphasis>
(
<emphasis id="B908EA89FFC5FFA7FE34FBF1FDCE7B3A" box="[445,550,1086,1110]" italics="true" pageId="1" pageNumber="23">Rivobates</emphasis>
)
<emphasis id="B908EA89FFC5FFA7FDB0FBF0FD2C7B3A" box="[569,708,1087,1110]" italics="true" pageId="1" pageNumber="23">primoryensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57F2FFC5FFA7FD46FBF0FCED7B3A" box="[719,773,1087,1110]" pageId="1" pageNumber="23" rank="species">sp.n.</taxonomicNameLabel>
: 1 – seta
<emphasis id="B908EA89FFC5FFA7FCF1FBF1FC497B3A" box="[888,929,1086,1110]" italics="true" pageId="1" pageNumber="23">Fch</emphasis>
, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm
</paragraph>
</caption>
<subSubSection id="C3666510FFC5FFA4FF1EFB0FFAD57ADB" lastPageId="2" lastPageNumber="24" pageId="1" pageNumber="23" type="description">
<paragraph id="8BC3369BFFC5FFA7FF1EFB0FFEC07BB6" blockId="1.[151,1437,1216,2052]" box="[151,296,1216,1242]" pageId="1" pageNumber="23">
<emphasis id="B908EA89FFC5FFA7FF1EFB0FFEC07BB6" bold="true" box="[151,296,1216,1242]" pageId="1" pageNumber="23">Description</emphasis>
</paragraph>
<paragraph id="8BC3369BFFC5FFA7FF1EFB2DFA807A2C" blockId="1.[151,1437,1216,2052]" pageId="1" pageNumber="23">
<emphasis id="B908EA89FFC5FFA7FF1EFB2DFF367B90" bold="true" box="[151,222,1250,1276]" pageId="1" pageNumber="23">Male.</emphasis>
Idiosoma oval and somewhat flattened dorsoventrally. Idiosomal setae associated with glandularia, ecept for trichobothria
<emphasis id="B908EA89FFC5FFA7FE3AFACBFE3C7A71" box="[435,468,1284,1309]" italics="true" pageId="1" pageNumber="23">Fp</emphasis>
,
<emphasis id="B908EA89FFC5FFA7FE6EFACBFDEC7A72" box="[487,516,1284,1310]" italics="true" pageId="1" pageNumber="23">Oi</emphasis>
and setae
<emphasis id="B908EA89FFC5FFA7FD07FACBFD407A71" box="[654,680,1284,1309]" italics="true" pageId="1" pageNumber="23">Pi</emphasis>
. Setae
<emphasis id="B908EA89FFC5FFA7FC8EFACCFCDC7A72" box="[775,820,1283,1310]" italics="true" pageId="1" pageNumber="23">Fch</emphasis>
(
<figureCitation id="13472A1EFFC5FFA7FCC0FACCFC7A7A72" box="[841,914,1283,1310]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="1" pageNumber="23">Fig. 1</figureCitation>
) much thicker than other idiosomal setae. Ventral view:
<figureCitation id="13472A1EFFC5FFA7FEB6FAEAFE6F7A2C" box="[319,391,1317,1344]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="1" pageNumber="23">Fig. 2</figureCitation>
. Coxal plate IV with two subcutaneous projections, in medial and posterior position.
</paragraph>
<paragraph id="8BC3369BFFC5FFA7FF1EFA89FEB17AEF" blockId="1.[151,1437,1216,2052]" pageId="1" pageNumber="23">
Genital plate inverse heart-shaped, wider than long (L/W ratio 0.8-0.9), anterior margin convex to obtuse angle (
<figureCitation id="13472A1EFFC5FFA7FF6FFAA7FEA47AEF" box="[230,332,1384,1411]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="1" pageNumber="23">Figs 3-4</figureCitation>
).
