462 lines
66 KiB
XML
462 lines
66 KiB
XML
<document id="CA3BA204F3B1828B0B05AB520987DD5E" ID-DOI="10.1093/zoolinnean/zlae162" ID-ISSN="0024-4082" ID-Zenodo-Dep="14832689" ID-ZooBank="7-01-2184" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1738158242914" checkinUser="plazi" docAuthor="Ortega, Jhael A., Cisneros-Heredia, Diego F., Camper, Jeffrey D., Romero-Carvajal, Andrés, Negrete, Leonardo & Ron, Santiago R." docDate="2025" docId="03D58787FFA9FFD6C033C0EF96A54EBD" docLanguage="en" docName="zlae162.pdf" docOrigin="Zoological Journal of the Linnean Society 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae162" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Phyllonastes dicaprioi Ortega, Cisneros-Heredia, Camper, Romero-Carvajal, Negrete & Ron, 2025, sp.nov." docType="treatment" docVersion="4" lastPageNumber="31" masterDocId="FFECFFFFFFB2FFC8C12CC61F94654B3C" masterDocTitle="Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography" masterLastPageNumber="60" masterPageNumber="1" pageNumber="28" updateTime="1738929907134" updateUser="ExternalLinkService">
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<mods:title id="0AC8A81DBE514DE4C541940E4D598545">Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography</mods:title>
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<mods:namePart id="DC4C4923E7686F13883030CE65FC703A">Ortega, Jhael A.</mods:namePart>
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<mods:affiliation id="9ED7B70A03AB1B3B2D8FBDA93997F236">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:roleTerm id="F61CE3E8223C92A8E5AFA8C50A94B65C">Author</mods:roleTerm>
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<mods:namePart id="972D45973729322C2AAB8990ACB2CC88">Cisneros-Heredia, Diego F.</mods:namePart>
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<mods:affiliation id="F2FA254809DD7BE6F90D3A65937725A0">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Museo de Zoología / Laboratorio de Zoología Terrestre, Quito, Ecuador & Instituto Nacional de Biodiversidad INABIO, División de Herpetología, Quito, Ecuador</mods:affiliation>
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<mods:roleTerm id="AA8D9E4D836822714E98081636860CC7">Author</mods:roleTerm>
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<mods:namePart id="CC3DD63229A9512E5B884F8ECD2B271B">Camper, Jeffrey D.</mods:namePart>
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<mods:affiliation id="F282AC3D2A7227B4A35945A23C379EE0">Department of Biology, Francis Marion University, Florence, SC 29506, USA</mods:affiliation>
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</mods:name>
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<mods:namePart id="CE0189E7BF027C41363A9B4257105472">Romero-Carvajal, Andrés</mods:namePart>
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<mods:affiliation id="6DF0D828A92F79FD50354FBE297B4C60">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="9F345DD91AF1D632E23E6FE0595B5791">Negrete, Leonardo</mods:namePart>
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<mods:affiliation id="FBA0D166BB7A195D9FCF4FCC21F17374">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="11E2C2C69F9ABBF1F2ADDE4CA3DF56D9">Ron, Santiago R.</mods:namePart>
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<mods:nameIdentifier id="D206B63A538A6D84C3E83CF2F471F060" type="email">santiago.r.ron@gmail.com</mods:nameIdentifier>
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<mods:title id="D807D692348BD5FB8D1D022EE6F48003">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="E020689F4B592C2BE863D59A100EA006">
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<mods:date id="9ED06BAEFF4D20B3EE0E9B5D43A24CB0">2025</mods:date>
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<mods:number id="0AF2819F6BECC56D4B282713DB11790D">203</mods:number>
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<mods:url id="E0969F822A09D31DF97784BB44FD17AF">https://doi.org/10.1093/zoolinnean/zlae162</mods:url>
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<mods:identifier id="AEC3EDD6BA1FA16908A933EC186037C1" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03D58787FFA9FFD6C033C0EF96A54EBD" ID-DOI="http://doi.org/10.5281/zenodo.14832705" ID-Zenodo-Dep="14832705" LSID="urn:lsid:plazi:treatment:03D58787FFA9FFD6C033C0EF96A54EBD" httpUri="http://treatment.plazi.org/id/03D58787FFA9FFD6C033C0EF96A54EBD" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
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<subSubSection id="C366651AFFA9FFD3C033C0EF96294C36" box="[287,588,1776,1803]" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BC33691FFA9FFD3C033C0EF96294C36" blockId="27.[113,762,1776,1901]" box="[287,588,1776,1803]" pageId="27" pageNumber="28">
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<emphasis id="B908EA83FFA9FFD3C033C0EF96294C36" bold="true" box="[287,588,1776,1803]" pageId="27" pageNumber="28">
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<taxonomicName id="4C7C4D12FFA9FFD3C033C0EF959E4C37" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[287,507,1776,1803]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="dicaprioi" status="sp. nov.">
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<emphasis id="B908EA83FFA9FFD3C033C0EF959E4C37" bold="true" box="[287,507,1776,1803]" italics="true" pageId="27" pageNumber="28">Phyllonastes dicaprioi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23B57F8FFA9FFD3C32CC0EF96294C36" box="[512,588,1776,1802]" pageId="27" pageNumber="28" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C366651AFFA9FFD3C15DC10895784C72" pageId="27" pageNumber="28" type="description">
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<paragraph id="8BC33691FFA9FFD3C15DC10895784C72" blockId="27.[113,762,1776,1901]" pageId="27" pageNumber="28">
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LSID:
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urn:lsid:zoobank.org:act:
