treatments-xml/data/03/D5/87/03D58787FFA3FFDBC2C1C3D3956F4D05.xml

<document id="A3C795467169C62B6BD38F38373832B7" ID-DOI="10.1093/zoolinnean/zlae162" ID-ISSN="0024-4082" ID-Zenodo-Dep="14832689" ID-ZooBank="7-01-2184" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1738158242914" checkinUser="plazi" docAuthor="Ortega, Jhael A., Cisneros-Heredia, Diego F., Camper, Jeffrey D., Romero-Carvajal, Andrés, Negrete, Leonardo &amp; Ron, Santiago R." docDate="2025" docId="03D58787FFA3FFDBC2C1C3D3956F4D05" docLanguage="en" docName="zlae162.pdf" docOrigin="Zoological Journal of the Linnean Society 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae162" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Phyllonastes Heyer 1977" docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="FFECFFFFFFB2FFC8C12CC61F94654B3C" masterDocTitle="Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography" masterLastPageNumber="60" masterPageNumber="1" pageNumber="18" updateTime="1738929907134" updateUser="ExternalLinkService">
<mods:mods id="E5AB48B97B17CE50FAA4B3351985C975" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C0ABE8E72E51F0D2E0779EAA828EA495">
<mods:title id="7C931A6D8E7D2E51DAEDF82303F96080">Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography</mods:title>
</mods:titleInfo>
<mods:name id="DDC7741B4395BCE9D3AF694F29E2A68C" type="personal">
<mods:role id="0794EDEAE5FB019E9E8856CEB34A6C3D">
<mods:roleTerm id="1A0D5449C25EAF4C78BEFAE4AE1ACECA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5409381179A058BE9E772016A4428C2">Ortega, Jhael A.</mods:namePart>
<mods:affiliation id="D75849A90EAC3D76FB733ECB1CC72BDA">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name id="E48C8E1CC82A4C7269650F939AA64721" type="personal">
<mods:role id="E8F825631DEA475383FD1B71CB1BA822">
<mods:roleTerm id="4DAA68FE22C3DA84F73E417D4F8AB3AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4B990291FD8B7E1D929530C26709E95D">Cisneros-Heredia, Diego F.</mods:namePart>
<mods:affiliation id="603E3A4577C146F2781D43FA427FC2F0">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Museo de Zoología / Laboratorio de Zoología Terrestre, Quito, Ecuador &amp; Instituto Nacional de Biodiversidad INABIO, División de Herpetología, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name id="E90500807B96781BD0316829D5A0473A" type="personal">
<mods:role id="D3E40B2DC829427C3B4759A3B05FB62E">
<mods:roleTerm id="04AE573C56347244C159BF13938FEA27">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A80B1F1F428E246EEB492E9442CCC8AD">Camper, Jeffrey D.</mods:namePart>
<mods:affiliation id="2E18DF5C972300C50803F4F18CE76330">Department of Biology, Francis Marion University, Florence, SC 29506, USA</mods:affiliation>
</mods:name>
<mods:name id="10255280F660DEEF5C4D8B94A36E3683" type="personal">
<mods:role id="D610661000252F6F218C05C6A10E4E11">
<mods:roleTerm id="09290A3E42B9FBC5C4F1623F8DDDEE19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3B6667AAD62D302CDC29A2F47E1CE65">Romero-Carvajal, Andrés</mods:namePart>
<mods:affiliation id="95686D2EC59082752EC57B367542DD4C">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name id="E665014052BC0C618BFFD5FA0D14F8F7" type="personal">
<mods:role id="DBBBE01ED070E8C62B484F5412A90B32">
<mods:roleTerm id="C15F4B70B91A3EAEDB5F2E39D6BD7A40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6FFFDBDA7986EA91A8DD4E4710DEF86">Negrete, Leonardo</mods:namePart>
<mods:affiliation id="EE5C4D9C60A4F93E2B012FDC2C160AFD">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name id="02EB83361091868D906549DD4C12B8B5" type="personal">
<mods:role id="B3E74C3E6CF6967BA7655FCCF17583F2">
<mods:roleTerm id="F332EA9BF53D2B07A5254423D6767190">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC893266404090AA51D89D934E74A1CE">Ron, Santiago R.</mods:namePart>
<mods:affiliation id="8203AC0C205242ED3EFE90C83D3537A2">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
<mods:nameIdentifier id="3BD5379E6A69CD8C0EF5B17BB354CA33" type="email">santiago.r.ron@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A2D45EA9585274CDA0E7AB537C62F32A">text</mods:typeOfResource>
<mods:relatedItem id="CE3A1A23D11361EF993CEF39328862E2" type="host">
<mods:titleInfo id="A68932445DED0ACCCC20D6125AB141BA">
<mods:title id="00B6865535A08E1A84BD161EA3848C0A">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="3DE57AD492E2C7E481431C61A0BC8478">
<mods:date id="8D54846F7E2AF420C0D3D7C643AD4EBD">2025</mods:date>
<mods:detail id="475946B44E49CF8AEDECC802E0F24305" type="pubDate">
<mods:number id="197E05866666491744F6AE4CCA949088">2025-01-13</mods:number>
</mods:detail>
<mods:detail id="E52238A0235A00B853F7127665415405" type="volume">
<mods:number id="432CE8C97D870235D5A5EB07264B5D31">203</mods:number>
</mods:detail>
<mods:detail id="98BC1E779FA3655C374E1B5517777C51" type="issue">
<mods:number id="89DC1205960EBECE0E5A26C555055792">1</mods:number>
</mods:detail>
<mods:extent id="60A7FDC145B39723DF4D6CAF794B75C5" unit="page">
<mods:start id="B0646A6D0C8C70341D9C7C8E10DD62A2">1</mods:start>
<mods:end id="15868EFCC93A3AB538BA967201258EB8">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="7498E4A78A608084E02037E942D3D39B">
<mods:url id="838760A7D19DF2486550A9CC70975315">https://doi.org/10.1093/zoolinnean/zlae162</mods:url>
</mods:location>
<mods:classification id="B9C1DD1F2962794C47FDDFA47201BE47">journal article</mods:classification>
<mods:identifier id="FD2C1B18C7FA68A485E29D9BF987F01C" type="DOI">10.1093/zoolinnean/zlae162</mods:identifier>
<mods:identifier id="82EB6F961D83EB1D3CF8C626A3FF2C6E" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="64619E1375A8557C8C0AE4DE471243BF" type="Zenodo-Dep">14832689</mods:identifier>
<mods:identifier id="5A069B01496596FD62DF883B637E516C" type="ZooBank">7-01-2184</mods:identifier>
</mods:mods>
<treatment id="03D58787FFA3FFDBC2C1C3D3956F4D05" ID-DOI="http://doi.org/10.5281/zenodo.14832549" ID-Zenodo-Dep="14832549" LSID="urn:lsid:plazi:treatment:03D58787FFA3FFDBC2C1C3D3956F4D05" httpUri="http://treatment.plazi.org/id/03D58787FFA3FFDBC2C1C3D3956F4D05" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C366651AFFA3FFD9C2C1C3D390954EDA" box="[1005,1264,1484,1511]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BC33691FFA3FFD9C2C1C3D390954EDA" blockId="17.[1005,1264,1484,1511]" box="[1005,1264,1484,1511]" pageId="17" pageNumber="18">
<heading id="D08B81FDFFA3FFD9C2C1C3D390954EDA" bold="true" box="[1005,1264,1484,1511]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="4C7C4D12FFA3FFD9C2C1C3D390954EDA" ID-CoL="6PPL" authority="Heyer, 1977" authorityName="Heyer" authorityYear="1977" box="[1005,1264,1484,1511]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B908EA83FFA3FFD9C2C1C3D390954EDA" bold="true" box="[1005,1264,1484,1511]" pageId="17" pageNumber="18">
<emphasis id="B908EA83FFA3FFD9C2C1C3D3900F4EDB" bold="true" box="[1005,1130,1484,1511]" italics="true" pageId="17" pageNumber="18">Phyllonastes</emphasis>
<bibRefCitation id="EFED4B60FFA3FFD9C542C3D290954EDA" author="Heyer WR" box="[1134,1264,1484,1511]" pageId="17" pageNumber="18" pagination="141 - 62" refId="ref39319" refString="Heyer WR. Taxonomic notes on frogs from the Madeira and Purus rivers, Brazil. Papeis Avulsos de Zoologia 1977; 31: 141 - 62. https: // doi. org / 10.11606 / 0031 - 1049.1977.31. p 141 - 162" type="journal article" year="1977">Heyer, 1977</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C366651AFFA3FFD9C56BC3E390F34D2A" box="[1095,1174,1532,1558]" pageId="17" pageNumber="18" type="description">
<paragraph id="8BC33691FFA3FFD9C56BC3E390F34D2A" blockId="17.[1095,1174,1532,1558]" box="[1095,1174,1532,1558]" pageId="17" pageNumber="18">
<emphasis id="B908EA83FFA3FFD9C56BC3E390F34D2A" bold="true" box="[1095,1174,1532,1558]" pageId="17" pageNumber="18">
(
<figureCitation id="13472A14FFA3FFD9C57EC3E390EE4D2A" box="[1106,1163,1532,1558]" captionStart="Figure 4" captionStartId="10.[129,193,1519,1543]" captionTargetBox="[164,1431,147,1488]" captionTargetId="figure-150@10.[161,1441,144,1491]" captionTargetPageId="10" captionText="Figure 4. Phylogenetic relationships of Phyllonastes + Urkuphrynegen. nov.(= Noblella northern clade). Maximum likelihood tree obtained for genes 16S, 12S, TYR and RAG1 from a matrix of 3846 bp for 170 terminals.Support values are on the corresponding branches:aLRT values before the slash and non-parametric bootstrap after the slash; triangles on the branches represent aLRT or bootstrap support values of 100%. For each specimen the voucher number is shown, or in its absence, the GenBank accession number.The outgroup is not shown.Black bars represent species recovered by contrasting morphological and genetic information. Red boxes represent species recovered by bPTP analysis. Abbreviations:Morpho. = Morphology; Consens.= Consensus.Photography of Bahius bilineatus by Mauro Teixeira Jr.Photography of Phyllonastes coloma by Luis A.Coloma.Photography of Phyllonastes mindo by Matthijs Hollanders.Photography of Phyllonastes myrmecoides by Mauricio Ortega. Photography of Phyllonastes naturetrekii by Juan Pablo Reyes.Photography of Phyllonastes worleyae by David Brito." figureDoi="http://doi.org/10.5281/zenodo.14832699" httpUri="https://zenodo.org/record/14832699/files/figure.png" pageId="17" pageNumber="18">Fig. 4</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C366651AFFA3FFD9C206C02697CC4D53" pageId="17" pageNumber="18" type="type_taxon">
<paragraph id="8BC33691FFA3FFD9C206C02697CC4D53" blockId="17.[810,1459,1592,1647]" pageId="17" pageNumber="18">
<emphasis id="B908EA83FFA3FFD9C206C02697CD4D73" box="[810,936,1592,1616]" italics="true" pageId="17" pageNumber="18">Type species:</emphasis>
<taxonomicName id="4C7C4D12FFA3FFD9C283C027915D4D6C" authority="Lynch, 1976" authorityName="Lynch" authorityYear="1976" box="[943,1336,1592,1616]" class="Amphibia" family="Craugastoridae" genus="Euparkerella" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="myrmecoides">
<emphasis id="B908EA83FFA3FFD9C283C02790CA4D6C" box="[943,1199,1592,1616]" italics="true" pageId="17" pageNumber="18">Euparkerella myrmecoides</emphasis>
<bibRefCitation id="EFED4B60FFA3FFD9C59AC027915D4D6C" author="Lynch JD" box="[1206,1336,1592,1616]" pageId="17" pageNumber="18" pagination="48 - 53" refId="ref40160" refString="Lynch JD. Two new species of frogs of the genus Euparkerella (Amphibia: Leptodactylidae) from Ecuador and Peru. Herpetologica 1976; 32: 48 - 53." type="journal article" year="1976">Lynch, 1976</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C366651AFFA3FFDBC206C08E94874829" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="description">
<paragraph id="8BC33691FFA3FFDBC206C08E954D4A9E" blockId="17.[810,1460,1681,1987]" lastBlockId="19.[113,764,144,419]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<emphasis id="B908EA83FFA3FFD9C206C08E97EE4D95" box="[810,907,1681,1705]" italics="true" pageId="17" pageNumber="18">Definition</emphasis>
(
<tableCitation id="C6FE032AFFA3FFD9C2B7C08E978B4D95" box="[923,1006,1681,1705]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="17" pageNumber="18" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
and 
<tableCitation id="C6FE032AFFA3FFD9C531C08E904F4D95" box="[1053,1066,1681,1705]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="17" pageNumber="18" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
<emphasis id="B908EA83FFA3FFD9C507C08E90584D94" box="[1067,1085,1681,1704]" italics="true" pageId="17" pageNumber="18">):</emphasis>
(1) Adult female average SVL &lt;
<quantity id="4C849B74FFA3FFD9C4AAC08E97304DF4" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="17" pageNumber="18" unit="mm" value="17.5">17.5 mm</quantity>
, adult male &lt;
<quantity id="4C849B74FFA3FFD9C2C3C0AF902A4DF4" box="[1007,1103,1712,1736]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="17" pageNumber="18" unit="mm" value="15.6">15.6 mm</quantity>
, 
<tableCitation id="C6FE032AFFA3FFD9C570C0AF90CF4DF5" box="[1116,1194,1712,1737]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="17" pageNumber="18" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
; (2) head narrower than body, wide body; (3) tympanic annulus present; (4) cranial crest absent; (5) finger I &lt;finger II; (6) distal phalanges of fingers and toes narrowly T-shaped; (7) knees and heels without tubercles; (8) presence of inner tarsal tubercle; (9) toes bearing circumferential grooves; (10) toe V &lt;toe III; (11) foot phalangeal formula 2-2-3-4-3; (12) dark facial mask extending at least to the middle of the flanks, triangular cloacal blotch present; (13) frontoparietals longer than wide, narrower at the anterior end; (14) premaxillary and maxillary teeth present; (15) nasal bones small, separate from maxilla; (16) zygomatic branch of squamosal much shorter than otic branch; (17) vomers reduced, broadly separated from each other, vomerine teeth absent; (18) occipital condyles widely separated from each other; (19) medial ridges present at least on presacral vertebrae III–IV; (20) urostyle crest very prominent; (21) expanded to broadly expanded sacral diapophyses; and (22) small prepollex and prehallux present.