</paragraph>
<paragraph id="8BC3369BFFC5FFA7FF7FFA45FDDF7922" blockId="1.[151,1437,1216,2052]" pageId="1" pageNumber="23">
Pedipalp (
<figureCitation id="13472A1EFFC5FFA7FEF9FA45FE537AC9" box="[368,443,1418,1445]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="1" pageNumber="23">Fig. 5</figureCitation>
): P-1 with a single dorsodistal seta; P-2 with six short, thick dorsal setae, ventral margin proximally concave, distoventrally protruding in a short rounded projection covered by a few denticles; P-3 with two dorsocentral, two dorsodistal short, thick setae and a single thin dorsodistal seta, ventral margin straight or slightly concave proximally, with a few denticles covering distal 2/3 of ventral surface; P-4 longer than P-2 (P-4/P-2 L ratio 1.24-1.42), ventral setae subequal in length and located on the same level near middle of segment.
</paragraph>
<paragraph id="8BC3369BFFC5FFA7FF7FF99AFE4179B9" blockId="1.[151,1437,1216,2052]" pageId="1" pageNumber="23">
I-Leg-5 with two subequal rather long distoventral setae (
<figureCitation id="13472A1EFFC5FFA7FC30F99BFBED7903" box="[953,1029,1620,1647]" captionStart="Figures 7-10" captionStartId="2.[151,237,1138,1162]" captionTargetBox="[198,1414,202,1120]" captionTargetId="figure-8@2.[151,1427,189,1120]" captionTargetPageId="2" captionText="Figures 7-10. Hygrobates (Rivobates) primoryensis sp. n.: 7 – I-Leg-4-6, 8 – IV-Leg-4-6, 9 – leg claw; 10 – genital field; 7-9 male, 10 – female. Scale bars: 7, 8, 10 = 100 μm, 9 = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13236528" httpUri="https://zenodo.org/record/13236528/files/figure.png" pageId="1" pageNumber="23">Fig. 7</figureCitation>
); IV-Leg-4 with five short, thick unequal distal setae, IV-Leg-5 with four unequal distal thick setae, IV-Leg-6 usually with three thick setae (
<figureCitation id="13472A1EFFC5FFA7FF29F957FF0279DF" box="[160,234,1688,1715]" captionStart="Figures 7-10" captionStartId="2.[151,237,1138,1162]" captionTargetBox="[198,1414,202,1120]" captionTargetId="figure-8@2.[151,1427,189,1120]" captionTargetPageId="2" captionText="Figures 7-10. Hygrobates (Rivobates) primoryensis sp. n.: 7 – I-Leg-4-6, 8 – IV-Leg-4-6, 9 – leg claw; 10 – genital field; 7-9 male, 10 – female. Scale bars: 7, 8, 10 = 100 μm, 9 = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13236528" httpUri="https://zenodo.org/record/13236528/files/figure.png" pageId="1" pageNumber="23">Fig. 8</figureCitation>
). Claws of all legs with long external clawlet and short internal one, lamella with slightly concave ventral margin (
<figureCitation id="13472A1EFFC5FFA7FEDAF975FE7379B9" box="[339,411,1722,1749]" captionStart="Figures 7-10" captionStartId="2.[151,237,1138,1162]" captionTargetBox="[198,1414,202,1120]" captionTargetId="figure-8@2.[151,1427,189,1120]" captionTargetPageId="2" captionText="Figures 7-10. Hygrobates (Rivobates) primoryensis sp. n.: 7 – I-Leg-4-6, 8 – IV-Leg-4-6, 9 – leg claw; 10 – genital field; 7-9 male, 10 – female. Scale bars: 7, 8, 10 = 100 μm, 9 = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13236528" httpUri="https://zenodo.org/record/13236528/files/figure.png" pageId="1" pageNumber="23">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BC3369BFFC5FFA7FF74F913FA9B7811" blockId="1.[151,1437,1216,2052]" pageId="1" pageNumber="23">
Measurements (n=4). Idiosoma L 430-600; seta
<emphasis id="B908EA89FFC5FFA7FCCDF914FC9A799A" box="[836,882,1755,1782]" italics="true" pageId="1" pageNumber="23">Fch</emphasis>
L 42-48; coxal plates I + capitulum mL 185- 215; genital plates L 125-135, W 135-170, cheliceral segments L: base 120-132, chela 72-78; pedipalp segments (P–1–5) L: 18-24, 71-73, 60-66, 95-102, 30-36; legs segments L: I-Leg-1-6: 48-55, 54-60, 84-90, 120- 125, 120-125, 115-120; II-Leg-1-6: 48-55, 54-60, 89-92, 115-125, 125-135, 105-125; III-Leg-1-6: 54-60, 60- 65, 95-102, 130-150, 150-170, 150-160; IV-Leg-1-6: 100-110, 84-95, 130-140, 170-185, 175-195, 160-175.