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<paragraph id="8BC33691FFA9FFD3C1A1C14A96E24C51" blockId="27.[113,762,1776,1901]" box="[141,647,1877,1901]" pageId="27" pageNumber="28">
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<taxonomicName id="4C7C4D12FFA9FFD3C1A1C14A957B4C51" baseAuthorityName="Lynch" baseAuthorityYear="1986" box="[141,286,1877,1901]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="heyeri">
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||
<emphasis id="B908EA83FFA9FFD3C1A1C14A957B4C51" box="[141,286,1877,1901]" italics="true" pageId="27" pageNumber="28">Noblella heyeri</emphasis>
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</taxonomicName>
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||
—
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||
<bibRefCitation id="EFED4B60FFA9FFD3C016C14A96E44C51" author="Garzon-Santomaro C & Sanchez-Nivicela JC & Mena Valenzuela PR" box="[314,641,1877,1901]" pageId="27" pageNumber="28" refId="ref38375" refString="Garzon-Santomaro C, Sanchez-Nivicela JC, Mena Valenzuela PR et al. Reptiles y Aves de la provincia de El Oro: Una guia de identificacion de especies del paramo al manglar. Quito, Ecuador: Serie de Publicaciones GADPEO - INABIO, 2019." type="book" year="2019">
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||
Garzón-Santomaro
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||
<emphasis id="B908EA83FFA9FFD3C328C14996564C51" box="[516,563,1877,1901]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
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||
(2019)
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||
</bibRefCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C366651AFFA9FFD3C15DC19091D64A42" pageId="27" pageNumber="28" type="materials_examined">
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<paragraph id="8BC33691FFA9FFD3C15DC19091FC4BFB" blockId="27.[113,763,1935,1990]" lastBlockId="27.[810,1459,144,199]" pageId="27" pageNumber="28">
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<materialsCitation id="3B143CCCFFA9FFD3C15DC19091FC4BFB" collectingDate="2017-07-03" collectionCode="DHMECN, SRJ" collectorName="Filemon Benitez & Miguel Urgiles" country="Ecuador" elevation="1705" latitude="-3.206528" location="province of El Oro" longLatPrecision="1" longitude="-79.67977" pageId="27" pageNumber="28" specimenCode="DHMECN 13729" specimenCount="1" stateProvince="province of El Oro" typeStatus="holotype">
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<emphasis id="B908EA83FFA9FFD3C15DC19096414C9B" box="[113,548,1935,1959]" italics="true" pageId="27" pageNumber="28">
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<typeStatus id="54C78833FFA9FFD3C15DC19094AF4C9B" box="[113,202,1935,1959]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="13472A14FFA9FFD3C1F0C19095414C9B" box="[220,292,1935,1959]" captionStart="Figure 16" captionStartId="28.[129,194,1713,1737]" captionTargetBox="[165,1433,153,1679]" captionTargetId="figure-95@28.[161,1441,144,1685]" captionTargetPageId="28" captionText="Figure 16. Preserved specimens of Phyllonastes dicaprioi. Lateral, dorsal, and ventral views of: A, holotype DHMECN 13729, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm." figureDoi="http://doi.org/10.5281/zenodo.14832731" httpUri="https://zenodo.org/record/14832731/files/figure.png" pageId="27" pageNumber="28">Figs 16</figureCitation>
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,
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||
<figureCitation id="13472A14FFA9FFD3C01CC190952F4C9B" box="[304,330,1935,1959]" captionStart="Figure 17" captionStartId="29.[112,177,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-482@29.[306,1266,144,1064]" captionTargetPageId="29" captionText="Figure 17. Palmar (A) and plantar (B) surfaces of Phyllonastes dicaprioi. Photographs of right hand and left foot of the holotype DHMECN 13729." figureDoi="http://doi.org/10.5281/zenodo.14832735" httpUri="https://zenodo.org/record/14832735/files/figure.png" pageId="27" pageNumber="28">17</figureCitation>
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):
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||
<specimenCode id="DBDA9EEAFFA9FFD3C047C19096414C9B" box="[363,548,1935,1959]" collectionCode="DHMECN" pageId="27" pageNumber="28">DHMECN 13729</specimenCode>
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</emphasis>
|
||
(field no. SRJ 2017- 364),
|
||
<specimenCount id="9D7AFD18FFA9FFD3C19EC1B195444CFA" box="[178,289,1966,1990]" pageId="27" pageNumber="28" type="adult">adult male</specimenCount>
|
||
from
|
||
<collectingCountry id="F36B7601FFA9FFD3C04AC1B196334CFA" box="[358,598,1966,1990]" name="Ecuador" pageId="27" pageNumber="28">República del Ecuador</collectingCountry>
|
||
,
|
||
<collectingRegion id="49B8F873FFA9FFD3C34FC1B197304B94" box="[611,853,144,1990]" pageId="27" pageNumber="28">province of El Oro</collectingRegion>
|
||
, El Retiro (
|
||
<geoCoordinate id="EE485056FFA9FFD3C2F8C68F902A4B94" box="[980,1103,144,168]" degrees="3.206528" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="-3.206528">3.206528°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE485056FFA9FFD3C571C68F90B84B94" box="[1117,1245,144,168]" degrees="79.67977" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-79.67977">79.67977°W</geoCoordinate>
|
||
,
|
||
<quantity id="4C849B74FFA9FFD3C5C6C68F91584B9B" box="[1258,1341,144,168]" metricMagnitude="3" metricUnit="m" metricValue="1.705" pageId="27" pageNumber="28" unit="m" value="1705.0">
|
||
<elevation id="0051D1A2FFA9FFD3C5C6C68F91584B9B" box="[1258,1341,144,168]" metricMagnitude="3" metricUnit="m" metricValue="1.705" pageId="27" pageNumber="28" unit="m" value="1705.0">1705 m</elevation>
|
||
</quantity>
|
||
a.s.l.), collected by
|
||
<collectorName id="26895347FFA9FFD3C2A7C6B090564BFB" box="[907,1075,175,199]" pageId="27" pageNumber="28">Filemón Benítez</collectorName>
|
||
and
|
||
<collectorName id="26895347FFA9FFD3C548C6B0909F4BFB" box="[1124,1274,175,199]" pageId="27" pageNumber="28">Miguel Urgilés</collectorName>
|
||
on
|
||