</paragraph>
<caption id="DF036619FFA0FFDAC141C47390344811" ID-Table-UUID="DF036619FFA0FFDAC141C47390344811" httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="18" pageNumber="19" startId="18.[109,164,620,645]" targetBox="[109,1937,834,1498]" targetIsTable="true" targetPageId="18" targetType="table">
<paragraph id="8BC33691FFA0FFDAC141C47390344811" blockId="18.[109,1936,620,813]" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC141C47394DE49B9" bold="true" box="[109,187,620,645]" pageId="18" pageNumber="19">Table 3.</emphasis>
General morphological characters of 
<taxonomicName id="4C7C4D12FFA0FFDAC331C47396E949B9" authorityName="Heyer" authorityYear="1977" box="[541,652,620,645]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B908EA83FFA0FFDAC331C47396E949B9" box="[541,652,620,645]" italics="true" pageId="18" pageNumber="19">Phyllonastes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7C4D12FFA0FFDAC394C473971149B9" box="[696,884,620,645]" family="Strabomantidae" genus="Urkuphryne" pageId="18" pageNumber="19" rank="species" species="undetermined">
<emphasis id="B908EA83FFA0FFDAC394C473974349B9" box="[696,806,620,645]" italics="true" pageId="18" pageNumber="19">Urkuphryne</emphasis>
species.
</taxonomicName>
Finger and toe characters of 
<taxonomicName id="4C7C4D12FFA0FFDAC5ADC473909549B9" authorityName="Heyer" authorityYear="1977" box="[1153,1264,620,645]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B908EA83FFA0FFDAC5ADC473909549B9" box="[1153,1264,620,645]" italics="true" pageId="18" pageNumber="19">Phyllonastes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7C4D12FFA0FFDAC431C47391BC49B9" box="[1309,1497,620,645]" family="Strabomantidae" genus="Urkuphryne" pageId="18" pageNumber="19" rank="species" species="undetermined">
<emphasis id="B908EA83FFA0FFDAC431C47391EE49B9" box="[1309,1419,620,645]" italics="true" pageId="18" pageNumber="19">Urkuphryne</emphasis>
species.
</taxonomicName>
Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: 
<bibRefCitation id="EFED4B60FFA0FFDAC291C4BB900D4981" author="Boulenger GA" box="[957,1128,676,701]" pageId="18" pageNumber="19" pagination="107 - 28" refId="ref36711" refString="Boulenger GA. An account of the reptiles and batrachians collected by Mr. W. F. H. Rosenberg in western Ecuador. Proceedings of the Zoological Society of London 1898; 66: 107 - 28. https: // doi. org / 10.1111 / j. 1096 - 3642.1898. tb 03134. x" type="journal article" year="1898">Boulenger (1898)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC55EC4BB913F4981" author="Miranda-Ribeiro A" box="[1138,1370,676,701]" pageId="18" pageNumber="19" pagination="319 - 20" refId="ref40389" refString="Miranda-Ribeiro A. Algumas consideracoes sobre Holoaden luderwaldti e generos correlatos. Revista do Museu Paulista 1920; 12: 319 - 20." type="journal article" year="1920">Miranda-Ribeiro (1920)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC449C4BB91844981" author="Parker HW" box="[1381,1505,676,701]" pageId="18" pageNumber="19" pagination="201 - 3" refId="ref41177" refString="Parker HW. A new brachycephalid frog from Brazil. Annals and Magazine of Natural History 1926; 18: 201 - 3." type="journal article" year="1926">Parker (1926</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC4C7C4BB927E4981" author="Parker HW" box="[1515,1563,676,701]" pageId="18" pageNumber="19" pagination="341 - 4" refId="ref41201" refString="Parker HW. The systematic status of some frogs in the Vienna Museum. Annals and Magazine of Natural History 1932; 10: 341 - 4. https: // doi. org / 10.1080 / 00222933208673582" type="journal article" year="1932">1932</bibRefCitation>
), 
<bibRefCitation id="EFED4B60FFA0FFDAC703C4BB937D4981" author="Miranda-Ribeiro A" box="[1583,1816,676,701]" pageId="18" pageNumber="19" pagination="66 - 9" refId="ref40413" refString="Miranda-Ribeiro A. Alguns batrachios novos das colleccoes do Museo Nacional. O campo 1937; 8: 66 - 9." type="journal article" year="1937">Miranda-Ribeiro (1937)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC60EC4BB94D749E5" author="Cochran DM" pageId="18" pageNumber="19" pagination="1 - 423" refId="ref37350" refString="Cochran DM. Frogs of southeastern Brazil. Bulletin of the United States National Museum 1955; 206: 1 - 423. https: // doi. org / 10.5479 / si. 03629236.206.1" type="journal article" year="1955">Cochran (1955)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC190C4DF955449E5" author="Lutz B" box="[188,305,704,729]" pageId="18" pageNumber="19" pagination="373 - 99" refId="ref40052" refString="Lutz B. Anfibios novos e raros das Serras Costeiras do Brasil (New or rare frogs from the Coastal Ranges of Brazil). Memorias do Instituto Oswaldo Cruz 1958; 56: 373 - 99. https: // doi. org / 10.1590 / s 0074 - 02761958000200002" type="journal article" year="1958">Lutz (1958)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC016C4DF95B649E5" author="Griffiths I" box="[314,467,704,729]" pageId="18" pageNumber="19" pagination="457 - 87" refId="ref38668" refString="Griffiths I. The phylogeny of Smithillus limbatus and that status of the Brachycephalidae (Amphibia, Salientia). Proceedings of the Zoological Society of London 1959; 132: 457 - 87. https: // doi. org / 10.1111 / j. 1469 - 7998.1959. tb 05531. x" type="journal article" year="1959">Griffiths (1959)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC0F2C4DF96F349E5" author="Bokermann WCA" box="[478,662,704,729]" pageId="18" pageNumber="19" refId="ref36597" refString="Bokermann WCA. Lista Anotada das Localidades Tipo de Anfibios Brasileiros. Sao Paulo: Servicio de Documentacao, Universidade Rural Sao Paulo, 1966." type="book" year="1966">Bokermann (1966)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC38DC4DF974149E5" author="Heyer WR" box="[673,804,704,729]" pageId="18" pageNumber="19" pagination="1 - 14" refId="ref39264" refString="Heyer WR. Studies on the genus Leptodactylus (Amphibia, Leptodactylidae) III. A redefinition of the genus Leptodactylus and a description of a new genus of leptodactylid frogs. Contributions in Science 1969; 155: 1 - 14. https: // doi. org / 10.5962 / p. 241143" type="journal article" year="1969">Heyer (1969)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC203C4DF978D49E5" author="Bokermann WCA" box="[815,1000,704,729]" pageId="18" pageNumber="19" pagination="11 - 8" refId="ref36623" refString="Bokermann WCA. Tres especies novas de Eleutherodactylus do sudeste da Bahia, Brasil (Anura, Leptodactylidae). Revista Brasileira de Biologia 1975; 34: 11 - 8." type="journal article" year="1975">Bokermann (1975)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC2DEC4DF900849E5" author="Lynch JD" box="[1010,1133,704,729]" pageId="18" pageNumber="19" pagination="1 - 51" refId="ref40131" refString="Lynch JD. A review of the Andean leptodactylid frog genus Phrynopus. Occasional Papers of the Museum of Natural History 1975; 35: 1 - 51." type="journal article" year="1975">Lynch (1975</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC55BC4DF90C249E5" author="Lynch JD" box="[1143,1191,704,729]" pageId="18" pageNumber="19" pagination="48 - 53" refId="ref40160" refString="Lynch JD. Two new species of frogs of the genus Euparkerella (Amphibia: Leptodactylidae) from Ecuador and Peru. Herpetologica 1976; 32: 48 - 53." type="journal article" year="1976">1976</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC59DC4DF908449E5" author="Lynch JD" box="[1201,1249,704,729]" pageId="18" pageNumber="19" pagination="423 - 31" refId="ref40191" refString="Lynch JD. New species of minute leptodactylid frogs from the Andes of Ecuador and Peru. Journal of Herpetology 1986; 20: 423 - 31. https: // doi. org / 10.2307 / 1564505" type="journal article" year="1986">1986</bibRefCitation>
), 
<bibRefCitation id="EFED4B60FFA0FFDAC5DAC4DF911C49E5" author="Heyer WR" box="[1270,1401,704,729]" pageId="18" pageNumber="19" pagination="141 - 62" refId="ref39319" refString="Heyer WR. Taxonomic notes on frogs from the Madeira and Purus rivers, Brazil. Papeis Avulsos de Zoologia 1977; 31: 141 - 62. https: // doi. org / 10.11606 / 0031 - 1049.1977.31. p 141 - 162" type="journal article" year="1977">Heyer (1977)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC4A8C4DF924649E5" author="De la Riva I" box="[1412,1571,704,729]" pageId="18" pageNumber="19" pagination="111 - 4" refId="ref37469" refString="De la Riva I. A new species of Phrynopus from Bolivia (Anura: Leptodactylidae). Herpetologica 1992; 48: 111 - 4." type="journal article" year="1992">De la Riva (1992</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC702C4DF923B49E5" author="De la Riva I" box="[1582,1630,704,729]" pageId="18" pageNumber="19" pagination="241 - 77" refId="ref37496" refString="De la Riva I. Bolivian frogs of the genus Phrynopus, with the description of twelve new species (Anura: Brachycephalidae). Herpetological Monographs 2007; 21: 241 - 77. https: // doi. org / 10.1655 / 07 - 011.1" type="journal article" year="2007">2007</bibRefCitation>
), 
<bibRefCitation id="EFED4B60FFA0FFDAC75EC4DF929B49E5" author="Kohler JA" box="[1650,1790,704,729]" pageId="18" pageNumber="19" pagination="47 - 53" refId="ref39718" refString="Kohler JA. new species of Phyllonastes Heyer from the Chapare region of Bolivia, with notes on Phyllonastes carrascoicola. Spixiana 2000; 23: 47 - 53." type="journal article" year="2000">Köhler (2000)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC624C4DF954149C9" author="Aguayo-Vedia CR &amp; Harvey MB" pageId="18" pageNumber="19" pagination="333 - 45" refId="ref36307" refString="Aguayo-Vedia CR, Harvey MB. Dos nuevas especies de Phrynopus (Anura: Leptodactylidae) de los bosques nublados de Bolivia. Revista de Biologia Tropical 2001; 49: 333 - 45." type="journal article" year="2001">Aguayo-Vedia and Harvey (2001)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC002C4C3963849C9" author="Caramaschi U &amp; Pombal JP" box="[302,605,732,757]" pageId="18" pageNumber="19" pagination="1261 - 3" refId="ref36827" refString="Caramaschi U, Pombal JP. Notes on the type-series of Holoaden bradei B. Lutz and Holoaden luederwaldti Miranda-Ribeiro (Anura, Brachycephalidae). Revista Brasileira de Zoologia 2006; 23: 1261 - 3." type="journal article" year="2006">Caramaschi and Pombal (2006)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC34BC4C396BB49C9" author="Lehr E" box="[615,734,732,757]" pageId="18" pageNumber="19" pagination="331 - 47" refId="ref39748" refString="Lehr E. Taxonomic status of some species of Peruvian Phrynopus (Anura: Leptodactylidae), with the description of a new species from the Andes of southern Peru. Herpetologica 2006; 62: 331 - 47. https: // doi. org / 10.1655 / 0018 - 0831 (2006) 62 [331: tsosso] 2.0. co; 2" type="journal article" year="2006">Lehr (2006)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC3C4C4C397D349C9" author="Heinicke MP &amp; Duellman WE &amp; Hedges SB" box="[744,950,732,757]" pageId="18" pageNumber="19" pagination="10092 - 7" refId="ref39114" refString="Heinicke MP, Duellman WE, Hedges SB. Major Caribbean and Central American frog faunas originated by ancient oceanic dispersal. Proceedings of the National Academy of Sciences of the United States of America 2007; 104: 10092 - 7. https: // doi. org / 10.1073 / pnas. 0611051104" type="journal article" year="2007">