</paragraph>
<paragraph id="8BC3369BFFC5FFA7FF7FF84BFEC97768" blockId="1.[151,1437,1216,2052]" pageId="1" pageNumber="23">
<emphasis id="B908EA89FFC5FFA7FF7FF84BFEB078F2" bold="true" box="[246,344,1924,1950]" pageId="1" pageNumber="23">Female.</emphasis>
Genital plate longer than width (L/W ratio 1.7-1.8), with 6-7 acetabula and 13-16 setae; medial margin straight and slightly indented anteriorly to middle of plate (
<figureCitation id="13472A1EFFC5FFA7FB8CF869FBBA78AD" box="[1029,1106,1958,1985]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="1" pageNumber="23">Figs 6</figureCitation>
,
<figureCitation id="13472A1EFFC5FFA7FBD6F869FB6878AD" box="[1119,1152,1958,1985]" captionStart="Figures 7-10" captionStartId="2.[151,237,1138,1162]" captionTargetBox="[198,1414,202,1120]" captionTargetId="figure-8@2.[151,1427,189,1120]" captionTargetPageId="2" captionText="Figures 7-10. Hygrobates (Rivobates) primoryensis sp. n.: 7 – I-Leg-4-6, 8 – IV-Leg-4-6, 9 – leg claw; 10 – genital field; 7-9 male, 10 – female. Scale bars: 7, 8, 10 = 100 μm, 9 = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13236528" httpUri="https://zenodo.org/record/13236528/files/figure.png" pageId="1" pageNumber="23">10</figureCitation>
). Anterior and posterior genital sclerites small and subequal in size. Excretory pore unsclerotized and located between flankig setae (
<emphasis id="B908EA89FFC5FFA7FF28F825FF557768" box="[161,189,2026,2052]" italics="true" pageId="1" pageNumber="23">Ci</emphasis>
and
<emphasis id="B908EA89FFC5FFA7FF7FF825FEF8776F" box="[246,272,2026,2051]" italics="true" pageId="1" pageNumber="23">Pi</emphasis>
).
</paragraph>
<caption id="DF036613FFC6FFA4FF1EFBBDFC667BC5" ID-DOI="http://doi.org/10.5281/zenodo.13236528" ID-Zenodo-Dep="13236528" httpUri="https://zenodo.org/record/13236528/files/figure.png" pageId="2" pageNumber="24" startId="2.[151,237,1138,1162]" targetBox="[198,1414,202,1120]" targetPageId="2" targetType="figure">
<paragraph id="8BC3369BFFC6FFA4FF1EFBBDFC667BC5" blockId="2.[151,1386,1138,1193]" pageId="2" pageNumber="24">
<emphasis id="B908EA89FFC6FFA4FF1EFBBDFECD7BE6" bold="true" box="[151,293,1138,1162]" pageId="2" pageNumber="24">Figures 7-10</emphasis>
.