<date id="FFC21051FFA9FFD3C40CC6B091F04BFB" box="[1312,1429,175,199]" pageId="27" pageNumber="28" value="2017-07-03">
|
||
<collectingDate id="EF86E9B9FFA9FFD3C40CC6B091F04BFB" box="[1312,1429,175,199]" pageId="27" pageNumber="28" value="2017-07-03">3 July 2017</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA9FFD3C206C6F691D64A42" blockId="27.[810,1461,233,382]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B143CCCFFA9FFD3C206C6F691D04A1C" collectionCode="DHMECN" collectorName="El Retiro" country="Ecuador" elevation="1330" latitude="-3.61847" location="El Oro Province" longLatPrecision="1" longitude="-79.58072" pageId="27" pageNumber="28" specimenCode="DHMECN 13723, DHMECN 13730" specimenCount="1" specimenCount-adult="25" stateProvince="El Oro Province" typeStatus="paratype">
|
||
<emphasis id="B908EA83FFA9FFD3C206C6F6905C4A3C" box="[810,1081,233,257]" italics="true" pageId="27" pageNumber="28">
|
||
<typeStatus id="54C78833FFA9FFD3C206C6F697ED4A3C" box="[810,904,233,256]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
|
||
(N = 9;
|
||
<figureCitation id="13472A14FFA9FFD3C2C8C6F690424A3D" box="[996,1063,233,257]" captionStart="Figure 16" captionStartId="28.[129,194,1713,1737]" captionTargetBox="[165,1433,153,1679]" captionTargetId="figure-95@28.[161,1441,144,1685]" captionTargetPageId="28" captionText="Figure 16. Preserved specimens of Phyllonastes dicaprioi. Lateral, dorsal, and ventral views of: A, holotype DHMECN 13729, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm." figureDoi="http://doi.org/10.5281/zenodo.14832731" httpUri="https://zenodo.org/record/14832731/files/figure.png" pageId="27" pageNumber="28">Fig. 16</figureCitation>
|
||
):
|
||
</emphasis>
|
||
All from
|
||
<collectingCountry id="F36B7601FFA9FFD3C588C6F6909F4A3D" box="[1188,1274,233,257]" name="Ecuador" pageId="27" pageNumber="28">Ecuador</collectingCountry>
|
||
,
|
||
<collectingRegion id="49B8F873FFA9FFD3C429C6F691D54A3D" box="[1285,1456,233,257]" country="Ecuador" name="El Oro" pageId="27" pageNumber="28">El Oro Province</collectingRegion>
|
||
:
|
||
<collectorName id="26895347FFA9FFD3C200C717905F4A63" pageId="27" pageNumber="28">
|
||
El Retiro,
|
||
<specimenCode id="DBDA9EEAFFA9FFD3C2BBC71790304A1C" box="[919,1109,264,288]" collectionCode="DHMECN" pageId="27" pageNumber="28">DHMECN 13723</specimenCode>
|
||
,
|
||
<specimenCode id="DBDA9EEAFFA9FFD3C54DC71791784A1C" box="[1121,1309,264,288]" collectionCode="DHMECN" pageId="27" pageNumber="28">DHMECN 13730</specimenCode>
|
||
<specimenCount id="9D7AFD18FFA9FFD3C409C71791D44A1C" box="[1317,1457,264,288]" pageId="27" pageNumber="28" type="adult">adult females</specimenCount>
|
||
;
|
||
<materialsCitation id="3B143CCCFFA9FFD3C206C738905B4A63" collectingDate="2017-07-03" collectionCode="DHMECN" collectorName="El Retiro" country="Ecuador" elevation="1330" latitude="-3.61847" location="All" longLatPrecision="1" longitude="-79.58072" pageId="27" pageNumber="28" specimenCode="DHMECN 13724-26, DHMECN 13728" specimenCount="1" specimenCount-adult="25" stateProvince="El Oro" typeStatus="paratype">
|
||
<specimenCode id="DBDA9EEAFFA9FFD3C206C73890684A03" box="[810,1037,295,319]" collectionCode="DHMECN" pageId="27" pageNumber="28">DHMECN 13724–26</specimenCode>
|
||
,
|
||
<specimenCode id="DBDA9EEAFFA9FFD3C53BC73890B44A03" box="[1047,1233,295,319]" collectionCode="DHMECN" pageId="27" pageNumber="28">DHMECN 13728</specimenCode>
|
||
<specimenCount id="9D7AFD18FFA9FFD3C5FBC73891284A03" box="[1239,1357,295,319]" pageId="27" pageNumber="28" type="adult">adult males</specimenCount>
|
||
; collected with the holotype. Sinsao,
|
||
</materialsCitation>
|
||
</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B143CCCFFA9FFD3C56BC75891D64A42" collectionCode="DHMECN" collectorName="El Retiro" country="Ecuador" elevation="1330" latitude="-3.61847" location="All" longLatPrecision="1" longitude="-79.58072" pageId="27" pageNumber="28" specimenCode="DHMECN 12527" specimenCount="26" specimenCount-adult="25" stateProvince="El Oro" typeStatus="paratype">
|
||
<collectionCode id="ED6DAE54FFA9FFD3C56BC75890DF4A63" box="[1095,1210,327,351]" pageId="27" pageNumber="28">DHMECN</collectionCode>
|
||
12524–
|
||
<specimenCount id="9D7AFD18FFA9FFD3C43FC759910C4A63" box="[1299,1385,326,351]" count="25" pageId="27" pageNumber="28" type="adult">25 adult</specimenCount>
|
||
males;
|
||
<specimenCode id="DBDA9EEAFFA9FFD3C206C77997864A42" box="[810,995,358,382]" collectionCode="DHMECN" pageId="27" pageNumber="28">DHMECN 12527</specimenCode>
|
||
<specimenCount id="9D7AFD18FFA9FFD3C2CAC779904C4A42" box="[998,1065,358,382]" pageId="27" pageNumber="28" type="female">female</specimenCount>
|
||
(
|
||
<geoCoordinate id="EE485056FFA9FFD3C518C77990C74A42" box="[1076,1186,358,382]" degrees="3.61847" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="1" value="-3.61847">3.61847°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE485056FFA9FFD3C585C77991484A42" box="[1193,1325,358,382]" degrees="79.58072" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="1" value="-79.58072">79.58072°W</geoCoordinate>
|
||
)
|
||
<elevation id="0051D1A2FFA9FFD3C414C77991D64A42" box="[1336,1459,358,382]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="27" pageNumber="28" unit="m" value="1330.0">
|
||
<quantity id="4C849B74FFA9FFD3C414C77991E14A41" box="[1336,1412,358,382]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="27" pageNumber="28" unit="m" value="1330.0">1330 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA9FFD3C206C78097F4492C" pageId="27" pageNumber="28" type="vernacular_names">
|
||
<paragraph id="8BC33691FFA9FFD3C206C78091094A8B" blockId="27.[810,1388,415,439]" box="[810,1388,415,439]" pageId="27" pageNumber="28">
|
||
<emphasis id="B908EA83FFA9FFD3C206C780900E4A8B" box="[810,1131,415,439]" italics="true" pageId="27" pageNumber="28">Proposed standard English name:</emphasis>
|
||
DiCaprio’s leaflitter frog.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA9FFD3C206C7C697F4492C" blockId="27.[810,1459,473,528]" pageId="27" pageNumber="28">
|
||