Heinicke 
<emphasis id="B908EA83FFA0FFDAC26FC4C3970B49C9" box="[835,878,732,757]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2007)
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC2EDC4C390FE49C9" author="De la Riva I &amp; Chaparro JC &amp; Padial JM" box="[961,1179,732,757]" pageId="18" pageNumber="19" pagination="42 - 50" refId="ref37661" refString="De la Riva I, Chaparro JC, Padial JM. A new, long-standing misidentified species of Psychrophrynella Hedges, Duellman &amp; Heinicke from Departamento Cusco, Peru (Anura: Strabomantidae). Zootaxa 2008 a; 1823: 42 - 50." type="journal article" year="2008">
De la Riva 
<emphasis id="B908EA83FFA0FFDAC50AC4C3902949C9" box="[1062,1100,732,757]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (2008a
</bibRefCitation>
, b, 2018), 
<bibRefCitation id="EFED4B60FFA0FFDAC5D3C4C391DA49C9" author="Hedges SB &amp; Duellman WE &amp; Heinicke MP" box="[1279,1471,732,757]" pageId="18" pageNumber="19" pagination="1 - 182" refId="ref38978" refString="Hedges SB, Duellman WE, Heinicke MP. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 2008; 1737: 1 - 182." type="journal article" year="2008">
Hedges 
<emphasis id="B908EA83FFA0FFDAC467C4C3911249C9" box="[1355,1399,732,757]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2008)
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC4E5C4C3935549C9" author="Guayasamin JM &amp; Teran-Valdez A" box="[1481,1840,732,757]" pageId="18" pageNumber="19" pagination="47 - 59" refId="ref38722" refString="Guayasamin JM, Teran-Valdez A. A new species of Noblella (Amphibia: Strabomantidae) from the western slopes of the Andes of Ecuador. Zootaxa 2009; 2161: 47 - 59." type="journal article" year="2009">Guayasamin and Terán-Valdez (2009)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC616C4C39571482D" author="Lehr E &amp; Catenazzi A" pageId="18" pageNumber="19" pagination="148 - 56" refId="ref39862" refString="Lehr E, Catenazzi A. A new species of minute Noblella (Anura: Strabomantidae) from southern Peru: the smallest frog of the Andes. Copeia 2009 a; 2009: 148 - 56. https: // doi. org / 10.1643 / ch- 07 - 270" type="journal article" year="2009">Lehr and Catenazzi (2009a</bibRefCitation>
, b, 2010), 
<bibRefCitation id="EFED4B60FFA0FFDAC05BC4E79650482D" author="Harvey MB &amp; Almendariz A" box="[375,565,760,785]" pageId="18" pageNumber="19" refId="ref38931" refString="Harvey MB, Almendariz A, Brito-M J et al. A new species of Noblella (Anura: Craugastoridae) from the Amazonian Slopes of the Ecuadorian Andes with Comments on Noblella lochites (Lynch). Zootaxa 2013; 3635: 001 - 14." type="journal volume" year="2013">
Harvey 
<emphasis id="B908EA83FFA0FFDAC0EEC4E79588482D" box="[450,493,760,785]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2013)
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC313C4E79714482D" author="De la Riva I &amp; Burrowes PA" box="[575,881,760,785]" pageId="18" pageNumber="19" pagination="459 - 70" refId="ref37543" refString="De la Riva I, Burrowes PA. A new species of Psychrophrynella (Anura: Craugastoridae) from the Cordillera Real, Department La Paz, Bolivia. Zootaxa 2014; 3887: 459 - 70. https: // doi. org / 10.11646 / zootaxa. 3887.4.4" type="journal article" year="2014">De la Riva and Burrowes (2014)</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC257C4E79022482D" author="Catenazzi A &amp; Uscapi V &amp; von May R" box="[891,1095,760,785]" pageId="18" pageNumber="19" pagination="71 - 84" refId="ref37190" refString="Catenazzi A, Uscapi V, von May R. A new species of Noblella (Amphibia, Anura, Craugastoridae) from the humid montane forests of Cusco, Peru. ZooKeys 2015; 516: 71 - 84. https: // doi. org / 10.3897 / zookeys. 516.9776" type="journal article" year="2015">
Catenazzi 
<emphasis id="B908EA83FFA0FFDAC2F1C4E7906D482D" box="[989,1032,760,785]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2015
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC57CC4E790E5482D" author="Catenazzi A &amp; Mamani L &amp; Lehr E" box="[1104,1152,760,785]" pageId="18" pageNumber="19" refId="ref36916" refString="Catenazzi A, Mamani L, Lehr E et al. New genus of terrestrial-breeding frogs (Holoadeninae, Strabomantidae, Terrarana) from Southern Peru. Diversity 2020; 12: 184. https: // doi. org / 10.3390 / d 12050184" type="journal volume" year="2020">2020</bibRefCitation>
), 
<bibRefCitation id="EFED4B60FFA0FFDAC5B9C4E791EB482D" author="Catenazzi A &amp; Ttito A" box="[1173,1422,760,785]" pageId="18" pageNumber="19" refId="ref36961" refString="Catenazzi A, Ttito A. A new species of Psychrophrynella (Amphibia, Anura, Craugastoridae) from the humid montane forests of Cusco, eastern slopes of the Peruvian Andes. PeerJ 2016; 4: e 1807." type="journal volume" year="2016">Catenazzi and Ttito (2016</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC4B4C4E791AD482D" author="Catenazzi A &amp; Ttito A" box="[1432,1480,760,785]" pageId="18" pageNumber="19" refId="ref37002" refString="Catenazzi A, Ttito A. Psychrophrynella glauca sp. n., a new species of terrestrial-breeding frogs (Amphibia, Anura, Strabomantidae) from the montane forests of the Amazonian Andes of Puno, Peru. PeerJ 2018; 6: e 4444. https: // doi. org / 10.7717 / peerj. 4444" type="journal volume" year="2018">2018</bibRefCitation>
, 2019), 
<bibRefCitation id="EFED4B60FFA0FFDAC73AC4E792BB482D" author="Mamani L &amp; Catenazzi A &amp; Ttito A" box="[1558,1758,760,785]" pageId="18" pageNumber="19" pagination="129 - 41" refId="ref40299" refString="Mamani L, Catenazzi A, Ttito A et al. A new species of Bryophryne (Anura: Strabomantidae) from the Cordillera de Vilcabamba, southeastern Peruvian Andes. Phyllomedusa 2017; 16: 129 - 41. https: // doi. org / 10.11606 / issn. 2316 - 9079. v 16 i 2 p 129 - 141" type="journal article" year="2017">
Mamani 
<emphasis id="B908EA83FFA0FFDAC746C4E792F3482D" box="[1642,1686,760,785]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2017)
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC7C4C4E794CC4811" author="Reyes-Puig JP &amp; Reyes-Puig C &amp; Ron SR" pageId="18" pageNumber="19" refId="ref41732" refString="Reyes-Puig JP, Reyes-Puig C, Ron SR et al. A new species of terrestrial frog of the genus Noblella Barbour, 1930 (Amphibia: Craugastoridae) from the Llanganates-Sangay Ecological Corridor, Tungurahua, Ecuador. PeerJ 2019; 7: e 7405. https: // doi. org / 10.7717 / peerj. 7405" type="journal volume" year="2019">
Reyes-Puig 
<emphasis id="B908EA83FFA0FFDAC674C4E793E6482D" box="[1880,1923,760,785]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2019
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC19DC50B94824811" author="Reyes-Puig C &amp; Maynard RJ &amp; Trageser SJ" box="[177,231,788,813]" pageId="18" pageNumber="19" pagination="162 - 71" refId="ref41678" refString="Reyes-Puig C, Maynard RJ, Trageser SJ et al. A new species of Noblella (Amphibia: Strabomantidae) from the Rio Manduriacu Reserve on the Pacific slopes from Ecuadorian Andes. Neotropical Biodiversity 2020; 6: 162 - 71. https: // doi. org / 10.1080 / 23766808.2020.1809287" type="journal article" year="2020">2020</bibRefCitation>
); Santa Cruz 
<emphasis id="B908EA83FFA0FFDAC049C50B95F54811" box="[357,400,788,813]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2019), 
<bibRefCitation id="EFED4B60FFA0FFDAC0CFC50B96C44811" author="Dubois A &amp; Ohler A &amp; Pyron RA" box="[483,673,788,813]" pageId="18" pageNumber="19" pagination="1 - 738" refId="ref37874" refString="Dubois A, Ohler A, Pyron RA. New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians (Lissamphibia). Megataxa 2021; 5: 1 - 738. https: // doi. org / 10.11646 / megataxa. 5.1.1" type="journal article" year="2021">
Dubois 
<emphasis id="B908EA83FFA0FFDAC302C50B963C4811" box="[558,601,788,813]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2021)
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC380C50B973B4811" author="Motta AP &amp; Taucce GPP &amp; Haddad BCF" box="[684,862,788,813]" pageId="18" pageNumber="19" pagination="663 - 79" refId="ref40525" refString="Motta AP, Taucce GPP, Haddad BCF et al. A new terraranan genus from the Brazilian Atlantic Forest with comments on the systematics of Brachycephaloidea (Amphibia: Anura). Journal of Zoological Systematics and Evolutionary Research 2021; 59: 663 - 79. https: // doi. org / 10.1111 / jzs. 12452" type="journal article" year="2021">
Motta 
<emphasis id="B908EA83FFA0FFDAC3C6C50B97734811" box="[746,790,788,813]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2021)
</bibRefCitation>
, 
<bibRefCitation id="EFED4B60FFA0FFDAC244C50B902E4811" author="Reyes-Puig C &amp; Guayasamin JM &amp; Koch C" box="[872,1099,788,813]" pageId="18" pageNumber="19" pagination="63 - 87" refId="ref41622" refString="Reyes-Puig C, Guayasamin JM, Koch C et al. A new species of the genus Noblella (Amphibia: Strabomantidae) from Ecuador, with new information for Noblella worleyae. Acta Herpetologica 2021; 16: 63 - 87. https: // doi. org / 10.36253 / a _ h- 10742" type="journal article" year="2021">
Reyes-Puig 
<emphasis id="B908EA83FFA0FFDAC2F4C50B90664811" box="[984,1027,788,813]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2021)
</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BC33691FFA0FFDAC141C55D929E4E80" pageId="18" pageNumber="19">
<table id="F97CC431FFA00037C141C55D93F44EE6" box="[109,1937,834,1498]" gridcols="14" gridrows="16" pageId="18" pageNumber="19">
<tr id="354C34D3FFA00037C141C55D93F44844" box="[109,1937,834,888]" gridrow="0" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C55D95774844" box="[109,274,834,888]" gridcol="0" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC141C55D94D24867" bold="true" box="[109,183,834,859]" pageId="18" pageNumber="19">Species</emphasis>
</th>
<th id="769D5DAFFFA00037C008C55D95AE4844" box="[292,459,834,888]" gridcol="1" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC008C55D95E04867" bold="true" box="[292,389,834,859]" pageId="18" pageNumber="19">Finger tip</emphasis>
</th>
<th id="769D5DAFFFA00037C0F0C55D96244844" box="[476,577,834,888]" gridcol="2" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC0F0C55D96244844" bold="true" box="[476,577,834,888]" pageId="18" pageNumber="19">Finger expansion</emphasis>
</th>