<taxonomicName id="4C7C4D18FFC6FFA4FEBBFBBDFD2A7BE6" authorityName=". &amp; . &amp; P- &amp; I- &amp; - &amp; -" authorityYear="2017" box="[306,706,1138,1162]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="24" phylum="Arthropoda" rank="species" species="primoryensis" status="sp. nov." subGenus="Rivobates">
<emphasis id="B908EA89FFC6FFA4FEBBFBBDFE467BE6" box="[306,430,1138,1162]" italics="true" pageId="2" pageNumber="24">Hygrobates</emphasis>
(
<emphasis id="B908EA89FFC6FFA4FE37FBBDFDCF7BE6" box="[446,551,1138,1162]" italics="true" pageId="2" pageNumber="24">Rivobates</emphasis>
)
<emphasis id="B908EA89FFC6FFA4FDBEFBBCFD2A7BE6" box="[567,706,1139,1162]" italics="true" pageId="2" pageNumber="24">primoryensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57F2FFC6FFA4FD41FBBCFCEC7BE6" box="[712,772,1139,1162]" pageId="2" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
: 7 – I-Leg-4-6, 8 – IV-Leg-4-6, 9 – leg claw; 10 – genital field; 7-9 male, 10 – female. Scale bars: 7, 8, 10 = 100 μm, 9 = 50 μm.
</paragraph>
</caption>
<paragraph id="8BC3369BFFC6FFA4FF74FAD9FAD57ADB" blockId="2.[151,1438,1301,1463]" pageId="2" pageNumber="24">
Measurements (n=2). Idiosoma L 480-650; seta
<emphasis id="B908EA89FFC6FFA4FCCDFADAFC9A7A5C" box="[836,882,1301,1328]" italics="true" pageId="2" pageNumber="24">Fch</emphasis>
L 42-48; coxal plates I + capitulum mL 230- 235; genital plate L 114-120, W 65-68, cheliceral segments L: base 160-165, chela 90; pedipalp segments (P– 1–5) L: 30-35, 78-90, 65-72, 108-115, 36-42; legs segments L: I-Leg-1-6: 50-55, 65-70, 95-102, 135-140, 135- 140, 125-140; II-Leg-1-6: 50-55, 65-78, 95-102, 135-140, 145-152, 130-145; III-Leg-1-6: 54-60, 72-84, 105- 110, 155-165, 180-186, 160-170; IV-Leg-1-6: 114-120, 100-108, 150-165, 204-210, 215-225, 190-200.
</paragraph>
</subSubSection>
<caption id="DF036613FFC6FFA4FF1EF865FF35788C" ID-DOI="http://doi.org/10.5281/zenodo.13236530" ID-Zenodo-Dep="13236530" httpUri="https://zenodo.org/record/13236530/files/figure.png" pageId="2" pageNumber="24" startId="2.[151,237,1962,1986]" targetBox="[207,1375,1543,1898]" targetPageId="2" targetType="figure">
<paragraph id="8BC3369BFFC6FFA4FF1EF865FF35788C" blockId="2.[151,1409,1962,2016]" pageId="2" pageNumber="24">
<emphasis id="B908EA89FFC6FFA4FF1EF865FEDA78AE" bold="true" box="[151,306,1962,1986]" pageId="2" pageNumber="24">Figures 11-12</emphasis>
.
<taxonomicName id="4C7C4D18FFC6FFA4FEB6F865FD7578AE" baseAuthorityName="Uchida" baseAuthorityYear="1934" box="[319,669,1962,1986]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="24" phylum="Arthropoda" rank="species" species="ezoensis" subGenus="Rivobates">
<emphasis id="B908EA89FFC6FFA4FEB6F865FE5378AE" box="[319,443,1962,1986]" italics="true" pageId="2" pageNumber="24">Hygrobates</emphasis>
(
<emphasis id="B908EA89FFC6FFA4FE45F865FDDD78AE" box="[460,565,1962,1986]" italics="true" pageId="2" pageNumber="24">Rivobates</emphasis>
)
<emphasis id="B908EA89FFC6FFA4FDCDF864FD7578AE" box="[580,669,1963,1986]" italics="true" pageId="2" pageNumber="24">ezoensis</emphasis>
</taxonomicName>
: 11-12 – genital field, 11 – female, 12 – male (after
<bibRefCitation id="EFED4B6AFFC6FFA4FB44F865FF39788C" author="Matsumoto, N. &amp; Dick, M. H. &amp; Mawatari S. F." pageId="2" pageNumber="24" pagination="1893 - 1945" refId="ref1583" refString="Matsumoto, N., Dick, M. H., &amp; Mawatari S. F. (2005) Water mites of the genus Hygrobates Koch, 1837 (Acari: Parasitengona: Hygrobatidae) from Hokkaido, northern Japan. Journal of Natural History, 39 (21), 1893 - 1945." type="journal article" year="2005">Matsumoto et al. 2005</bibRefCitation>
).
</paragraph>
</caption>
<subSubSection id="C3666510FFC7FFA5FF1EFF0BFCC97A3C" pageId="3" pageNumber="25" type="discussion">
<paragraph id="8BC3369BFFC7FFA5FF1EFF0BFE697EC5" blockId="3.[151,1436,195,425]" pageId="3" pageNumber="25">
<emphasis id="B908EA89FFC7FFA5FF1EFF0BFEF87FB2" bold="true" box="[151,272,196,222]" pageId="3" pageNumber="25">Remarks.</emphasis>
The present species is similar to
<taxonomicName id="4C7C4D18FFC7FFA5FD2EFF0CFBBC7FB2" authority="(Uchida, 1934)" baseAuthorityName="Uchida" baseAuthorityYear="1934" box="[679,1108,195,222]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="25" phylum="Arthropoda" rank="species" species="ezoensis">
<emphasis id="B908EA89FFC7FFA5FD2EFF0CFC717FB2" box="[679,921,195,222]" italics="true" pageId="3" pageNumber="25">Hygrobates ezoensis</emphasis>
(Uchida, 1934)
</taxonomicName>
but differs in the following characters (character states of adult
<taxonomicName id="4C7C4D18FFC7FFA5FDDDFF29FD377F93" baseAuthorityName="Uchida" baseAuthorityYear="1934" box="[596,735,230,255]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="25" phylum="Arthropoda" rank="species" species="ezoensis">
<emphasis id="B908EA89FFC7FFA5FDDDFF29FD377F93" box="[596,735,230,255]" italics="true" pageId="3" pageNumber="25">H. ezoensis</emphasis>
</taxonomicName>
given in parenthesis, data from
<bibRefCitation id="EFED4B6AFFC7FFA5FBFAFF29FA677E6C" author="Matsumoto, N. &amp; Dick, M. H. &amp; Mawatari S. F." box="[1139,1423,229,256]" pageId="3" pageNumber="25" pagination="1893 - 1945" refId="ref1583" refString="Matsumoto, N., Dick, M. H., &amp; Mawatari S. F. (2005) Water mites of the genus Hygrobates Koch, 1837 (Acari: Parasitengona: Hygrobatidae) from Hokkaido, northern Japan. Journal of Natural History, 39 (21), 1893 - 1945." type="journal article" year="2005">Matsumoto et al. 2005</bibRefCitation>
): integument smooth (finely striated); coxal plate IV with two medial projections,
<figureCitation id="13472A1EFFC7FFA5FBD9FEC8FB7D7E4E" box="[1104,1173,263,290]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="3" pageNumber="25">Fig. 2</figureCitation>
(without projections); anterior and posterior genital sclerites in females relatively narrow,
<figureCitation id="13472A1EFFC7FFA5FC54FEE7FBC57E2F" box="[989,1069,296,323]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="3" pageNumber="25">Figs 6</figureCitation>
,
<figureCitation id="13472A1EFFC7FFA5FBB7FEE7FBB37E2F" box="[1086,1115,296,323]" captionStart="Figures 7-10" captionStartId="2.[151,237,1138,1162]" captionTargetBox="[198,1414,202,1120]" captionTargetId="figure-8@2.[151,1427,189,1120]" captionTargetPageId="2" captionText="Figures 7-10. Hygrobates (Rivobates) primoryensis sp. n.: 7 – I-Leg-4-6, 8 – IV-Leg-4-6, 9 – leg claw; 10 – genital field; 7-9 male, 10 – female. Scale bars: 7, 8, 10 = 100 μm, 9 = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.13236528" httpUri="https://zenodo.org/record/13236528/files/figure.png" pageId="3" pageNumber="25">10</figureCitation>
(wide,
<figureCitation id="13472A1EFFC7FFA5FB36FEE7FAF07E2F" box="[1215,1304,296,323]" captionStart="Figures 11-12" captionStartId="2.[151,237,1962,1986]" captionTargetBox="[207,1375,1543,1898]" captionTargetId="figure-237@2.[151,1428,1543,1931]" captionTargetPageId="2" captionText="Figures 11-12. Hygrobates (Rivobates) ezoensis: 11-12 – genital field, 11 – female, 12 – male (after Matsumoto et al. 2005)." figureDoi="http://doi.org/10.5281/zenodo.13236530" httpUri="https://zenodo.org/record/13236530/files/figure.png" pageId="3" pageNumber="25">Fig. 11</figureCitation>
), posterior margin of the genital plate of male deeply indented,
<figureCitation id="13472A1EFFC7FFA5FC99FE85FC9B7E09" box="[784,883,330,357]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="3" pageNumber="25">Figs 3-4</figureCitation>
(with straight to convex posterior margin and without the indentation,
<figureCitation id="13472A1EFFC7FFA5FE32FEA3FDFB7EEB" box="[443,531,364,391]" captionStart="Figures 11-12" captionStartId="2.[151,237,1962,1986]" captionTargetBox="[207,1375,1543,1898]" captionTargetId="figure-237@2.[151,1428,1543,1931]" captionTargetPageId="2" captionText="Figures 11-12. Hygrobates (Rivobates) ezoensis: 11-12 – genital field, 11 – female, 12 – male (after Matsumoto et al. 2005)." figureDoi="http://doi.org/10.5281/zenodo.13236530" httpUri="https://zenodo.org/record/13236530/files/figure.png" pageId="3" pageNumber="25">Fig. 12</figureCitation>
); P-2 with a short rounded projection,
<figureCitation id="13472A1EFFC7FFA5FC6AFEA3FBC17EEB" box="[995,1065,364,391]" captionStart="Figures 1-6" captionStartId="1.[151,237,1087,1111]" captionTargetBox="[153,1424,210,1050]" captionTargetId="figure-17@1.[151,1433,189,1050]" captionTargetPageId="1" captionText="Figures 1-6. Hygrobates (Rivobates) primoryensis sp.n.: 1 – seta Fch, 2 – ventral view, 3-4, 6 – genital field, 5 – pedipalp; 1-5 – male, 6 – female. Scale bars: 1, 5 = 50 μm, 2-4, 6 = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.13236526" httpUri="https://zenodo.org/record/13236526/files/figure.png" pageId="3" pageNumber="25">Fig. 5</figureCitation>
(with tapering, pointed ventral projection,
<figureCitation id="13472A1EFFC7FFA5FE94FE41FE9B7EC5" box="[285,371,398,425]" captionStart="Figure 13" captionStartId="3.[151,226,1100,1124]" captionTargetBox="[343,1242,481,1051]" captionTargetId="figure-122@3.[303,1284,425,1094]" captionTargetPageId="3" captionText="Figure 13. Hygrobates (Rivobates) ezoensis: 13 – pedipalp (after Matsumoto et al. 2005)." figureDoi="http://doi.org/10.5281/zenodo.13236532" httpUri="https://zenodo.org/record/13236532/files/figure.png" pageId="3" pageNumber="25">Fig. 13</figureCitation>
).
</paragraph>
<caption id="DF036613FFC7FFA5FF1EFB83FBBD7B08" ID-DOI="http://doi.org/10.5281/zenodo.13236532" ID-Zenodo-Dep="13236532" box="[151,1109,1100,1124]" httpUri="https://zenodo.org/record/13236532/files/figure.png" pageId="3" pageNumber="25" startId="3.[151,226,1100,1124]" targetBox="[343,1242,481,1051]" targetPageId="3" targetType="figure">
<paragraph id="8BC3369BFFC7FFA5FF1EFB83FBBD7B08" blockId="3.[151,1109,1100,1124]" box="[151,1109,1100,1124]" pageId="3" pageNumber="25">
<emphasis id="B908EA89FFC7FFA5FF1EFB83FEE27B08" bold="true" box="[151,266,1100,1124]" pageId="3" pageNumber="25">Figure 13.</emphasis>
<taxonomicName id="4C7C4D18FFC7FFA5FE9BFB83FD877B08" baseAuthorityName="Uchida" baseAuthorityYear="1934" box="[274,623,1100,1124]" class="Arachnida" family="Hygrobatidae" genus="Hygrobates" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="25" phylum="Arthropoda" rank="species" species="ezoensis" subGenus="Rivobates">
<emphasis id="B908EA89FFC7FFA5FE9BFB83FE667B08" box="[274,398,1100,1124]" italics="true" pageId="3" pageNumber="25">Hygrobates</emphasis>
(
<emphasis id="B908EA89FFC7FFA5FE14FB83FDEE7B08" box="[413,518,1100,1124]" italics="true" pageId="3" pageNumber="25">Rivobates</emphasis>
)
<emphasis id="B908EA89FFC7FFA5FD9FFB82FD877B08" box="[534,623,1101,1124]" italics="true" pageId="3" pageNumber="25">ezoensis</emphasis>
</taxonomicName>
: 13 – pedipalp (after
<bibRefCitation id="EFED4B6AFFC7FFA5FCDCFB82FBA17B08" author="Matsumoto, N. &amp; Dick, M. H. &amp; Mawatari S. F." box="[853,1097,1100,1124]" pageId="3" pageNumber="25" pagination="1893 - 1945" refId="ref1583" refString="Matsumoto, N., Dick, M. H., &amp; Mawatari S. F. (2005) Water mites of the genus Hygrobates Koch, 1837 (Acari: Parasitengona: Hygrobatidae) from Hokkaido, northern Japan. Journal of Natural History, 39 (21), 1893 - 1945." type="journal article" year="2005">Matsumoto et al. 2005</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8BC3369BFFC7FFA5FF1EFB60FB657BA5" blockId="3.[151,1165,1198,1225]" box="[151,1165,1198,1225]" pageId="3" pageNumber="25">
<emphasis id="B908EA89FFC7FFA5FF1EFB60FEC77BA5" bold="true" box="[151,303,1199,1225]" pageId="3" pageNumber="25">Ethymology</emphasis>
. The species is named after the Primory Territory where it was collected.
</paragraph>
<paragraph id="8BC3369BFFC7FFA5FF1EFB3DFE2F7A61" blockId="3.[151,455,1266,1293]" box="[151,455,1266,1293]" pageId="3" pageNumber="25">
<emphasis id="B908EA89FFC7FFA5FF1EFB3DFF177A60" bold="true" box="[151,255,1266,1292]" pageId="3" pageNumber="25">Habitat.</emphasis>
Running waters.
</paragraph>
<paragraph id="8BC3369BFFC7FFA5FF1EFAF9FCC97A3C" blockId="3.[151,801,1333,1360]" box="[151,801,1333,1360]" pageId="3" pageNumber="25">
<emphasis id="B908EA89FFC7FFA5FF1EFAF9FED87A3C" bold="true" box="[151,304,1334,1360]" pageId="3" pageNumber="25">Distrbution.</emphasis>
Asia,
<collectingCountry id="F36B760BFFC7FFA5FEF5FAFAFE257A3C" box="[380,461,1333,1360]" name="Russia" pageId="3" pageNumber="25">Russia</collectingCountry>
, Far East, Primory Territory.
</paragraph>
</subSubSection>
</treatment>
</document>
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