<emphasis id="B908EA83FFA9FFD3C206C7C690FD4ACC" box="[810,1176,473,497]" italics="true" pageId="27" pageNumber="28">Proposed standard Spanish name:</emphasis>
|
||
Cutín de Hojarasca de DiCaprio.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA9FFD3C206C42E90054D08" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8BC33691FFA9FFD3C206C42E90054D08" blockId="27.[809,1461,561,1588]" pageId="27" pageNumber="28">
|
||
<emphasis id="B908EA83FFA9FFD3C206C42E97EB4975" box="[810,910,561,585]" italics="true" pageId="27" pageNumber="28">Definition</emphasis>
|
||
(
|
||
<figureCitation id="13472A14FFA9FFD3C28FC42E97944975" box="[931,1009,561,585]" captionStart="Figure 16" captionStartId="28.[129,194,1713,1737]" captionTargetBox="[165,1433,153,1679]" captionTargetId="figure-95@28.[161,1441,144,1685]" captionTargetPageId="28" captionText="Figure 16. Preserved specimens of Phyllonastes dicaprioi. Lateral, dorsal, and ventral views of: A, holotype DHMECN 13729, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm." figureDoi="http://doi.org/10.5281/zenodo.14832731" httpUri="https://zenodo.org/record/14832731/files/figure.png" pageId="27" pageNumber="28">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="13472A14FFA9FFD3C52CC42E907F4975" box="[1024,1050,561,585]" captionStart="Figure 17" captionStartId="29.[112,177,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-482@29.[306,1266,144,1064]" captionTargetPageId="29" captionText="Figure 17. Palmar (A) and plantar (B) surfaces of Phyllonastes dicaprioi. Photographs of right hand and left foot of the holotype DHMECN 13729." figureDoi="http://doi.org/10.5281/zenodo.14832735" httpUri="https://zenodo.org/record/14832735/files/figure.png" pageId="27" pageNumber="28">17</figureCitation>
|
||
;
|
||
<tableCitation id="C6FE032AFFA9FFD3C500C42E90E34975" box="[1068,1158,561,585]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="27" pageNumber="28" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="C6FE032AFFA9FFD3C593C42E90A94975" box="[1215,1228,561,585]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="27" pageNumber="28" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
|
||
<emphasis id="B908EA83FFA9FFD3C5E1C42D90BA4975" box="[1229,1247,562,585]" italics="true" pageId="27" pageNumber="28">):</emphasis>
|
||
We assign the new species to the genus
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C52FC44E901A4955" authorityName="Heyer" authorityYear="1977" box="[1027,1151,593,617]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FFA9FFD3C52FC44E901A4955" box="[1027,1151,593,617]" italics="true" pageId="27" pageNumber="28">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
based on its phylogenetic relationships. The new species is characterized by: (1) skin on head, dorsum and flanks shagreen, dorsolateral folds absent, dorsal limb skin smooth with dispersed low warts, ventral skin and posterior surface of legs areolate, discoidal fold present; (2) tympanic membrane undifferentiated, tympanic annulus present but weakly defined, small supratympanic fold present, row of postrictal tubercles present; (3) snout rounded in dorsal and lateral views; (4) upper eyelid without tubercles, cranial crests absent; (5) vomerine teeth absent; (6) vocal slits and nuptial pads absent; (7) fingers not expanded distally, finger tips acuminate without papillae, finger I shorter than finger II, supernumerary tubercles present; (8) fingers bearing narrow lateral fringes, well-defined pads without circumferential grooves; (9) distal phalanges blunt or T-shaped, phalangeal formula of hand 2-2-3-3; (10) ulnar tubercles usually present; (11) knees and heels without tubercles, outer edge of tarsus without tubercles, inner edge of tarsus bearing one conical tubercle; (12) inner metatarsal tubercle large, elongate in ventral view and rounded in lateral view, bigger than conical outer metatarsal tubercle; (13) toes expanded distally, with acuminate toe tips and lacking papillae; supernumerary tubercles, lateral fringes, and toe basal webbing absent; all toes bearing ill-defined circumferential grooves, toe V shorter than toe III; (14) in life, dorsum brown, with two suprainguinal spots, dark or light middorsal stripe and dark marks and chevrons in some specimens; dark mask (but faint in some specimens), extending onto darker flanks; arms brown, with dark marks usually forming bands; ventral surfaces of body and legs light brown, with small cream flecks and dots; and (15) SVL in adult males
|
||
<quantity id="4C849B74FFA9FFD3C2E2C3E2905D4D28" box="[974,1080,1533,1557]" metricMagnitude="-2" metricUnit="m" metricValue="1.262" pageId="27" pageNumber="28" unit="mm" value="12.62">12.62 mm</quantity>
|
||
(
|
||
<emphasis id="B908EA83FFA9FFD3C560C3E2903A4D28" box="[1100,1119,1533,1556]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
= 7) and SVL in adult females
|
||
<quantity id="4C849B74FFA9FFD3C206C00397F04D0F" box="[810,917,1564,1588]" metricMagnitude="-2" metricUnit="m" metricValue="1.293" pageId="27" pageNumber="28" unit="mm" value="12.93">12.93 mm</quantity>
|
||
(
|
||
<emphasis id="B908EA83FFA9FFD3C28AC00297DC4D08" box="[934,953,1565,1588]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
= 3) (
|
||
<tableCitation id="C6FE032AFFA9FFD3C52EC003902A4D08" box="[1026,1103,1564,1588]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="27" pageNumber="28" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA9FFD4C206C04990914C9A" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="diagnosis">
|
||
<paragraph id="8BC33691FFA9FFD4C206C04990914C9A" blockId="27.[810,1465,1621,1990]" lastBlockId="28.[825,1474,1841,1959]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
<emphasis id="B908EA83FFA9FFD3C206C04990FE4D51" box="[810,1179,1621,1645]" italics="true" pageId="27" pageNumber="28">
|
||
Diagnosis:
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C283C04A90FE4D51" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[943,1179,1621,1645]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="dicaprioi">Phyllonastes dicaprioi</taxonomicName>
|
||
</emphasis>
|
||
resembles the other