<th id="769D5DAFFFA00037C378C55D96FD4844" box="[596,664,834,888]" gridcol="3" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC378C55D96FD4844" bold="true" box="[596,664,834,888]" pageId="18" pageNumber="19">Finger groove</emphasis>
</th>
<th id="769D5DAFFFA00037C386C55D97604844" box="[682,773,834,888]" gridcol="4" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC386C55D97604844" bold="true" box="[682,773,834,888]" pageId="18" pageNumber="19">Papilla finger tip</emphasis>
</th>
<th id="769D5DAFFFA00037C234C55D973D4844" box="[792,856,834,888]" gridcol="5" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC234C55D972B4844" bold="true" box="[792,856,834,888]" pageId="18" pageNumber="19">Finger I &lt;II</emphasis>
</th>
<th id="769D5DAFFFA00037C247C55D90754844" box="[875,1040,834,888]" gridcol="6" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC247C55D90754844" bold="true" box="[875,1040,834,888]" pageId="18" pageNumber="19">Supernumerary palmar tubercles</emphasis>
</th>
<th id="769D5DAFFFA00037C50DC55D90AD4844" box="[1057,1224,834,888]" gridcol="7" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC50DC55D90154844" bold="true" pageId="18" pageNumber="19">Hand phalangeal formula</emphasis>
</th>
<th id="769D5DAFFFA00037C5F5C55D913A4844" box="[1241,1375,834,888]" gridcol="8" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC5F5C55D917A4867" bold="true" box="[1241,1311,834,859]" pageId="18" pageNumber="19">Toe tip</emphasis>
</th>
<th id="769D5DAFFFA00037C45CC55D91B04844" box="[1392,1493,834,888]" gridcol="9" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC45CC55D91B04844" bold="true" box="[1392,1493,834,888]" pageId="18" pageNumber="19">Toe expansion</emphasis>
</th>
<th id="769D5DAFFFA00037C4CBC55D924E4844" box="[1511,1579,834,888]" gridcol="10" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC4CBC55D924E4844" bold="true" box="[1511,1579,834,888]" pageId="18" pageNumber="19">Toes groove</emphasis>
</th>
<th id="769D5DAFFFA00037C711C55D92E44844" box="[1597,1665,834,888]" gridcol="11" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC711C55D92184844" bold="true" box="[1597,1665,834,888]" pageId="18" pageNumber="19">Papilla toe tip</emphasis>
</th>
<th id="769D5DAFFFA00037C7BFC55D92BC4844" box="[1683,1753,834,888]" gridcol="12" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC7BFC55D92BC4844" bold="true" box="[1683,1753,834,888]" pageId="18" pageNumber="19">Toe V &lt;III</emphasis>
</th>
<th id="769D5DAFFFA00037C7C7C55D93F44844" box="[1771,1937,834,888]" gridcol="13" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="B908EA83FFA0FFDAC7C7C55D93F44844" bold="true" box="[1771,1937,834,888]" pageId="18" pageNumber="19">Supernumerary plantar tubercles</emphasis>
</th>
</tr>
<tr id="354C34D3FFA00037C141C59093F44894" box="[109,1937,911,936]" gridrow="1" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C59095774894" box="[109,274,911,936]" gridcol="0" gridrow="1" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C59095624894" authority="Acuminate" authorityName="Acuminate" box="[109,263,911,936]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="ecuadoriensis">
<emphasis id="B908EA83FFA0FFDAC141C59095624894" bold="true" box="[109,263,911,936]" italics="true" pageId="18" pageNumber="19">P. ecuadoriensis</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C59095AE4894" box="[292,459,911,936]" gridcol="1" gridrow="1" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C59096244894" box="[476,577,911,936]" gridcol="2" gridrow="1" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C378C59096FD4894" box="[596,664,911,936]" gridcol="3" gridrow="1" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C59097604894" box="[682,773,911,936]" gridcol="4" gridrow="1" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C234C590973D4894" box="[792,856,911,936]" gridcol="5" gridrow="1" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C59090754894" box="[875,1040,911,936]" gridcol="6" gridrow="1" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C50DC59090AD4894" box="[1057,1224,911,936]" gridcol="7" gridrow="1" pageId="18" pageNumber="19">2-2-3-2</td>
<td id="769D5DAFFFA00037C5F5C590913A4894" box="[1241,1375,911,936]" gridcol="8" gridrow="1" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC59091B04894" box="[1392,1493,911,936]" gridcol="9" gridrow="1" pageId="18" pageNumber="19">Exp</td>
<td id="769D5DAFFFA00037C4CBC590924E4894" box="[1511,1579,911,936]" gridcol="10" gridrow="1" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C59092E44894" box="[1597,1665,911,936]" gridcol="11" gridrow="1" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7BFC59092BC4894" box="[1683,1753,911,936]" gridcol="12" gridrow="1" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C59093F44894" box="[1771,1937,911,936]" gridcol="13" gridrow="1" pageId="18" pageNumber="19">P</td>
</tr>
<tr id="354C34D3FFA00037C141C5AE93F448DB" box="[109,1937,945,999]" gridrow="2" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C5AE957748DB" box="[109,274,945,999]" gridcol="0" gridrow="2" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C5AE957748DB" authority="Acuminate" authorityName="Acuminate" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="cerrogolondrinas">
<emphasis id="B908EA83FFA0FFDAC141C5AE957748DB" bold="true" italics="true" pageId="18" pageNumber="19">P. cerrogolondrinas</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C5AE95AE48DB" box="[292,459,945,999]" gridcol="1" gridrow="2" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C5AE962448DB" box="[476,577,945,999]" gridcol="2" gridrow="2" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C5AE96FD48DB" box="[596,664,945,999]" gridcol="3" gridrow="2" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C386C5AE976048DB" box="[682,773,945,999]" gridcol="4" gridrow="2" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C5AE973D48DB" box="[792,856,945,999]" gridcol="5" gridrow="2" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C5AE907548DB" box="[875,1040,945,999]" gridcol="6" gridrow="2" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC5AE90AD48DB" box="[1057,1224,945,999]" gridcol="7" gridrow="2" pageId="18" pageNumber="19">2-2-3-2</td>
<td id="769D5DAFFFA00037C5F5C5AE913A48DB" box="[1241,1375,945,999]" gridcol="8" gridrow="2" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC5AE91B048DB" box="[1392,1493,945,999]" gridcol="9" gridrow="2" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC5AE924E48DB" box="[1511,1579,945,999]" gridcol="10" gridrow="2" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C5AE92E448DB" box="[1597,1665,945,999]" gridcol="11" gridrow="2" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC5AE92BC48DB" box="[1683,1753,945,999]" gridcol="12" gridrow="2" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C5AE93F448DB" box="[1771,1937,945,999]" gridcol="13" gridrow="2" pageId="18" pageNumber="19">P</td>
</tr>
<tr id="354C34D3FFA00037C141C5EF93F44F35" box="[109,1937,1008,1033]" gridrow="3" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C5EF95774F35" box="[109,274,1008,1033]" gridcol="0" gridrow="3" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C5EF94A44F35" authority="Acuminate" authorityName="Acuminate" box="[109,193,1008,1033]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="coloma">
<emphasis id="B908EA83FFA0FFDAC141C5EF94A44F35" box="[109,193,1008,1033]" italics="true" pageId="18" pageNumber="19">P.coloma</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C5EF95AE4F35" box="[292,459,1008,1033]" gridcol="1" gridrow="3" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C5EF96244F35" box="[476,577,1008,1033]" gridcol="2" gridrow="3" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C5EF96FD4F35" box="[596,664,1008,1033]" gridcol="3" gridrow="3" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C5EF97604F35" box="[682,773,1008,1033]" gridcol="4" gridrow="3" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C5EF973D4F35" box="[792,856,1008,1033]" gridcol="5" gridrow="3" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C5EF90754F35" box="[875,1040,1008,1033]" gridcol="6" gridrow="3" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC5EF90AD4F35" box="[1057,1224,1008,1033]" gridcol="7" gridrow="3" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C5EF913A4F35" box="[1241,1375,1008,1033]" gridcol="8" gridrow="3" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC5EF91B04F35" box="[1392,1493,1008,1033]" gridcol="9" gridrow="3" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C4CBC5EF924E4F35" box="[1511,1579,1008,1033]" gridcol="10" gridrow="3" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C5EF92E44F35" box="[1597,1665,1008,1033]" gridcol="11" gridrow="3" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC5EF92BC4F35" box="[1683,1753,1008,1033]" gridcol="12" gridrow="3" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C5EF93F44F35" box="[1771,1937,1008,1033]" gridcol="13" gridrow="3" pageId="18" pageNumber="19">A</td>
</tr>
<tr id="354C34D3FFA00037C141C20D93F44F17" box="[109,1937,1042,1067]" gridrow="4" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C20D95774F17" box="[109,274,1042,1067]" gridcol="0" gridrow="4" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C20D94B94F17" authority="Acuminate" authorityName="Acuminate" box="[109,220,1042,1067]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="dicaprioi">
<emphasis id="B908EA83FFA0FFDAC141C20D94B94F17" bold="true" box="[109,220,1042,1067]" italics="true" pageId="18" pageNumber="19">P. dicaprioi</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C20D95AE4F17" box="[292,459,1042,1067]" gridcol="1" gridrow="4" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C20D96244F17" box="[476,577,1042,1067]" gridcol="2" gridrow="4" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C20D96FD4F17" box="[596,664,1042,1067]" gridcol="3" gridrow="4" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C20D97604F17" box="[682,773,1042,1067]" gridcol="4" gridrow="4" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C20D973D4F17" box="[792,856,1042,1067]" gridcol="5" gridrow="4" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C20D90754F17" box="[875,1040,1042,1067]" gridcol="6" gridrow="4" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC20D90AD4F17" box="[1057,1224,1042,1067]" gridcol="7" gridrow="4" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C20D913A4F17" box="[1241,1375,1042,1067]" gridcol="8" gridrow="4" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC20D91B04F17" box="[1392,1493,1042,1067]" gridcol="9" gridrow="4" pageId="18" pageNumber="19">Exp</td>