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C206C06A97C44DB1" authorityName="Heyer" authorityYear="1977" box="[810,929,1653,1677]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FFA9FFD3C206C06A97C44DB1" box="[810,929,1653,1677]" italics="true" pageId="27" pageNumber="28">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
from the western foothills of Ecuadorian Andes,
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C488C06A97AE4D90" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA9FFD3C488C06A97AE4D90" italics="true" pageId="27" pageNumber="28">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C2FAC08A90574D90" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" box="[982,1074,1684,1708]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA9FFD3C2FAC08A90574D90" box="[982,1074,1684,1708]" italics="true" pageId="27" pageNumber="28">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C511C08A90F74D90" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[1085,1170,1684,1708]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA9FFD3C511C08A90F74D90" box="[1085,1170,1684,1708]" italics="true" pageId="27" pageNumber="28">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C5E6C08A91514D90" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[1226,1332,1684,1708]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA9FFD3C5E6C08A91514D90" box="[1226,1332,1684,1708]" italics="true" pageId="27" pageNumber="28">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
, by having a tympanic annulus, supernumerary palmar tubercles, fingers not expanded distally, and circumferential grooves on toes. It differs from all of them by lacking vocal slits. Besides,
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C435C0EC91ED4C36" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1305,1416,1778,1802]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA9FFD3C435C0EC91ED4C36" box="[1305,1416,1778,1802]" italics="true" pageId="27" pageNumber="28">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C2BCC10D902F4C15" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[912,1098,1809,1833]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA9FFD3C2BCC10D902F4C15" box="[912,1098,1809,1833]" italics="true" pageId="27" pageNumber="28">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
by having a round snout in dorsal view (snout truncate in dorsal view in
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C5EAC12E91E44C75" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1222,1409,1841,1865]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA9FFD3C5EAC12E91E44C75" box="[1222,1409,1841,1865]" italics="true" pageId="27" pageNumber="28">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
), by having a supratympanic fold (absent in
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C5E8C14E911E4C54" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1220,1403,1872,1896]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA9FFD3C5E8C14E911E4C54" box="[1220,1403,1872,1896]" italics="true" pageId="27" pageNumber="28">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
) and three phalanges in finger IV (finger IV with two phalanges in
|
||
<taxonomicName id="4C7C4D12FFA9FFD3C488C16F97AE4C9B" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA9FFD3C488C16F97AE4C9B" italics="true" pageId="27" pageNumber="28">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
), and by the absence of circumferential grooves in fingers and supernumerary plantar tubercles (both present in
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C1B0C12D95304C75" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[156,341,1841,1865]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFAEFFD4C1B0C12D95304C75" box="[156,341,1841,1865]" italics="true" pageId="28" pageNumber="29">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C041C12D96254C75" box="[365,576,1841,1865]" class="Amphibia" family="Craugastoridae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFAEFFD4C041C12D96254C75" box="[365,576,1841,1865]" italics="true" pageId="28" pageNumber="29">Pristimantis dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C3D5C12D94A34C54" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFAEFFD4C3D5C12D94A34C54" italics="true" pageId="28" pageNumber="29">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
by having toes distally expanded (slightly expanded in
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C3D5C14E94A34CBB" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFAEFFD4C3D5C14E94A34CBB" italics="true" pageId="28" pageNumber="29">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C03FC16F95034CBB" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[275,358,1903,1927]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFAEFFD4C03FC16F95034CBB" box="[275,358,1903,1927]" italics="true" pageId="28" pageNumber="29">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
by having acuminate toe tips (rounded in
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C1ACC18F94B34C9B" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[128,214,1935,1959]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFAEFFD4C1ACC18F94B34C9B" box="[128,214,1935,1959]" italics="true" pageId="28" pageNumber="29">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
), and from
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C07BC18F95A64C9B" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[343,451,1935,1959]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFAEFFD4C07BC18F95A64C9B" box="[343,451,1935,1959]" italics="true" pageId="28" pageNumber="29">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
by the absence supernumerary plantar tubercles (present in
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C547C12D90BD4C75" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[1131,1240,1841,1865]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFAEFFD4C547C12D90BD4C75" box="[1131,1240,1841,1865]" italics="true" pageId="28" pageNumber="29">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
) and by the mask extending to the groins (mask extending to flank half in
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C46BC14E91CA4C54" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[1351,1455,1872,1896]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFAEFFD4C46BC14E91CA4C54" box="[1351,1455,1872,1896]" italics="true" pageId="28" pageNumber="29">P.worleyae</emphasis>
|
||
</taxonomicName>
|
||
). For a comparison with other
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C547C17091544CBB" box="[1131,1329,1903,1927]" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EA83FFAEFFD4C547C17090874CBB" box="[1131,1250,1903,1927]" italics="true" pageId="28" pageNumber="29">Phyllonastes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
that are more phylogenetically distant, see
|
||
<tableCitation id="C6FE032AFFAEFFD4C571C19090D44C9B" box="[1117,1201,1935,1959]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="28" pageNumber="29" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="C6FE032AFFAEFFD4C5CDC190908B4C9B" box="[1249,1262,1935,1959]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="28" pageNumber="29" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF036619FFAEFFD4C1ADC0AE969E4C3C" ID-DOI="http://doi.org/10.5281/zenodo.14832731" ID-Zenodo-Dep="14832731" httpUri="https://zenodo.org/record/14832731/files/figure.png" pageId="28" pageNumber="29" startId="28.[129,194,1713,1737]" targetBox="[165,1433,153,1679]" targetPageId="28" targetType="figure">
|
||
<paragraph id="8BC33691FFAEFFD4C1ADC0AE969E4C3C" blockId="28.[129,1467,1713,1793]" pageId="28" pageNumber="29">
|
||
<emphasis id="B908EA83FFAEFFD4C1ADC0AE94804DF5" bold="true" box="[129,229,1713,1737]" pageId="28" pageNumber="29">Figure 16.</emphasis>
|
||
Preserved specimens of
|
||
<taxonomicName id="4C7C4D12FFAEFFD4C0E0C0AE96C64DF5" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[460,675,1713,1737]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFAEFFD4C0E0C0AE96C64DF5" bold="true" box="[460,675,1713,1737]" italics="true" pageId="28" pageNumber="29">Phyllonastes dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
. Lateral, dorsal, and ventral views of: A, holotype
|
||
<emphasis id="B908EA83FFAEFFD4C555C0AE91474DF5" box="[1145,1314,1713,1737]" italics="true" pageId="28" pageNumber="29">DHMECN 13729</emphasis>
|
||
, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF036619FFAFFFD5C15CC25C94D64F4B" ID-DOI="http://doi.org/10.5281/zenodo.14832735" ID-Zenodo-Dep="14832735" httpUri="https://zenodo.org/record/14832735/files/figure.png" pageId="29" pageNumber="30" startId="29.[112,177,1091,1115]" targetBox="[306,1266,144,1064]" targetPageId="29" targetType="figure">
|
||
<paragraph id="8BC33691FFAFFFD5C15CC25C94D64F4B" blockId="29.[112,1434,1091,1144]" pageId="29" pageNumber="30">
|
||
<emphasis id="B908EA83FFAFFFD5C15CC25C94B14F67" bold="true" box="[112,212,1091,1115]" pageId="29" pageNumber="30">Figure 17.</emphasis>
|
||
Palmar (A) and plantar (B) surfaces of
|
||
<taxonomicName id="4C7C4D12FFAFFFD5C364C25C977A4F67" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[584,799,1091,1115]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFAFFFD5C364C25C977A4F67" bold="true" box="[584,799,1091,1115]" italics="true" pageId="29" pageNumber="30">Phyllonastes dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
. Photographs of right hand and left foot of the holotype
|
||
<emphasis id="B908EA83FFAFFFD5C41EC25B94C84F44" italics="true" pageId="29" pageNumber="30">DHMECN 13729</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C366651AFFAFFFD6C15DC2B795F44889" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
|
||
<paragraph id="8BC33691FFAFFFD5C15DC2B794944E60" blockId="29.[112,763,1191,1968]" pageId="29" pageNumber="30">
|
||
<emphasis id="B908EA83FFAFFFD5C15DC2B795954F83" box="[113,496,1191,1216]" italics="true" pageId="29" pageNumber="30">
|
||
Description of
|
||
<typeStatus id="54C78833FFAFFFD5C029C2B895324F83" box="[261,343,1191,1215]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13472A14FFAFFFD5C040C2B795D04F83" box="[364,437,1191,1215]" captionStart="Figure 16" captionStartId="28.[129,194,1713,1737]" captionTargetBox="[165,1433,153,1679]" captionTargetId="figure-95@28.[161,1441,144,1685]" captionTargetPageId="28" captionText="Figure 16. Preserved specimens of Phyllonastes dicaprioi. Lateral, dorsal, and ventral views of: A, holotype DHMECN 13729, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm." figureDoi="http://doi.org/10.5281/zenodo.14832731" httpUri="https://zenodo.org/record/14832731/files/figure.png" pageId="29" pageNumber="30">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation id="13472A14FFAFFFD5C0EFC2B795B84FFC" box="[451,477,1192,1216]" captionStart="Figure 17" captionStartId="29.[112,177,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-482@29.[306,1266,144,1064]" captionTargetPageId="29" captionText="Figure 17. Palmar (A) and plantar (B) surfaces of Phyllonastes dicaprioi. Photographs of right hand and left foot of the holotype DHMECN 13729." figureDoi="http://doi.org/10.5281/zenodo.14832735" httpUri="https://zenodo.org/record/14832735/files/figure.png" pageId="29" pageNumber="30">17</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Adult male (
|
||
<emphasis id="B908EA83FFAFFFD5C3A6C2B794D74FE3" italics="true" pageId="29" pageNumber="30">DHMECN 13729</emphasis>
|
||
). Measurements (in millimetres): SVL, 12.9; tibia length, 5.5; foot length, 4.9; head length, 3.0; head width, 3.5; eye diameter, 1.5; tympanum diameter, 0.5; interorbital distance, 1.3; upper eyelid width, 0.8; internarial distance, 1.3; eye–nostril distance 0.8.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFAFFFD5C1A1C37C95AA4D4A" blockId="29.[112,763,1191,1968]" pageId="29" pageNumber="30">Head wider than long, head slightly narrower than body; canthus rostralis weakly defined; loreal region slightly concave in dorsal view; cranial crests absent; upper eyelid bearing no tubercles. Tympanic annulus weakly defined, differentiated in its anterior and inferior portions; tympanic membrane undifferentiated from the surrounding skin; small supratympanic fold present; postrictal tubercles present. Snout round in dorsal and lateral views, without rostral papilla. Vomerine teeth absent; vocal slits and nuptial pads absent.</paragraph>