<td id="769D5DAFFFA00037C4CBC20D924E4F17" box="[1511,1579,1042,1067]" gridcol="10" gridrow="4" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C20D92E44F17" box="[1597,1665,1042,1067]" gridcol="11" gridrow="4" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC20D92BC4F17" box="[1683,1753,1042,1067]" gridcol="12" gridrow="4" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C20D93F44F17" box="[1771,1937,1042,1067]" gridcol="13" gridrow="4" pageId="18" pageNumber="19">A</td>
</tr>
<tr id="354C34D3FFA00037C141C22C93F44F70" box="[109,1937,1075,1100]" gridrow="5" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C22C95774F70" box="[109,274,1075,1100]" gridcol="0" gridrow="5" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C22C94D24F70" authority="Round" authorityName="Round" box="[109,183,1075,1100]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FFA0FFDAC141C22C94D24F70" box="[109,183,1075,1100]" italics="true" pageId="18" pageNumber="19">P.heyeri</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C22C95AE4F70" box="[292,459,1075,1100]" gridcol="1" gridrow="5" pageId="18" pageNumber="19">Round</td>
<td id="769D5DAFFFA00037C0F0C22C96244F70" box="[476,577,1075,1100]" gridcol="2" gridrow="5" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C378C22C96FD4F70" box="[596,664,1075,1100]" gridcol="3" gridrow="5" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C22C97604F70" box="[682,773,1075,1100]" gridcol="4" gridrow="5" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C22C973D4F70" box="[792,856,1075,1100]" gridcol="5" gridrow="5" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C22C90754F70" box="[875,1040,1075,1100]" gridcol="6" gridrow="5" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC22C90AD4F70" box="[1057,1224,1075,1100]" gridcol="7" gridrow="5" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C22C913A4F70" box="[1241,1375,1075,1100]" gridcol="8" gridrow="5" pageId="18" pageNumber="19">Subacuminate</td>
<td id="769D5DAFFFA00037C45CC22C91B04F70" box="[1392,1493,1075,1100]" gridcol="9" gridrow="5" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC22C924E4F70" box="[1511,1579,1075,1100]" gridcol="10" gridrow="5" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C22C92E44F70" box="[1597,1665,1075,1100]" gridcol="11" gridrow="5" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC22C92BC4F70" box="[1683,1753,1075,1100]" gridcol="12" gridrow="5" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C22C93F44F70" box="[1771,1937,1075,1100]" gridcol="13" gridrow="5" pageId="18" pageNumber="19">A</td>
</tr>
<tr id="354C34D3FFA00037C141C24A93F44F53" box="[109,1937,1109,1135]" gridrow="6" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C24A95774F53" box="[109,274,1109,1135]" gridcol="0" gridrow="6" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C24994994F53" authority="Subacuminate" authorityName="Subacuminate" box="[109,252,1110,1135]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="plateadensis">
<emphasis id="B908EA83FFA0FFDAC141C24994994F53" bold="true" box="[109,252,1110,1135]" italics="true" pageId="18" pageNumber="19">P. plateadensis</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C24A95AE4F53" box="[292,459,1109,1135]" gridcol="1" gridrow="6" pageId="18" pageNumber="19">Subacuminate</td>
<td id="769D5DAFFFA00037C0F0C24A96244F53" box="[476,577,1109,1135]" gridcol="2" gridrow="6" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C24A96FD4F53" box="[596,664,1109,1135]" gridcol="3" gridrow="6" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C386C24A97604F53" box="[682,773,1109,1135]" gridcol="4" gridrow="6" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C24A973D4F53" box="[792,856,1109,1135]" gridcol="5" gridrow="6" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C24A90754F53" box="[875,1040,1109,1135]" gridcol="6" gridrow="6" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC24A90AD4F53" box="[1057,1224,1109,1135]" gridcol="7" gridrow="6" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C24A913A4F53" box="[1241,1375,1109,1135]" gridcol="8" gridrow="6" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC24A91B04F53" box="[1392,1493,1109,1135]" gridcol="9" gridrow="6" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC24A924E4F53" box="[1511,1579,1109,1135]" gridcol="10" gridrow="6" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C24A92E44F53" box="[1597,1665,1109,1135]" gridcol="11" gridrow="6" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC24A92BC4F53" box="[1683,1753,1109,1135]" gridcol="12" gridrow="6" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C24A93F44F53" box="[1771,1937,1109,1135]" gridcol="13" gridrow="6" pageId="18" pageNumber="19">P</td>
</tr>
<tr id="354C34D3FFA00037C141C26893F44FAC" box="[109,1937,1143,1168]" gridrow="7" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C26895774FAC" box="[109,274,1143,1168]" gridcol="0" gridrow="7" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C26894A64FAC" authority="Acuminate" authorityName="Acuminate" box="[109,195,1143,1168]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="lochites">
<emphasis id="B908EA83FFA0FFDAC141C26894A64FAC" box="[109,195,1143,1168]" italics="true" pageId="18" pageNumber="19">P.lochites</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C26895AE4FAC" box="[292,459,1143,1168]" gridcol="1" gridrow="7" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C26896244FAC" box="[476,577,1143,1168]" gridcol="2" gridrow="7" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C26896FD4FAC" box="[596,664,1143,1168]" gridcol="3" gridrow="7" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C26897604FAC" box="[682,773,1143,1168]" gridcol="4" gridrow="7" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C268973D4FAC" box="[792,856,1143,1168]" gridcol="5" gridrow="7" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C26890754FAC" box="[875,1040,1143,1168]" gridcol="6" gridrow="7" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC26890AD4FAC" box="[1057,1224,1143,1168]" gridcol="7" gridrow="7" pageId="18" pageNumber="19">2-2-3-2</td>
<td id="769D5DAFFFA00037C5F5C268913A4FAC" box="[1241,1375,1143,1168]" gridcol="8" gridrow="7" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC26891B04FAC" box="[1392,1493,1143,1168]" gridcol="9" gridrow="7" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC268924E4FAC" box="[1511,1579,1143,1168]" gridcol="10" gridrow="7" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C26892E44FAC" box="[1597,1665,1143,1168]" gridcol="11" gridrow="7" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC26892BC4FAC" box="[1683,1753,1143,1168]" gridcol="12" gridrow="7" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C26893F44FAC" box="[1771,1937,1143,1168]" gridcol="13" gridrow="7" pageId="18" pageNumber="19">P</td>
</tr>
<tr id="354C34D3FFA00037C141C28693F44F8E" box="[109,1937,1177,1202]" gridrow="8" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C28695774F8E" box="[109,274,1177,1202]" gridcol="0" gridrow="8" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C28694BD4F8E" authority="Acuminate" authorityName="Acuminate" box="[109,216,1177,1202]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="macuma">
<emphasis id="B908EA83FFA0FFDAC141C28694BD4F8E" bold="true" box="[109,216,1177,1202]" italics="true" pageId="18" pageNumber="19">P. macuma</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C28695AE4F8E" box="[292,459,1177,1202]" gridcol="1" gridrow="8" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C28696244F8E" box="[476,577,1177,1202]" gridcol="2" gridrow="8" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C28696FD4F8E" box="[596,664,1177,1202]" gridcol="3" gridrow="8" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C28697604F8E" box="[682,773,1177,1202]" gridcol="4" gridrow="8" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C286973D4F8E" box="[792,856,1177,1202]" gridcol="5" gridrow="8" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C28690754F8E" box="[875,1040,1177,1202]" gridcol="6" gridrow="8" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC28690AD4F8E" box="[1057,1224,1177,1202]" gridcol="7" gridrow="8" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C286913A4F8E" box="[1241,1375,1177,1202]" gridcol="8" gridrow="8" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC28691B04F8E" box="[1392,1493,1177,1202]" gridcol="9" gridrow="8" pageId="18" pageNumber="19">Exp</td>
<td id="769D5DAFFFA00037C4CBC286924E4F8E" box="[1511,1579,1177,1202]" gridcol="10" gridrow="8" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C28692E44F8E" box="[1597,1665,1177,1202]" gridcol="11" gridrow="8" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7BFC28692BC4F8E" box="[1683,1753,1177,1202]" gridcol="12" gridrow="8" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C28693F44F8E" box="[1771,1937,1177,1202]" gridcol="13" gridrow="8" pageId="18" pageNumber="19">P</td>
</tr>
<tr id="354C34D3FFA00037C141C2A493F44FE8" box="[109,1937,1211,1236]" gridrow="9" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C2A495774FE8" box="[109,274,1211,1236]" gridcol="0" gridrow="9" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C2A494DE4FE8" authority="Round" authorityName="Round" box="[109,187,1211,1236]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="mindo">
<emphasis id="B908EA83FFA0FFDAC141C2A494DE4FE8" box="[109,187,1211,1236]" italics="true" pageId="18" pageNumber="19">P.mindo</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C2A495AE4FE8" box="[292,459,1211,1236]" gridcol="1" gridrow="9" pageId="18" pageNumber="19">Round</td>
<td id="769D5DAFFFA00037C0F0C2A496244FE8" box="[476,577,1211,1236]" gridcol="2" gridrow="9" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C2A496FD4FE8" box="[596,664,1211,1236]" gridcol="3" gridrow="9" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C2A497604FE8" box="[682,773,1211,1236]" gridcol="4" gridrow="9" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C2A4973D4FE8" box="[792,856,1211,1236]" gridcol="5" gridrow="9" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C2A490754FE8" box="[875,1040,1211,1236]" gridcol="6" gridrow="9" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC2A490AD4FE8" box="[1057,1224,1211,1236]" gridcol="7" gridrow="9" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C2A4913A4FE8" box="[1241,1375,1211,1236]" gridcol="8" gridrow="9" pageId="18" pageNumber="19">Round</td>
<td id="769D5DAFFFA00037C45CC2A491B04FE8" box="[1392,1493,1211,1236]" gridcol="9" gridrow="9" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC2A4924E4FE8" box="[1511,1579,1211,1236]" gridcol="10" gridrow="9" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C2A492E44FE8" box="[1597,1665,1211,1236]" gridcol="11" gridrow="9" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC2A492BC4FE8" box="[1683,1753,1211,1236]" gridcol="12" gridrow="9" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C2A493F44FE8" box="[1771,1937,1211,1236]" gridcol="13" gridrow="9" pageId="18" pageNumber="19">A</td>
</tr>
<tr id="354C34D3FFA00037C141C2C293F44FCA" box="[109,1937,1245,1270]" gridrow="10" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C2C295774FCA" box="[109,274,1245,1270]" gridcol="0" gridrow="10" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C2C294904FCA" authority="Acuminate" box="[109,245,1245,1270]" class="Amphibia" family="Craugastoridae" genus="Euparkerella" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="myrmecoides">
<emphasis id="B908EA83FFA0FFDAC141C2C294904FCA" box="[109,245,1245,1270]" italics="true" pageId="18" pageNumber="19">P.myrmecoides</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C2C295AE4FCA" box="[292,459,1245,1270]" gridcol="1" gridrow="10" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C2C296244FCA" box="[476,577,1245,1270]" gridcol="2" gridrow="10" pageId="18" pageNumber="19">Exp</td>
<td id="769D5DAFFFA00037C378C2C296FD4FCA" box="[596,664,1245,1270]" gridcol="3" gridrow="10" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C2C297604FCA" box="[682,773,1245,1270]" gridcol="4" gridrow="10" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C234C2C2973D4FCA" box="[792,856,1245,1270]" gridcol="5" gridrow="10" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C2C290754FCA" box="[875,1040,1245,1270]" gridcol="6" gridrow="10" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC2C290AD4FCA" box="[1057,1224,1245,1270]" gridcol="7" gridrow="10" pageId="18" pageNumber="19">2-2-3-2</td>
<td id="769D5DAFFFA00037C5F5C2C2913A4FCA" box="[1241,1375,1245,1270]" gridcol="8" gridrow="10" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC2C291B04FCA" box="[1392,1493,1245,1270]" gridcol="9" gridrow="10" pageId="18" pageNumber="19">Exp</td>
<td id="769D5DAFFFA00037C4CBC2C2924E4FCA" box="[1511,1579,1245,1270]" gridcol="10" gridrow="10" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C2C292E44FCA" box="[1597,1665,1245,1270]" gridcol="11" gridrow="10" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7BFC2C292BC4FCA" box="[1683,1753,1245,1270]" gridcol="12" gridrow="10" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C2C293F44FCA" box="[1771,1937,1245,1270]" gridcol="13" gridrow="10" pageId="18" pageNumber="19">A</td>
</tr>
<tr id="354C34D3FFA00037C141C2E093F44E24" box="[109,1937,1279,1304]" gridrow="11" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C2E095774E24" box="[109,274,1279,1304]" gridcol="0" gridrow="11" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C2E0948F4E24" authority="Acuminate" authorityName="Acuminate" box="[109,234,1279,1304]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="naturetrekii">
<emphasis id="B908EA83FFA0FFDAC141C2E0948F4E24" box="[109,234,1279,1304]" italics="true" pageId="18" pageNumber="19">P.naturetrekii</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C2E095AE4E24" box="[292,459,1279,1304]" gridcol="1" gridrow="11" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C2E096244E24" box="[476,577,1279,1304]" gridcol="2" gridrow="11" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C2E096FD4E24" box="[596,664,1279,1304]" gridcol="3" gridrow="11" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C2E097604E24" box="[682,773,1279,1304]" gridcol="4" gridrow="11" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C2E0973D4E24" box="[792,856,1279,1304]" gridcol="5" gridrow="11" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C2E090754E24" box="[875,1040,1279,1304]" gridcol="6" gridrow="11" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C50DC2E090AD4E24" box="[1057,1224,1279,1304]" gridcol="7" gridrow="11" pageId="18" pageNumber="19">2-2-3-2</td>
<td id="769D5DAFFFA00037C5F5C2E0913A4E24" box="[1241,1375,1279,1304]" gridcol="8" gridrow="11" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC2E091B04E24" box="[1392,1493,1279,1304]" gridcol="9" gridrow="11" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC2E0924E4E24" box="[1511,1579,1279,1304]" gridcol="10" gridrow="11" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C2E092E44E24" box="[1597,1665,1279,1304]" gridcol="11" gridrow="11" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC2E092BC4E24" box="[1683,1753,1279,1304]" gridcol="12" gridrow="11" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C2E093F44E24" box="[1771,1937,1279,1304]" gridcol="13" gridrow="11" pageId="18" pageNumber="19">A</td>
</tr>
<tr id="354C34D3FFA00037C141C33F93F44E05" box="[109,1937,1312,1337]" gridrow="12" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C33F95774E05" box="[109,274,1312,1337]" gridcol="0" gridrow="12" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C33F94854E05" authority="Acuminate" authorityName="Acuminate" box="[109,224,1312,1337]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="personinus">
<emphasis id="B908EA83FFA0FFDAC141C33F94854E05" box="[109,224,1312,1337]" italics="true" pageId="18" pageNumber="19">P.personinus</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C33F95AE4E05" box="[292,459,1312,1337]" gridcol="1" gridrow="12" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C0F0C33F96244E05" box="[476,577,1312,1337]" gridcol="2" gridrow="12" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C33F96FD4E05" box="[596,664,1312,1337]" gridcol="3" gridrow="12" pageId="18" pageNumber="19">I</td>
<td id="769D5DAFFFA00037C386C33F97604E05" box="[682,773,1312,1337]" gridcol="4" gridrow="12" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C33F973D4E05" box="[792,856,1312,1337]" gridcol="5" gridrow="12" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C33F90754E05" box="[875,1040,1312,1337]" gridcol="6" gridrow="12" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC33F90AD4E05" box="[1057,1224,1312,1337]" gridcol="7" gridrow="12" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C33F913A4E05" box="[1241,1375,1312,1337]" gridcol="8" gridrow="12" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC33F91B04E05" box="[1392,1493,1312,1337]" gridcol="9" gridrow="12" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC33F924E4E05" box="[1511,1579,1312,1337]" gridcol="10" gridrow="12" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C33F92E44E05" box="[1597,1665,1312,1337]" gridcol="11" gridrow="12" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC33F92BC4E05" box="[1683,1753,1312,1337]" gridcol="12" gridrow="12" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C33F93F44E05" box="[1771,1937,1312,1337]" gridcol="13" gridrow="12" pageId="18" pageNumber="19">A</td>
</tr>
<tr id="354C34D3FFA00037C141C35D93F44E67" box="[109,1937,1346,1371]" gridrow="13" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C35D95774E67" box="[109,274,1346,1371]" gridcol="0" gridrow="13" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C35D949E4E67" authority="Subacuminate" authorityName="Subacuminate" box="[109,251,1346,1371]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="sardinayacu">
<emphasis id="B908EA83FFA0FFDAC141C35D949E4E67" bold="true" box="[109,251,1346,1371]" italics="true" pageId="18" pageNumber="19">P. sardinayacu</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C35D95AE4E67" box="[292,459,1346,1371]" gridcol="1" gridrow="13" pageId="18" pageNumber="19">Subacuminate</td>
<td id="769D5DAFFFA00037C0F0C35D96244E67" box="[476,577,1346,1371]" gridcol="2" gridrow="13" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C35D96FD4E67" box="[596,664,1346,1371]" gridcol="3" gridrow="13" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C35D97604E67" box="[682,773,1346,1371]" gridcol="4" gridrow="13" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C35D973D4E67" box="[792,856,1346,1371]" gridcol="5" gridrow="13" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C35D90754E67" box="[875,1040,1346,1371]" gridcol="6" gridrow="13" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC35D90AD4E67" box="[1057,1224,1346,1371]" gridcol="7" gridrow="13" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C35D913A4E67" box="[1241,1375,1346,1371]" gridcol="8" gridrow="13" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC35D91B04E67" box="[1392,1493,1346,1371]" gridcol="9" gridrow="13" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC35D924E4E67" box="[1511,1579,1346,1371]" gridcol="10" gridrow="13" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C35D92E44E67" box="[1597,1665,1346,1371]" gridcol="11" gridrow="13" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC35D92BC4E67" box="[1683,1753,1346,1371]" gridcol="12" gridrow="13" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C35D93F44E67" box="[1771,1937,1346,1371]" gridcol="13" gridrow="13" pageId="18" pageNumber="19">P</td>
</tr>
<tr id="354C34D3FFA00037C141C37B93F44EA6" box="[109,1937,1380,1434]" gridrow="14" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C37B95774EA6" box="[109,274,1380,1434]" gridcol="0" gridrow="14" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C37B94AA4E41" authority="Subacuminate" authorityName="Subacuminate" box="[109,207,1380,1405]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="worleyae">
<emphasis id="B908EA83FFA0FFDAC141C37B94AA4E41" box="[109,207,1380,1405]" italics="true" pageId="18" pageNumber="19">P.worleyae</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C37B95AE4EA6" box="[292,459,1380,1434]" gridcol="1" gridrow="14" pageId="18" pageNumber="19">Subacuminate/ acuminate</td>
<td id="769D5DAFFFA00037C0F0C37B96244EA6" box="[476,577,1380,1434]" gridcol="2" gridrow="14" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C37B96FD4EA6" box="[596,664,1380,1434]" gridcol="3" gridrow="14" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C37B97604EA6" box="[682,773,1380,1434]" gridcol="4" gridrow="14" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C37B973D4EA6" box="[792,856,1380,1434]" gridcol="5" gridrow="14" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C37B90754EA6" box="[875,1040,1380,1434]" gridcol="6" gridrow="14" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC37B90AD4EA6" box="[1057,1224,1380,1434]" gridcol="7" gridrow="14" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C37B913A4EA6" box="[1241,1375,1380,1434]" gridcol="8" gridrow="14" pageId="18" pageNumber="19">Acuminate</td>
<td id="769D5DAFFFA00037C45CC37B91B04EA6" box="[1392,1493,1380,1434]" gridcol="9" gridrow="14" pageId="18" pageNumber="19">SExp</td>
<td id="769D5DAFFFA00037C4CBC37B924E4EA6" box="[1511,1579,1380,1434]" gridcol="10" gridrow="14" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C711C37B92E44EA6" box="[1597,1665,1380,1434]" gridcol="11" gridrow="14" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC37B92BC4EA6" box="[1683,1753,1380,1434]" gridcol="12" gridrow="14" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C37B93F44EA6" box="[1771,1937,1380,1434]" gridcol="13" gridrow="14" pageId="18" pageNumber="19">P</td>
</tr>
<tr id="354C34D3FFA00037C141C3BC93F44EE6" box="[109,1937,1443,1498]" gridrow="15" pageId="18" pageNumber="19">
<th id="769D5DAFFFA00037C141C3BC95774EE6" box="[109,274,1443,1498]" gridcol="0" gridrow="15" pageId="18" pageNumber="19">
<taxonomicName id="4C7C4D12FFA0FFDAC141C3BB94B64E81" authority="Round" authorityName="Round" box="[109,211,1444,1469]" family="Strabomantidae" genus="Urkuphryne" pageId="18" pageNumber="19" rank="species" species="merinoi">
<emphasis id="B908EA83FFA0FFDAC141C3BB94B64E81" bold="true" box="[109,211,1444,1469]" italics="true" pageId="18" pageNumber="19">U. merinoi</emphasis>
</taxonomicName>
</th>
<td id="769D5DAFFFA00037C008C3BC95AE4EE6" box="[292,459,1443,1498]" gridcol="1" gridrow="15" pageId="18" pageNumber="19">Round to broadly rounded</td>
<td id="769D5DAFFFA00037C0F0C3BC96244EE6" box="[476,577,1443,1498]" gridcol="2" gridrow="15" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C378C3BC96FD4EE6" box="[596,664,1443,1498]" gridcol="3" gridrow="15" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C386C3BC97604EE6" box="[682,773,1443,1498]" gridcol="4" gridrow="15" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C234C3BC973D4EE6" box="[792,856,1443,1498]" gridcol="5" gridrow="15" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C247C3BC90754EE6" box="[875,1040,1443,1498]" gridcol="6" gridrow="15" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C50DC3BC90AD4EE6" box="[1057,1224,1443,1498]" gridcol="7" gridrow="15" pageId="18" pageNumber="19">2-2-3-3</td>
<td id="769D5DAFFFA00037C5F5C3BC913A4EE6" box="[1241,1375,1443,1498]" gridcol="8" gridrow="15" pageId="18" pageNumber="19">Round</td>
<td id="769D5DAFFFA00037C45CC3BC91B04EE6" box="[1392,1493,1443,1498]" gridcol="9" gridrow="15" pageId="18" pageNumber="19">NExp</td>
<td id="769D5DAFFFA00037C4CBC3BC924E4EE6" box="[1511,1579,1443,1498]" gridcol="10" gridrow="15" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C711C3BC92E44EE6" box="[1597,1665,1443,1498]" gridcol="11" gridrow="15" pageId="18" pageNumber="19">A</td>
<td id="769D5DAFFFA00037C7BFC3BC92BC4EE6" box="[1683,1753,1443,1498]" gridcol="12" gridrow="15" pageId="18" pageNumber="19">P</td>
<td id="769D5DAFFFA00037C7C7C3BC93F44EE6" box="[1771,1937,1443,1498]" gridcol="13" gridrow="15" pageId="18" pageNumber="19">A</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC33691FFA1FFDBC15DC7DB94874829" blockId="19.[113,762,452,789]" pageId="19" pageNumber="20">
<emphasis id="B908EA83FFA1FFDBC15DC7DB96114AE0" box="[113,628,452,476]" italics="true" pageId="19" pageNumber="20">
Content (15 species): 
<taxonomicName id="4C7C4D12FFA1FFDBC07EC7DB96114AE0" box="[338,628,452,476]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="cerrogolondrinas">Phyllonastes cerrogolondrinas</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC3AFC7DB96034AC7" authority="(Guayasamin &amp; Teran-Valdez, 2009)" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" baseAuthorityName="Guayasamin &amp; Teran-Valdez" baseAuthorityYear="2009" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma" status="comb. nov.">
<emphasis id="B908EA83FFA1FFDBC3AFC7DB94D34AC7" italics="true" pageId="19" pageNumber="20">Phyllonastes coloma</emphasis>
(
<bibRefCitation id="EFED4B60FFA1FFDBC1FCC7FC96054AC7" author="Guayasamin JM &amp; Teran-Valdez A" box="[208,608,483,507]" pageId="19" pageNumber="20" pagination="47 - 59" refId="ref38722" refString="Guayasamin JM, Teran-Valdez A. A new species of Noblella (Amphibia: Strabomantidae) from the western slopes of the Andes of Ecuador. Zootaxa 2009; 2161: 47 - 59." type="journal article" year="2009">Guayasamin &amp; Terán-Valdez, 2009</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23B57F8FFA1FFDBC355C7FC96924AC7" box="[633,759,483,507]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC15DC41C951C4927" box="[113,377,515,539]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="ecuadoriensis">
<emphasis id="B908EA83FFA1FFDBC15DC41C951C4927" box="[113,377,515,539]" italics="true" pageId="19" pageNumber="20">Phyllonastes ecuadoriensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC0A3C41C96084927" box="[399,621,515,539]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="dicaprioi">
<emphasis id="B908EA83FFA1FFDBC0A3C41C96084927" box="[399,621,515,539]" italics="true" pageId="19" pageNumber="20">Phyllonastes dicaprioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC3AFC41C952B4906" authority="Lynch, 1986" authorityName="Lynch" authorityYear="1986" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FFA1FFDBC3AFC41C94CE4906" italics="true" pageId="19" pageNumber="20">Phyllonastes heyeri</emphasis>
<bibRefCitation id="EFED4B60FFA1FFDBC1ECC43D952B4906" author="Lynch JD" box="[192,334,546,570]" pageId="19" pageNumber="20" pagination="423 - 31" refId="ref40191" refString="Lynch JD. New species of minute leptodactylid frogs from the Andes of Ecuador and Peru. Journal of Herpetology 1986; 20: 423 - 31. https: // doi. org / 10.2307 / 1564505" type="journal article" year="1986">Lynch, 1986</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC044C43D96934906" authority="Duellman, 1991" authorityName="Duellman" authorityYear="1991" box="[360,758,546,570]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B908EA83FFA1FFDBC044C43D96494906" box="[360,556,546,570]" italics="true" pageId="19" pageNumber="20">Phyllonastes lynchi</emphasis>
<bibRefCitation id="EFED4B60FFA1FFDBC36DC43D96934906" author="Duellman WE" box="[577,758,546,570]" pageId="19" pageNumber="20" pagination="9 - 13" refId="ref37929" refString="Duellman WE. A new species of leptodactylid frog, genus Phyllonastes, from Peru. Herpetologica 1991; 47: 9 - 13." type="journal article" year="1991">Duellman, 1991</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC15DC45E95BC4965" authority="(Lynch, 1976)" baseAuthorityName="Lynch" baseAuthorityYear="1976" box="[113,473,577,601]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="lochites">
<emphasis id="B908EA83FFA1FFDBC15DC45E955D4965" box="[113,312,577,601]" italics="true" pageId="19" pageNumber="20">Phyllonastes lochites</emphasis>
(
<bibRefCitation id="EFED4B60FFA1FFDBC06BC45E95B44965" author="Lynch JD" box="[327,465,577,601]" pageId="19" pageNumber="20" pagination="48 - 53" refId="ref40160" refString="Lynch JD. Two new species of frogs of the genus Euparkerella (Amphibia: Leptodactylidae) from Ecuador and Peru. Herpetologica 1976; 32: 48 - 53." type="journal article" year="1976">Lynch, 1976</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC0C9C45E95474945" authority="(Reyes-Puig et al., 2021)" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" baseAuthorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" baseAuthorityYear="2021" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="mindo" status="comb. nov.">
<emphasis id="B908EA83FFA1FFDBC0C9C45E96C74965" box="[485,674,577,601]" italics="true" pageId="19" pageNumber="20">Phyllonastes mindo</emphasis>
(
<bibRefCitation id="EFED4B60FFA1FFDBC39DC45E957C4945" author="Reyes-Puig C &amp; Guayasamin JM &amp; Koch C" pageId="19" pageNumber="20" pagination="63 - 87" refId="ref41622" refString="Reyes-Puig C, Guayasamin JM, Koch C et al. A new species of the genus Noblella (Amphibia: Strabomantidae) from Ecuador, with new information for Noblella worleyae. Acta Herpetologica 2021; 16: 63 - 87. https: // doi. org / 10.36253 / a _ h- 10742" type="journal article" year="2021">
Reyes-Puig 
<emphasis id="B908EA83FFA1FFDBC18BC47E94B24945" box="[167,215,609,633]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2021
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23B57F8FFA1FFDBC004C47E95FF4944" box="[296,410,609,633]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC08AC47E96134944" box="[422,630,609,633]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="macuma">
<emphasis id="B908EA83FFA1FFDBC08AC47E96134944" box="[422,630,609,633]" italics="true" pageId="19" pageNumber="20">Phyllonastes macuma</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC3AFC47E95FC49A4" authority="(Lynch, 1976)" baseAuthorityName="Lynch" baseAuthorityYear="1976" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="myrmecoides">
<emphasis id="B908EA83FFA1FFDBC3AFC47E948B49A4" italics="true" pageId="19" pageNumber="20">Phyllonastes myrmecoides</emphasis>
(
<bibRefCitation id="EFED4B60FFA1FFDBC02DC49F95F449A4" author="Lynch JD" box="[257,401,640,664]" pageId="19" pageNumber="20" pagination="48 - 53" refId="ref40160" refString="Lynch JD. Two new species of frogs of the genus Euparkerella (Amphibia: Leptodactylidae) from Ecuador and Peru. Herpetologica 1976; 32: 48 - 53." type="journal article" year="1976">Lynch, 1976</bibRefCitation>
)
</taxonomicName>
; 
<taxonomicName id="4C7C4D12FFA1FFDBC086C49F954C498B" authority="(Reyes-Puig et al., 2019)" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" baseAuthorityName="Reyes-Puig et al." baseAuthorityYear="2019" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="naturetrekii" status="comb. nov.">
<emphasis id="B908EA83FFA1FFDBC086C49F96FA49A4" box="[426,671,640,664]" italics="true" pageId="19" pageNumber="20">Phyllonastes naturetrekii</emphasis>
(
<bibRefCitation id="EFED4B60FFA1FFDBC39DC49F9547498B" author="Reyes-Puig JP &amp; Reyes-Puig C &amp; Ron SR" pageId="19" pageNumber="20" refId="ref41732" refString="Reyes-Puig JP, Reyes-Puig C, Ron SR et al. A new species of terrestrial frog of the genus Noblella Barbour, 1930 (Amphibia: Craugastoridae) from the Llanganates-Sangay Ecological Corridor, Tungurahua, Ecuador. PeerJ 2019; 7: e 7405. https: // doi. org / 10.7717 / peerj. 7405" type="journal volume" year="2019">
Reyes-Puig 
<emphasis id="B908EA83FFA1FFDBC185C4BF94B9498B" box="[169,220,671,695]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2019
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23B57F8FFA1FFDBC01EC48095C2498B" box="[306,423,671,695]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC098C480949449EB" authority="(Harvey et al., 2013)" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" baseAuthorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" baseAuthorityYear="2013" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="personinus" status="comb. nov.">
<emphasis id="B908EA83FFA1FFDBC098C48096FE498B" box="[436,667,671,695]" italics="true" pageId="19" pageNumber="20">Phyllonastes personinus</emphasis>
(
<bibRefCitation id="EFED4B60FFA1FFDBC387C480948E49EB" author="Harvey MB &amp; Almendariz A" pageId="19" pageNumber="20" refId="ref38931" refString="Harvey MB, Almendariz A, Brito-M J et al. A new species of Noblella (Anura: Craugastoridae) from the Amazonian Slopes of the Ecuadorian Andes with Comments on Noblella lochites (Lynch). Zootaxa 2013; 3635: 001 - 14." type="journal volume" year="2013">
Harvey 
<emphasis id="B908EA83FFA1FFDBC15DC4A094C049EB" box="[113,165,703,727]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2013
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23B57F8FFA1FFDBC1D7C4A0951549EA" box="[251,368,703,727]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC051C4A0961649EB" box="[381,627,703,727]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="plateadensis">
<emphasis id="B908EA83FFA1FFDBC051C4A0961649EB" box="[381,627,703,727]" italics="true" pageId="19" pageNumber="20">Phyllonastes plateadensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7C4D12FFA1FFDBC3AFC4A0948049CA" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="sardinayacu">
<emphasis id="B908EA83FFA1FFDBC3AFC4A0948049CA" italics="true" pageId="19" pageNumber="20">Phyllonastes sardinayacu</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C7C4D12FFA1FFDBC00EC4C1969F49CA" authority="(Reyes-Puig et al., 2020)" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" baseAuthorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" baseAuthorityYear="2020" box="[290,762,734,758]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="worleyae" status="comb. nov.">
<emphasis id="B908EA83FFA1FFDBC00EC4C1959149CA" box="[290,500,734,758]" italics="true" pageId="19" pageNumber="20">Phyllonastes worleyae</emphasis>
(
<bibRefCitation id="EFED4B60FFA1FFDBC32DC4C1969649CA" author="Reyes-Puig C &amp; Maynard RJ &amp; Trageser SJ" box="[513,755,734,758]" pageId="19" pageNumber="20" pagination="162 - 71" refId="ref41678" refString="Reyes-Puig C, Maynard RJ, Trageser SJ et al. A new species of Noblella (Amphibia: Strabomantidae) from the Rio Manduriacu Reserve on the Pacific slopes from Ecuadorian Andes. Neotropical Biodiversity 2020; 6: 162 - 71. https: // doi. org / 10.1080 / 23766808.2020.1809287" type="journal article" year="2020">
Reyes-Puig 
<emphasis id="B908EA83FFA1FFDBC3ADC4C096D549CA" box="[641,688,734,758]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2020
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23B57F8FFA1FFDBC15FC4E294874829" box="[115,226,765,789]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C366651AFFA1FFDBC15DC52896DE4F37" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BC33691FFA1FFDBC15DC52896DE4F37" blockId="19.[112,764,823,1035]" pageId="19" pageNumber="20">
<emphasis id="B908EA83FFA1FFDBC15DC528948B4873" box="[113,238,823,847]" italics="true" pageId="19" pageNumber="20">Distribution:</emphasis>
Pacific foothills and western slopes of the Andes of 
<collectingCountry id="F36B7601FFA1FFDBC15EC54994AD4852" box="[114,200,854,878]" name="Ecuador" pageId="19" pageNumber="20">Ecuador</collectingCountry>
, in Western Foothill Forest and Western Montane Forest between 1100 and 
<quantity id="4C849B74FFA1FFDBC018C56A95E648B1" box="[308,387,885,909]" metricMagnitude="3" metricUnit="m" metricValue="2.554" pageId="19" pageNumber="20" unit="m" value="2554.0">2554 m</quantity>
a.s.l.; on the western Amazon Basin, eastern Andean slopes of 
<collectingCountry id="F36B7601FFA1FFDBC053C58A95B04891" box="[383,469,917,941]" name="Ecuador" pageId="19" pageNumber="20">Ecuador</collectingCountry>
, in Eastern Montane Forest, Eastern Foothill Forest ≤ 
<quantity id="4C849B74FFA1FFDBC045C5AB95DF48F0" box="[361,442,948,972]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="20" unit="m" value="2400.0">2400 m</quantity>
.a.s.l. in 
<collectingCountry id="F36B7601FFA1FFDBC32FC5AB963D48F0" box="[515,600,948,972]" name="Ecuador" pageId="19" pageNumber="20">Ecuador</collectingCountry>
and Amazonian Tropical Rainforest from southeastern 
<collectingCountry id="F36B7601FFA1FFDBC0DEC5CC963D48D7" box="[498,600,979,1003]" name="Colombia" pageId="19" pageNumber="20">Colombia</collectingCountry>
to northern and central 
<collectingCountry id="F36B7601FFA1FFDBC196C5EC948E4F36" box="[186,235,1011,1034]" name="Peru" pageId="19" pageNumber="20">Peru</collectingCountry>
, and northwestern 
<collectingCountry id="F36B7601FFA1FFDBC080C5EC958D4F37" box="[428,488,1011,1035]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
, ≥ 
<quantity id="4C849B74FFA1FFDBC0D3C5EC96564F36" box="[511,563,1011,1035]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="19" pageNumber="20" unit="m" value="90.0">90 m</quantity>
a.s.l. (
<figureCitation id="13472A14FFA1FFDBC35FC5EC96CE4F37" box="[627,683,1011,1035]" captionStart="Figure 3" captionStartId="9.[113,175,1807,1831]" captionTargetBox="[149,1423,152,1776]" captionTargetId="figure-7@9.[146,1426,149,1779]" captionTargetPageId="9" captionText="Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed." figureDoi="http://doi.org/10.5281/zenodo.14832695" httpUri="https://zenodo.org/record/14832695/files/figure.png" pageId="19" pageNumber="20">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366651AFFA1FFDBC15DC233956F4D05" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BC33691FFA1FFDBC15DC233956F4D05" blockId="19.[113,766,1068,1593]" pageId="19" pageNumber="20">
<emphasis id="B908EA83FFA1FFDBC15DC23394AA4F78" box="[113,207,1068,1092]" italics="true" pageId="19" pageNumber="20">Remarks:</emphasis>
The specific epithet 
<taxonomicName id="4C7C4D12FFA1FFDBC09CC232966B4F78" box="[432,526,1069,1092]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="personina">
<emphasis id="B908EA83FFA1FFDBC09CC232966B4F78" box="[432,526,1069,1092]" italics="true" pageId="19" pageNumber="20">personina</emphasis>
</taxonomicName>
is replaced by the specific epithet 
<taxonomicName id="4C7C4D12FFA1FFDBC1DBC253953B4F5F" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" baseAuthorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" baseAuthorityYear="2013" box="[247,350,1100,1123]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="personinus">
<emphasis id="B908EA83FFA1FFDBC1DBC253953B4F5F" box="[247,350,1100,1123]" italics="true" pageId="19" pageNumber="20">personinus</emphasis>
</taxonomicName>
to match the masculine gender of the genus 
<taxonomicName id="4C7C4D12FFA1FFDBC19CC27495424FBF" authorityName="Heyer" authorityYear="1977" box="[176,295,1131,1155]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="B908EA83FFA1FFDBC19CC27495424FBF" box="[176,295,1131,1155]" italics="true" pageId="19" pageNumber="20">Phyllonastes</emphasis>
</taxonomicName>
. Thus, 
<taxonomicName id="4C7C4D12FFA1FFDBC047C274967A4FBE" box="[363,543,1131,1155]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="personina">
<emphasis id="B908EA83FFA1FFDBC047C274967A4FBE" box="[363,543,1131,1155]" italics="true" pageId="19" pageNumber="20">Noblella personina</emphasis>
</taxonomicName>
becomes 
<taxonomicName id="4C7C4D12FFA1FFDBC3ADC27494BD4F9E" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" baseAuthorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" baseAuthorityYear="2013" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="personinus">
<emphasis id="B908EA83FFA1FFDBC3ADC27494BD4F9E" italics="true" pageId="19" pageNumber="20">Phyllonastes personinus</emphasis>
</taxonomicName>
. In the new specific epithet, the root 
<emphasis id="B908EA83FFA1FFDBC370C29496C74F9E" box="[604,674,1163,1186]" italics="true" pageId="19" pageNumber="20">person-</emphasis>
is maintained, and the diminutive suffix 
<emphasis id="B908EA83FFA1FFDBC0FDC2B595924FFD" box="[465,503,1194,1217]" italics="true" pageId="19" pageNumber="20">-ina</emphasis>
is replaced by the Latin suffix - 
<emphasis id="B908EA83FFA1FFDBC198C2D694AC4FDC" box="[180,201,1225,1248]" italics="true" pageId="19" pageNumber="20">us</emphasis>
. We include 
<taxonomicName id="4C7C4D12FFA1FFDBC065C2D695F24FDD" authorityName="Duellman" authorityYear="1991" box="[329,407,1225,1249]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B908EA83FFA1FFDBC065C2D695F24FDD" box="[329,407,1225,1249]" italics="true" pageId="19" pageNumber="20">P. lynchi</emphasis>
</taxonomicName>
in the genus 
<taxonomicName id="4C7C4D12FFA1FFDBC336C2D696F44FDD" authorityName="Heyer" authorityYear="1977" box="[538,657,1225,1249]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="B908EA83FFA1FFDBC336C2D696F44FDD" box="[538,657,1225,1249]" italics="true" pageId="19" pageNumber="20">Phyllonastes</emphasis>
</taxonomicName>
because it has the synapomorphies of the group: (1) absence of vomerine teeth; (2) subacuminate toe tips; (3) toes distally expanded; and (4) presence of circumferential grooves in toes (
<bibRefCitation id="EFED4B60FFA1FFDBC3B8C33894C04E62" author="Duellman WE" pageId="19" pageNumber="20" pagination="9 - 13" refId="ref37929" refString="Duellman WE. A new species of leptodactylid frog, genus Phyllonastes, from Peru. Herpetologica 1991; 47: 9 - 13." type="journal article" year="1991">Duellman 1991</bibRefCitation>
). Additionally, it presents characteristics more frequently found within the northern clade: (1) presence of tympanic membrane; and (2) presence of inner tarsal tubercle. In addition, its distribution range is closer to the northern clade (
<figureCitation id="13472A14FFA1FFDBC3F8C3BB941B4EE7" captionStart="Figure 3" captionStartId="9.[113,175,1807,1831]" captionTargetBox="[149,1423,152,1776]" captionTargetId="figure-7@9.[146,1426,149,1779]" captionTargetPageId="9" captionText="Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed." figureDoi="http://doi.org/10.5281/zenodo.14832695" httpUri="https://zenodo.org/record/14832695/files/figure.png" pageId="19" pageNumber="20">Fig. 3</figureCitation>
). It is ~ 
<quantity id="4C849B74FFA1FFDBC1E3C3DC957B4EE7" box="[207,286,1475,1499]" metricMagnitude="5" metricUnit="m" metricValue="2.18" pageId="19" pageNumber="20" unit="km" value="218.0">218 km</quantity>
in a straight line from the confirmed record of 
<taxonomicName id="4C7C4D12FFA1FFDBC15EC3FC94A54EC7" authorityName="Duellman" authorityYear="1991" box="[114,192,1507,1531]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B908EA83FFA1FFDBC15EC3FC94A54EC7" box="[114,192,1507,1531]" italics="true" pageId="19" pageNumber="20">P. lynchi</emphasis>
</taxonomicName>
to the nearest confirmed record of the northern clade (
<taxonomicName id="4C7C4D12FFA1FFDBC3C6C3FC94CE4D26" authorityName="Lynch" authorityYear="1986" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FFA1FFDBC3C6C3FC94CE4D26" italics="true" pageId="19" pageNumber="20">P. heyeri</emphasis>
</taxonomicName>
) and ~ 
<quantity id="4C849B74FFA1FFDBC1D9C01D95204D26" box="[245,325,1538,1562]" metricMagnitude="5" metricUnit="m" metricValue="4.9" pageId="19" pageNumber="20" unit="km" value="490.0">490 km</quantity>
to the nearest record of the southern clade (
<taxonomicName id="4C7C4D12FFA1FFDBC150C03D949F4D05" box="[124,250,1569,1593]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="duellmani">
<emphasis id="B908EA83FFA1FFDBC150C03D949F4D05" box="[124,250,1569,1593]" italics="true" pageId="19" pageNumber="20">N. duellmani</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>