|
||
<paragraph id="8BC33691FFAFFFD5C1A1C06291584FE3" blockId="29.[112,763,1191,1968]" lastBlockId="29.[809,1461,1191,1686]" pageId="29" pageNumber="30">Skin on head, dorsum, and flanks shagreen, limbs smooth dorsally, with dispersed low warts; skin on posterior surfaces of legs and venter areolate; dorsolateral folds absent; discoidal fold present. Ulnar tubercle small and low; palmar tubercles prominent, outer palmar tubercle slightly elongate, thenar tubercle large and elongate, both rounded in lateral view; subarticular tubercles well defined, round in ventral and lateral views; distal subarticular tubercles not visible; supernumerary tubercles visible; thin lateral dermal fringes present, discs bearing well-defined pads without circumferential grooves, fingers not expanded distally, tip of fingers acuminate, without papillae; relative length of fingers I <II <IV <III; phalangeal formula of 2-2-3-3.</paragraph>
|
||
<paragraph id="8BC33691FFAFFFD5C269C2F991C74DAA" blockId="29.[809,1461,1191,1686]" pageId="29" pageNumber="30">Hindlimbs robust; heel and knee without tubercles; tarsal folds (inner and outer) absent, outer tarsal tubercles absent; inner tarsal tubercle present, prominent, and conical; inner metatarsal tubercle large, elongate in ventral view, rounded in lateral view; outer metatarsal tubercle small, well defined, prominent, rounded in ventral view, conical in lateral view; supernumerary plantar tubercles absent; subarticular tubercles well defined, round and prominent in dorsal view, toes without lateral fringes; basal webbing absent; discs rounded and bearing well-defined rounded pads; circumferential grooves present, ill defined; toes expanded distally, tip of toes acuminate without papillae; relative length of toes is I <II <V <III <IV; toe III longer than toe V (toe III surpasses the distal border of the second subarticular tubercle of toe IV; toe V reaches the proximal border of the second tubercle of toe IV).</paragraph>
|
||
<paragraph id="8BC33691FFAFFFD6C206C0A896884A19" blockId="29.[809,1461,1719,1962]" lastBlockId="30.[128,779,144,293]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<emphasis id="B908EA83FFAFFFD5C206C0A890474DF3" box="[810,1058,1719,1743]" italics="true" pageId="29" pageNumber="30">
|
||
Colour of
|
||
<typeStatus id="54C78833FFAFFFD5C2A0C0A897BB4DF3" box="[908,990,1719,1743]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
in life:
|
||
</emphasis>
|
||
Dorsum brown, with darker, scattered marks and chevrons, two suprainguinal spots and a middorsal cream stripe extending to the vent. Cloacal region dark brown, with a cream stripe extending towards each hindlimb along the posterior surface of the thigh, ventral surface of shank, and posterior surface of pes, ending at the inner metatarsal tubercle. Faint dark mask present, extending from the tip of the snout to the groins, bordering dorsally dark flanks.Lips with dark vertical bars. Arms and legs brown, with dark marks usually forming bands. Ventral surfaces of body and legs light brown, with small cream flecks and dots. A mid-ventral pale stripe extends from the throat to the vent; a pale stripe extends ventrally along each hindlimb, joining the midventral pale stripe to form a cross on the chest.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFACFFD6C1ADC75094DC4ABB" blockId="30.[129,778,335,391]" pageId="30" pageNumber="31">
|
||
<emphasis id="B908EA83FFACFFD6C1ADC75095AE4A5B" box="[129,459,335,359]" italics="true" pageId="30" pageNumber="31">
|
||
Colour of
|
||
<typeStatus id="54C78833FFACFFD6C1CDC75095564A5B" box="[225,307,335,359]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
|
||
in preservative:
|
||
</emphasis>
|
||
Similar to coloration in life but paler.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFACFFD6C1ADC7B695F44889" blockId="30.[127,779,424,949]" pageId="30" pageNumber="31">
|
||
<emphasis id="B908EA83FFACFFD6C1ADC7B695234A83" box="[129,326,424,448]" italics="true" pageId="30" pageNumber="31">
|
||
Variation (
|
||
<figureCitation id="13472A14FFACFFD6C1C3C7B695564AFC" box="[239,307,424,448]" captionStart="Figure 16" captionStartId="28.[129,194,1713,1737]" captionTargetBox="[165,1433,153,1679]" captionTargetId="figure-95@28.[161,1441,144,1685]" captionTargetPageId="28" captionText="Figure 16. Preserved specimens of Phyllonastes dicaprioi. Lateral, dorsal, and ventral views of: A, holotype DHMECN 13729, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm." figureDoi="http://doi.org/10.5281/zenodo.14832731" httpUri="https://zenodo.org/record/14832731/files/figure.png" pageId="30" pageNumber="31">Fig. 16</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<specimenCount id="9D7AFD18FFACFFD6C07AC7B7959F4AFC" box="[342,506,424,448]" count="4" pageId="30" pageNumber="31" type="generic">Four specimens</specimenCount>
|
||
have an almost unnoticeable tympanic membrane and annulus, on one or both sides (DHMECN 12524–5 and 13724–25). Ulnar tubercles absent in all
|
||
<typeStatus id="54C78833FFACFFD6C192C41995464922" box="[190,291,518,542]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
|
||
.
|
||
<specimenCount id="9D7AFD18FFACFFD6C003C41995BB4922" box="[303,478,518,542]" count="3" pageId="30" pageNumber="31" type="generic">Three specimens</specimenCount>
|
||
have the dorsum paler than the
|
||
<typeStatus id="54C78833FFACFFD6C183C43A95684901" box="[175,269,549,573]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
|
||
, without dark marks except for suprainguinal spots (DHMECN 13723, 13726, and 12524). One individual is similar to the
|
||
<typeStatus id="54C78833FFACFFD6C00AC47B95E74940" box="[294,386,612,636]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
|
||
in having a pale mid-dorsal stripe but lacks dark marks, except for suprainguinal spots.
|
||
<specimenCount id="9D7AFD18FFACFFD6C341C49B976E49A7" box="[621,779,643,667]" count="2" pageId="30" pageNumber="31" type="generic">Two specimens</specimenCount>
|
||
have a darker brown dorsum than the
|
||
<typeStatus id="54C78833FFACFFD6C32BC4BC96064987" box="[519,611,675,699]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
|
||
and a faint dark mid-dorsal stripe and inverted-V chevrons (DHMECN 12527 and 13720).
|
||
<specimenCount id="9D7AFD18FFACFFD6C026C4FE95C449C5" box="[266,417,737,761]" count="1" pageId="30" pageNumber="31" type="generic">One specimen</specimenCount>
|
||
has a dark hourglass-shaped large mark on the dorsum DHMECN 13725.
|
||
<specimenCount id="9D7AFD18FFACFFD6C33DC51E96C64825" box="[529,675,769,793]" count="1" pageId="30" pageNumber="31" type="generic">One specimen</specimenCount>
|
||
has a faint midventral pale stripe, but no stripes on ventral surfaces of the arms (DHMECN 13728).
|
||
<specimenCount id="9D7AFD18FFACFFD6C0BEC55F9655486B" box="[402,560,831,855]" count="2" pageId="30" pageNumber="31" type="generic">Two specimens</specimenCount>
|
||
have faint pale stripes along hindlegs (DHMECN 13724 and 13728). Midventral and hindleg stripes are absent in all
|
||
<typeStatus id="54C78833FFACFFD6C0E1C561965748AA" box="[461,562,894,918]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
|
||
. Morphometric variation is detailed in
|
||
<tableCitation id="C6FE032AFFACFFD6C06CC58295EE4889" box="[320,395,925,949]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="30" pageNumber="31" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFACFFD6C1ADC5C896534FF6" pageId="30" pageNumber="31" type="distribution">
|
||
<paragraph id="8BC33691FFACFFD6C1ADC5C896534FF6" blockId="30.[128,778,983,1226]" pageId="30" pageNumber="31">
|
||
<emphasis id="B908EA83FFACFFD6C1ADC5C895394F32" italics="true" pageId="30" pageNumber="31">
|
||
Distribution, natural history, and conservation status (
|
||
<figureCitation id="13472A14FFACFFD6C396C5C8969248D3" box="[698,759,983,1007]" captionStart="Figure 3" captionStartId="9.[113,175,1807,1831]" captionTargetBox="[149,1423,152,1776]" captionTargetId="figure-7@9.[146,1426,149,1779]" captionTargetPageId="9" captionText="Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed." figureDoi="http://doi.org/10.5281/zenodo.14832695" httpUri="https://zenodo.org/record/14832695/files/figure.png" pageId="30" pageNumber="31">Fig. 3</figureCitation>
|
||
):
|
||
<taxonomicName id="4C7C4D12FFACFFD6C1ADC5E995394F32" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[129,348,1014,1038]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="dicaprioi">Phyllonastes dicaprioi</taxonomicName>
|
||
</emphasis>
|
||
is known from the Western Montane Forest of the province
|
||
<collectingRegion id="49B8F873FFACFFD6C049C20A95CE4F11" box="[357,427,1045,1069]" country="Ecuador" name="El Oro" pageId="30" pageNumber="31">El Oro</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36B7601FFACFFD6C098C20A966E4F11" box="[436,523,1045,1069]" name="Ecuador" pageId="30" pageNumber="31">Ecuador</collectingCountry>
|
||
, between 1330 and
|
||
<quantity id="4C849B74FFACFFD6C3F9C20A94F34F70" metricMagnitude="3" metricUnit="m" metricValue="1.705" pageId="30" pageNumber="31" unit="m" value="1705.0">1705 m</quantity>
|
||
a.s.l. Individuals were collected at night on the ground, buried in leaf litter. Because of the lack of information on population size and geographical range, we suggest assigning
|
||
<taxonomicName id="4C7C4D12FFACFFD6C3ADC26B968A4FB7" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[641,751,1139,1163]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFACFFD6C3ADC26B968A4FB7" box="[641,751,1139,1163]" italics="true" pageId="30" pageNumber="31">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
to the Data Deficient IUCN Red List Category (based on
|
||
<bibRefCitation id="EFED4B60FFACFFD6C3EBC28C96404FF6" author="IUCN Standards and Petitions Committee & Using the IUCN Red List Categories" pageId="30" pageNumber="31" refId="ref39535" refString="IUCN Standards and Petitions Committee. Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee, 2023. https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (25 January 2024, date last accessed)." type="url" year="2023">IUCN Standards and Petitions Committee 2023</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFACFFD6C1ADC2F496A54EBD" pageId="30" pageNumber="31" type="etymology">
|
||
<paragraph id="8BC33691FFACFFD6C1ADC2F496A54EBD" blockId="30.[128,778,1259,1409]" pageId="30" pageNumber="31">
|
||
<emphasis id="B908EA83FFACFFD6C1ADC2F494954E3F" box="[129,240,1259,1283]" italics="true" pageId="30" pageNumber="31">Etymology:</emphasis>
|
||
The specific name
|
||
<taxonomicName id="4C7C4D12FFACFFD6C098C2F4966F4E3F" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[436,522,1259,1283]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFACFFD6C098C2F4966F4E3F" box="[436,522,1259,1283]" italics="true" pageId="30" pageNumber="31">dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
is a patronym honouring Leonardo DiCaprio, actor, film producer, and conservationist. He has voiced his support for several conservation initiatives in
|
||
<collectingCountry id="F36B7601FFACFFD6C1AEC35694B94E5D" box="[130,220,1353,1377]" name="Ecuador" pageId="30" pageNumber="31">Ecuador</collectingCountry>
|
||
; and in 2021, he pledged funds for the conservation and ecosystem restoration of the
|
||
<collectingRegion id="49B8F873FFACFFD6C08FC37696394EBD" box="[419,604,1385,1409]" country="Ecuador" name="Galapagos" pageId="30" pageNumber="31">Galapagos Islands</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36B7601FFACFFD6C349C37696D84EBD" box="[613,701,1385,1409]" name="Ecuador" pageId="30" pageNumber="31">Ecuador</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |