658 lines
95 KiB
XML
658 lines
95 KiB
XML
<document id="937475B6469BAEA5ABD5A5ED0C35C078" ID-DOI="10.1093/zoolinnean/zlae162" ID-ISSN="0024-4082" ID-Zenodo-Dep="14832689" ID-ZooBank="7-01-2184" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1738158242914" checkinUser="plazi" docAuthor="Ortega, Jhael A., Cisneros-Heredia, Diego F., Camper, Jeffrey D., Romero-Carvajal, Andrés, Negrete, Leonardo & Ron, Santiago R." docDate="2025" docId="03D58787FFA1FFDEC1D4C04990904829" docLanguage="en" docName="zlae162.pdf" docOrigin="Zoological Journal of the Linnean Society 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae162" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Phyllonastes cerrogolondrinas Ortega, Cisneros-Heredia, Camper, Romero-Carvajal, Negrete & Ron, 2025, sp.nov." docType="treatment" docVersion="4" lastPageNumber="23" masterDocId="FFECFFFFFFB2FFC8C12CC61F94654B3C" masterDocTitle="Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography" masterLastPageNumber="60" masterPageNumber="1" pageNumber="20" updateTime="1738929907134" updateUser="ExternalLinkService">
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<mods:title id="49BC3528E5FFA63C625D6AB39B12FF79">Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography</mods:title>
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<mods:namePart id="353FD706E3D168EE3EE693BB4EBBE5F3">Ortega, Jhael A.</mods:namePart>
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<mods:affiliation id="52EFF4908E78B3EA570EC512E14E4E9C">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="24C98B75C3617EDBA1871C92A94761C8">Cisneros-Heredia, Diego F.</mods:namePart>
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<mods:affiliation id="3FFCED25A542FC721780D246F8121F32">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Museo de Zoología / Laboratorio de Zoología Terrestre, Quito, Ecuador & Instituto Nacional de Biodiversidad INABIO, División de Herpetología, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="2C37960DF9496FFC9507DFC239AADDAA">Camper, Jeffrey D.</mods:namePart>
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<mods:affiliation id="0C954F232DCCE0FF5114702297364710">Department of Biology, Francis Marion University, Florence, SC 29506, USA</mods:affiliation>
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<mods:namePart id="8BC0BDAFB80C8A662FF85B6ACF493C95">Romero-Carvajal, Andrés</mods:namePart>
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<mods:affiliation id="7EABAE014E1E9ED1FD426BB8C9ADB96C">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="7F19E100B1AB3F2BFFBD634244ACDB45">Negrete, Leonardo</mods:namePart>
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<mods:affiliation id="C18785679835C8239813F88B180FCC4C">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="A37BAFC47D9E22BF8E7E04E16B000C5F">Ron, Santiago R.</mods:namePart>
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<mods:affiliation id="7EF17193DF308806AE9F42FC3EA2573F">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:nameIdentifier id="460B53BBC6B33022CC8ED662E09EF5D4" type="email">santiago.r.ron@gmail.com</mods:nameIdentifier>
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<mods:date id="1D801BA46181889B54AA0B319813417B">2025</mods:date>
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<treatment id="03D58787FFA1FFDEC1D4C04990904829" ID-DOI="http://doi.org/10.5281/zenodo.14832697" ID-Zenodo-Dep="14832697" LSID="urn:lsid:plazi:treatment:03D58787FFA1FFDEC1D4C04990904829" httpUri="http://treatment.plazi.org/id/03D58787FFA1FFDEC1D4C04990904829" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20">
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<subSubSection id="C366651AFFA1FFDBC1D4C04996164D4D" box="[248,627,1622,1649]" pageId="19" pageNumber="20" type="nomenclature">
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<taxonomicName id="4C7C4D12FFA1FFDBC1D4C04996474D4D" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[248,546,1622,1649]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="cerrogolondrinas" status="sp. nov.">
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<emphasis id="B908EA83FFA1FFDBC1D4C04996474D4D" bold="true" box="[248,546,1622,1649]" italics="true" pageId="19" pageNumber="20">Phyllonastes cerrogolondrinas</emphasis>
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<paragraph id="8BC33691FFA1FFDBC15DC0C996FE4C96" blockId="19.[113,763,1750,1962]" pageId="19" pageNumber="20">
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<materialsCitation id="3B143CCCFFA1FFDBC15DC0C996FE4C96" collectingDate="2016-12-14" collectionCode="QCAZ" collectorName="Diego Almeida & Kunam Nusirquia & Darwin Nunez & Fernando Ayala & David Mantilla & Santiago Recalde & Carlos Castro & Polibio Malte & Josue Quintanchala" country="Ecuador" elevation="2554" latitude="0.8224" location="El Goaltal Parish" longLatPrecision="7" longitude="-78.0923" municipality="Canton Espejo" pageId="19" pageNumber="20" specimenCode="QCAZ 66090" specimenCount="1" stateProvince="Carchi Province" typeStatus="holotype">
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<emphasis id="B908EA83FFA1FFDBC15DC0C996704DD2" box="[113,533,1750,1774]" italics="true" pageId="19" pageNumber="20">
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<typeStatus id="54C78833FFA1FFDBC15DC0C994AF4DD2" box="[113,202,1750,1774]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="13472A14FFA1FFDBC1CDC0C895484DD2" box="[225,301,1750,1774]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="19" pageNumber="20">Figs 10</figureCitation>
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,
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<figureCitation id="13472A14FFA1FFDBC013C0C9953C4DD2" box="[319,345,1750,1774]" captionStart="Figure 11" captionStartId="21.[113,178,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-97@21.[306,1266,144,1064]" captionTargetPageId="21" captionText="Figure 11. Palmar (A) and plantar (B) surfaces of Phyllonastes cerrogolondrinas. Photographs of right hand and right foot of the holotype QCAZ 66090. Finger IV of the hand shows some raised skin at the terminal end." figureDoi="http://doi.org/10.5281/zenodo.14832717" httpUri="https://zenodo.org/record/14832717/files/figure.png" pageId="19" pageNumber="20">11</figureCitation>
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):
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<specimenCode id="DBDA9EEAFFA1FFDBC0AFC0C996704DD2" box="[387,533,1750,1774]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="19" pageNumber="20" type="Museum">QCAZ 66090</specimenCode>
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</emphasis>
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(field no. SC-PUCE 48711)
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<specimenCount id="9D7AFD18FFA1FFDBC1E6C0EA95594C31" box="[202,316,1781,1805]" pageId="19" pageNumber="20" type="adult">adult male</specimenCount>
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from
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<collectingCountry id="F36B7601FFA1FFDBC0A4C0EA95BA4C31" box="[392,479,1781,1805]" name="Ecuador" pageId="19" pageNumber="20">Ecuador</collectingCountry>
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,
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<collectingRegion id="49B8F873FFA1FFDBC0C3C0EA96F94C31" box="[495,668,1781,1805]" country="Ecuador" name="Carchi" pageId="19" pageNumber="20">Carchi Province</collectingRegion>
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,
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<collectingMunicipality id="6BA7ACEBFFA1FFDBC381C0EA94D24C11" pageId="19" pageNumber="20">Canton Espejo</collectingMunicipality>
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,
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<location id="8EA3604AFFA1FFDBC1EDC10A950A4C11" LSID="urn:lsid:plazi:treatment:03D58787FFA1FFDEC1D4C04990904829:8EA3604AFFA1FFDBC1EDC10A950A4C11" box="[193,367,1813,1837]" country="Ecuador" latitude="0.8224" longLatPrecision="7" longitude="-78.0923" municipality="Canton Espejo" name="El Goaltal Parish" pageId="19" pageNumber="20" stateProvince="Carchi Province">El Goaltal Parish</location>
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,
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<location id="8EA3604AFFA1FFDBC055C10A96934C11" LSID="urn:lsid:plazi:treatment:03D58787FFA1FFDEC1D4C04990904829:8EA3604AFFA1FFDBC055C10A96934C11" box="[377,758,1813,1837]" country="Ecuador" latitude="0.8224" longLatPrecision="7" longitude="-78.0923" municipality="Canton Espejo" name="Bosque Protector Cerro Golondrinas" pageId="19" pageNumber="20" stateProvince="Carchi Province">Bosque Protector Cerro Golondrinas</location>
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,
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<location id="8EA3604AFFA1FFDBC15EC12B955F4C70" LSID="urn:lsid:plazi:treatment:03D58787FFA1FFDEC1D4C04990904829:8EA3604AFFA1FFDBC15EC12B955F4C70" box="[114,314,1844,1868]" country="Ecuador" latitude="0.8224" longLatPrecision="7" longitude="-78.0923" municipality="Canton Espejo" name="trail to Santa Blanca" pageId="19" pageNumber="20" stateProvince="Carchi Province">trail to Santa Blanca</location>
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(
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<geoCoordinate id="EE485056FFA1FFDBC06BC12B95CB4C70" box="[327,430,1844,1868]" degrees="0.8224" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="5" value="0.8224">0.8224°N</geoCoordinate>
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,
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<geoCoordinate id="EE485056FFA1FFDBC09BC12B96544C70" box="[439,561,1844,1868]" degrees="78.0923" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="5" value="-78.0923">78.0923°W</geoCoordinate>
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), 2554 m. Collected by
|
||
<collectorName id="26895347FFA1FFDBC1A3C14C954E4C57" box="[143,299,1875,1899]" pageId="19" pageNumber="20">Diego Almeida</collectorName>
|
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,
|
||
<collectorName id="26895347FFA1FFDBC01FC14B95894C57" box="[307,492,1875,1899]" pageId="19" pageNumber="20">Kunam Nusirquia</collectorName>
|
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,
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<collectorName id="26895347FFA1FFDBC0D8C14C96E84C57" box="[500,653,1875,1899]" pageId="19" pageNumber="20">Darwin Núñez</collectorName>
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||
,
|
||
<collectorName id="26895347FFA1FFDBC3BBC14C94CE4CB7" pageId="19" pageNumber="20">Fernando Ayala</collectorName>
|
||
,
|
||
<collectorName id="26895347FFA1FFDBC19AC16C95364CB7" box="[182,339,1907,1931]" pageId="19" pageNumber="20">David Mantilla</collectorName>
|
||
,
|
||
<collectorName id="26895347FFA1FFDBC070C16C96684CB7" box="[348,525,1907,1931]" pageId="19" pageNumber="20">Santiago Recalde</collectorName>
|
||
,
|
||
<collectorName id="26895347FFA1FFDBC334C16C96C34CB7" box="[536,678,1907,1931]" pageId="19" pageNumber="20">Carlos Castro</collectorName>
|
||
,
|
||
<collectorName id="26895347FFA1FFDBC39CC16C94D54C96" pageId="19" pageNumber="20">Polibio Malte</collectorName>
|
||
, and
|
||
<collectorName id="26895347FFA1FFDBC1C8C18D95CE4C96" box="[228,427,1938,1962]" pageId="19" pageNumber="20">Josué Quintanchala</collectorName>
|
||
on
|
||
<date id="FFC21051FFA1FFDBC0FDC18D96F24C96" box="[465,663,1938,1962]" pageId="19" pageNumber="20" value="2016-12-14">
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||
<collectingDate id="EF86E9B9FFA1FFDBC0FDC18D96F24C96" box="[465,663,1938,1962]" pageId="19" pageNumber="20" value="2016-12-14">14 December 2016</collectingDate>
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||
</date>
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||
.
|
||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BC33691FFA1FFDBC206C68E97004A3A" blockId="19.[810,1459,144,262]" pageId="19" pageNumber="20">
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||
<materialsCitation id="3B143CCCFFA1FFDBC206C68E90D04BFB" collectingDate="2016-12-14" collectionCode="QCAZ" collectorName="Diego Almeida & Kunam Nusirquia & Darwin Nunez & Fernando Ayala & David Mantilla & Santiago Recalde & Carlos Castro & Polibio Malte & Josue Quintanchala" country="Ecuador" elevation="0" latitude="0.8224" location="El Goaltal Parish" longLatPrecision="7" longitude="-78.0923" municipality="Canton Espejo" pageId="19" pageNumber="20" specimenCode="QCAZ 66086" specimenCount="1" specimenCount-adult="87" stateProvince="Carchi Province" typeStatus="paratype">
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<emphasis id="B908EA83FFA1FFDBC206C68E90524B9B" box="[810,1079,144,168]" italics="true" pageId="19" pageNumber="20">
|
||
<typeStatus id="54C78833FFA1FFDBC206C68E97EC4B94" box="[810,905,145,168]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
|
||
(N = 4;
|
||
<figureCitation id="13472A14FFA1FFDBC2CFC68E90404B94" box="[995,1061,144,168]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Same locality and collectors as holotype,
|
||
<specimenCode id="DBDA9EEAFFA1FFDBC249C6B0979C4BFB" box="[869,1017,175,199]" pageId="19" pageNumber="20">QCAZ 66086</specimenCode>
|
||
–87 adult females
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B143CCCFFA1FFDBC5ECC6B091D54BFB" box="[1216,1456,175,199]" collectionCode="QCAZ" pageId="19" pageNumber="20" specimenCode="QCAZ 66088" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBDA9EEAFFA1FFDBC5ECC6B091374BFB" box="[1216,1362,175,199]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="19" pageNumber="20" type="Museum">QCAZ 66088</specimenCode>
|
||
juvenile
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B143CCCFFA1FFDBC206C6D097044A3A" collectingDate="2016-12-12" collectingDateMax="2016-12-14" collectingDateMin="2016-12-12" collectionCode="QCAZ" pageId="19" pageNumber="20" specimenCode="QCAZ 66089" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBDA9EEAFFA1FFDBC206C6D097D84BDB" box="[810,957,207,231]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="19" pageNumber="20" type="Museum">QCAZ 66089</specimenCode>
|
||
<specimenCount id="9D7AFD18FFA1FFDBC2EAC6D090534BDB" box="[966,1078,207,231]" pageId="19" pageNumber="20" type="adult">adult male</specimenCount>
|
||
. Collected on
|
||
<date id="FFC21051FFA1FFDBC5FFC6D097044A3A" pageId="19" pageNumber="20" value="2016-12-12" valueMax="2016-12-14" valueMin="2016-12-12">
|
||
<collectingDate id="EF86E9B9FFA1FFDBC5FFC6D097044A3A" pageId="19" pageNumber="20" value="2016-12-12" valueMax="2016-12-14" valueMin="2016-12-12">12 and 14 December 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA1FFDBC206C73897D74AA4" pageId="19" pageNumber="20" type="vernacular_names">
|
||
<paragraph id="8BC33691FFA1FFDBC206C73891E54A03" blockId="19.[810,1408,295,319]" box="[810,1408,295,319]" pageId="19" pageNumber="20">
|
||
<emphasis id="B908EA83FFA1FFDBC206C738900E4A03" box="[810,1131,295,319]" italics="true" pageId="19" pageNumber="20">Proposed standard English name:</emphasis>
|
||
Golondrinas leaflitter frog.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA1FFDBC206C77E97D74AA4" blockId="19.[810,1459,353,408]" pageId="19" pageNumber="20">
|
||
<emphasis id="B908EA83FFA1FFDBC206C77E90FD4A44" box="[810,1176,353,377]" italics="true" pageId="19" pageNumber="20">Proposed standard Spanish name:</emphasis>
|
||
Cutín de Hojarasca de Golondrinas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA1FFDBC206C7A6978C4E62" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="8BC33691FFA1FFDBC206C7A6978C4E62" blockId="19.[808,1461,441,1374]" pageId="19" pageNumber="20">
|
||
<emphasis id="B908EA83FFA1FFDBC206C7A697EE4AED" box="[810,907,441,465]" italics="true" pageId="19" pageNumber="20">Definition</emphasis>
|
||
(
|
||
<figureCitation id="13472A14FFA1FFDBC2B5C7A697844AED" box="[921,993,441,465]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="19" pageNumber="20">Figs 10</figureCitation>
|
||
,
|
||
<figureCitation id="13472A14FFA1FFDBC2C6C7A690614AED" box="[1002,1028,441,465]" captionStart="Figure 11" captionStartId="21.[113,178,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-97@21.[306,1266,144,1064]" captionTargetPageId="21" captionText="Figure 11. Palmar (A) and plantar (B) surfaces of Phyllonastes cerrogolondrinas. Photographs of right hand and right foot of the holotype QCAZ 66090. Finger IV of the hand shows some raised skin at the terminal end." figureDoi="http://doi.org/10.5281/zenodo.14832717" httpUri="https://zenodo.org/record/14832717/files/figure.png" pageId="19" pageNumber="20">11</figureCitation>
|
||
;
|
||
<tableCitation id="C6FE032AFFA1FFDBC523C7A690044AED" box="[1039,1121,441,465]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="19" pageNumber="20" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="C6FE032AFFA1FFDBC5A1C7A690FF4AED" box="[1165,1178,441,465]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="19" pageNumber="20" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
|
||
<emphasis id="B908EA83FFA1FFDBC5B7C7A590CB4AED" box="[1179,1198,442,465]" italics="true" pageId="19" pageNumber="20">):</emphasis>
|
||
We assign the new species to the genus
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC285C7C690454ACD" authorityName="Heyer" authorityYear="1977" box="[937,1056,473,497]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FFA1FFDBC285C7C690454ACD" box="[937,1056,473,497]" italics="true" pageId="19" pageNumber="20">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
based on its phylogenetic relationships. The new species is characterized by: (1) dorsal skin smooth to finely shagreen, dorsolateral folds absent, ventral surfaces weakly areolate, flanks finely shagreen, discoidal fold present; (2) tympanic membrane and tympanic annulus defined, supratympanic fold and postrictal tubercle absent; (3) snout truncate in dorsal view, round in profile; (4) upper eyelid without tubercles, cranial crests absent; (5) vomerine teeth absent; (6) vocal slits present, nuptial pads absent; (7) fingers not expanded distally, with fingertips acuminate lacking papillae, finger I shorter than finger II, supernumerary tubercles present; (8) fingers bearing broad lateral fringes and ill-defined circumferential grooves; (9) distal phalanges blunt or T-shaped, phalangeal formula of hand 2-2- 3-2; (10) ulnar tubercles absent; (11) knees and heels without tubercles, outer edge of tarsus without tubercles, inner edge of tarsus bearing one rounded tubercle; (12) inner metatarsal tubercle elongate in ventral view and flat in lateral view, bigger than rounded outer metatarsal tubercle; (13) toes slightly expanded distally, with acuminate toe tips and lacking papillae, supernumerary tubercles well defined, toes with narrow lateral fringes, toe basal webbing present, all toes bearing ill-defined circumferential grooves, toe V shorter than toe III; (14) in life, dorsum light brown, with darker tubercles and a darker hourglass-shaped blotch, circular dark brown sacral lateral patches, facial mask dark brown extending from the tip of snout to the midflank, ventral surfaces bearing irregular white flecks, distal half of shanks and heels bearing yellowish brown spots; and (15) SVL in adult males
|
||
<quantity id="4C849B74FFA1FFDBC241C33997B04E02" box="[877,981,1318,1342]" metricMagnitude="-2" metricUnit="m" metricValue="1.153" pageId="19" pageNumber="20" unit="mm" value="11.53">11.53 mm</quantity>
|
||
(
|
||
<emphasis id="B908EA83FFA1FFDBC2C4C338979E4E02" box="[1000,1019,1319,1342]" italics="true" pageId="19" pageNumber="20">N</emphasis>
|
||
= 2) and SVL in adult females
|
||
<quantity id="4C849B74FFA1FFDBC467C33991D64E02" box="[1355,1459,1318,1342]" metricMagnitude="-2" metricUnit="m" metricValue="1.365" pageId="19" pageNumber="20" unit="mm" value="13.65">13.65 mm</quantity>
|
||
(
|
||
<emphasis id="B908EA83FFA1FFDBC219C358972D4E62" box="[821,840,1351,1374]" italics="true" pageId="19" pageNumber="20">N</emphasis>
|
||
= 2) (
|
||
<tableCitation id="C6FE032AFFA1FFDBC2A1C35997BD4E62" box="[909,984,1350,1374]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="19" pageNumber="20" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA1FFDCC206C39F962D4C2B" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="diagnosis">
|
||
<paragraph id="8BC33691FFA1FFDCC206C39F962D4C2B" blockId="19.[810,1463,1407,1964]" lastBlockId="20.[129,783,1447,1816]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
<emphasis id="B908EA83FFA1FFDBC206C39F90B74EAB" box="[810,1234,1407,1431]" italics="true" pageId="19" pageNumber="20">
|
||
Diagnosis:
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC284C36090B74EAB" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[936,1234,1407,1431]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="cerrogolondrinas">Phyllonastes cerrogolondrinas</taxonomicName>
|
||
</emphasis>
|
||
resembles the other
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC206C38097924E8B" box="[810,1015,1439,1463]" pageId="19" pageNumber="20">
|
||
<emphasis id="B908EA83FFA1FFDBC206C38097C14E8B" box="[810,932,1439,1463]" italics="true" pageId="19" pageNumber="20">Phyllonastes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from the western foothills of Ecuadorian Andes:
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC2ACC3A097864EEA" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" box="[896,995,1470,1494]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA1FFDBC2ACC3A097864EEA" box="[896,995,1470,1494]" italics="true" pageId="19" pageNumber="20">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC2D8C3A0900C4EEA" box="[1012,1129,1470,1494]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA1FFDBC2D8C3A0900C4EEA" box="[1012,1129,1470,1494]" italics="true" pageId="19" pageNumber="20">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC557C3A090B24EEA" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[1147,1239,1470,1494]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA1FFDBC557C3A090B24EEA" box="[1147,1239,1470,1494]" italics="true" pageId="19" pageNumber="20">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC437C3A091EB4EEA" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[1307,1422,1470,1494]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA1FFDBC437C3A091EB4EEA" box="[1307,1422,1470,1494]" italics="true" pageId="19" pageNumber="20">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
by having tympanic annulus, supernumerary palmar tubercles, fingers not expanded distally, and circumferential grooves on toes. It differs from all of them by having a dark brown belly with irregular white flecks, in life (belly bright orange in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC48EC02397144D4F" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA1FFDBC48EC02397144D4F" italics="true" pageId="19" pageNumber="20">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
, brown with brown marks in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC5E5C044915B4D4F" box="[1225,1342,1627,1651]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA1FFDBC5E5C044915B4D4F" box="[1225,1342,1627,1651]" italics="true" pageId="19" pageNumber="20">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
, yellowish cream in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC2A2C06497804DAE" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[910,997,1658,1682]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA1FFDBC2A2C06497804DAE" box="[910,997,1658,1682]" italics="true" pageId="19" pageNumber="20">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
and yellowish cream with minute speckling in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC268C08597D54D8D" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[836,944,1689,1713]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA1FFDBC268C08597D54D8D" box="[836,944,1689,1713]" italics="true" pageId="19" pageNumber="20">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
), truncate snout in dorsal view (snout rounded in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC206C0A697EC4DED" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" box="[810,905,1721,1745]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA1FFDBC206C0A697EC4DED" box="[810,905,1721,1745]" italics="true" pageId="19" pageNumber="20">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC2BAC0A690634DED" box="[918,1030,1721,1745]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA1FFDBC2BAC0A690634DED" box="[918,1030,1721,1745]" italics="true" pageId="19" pageNumber="20">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC53FC0A6900F4DED" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[1043,1130,1721,1745]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA1FFDBC53FC0A6900F4DED" box="[1043,1130,1721,1745]" italics="true" pageId="19" pageNumber="20">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC58BC0A691714DED" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[1191,1300,1721,1745]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA1FFDBC58BC0A691714DED" box="[1191,1300,1721,1745]" italics="true" pageId="19" pageNumber="20">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
), and rounded tarsal tubercle (subconic tarsal tubercle in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC5C0C0C6912C4DCC" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" box="[1260,1353,1752,1776]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA1FFDBC5C0C0C6912C4DCC" box="[1260,1353,1752,1776]" italics="true" pageId="19" pageNumber="20">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC47AC0C691C84DCC" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[1366,1453,1752,1776]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA1FFDBC47AC0C691C84DCC" box="[1366,1453,1752,1776]" italics="true" pageId="19" pageNumber="20">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC270C0E797AA4C33" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[860,975,1783,1807]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA1FFDBC270C0E797AA4C33" box="[860,975,1783,1807]" italics="true" pageId="19" pageNumber="20">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
, and conical tarsal tubercle in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC401C0E791C44C33" box="[1325,1441,1783,1807]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA1FFDBC401C0E791C44C33" box="[1325,1441,1783,1807]" italics="true" pageId="19" pageNumber="20">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC206C10890344C13" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[810,1105,1815,1839]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA1FFDBC206C10890344C13" box="[810,1105,1815,1839]" italics="true" pageId="19" pageNumber="20">Phyllonastes cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC421C10891094C13" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" box="[1293,1388,1815,1839]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA1FFDBC421C10891094C13" box="[1293,1388,1815,1839]" italics="true" pageId="19" pageNumber="20">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
by the presence of supernumerary plantar tubercles (supernumerary plantar tubercles absent in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC54DC14990A24C51" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" box="[1121,1223,1877,1901]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA1FFDBC54DC14990A24C51" box="[1121,1223,1877,1901]" italics="true" pageId="19" pageNumber="20">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
) and circumferential grooves in fingers (fingers without circumferential grooves in
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC48DC16A97144C90" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Guayasamin & Teran-Valdez" baseAuthorityYear="2009" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="coloma">
|
||
<emphasis id="B908EA83FFA1FFDBC48DC16A97144C90" italics="true" pageId="19" pageNumber="20">P. coloma</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C7C4D12FFA1FFDBC2ABC18B90CE4C90" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[903,1195,1940,1964]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA1FFDBC2ABC18B90CE4C90" box="[903,1195,1940,1964]" italics="true" pageId="19" pageNumber="20">Phyllonastes cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
can also be distinguished from
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC196C3B7954F4E83" box="[186,298,1447,1471]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA6FFDCC196C3B7954F4E83" box="[186,298,1447,1471]" italics="true" pageId="20" pageNumber="21">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
by the presence of vocal slits in males (males without vocal slits in
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC070C3D895AF4EE3" box="[348,458,1479,1503]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA6FFDCC070C3D895AF4EE3" box="[348,458,1479,1503]" italics="true" pageId="20" pageNumber="21">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
) and by the presence of supernumerary plantar tubercles (supernumerary plantar tubercles absent in
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC034C01995ED4D21" box="[280,392,1541,1565]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA6FFDCC034C01995ED4D21" box="[280,392,1541,1565]" italics="true" pageId="20" pageNumber="21">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
). It also differs from
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC341C01996A04D21" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[621,709,1541,1565]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA6FFDCC341C01996A04D21" box="[621,709,1541,1565]" italics="true" pageId="20" pageNumber="21">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC3D6C01994B24D01" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA6FFDCC3D6C01994B24D01" italics="true" pageId="20" pageNumber="21">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
by having weakly areolate ventral surfaces (ventral surfaces smooth in
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC078C05A95CB4D60" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[340,430,1604,1628]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA6FFDCC078C05A95CB4D60" box="[340,430,1604,1628]" italics="true" pageId="20" pageNumber="21">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC0CBC05A963D4D60" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[487,600,1604,1628]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA6FFDCC0CBC05A963D4D60" box="[487,600,1604,1628]" italics="true" pageId="20" pageNumber="21">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
), acuminate toe tips (toe tips rounded in
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC0B5C07B95914D47" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[409,500,1635,1659]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA6FFDCC0B5C07B95914D47" box="[409,500,1635,1659]" italics="true" pageId="20" pageNumber="21">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
), circumferential grooves in fingers (fingers lacking circumferential grooves in
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC39DC09C976C4DA7" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[689,777,1667,1691]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA6FFDCC39DC09C976C4DA7" box="[689,777,1667,1691]" italics="true" pageId="20" pageNumber="21">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC183C0BC957B4D86" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[175,286,1698,1722]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA6FFDCC183C0BC957B4D86" box="[175,286,1698,1722]" italics="true" pageId="20" pageNumber="21">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
), and by the absence of a supratympanic fold and ulnar tubercles (both present in
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC32EC0DD963D4DE5" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2021" box="[514,600,1729,1753]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="mindo">
|
||
<emphasis id="B908EA83FFA6FFDCC32EC0DD963D4DE5" box="[514,600,1729,1753]" italics="true" pageId="20" pageNumber="21">P. mindo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC3A7C0DD969C4DE5" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" baseAuthorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" baseAuthorityYear="2020" box="[651,761,1729,1753]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="worleyae">
|
||
<emphasis id="B908EA83FFA6FFDCC3A7C0DD969C4DE5" box="[651,761,1729,1753]" italics="true" pageId="20" pageNumber="21">P. worleyae</emphasis>
|
||
</taxonomicName>
|
||
). For a comparison with other
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC083C0FE961C4DC5" box="[431,633,1761,1785]" pageId="20" pageNumber="21">
|
||
<emphasis id="B908EA83FFA6FFDCC083C0FE964C4DC5" box="[431,553,1761,1785]" italics="true" pageId="20" pageNumber="21">Phyllonastes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
that are more phylogenetically distant, see
|
||
<tableCitation id="C6FE032AFFA6FFDCC081C11F96674C24" box="[429,514,1792,1816]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="20" pageNumber="21" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="C6FE032AFFA6FFDCC319C11F96274C24" box="[565,578,1792,1816]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="20" pageNumber="21" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF036619FFA6FFDCC1ADC2E195B54E56" ID-DOI="http://doi.org/10.5281/zenodo.14832715" ID-Zenodo-Dep="14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="20" pageNumber="21" startId="20.[129,194,1278,1302]" targetBox="[166,1436,151,1242]" targetPageId="20" targetType="figure">
|
||
<paragraph id="8BC33691FFA6FFDCC1ADC2E191164E29" blockId="20.[129,1465,1278,1386]" box="[129,1395,1278,1302]" pageId="20" pageNumber="21">
|
||
<emphasis id="B908EA83FFA6FFDCC1ADC2E194804E2A" bold="true" box="[129,229,1278,1302]" pageId="20" pageNumber="21">Figure 10.</emphasis>
|
||
Live and preserved specimens of
|
||
<taxonomicName id="4C7C4D12FFA6FFDCC30DC2E197204E2A" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[545,837,1278,1302]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA6FFDCC30DC2E197204E2A" bold="true" box="[545,837,1278,1302]" italics="true" pageId="20" pageNumber="21">Phyllonastes cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
. Lateral, dorsal, and ventral views of: A, B, holotype
|
||
<emphasis id="B908EA83FFA6FFDCC41FC2E191164E29" box="[1331,1395,1278,1301]" italics="true" pageId="20" pageNumber="21">QCAZ</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA6FFDCC1ADC30595B54E56" blockId="20.[129,1465,1278,1386]" pageId="20" pageNumber="21">
|
||
<emphasis id="B908EA83FFA6FFDCC1ADC30594D84E0E" box="[129,189,1306,1330]" italics="true" pageId="20" pageNumber="21">66090</emphasis>
|
||
, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm. A, in life. B–D, in preservative. Scale bars are given only for preserved adults. The depigmentation visible in the specimens is attributable to preservation defects.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C366651AFFA6FFDEC1ADC15F95D24DE9" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="8BC33691FFA6FFDCC1ADC15F90184EE3" blockId="20.[129,777,1855,1974]" lastBlockId="20.[824,1475,1447,1973]" pageId="20" pageNumber="21">
|
||
<emphasis id="B908EA83FFA6FFDCC1ADC15F964D4C6B" box="[129,552,1855,1879]" italics="true" pageId="20" pageNumber="21">
|
||
Description of the
|
||
<typeStatus id="54C78833FFA6FFDCC010C12095EB4C6B" box="[316,398,1855,1879]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13472A14FFA6FFDCC08FC15F95884C6B" box="[419,493,1855,1879]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="20" pageNumber="21">Figs 10</figureCitation>
|
||
,
|
||
<figureCitation id="13472A14FFA6FFDCC0D7C12096704C6B" box="[507,533,1855,1879]" captionStart="Figure 11" captionStartId="21.[113,178,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-97@21.[306,1266,144,1064]" captionTargetPageId="21" captionText="Figure 11. Palmar (A) and plantar (B) surfaces of Phyllonastes cerrogolondrinas. Photographs of right hand and right foot of the holotype QCAZ 66090. Finger IV of the hand shows some raised skin at the terminal end." figureDoi="http://doi.org/10.5281/zenodo.14832717" httpUri="https://zenodo.org/record/14832717/files/figure.png" pageId="20" pageNumber="21">11</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Adult male (
|
||
<emphasis id="B908EA83FFA6FFDCC3E8C15F94A74C4B" italics="true" pageId="20" pageNumber="21">QCAZ 66090</emphasis>
|
||
). Measurements (in millimetres): SVL, 10.89; tibia length, 6.07; foot length, 5.91; head length, 3.36; head width, 3.64; eye diameter, 1.32; tympanum diameter, 0.48; interorbital distance, 1.93; upper eyelid width, 1.31; internarial distance, 1.38; eye–nostril distance, 0.76.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA6FFDCC279C3F9970F4DC4" blockId="20.[824,1475,1447,1973]" pageId="20" pageNumber="21">Body robust, head slightly wider than long, narrower than body; snout truncate in dorsal view, short and rounded in lateral profile, without rostral papilla; canthus rostralis straight in dorsal view; loreal region slightly concave; eyelids without tubercles; interorbital space flat, with no cranial crests; tympanic membrane and annulus distinct (more conspicuous in its anterior half), without supratympanic fold; postrictal tubercles absent; vomerine teeth absent, vocal slits present; and nuptial pads absent.</paragraph>
|
||
<paragraph id="8BC33691FFA6FFDDC279C11F95964C9D" blockId="20.[824,1475,1447,1973]" lastBlockId="21.[113,763,1804,1953]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">Skin on dorsum and flanks finely shagreen; dorsolateral folds absent; ventral surfaces weakly areolate; skin in cloacal region areolate; discoidal fold present. Ulnar tubercles absent, palmar tubercles low, outer palmar tubercle circular, larger than elongate thenar tubercle; subarticular tubercles well defined, round in ventral and lateral view; well-defined rounded supernumerary tubercles; broad dermal fringes on fingers, fingers not expanded distally, discs narrow, bearing round and inconspicuous pads, with an ill-defined circumferential groove; fingertips acuminate, without papillae; relative lengths of fingers is I <II <IV <III; phalangeal formula of hand is 2-2-3-2.</paragraph>
|
||
<caption id="DF036619FFA7FFDDC15DC25C97064F44" ID-DOI="http://doi.org/10.5281/zenodo.14832717" ID-Zenodo-Dep="14832717" httpUri="https://zenodo.org/record/14832717/files/figure.png" pageId="21" pageNumber="22" startId="21.[113,178,1091,1115]" targetBox="[306,1266,144,1064]" targetPageId="21" targetType="figure">
|
||
<paragraph id="8BC33691FFA7FFDDC15DC25C97064F44" blockId="21.[113,1418,1091,1144]" pageId="21" pageNumber="22">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC25C94B04F67" bold="true" box="[113,213,1091,1115]" pageId="21" pageNumber="22">Figure 11.</emphasis>
|
||
Palmar (A) and plantar (B) surfaces of
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC364C25C97094F67" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[584,876,1091,1115]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA7FFDDC364C25C97094F67" bold="true" box="[584,876,1091,1115]" italics="true" pageId="21" pageNumber="22">Phyllonastes cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
. Photographs of right hand and right foot of the holotype
|
||
<emphasis id="B908EA83FFA7FFDDC15DC27F94974F4B" box="[113,242,1119,1143]" italics="true" pageId="21" pageNumber="22">QCAZ 66090</emphasis>
|
||
. Finger IV of the hand shows some raised skin at the terminal end.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF036619FFA7FFDDC15DC2B994D34FCA" ID-Table-UUID="DF036619FFA7FFDDC15DC2B994D34FCA" httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="21" pageNumber="22" startId="21.[113,168,1190,1214]" targetBox="[113,1459,1292,1736]" targetIsTable="true" targetPageId="21" targetType="table">
|
||
<paragraph id="8BC33691FFA7FFDDC15DC2B994D34FCA" blockId="21.[113,1440,1190,1270]" pageId="21" pageNumber="22">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC2B994DA4F82" bold="true" box="[113,191,1190,1214]" pageId="21" pageNumber="22">Table 4.</emphasis>
|
||
Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males. of
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC579C2B990A14F82" authorityName="Heyer" authorityYear="1977" box="[1109,1220,1190,1214]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FFA7FFDDC579C2B990A14F82" box="[1109,1220,1190,1214]" italics="true" pageId="21" pageNumber="22">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC5DDC2B9913A4F82" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1265,1375,1190,1214]" family="Strabomantidae" genus="Urkuphryne" pageId="21" pageNumber="22" rank="genus" status="new species">
|
||
<emphasis id="B908EA83FFA7FFDDC5DDC2B9913A4F82" box="[1265,1375,1190,1214]" italics="true" pageId="21" pageNumber="22">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B57F8FFA7FFDDC448C2B894D24FE6" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
|
||
. The number of individuals is given in parentheses after the sex. For each group, minimum (Min) and maximum (Max) values are also shown.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC33691FFA7FFDDC15DC31391E44DF4" pageId="21" pageNumber="22">
|
||
<table id="F97CC431FFA70037C15DC31391D64DF4" box="[113,1459,1292,1736]" gridcols="5" gridrows="13" pageId="21" pageNumber="22">
|
||
<tr id="354C34D3FFA70037C15DC31391D64E18" box="[113,1459,1292,1316]" gridrow="0" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC31395484E18" box="[113,301,1292,1316]" gridcol="0" gridrow="0" pageId="21" pageNumber="22">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC31394DE4E18" bold="true" box="[113,187,1292,1316]" pageId="21" pageNumber="22">Species</emphasis>
|
||
</th>
|
||
<th id="769D5DAFFFA70037C0C5C31396364E18" box="[489,595,1292,1316]" gridcol="1" gridrow="0" pageId="21" pageNumber="22">
|
||
<emphasis id="B908EA83FFA7FFDDC0C5C313966E4E18" bold="true" box="[489,523,1292,1316]" pageId="21" pageNumber="22">Sex</emphasis>
|
||
</th>
|
||
<th id="769D5DAFFFA70037C222C31397ED4E18" box="[782,904,1292,1316]" gridcol="2" gridrow="0" pageId="21" pageNumber="22">
|
||
<emphasis id="B908EA83FFA7FFDDC222C31397ED4E18" bold="true" box="[782,904,1292,1316]" pageId="21" pageNumber="22">SVL average</emphasis>
|
||
</th>
|
||
<th id="769D5DAFFFA70037C568C31390F94E18" box="[1092,1180,1292,1316]" gridcol="3" gridrow="0" pageId="21" pageNumber="22">
|
||
<emphasis id="B908EA83FFA7FFDDC568C31390F94E18" bold="true" box="[1092,1180,1292,1316]" pageId="21" pageNumber="22">SVL Min</emphasis>
|
||
</th>
|
||
<th id="769D5DAFFFA70037C474C31391D64E18" box="[1368,1459,1292,1316]" gridcol="4" gridrow="0" pageId="21" pageNumber="22">
|
||
<emphasis id="B908EA83FFA7FFDDC474C31391D64E18" bold="true" box="[1368,1459,1292,1316]" pageId="21" pageNumber="22">SVL Max</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC32491D64E68" box="[113,1459,1339,1364]" gridrow="1" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC32495484E68" box="[113,301,1339,1364]" gridcol="0" gridrow="1" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC32395484E6F" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[113,301,1339,1363]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC32395484E6F" bold="true" box="[113,301,1339,1363]" italics="true" pageId="21" pageNumber="22">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C32496364E68" box="[489,595,1339,1364]" gridcol="1" gridrow="1" pageId="21" pageNumber="22">Female (2)</td>
|
||
<td id="769D5DAFFFA70037C222C32497ED4E68" box="[782,904,1339,1364]" gridcol="2" gridrow="1" pageId="21" pageNumber="22">13.7</td>
|
||
<td id="769D5DAFFFA70037C568C32490F94E68" box="[1092,1180,1339,1364]" gridcol="3" gridrow="1" pageId="21" pageNumber="22">13.6</td>
|
||
<td id="769D5DAFFFA70037C474C32491D64E68" box="[1368,1459,1339,1364]" gridcol="4" gridrow="1" pageId="21" pageNumber="22">13.8</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC34291D64E4A" box="[113,1459,1373,1398]" gridrow="2" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC34295484E4A" box="[113,301,1373,1398]" gridcol="0" gridrow="2" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC34195484E49" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[113,301,1373,1397]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC34195484E49" bold="true" box="[113,301,1373,1397]" italics="true" pageId="21" pageNumber="22">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C34296364E4A" box="[489,595,1373,1398]" gridcol="1" gridrow="2" pageId="21" pageNumber="22">Male (2)</td>
|
||
<td id="769D5DAFFFA70037C222C34297ED4E4A" box="[782,904,1373,1398]" gridcol="2" gridrow="2" pageId="21" pageNumber="22">11.5</td>
|
||
<td id="769D5DAFFFA70037C568C34290F94E4A" box="[1092,1180,1373,1398]" gridcol="3" gridrow="2" pageId="21" pageNumber="22">10.9</td>
|
||
<td id="769D5DAFFFA70037C474C34291D64E4A" box="[1368,1459,1373,1398]" gridcol="4" gridrow="2" pageId="21" pageNumber="22">12.2</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC36091D64EA4" box="[113,1459,1407,1432]" gridrow="3" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC36095484EA4" box="[113,301,1407,1432]" gridcol="0" gridrow="3" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC39F956E4EAB" box="[113,267,1407,1431]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="ecuadoriensis">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC39F956E4EAB" bold="true" box="[113,267,1407,1431]" italics="true" pageId="21" pageNumber="22">P. ecuadoriensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C36096364EA4" box="[489,595,1407,1432]" gridcol="1" gridrow="3" pageId="21" pageNumber="22">Female (2)</td>
|
||
<td id="769D5DAFFFA70037C222C36097ED4EA4" box="[782,904,1407,1432]" gridcol="2" gridrow="3" pageId="21" pageNumber="22">17.4</td>
|
||
<td id="769D5DAFFFA70037C568C36090F94EA4" box="[1092,1180,1407,1432]" gridcol="3" gridrow="3" pageId="21" pageNumber="22">16.6</td>
|
||
<td id="769D5DAFFFA70037C474C36091D64EA4" box="[1368,1459,1407,1432]" gridcol="4" gridrow="3" pageId="21" pageNumber="22">18.1</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC3BE91D64E85" box="[113,1459,1441,1465]" gridrow="4" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC3BE95484E85" box="[113,301,1441,1465]" gridcol="0" gridrow="4" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC3BD956E4E85" box="[113,267,1441,1465]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="ecuadoriensis">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC3BD956E4E85" bold="true" box="[113,267,1441,1465]" italics="true" pageId="21" pageNumber="22">P. ecuadoriensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C3BE96364E85" box="[489,595,1441,1465]" gridcol="1" gridrow="4" pageId="21" pageNumber="22">Males (2)</td>
|
||
<td id="769D5DAFFFA70037C222C3BE97ED4E85" box="[782,904,1441,1465]" gridcol="2" gridrow="4" pageId="21" pageNumber="22">14.2</td>
|
||
<td id="769D5DAFFFA70037C568C3BE90F94E85" box="[1092,1180,1441,1465]" gridcol="3" gridrow="4" pageId="21" pageNumber="22">13.6</td>
|
||
<td id="769D5DAFFFA70037C474C3BE91D64E85" box="[1368,1459,1441,1465]" gridcol="4" gridrow="4" pageId="21" pageNumber="22">14.8</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC3DC91D64EE7" box="[113,1459,1475,1499]" gridrow="5" pageId="21" pageNumber="22" rowspan-3="1" rowspan-4="1">
|
||
<th id="769D5DAFFFA70037C15DC3DC95484EE7" box="[113,301,1475,1499]" gridcol="0" gridrow="5" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC3DC94854EE7" box="[113,224,1475,1499]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC3DC94854EE7" bold="true" box="[113,224,1475,1499]" italics="true" pageId="21" pageNumber="22">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C3DC96364EE7" box="[489,595,1475,1499]" gridcol="1" gridrow="5" pageId="21" pageNumber="22">Female (1)</td>
|
||
<td id="769D5DAFFFA70037C222C3DC97ED4EE7" box="[782,904,1475,1499]" gridcol="2" gridrow="5" pageId="21" pageNumber="22">15.2</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC3FA91D64EC1" box="[113,1459,1509,1533]" gridrow="6" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC3FA95484EC1" box="[113,301,1509,1533]" gridcol="0" gridrow="6" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC3FA94854EC1" box="[113,224,1509,1533]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="dicaprioi">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC3FA94854EC1" bold="true" box="[113,224,1509,1533]" italics="true" pageId="21" pageNumber="22">P. dicaprioi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C3FA96364EC1" box="[489,595,1509,1533]" gridcol="1" gridrow="6" pageId="21" pageNumber="22">Male (6)</td>
|
||
<td id="769D5DAFFFA70037C222C3FA97ED4EC1" box="[782,904,1509,1533]" gridcol="2" gridrow="6" pageId="21" pageNumber="22">12.2 (1.7)</td>
|
||
<td id="769D5DAFFFA70037C568C3FA90F94EC1" box="[1092,1180,1509,1533]" gridcol="3" gridrow="6" pageId="21" pageNumber="22">10.6</td>
|
||
<td id="769D5DAFFFA70037C474C3FA91D64EC1" box="[1368,1459,1509,1533]" gridcol="4" gridrow="6" pageId="21" pageNumber="22">14.6</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC01891D64D23" box="[113,1459,1543,1567]" gridrow="7" pageId="21" pageNumber="22" rowspan-3="1" rowspan-4="1">
|
||
<th id="769D5DAFFFA70037C15DC01895484D23" box="[113,301,1543,1567]" gridcol="0" gridrow="7" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC01894B94D22" box="[113,220,1543,1566]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="macuma">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC01894B94D22" bold="true" box="[113,220,1543,1566]" italics="true" pageId="21" pageNumber="22">P. macuma</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C01896364D23" box="[489,595,1543,1567]" gridcol="1" gridrow="7" pageId="21" pageNumber="22">Female (1)</td>
|
||
<td id="769D5DAFFFA70037C222C01897ED4D23" box="[782,904,1543,1567]" gridcol="2" gridrow="7" pageId="21" pageNumber="22">14.5</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC03691D64D7D" box="[113,1459,1577,1601]" gridrow="8" pageId="21" pageNumber="22" rowspan-3="1" rowspan-4="1">
|
||
<th id="769D5DAFFFA70037C15DC03695484D7D" box="[113,301,1577,1601]" gridcol="0" gridrow="8" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC03694B94D7C" box="[113,220,1577,1600]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="macuma">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC03694B94D7C" bold="true" box="[113,220,1577,1600]" italics="true" pageId="21" pageNumber="22">P. macuma</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C03696364D7D" box="[489,595,1577,1601]" gridcol="1" gridrow="8" pageId="21" pageNumber="22">Male (1)</td>
|
||
<td id="769D5DAFFFA70037C222C03697ED4D7D" box="[782,904,1577,1601]" gridcol="2" gridrow="8" pageId="21" pageNumber="22">11.9</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC05591D64D5F" box="[113,1459,1610,1635]" gridrow="9" pageId="21" pageNumber="22" rowspan-3="1" rowspan-4="1">
|
||
<th id="769D5DAFFFA70037C15DC05595484D5F" box="[113,301,1610,1635]" gridcol="0" gridrow="9" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC05495654D5E" box="[113,256,1610,1634]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="plateadensis">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC05495654D5E" bold="true" box="[113,256,1610,1634]" italics="true" pageId="21" pageNumber="22">P. plateadensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C05596364D5F" box="[489,595,1610,1635]" gridcol="1" gridrow="9" pageId="21" pageNumber="22">Female (1)</td>
|
||
<td id="769D5DAFFFA70037C222C05597ED4D5F" box="[782,904,1610,1635]" gridcol="2" gridrow="9" pageId="21" pageNumber="22">15.7</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC07391D64DB8" box="[113,1459,1644,1668]" gridrow="10" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC07395484DB8" box="[113,301,1644,1668]" gridcol="0" gridrow="10" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC072949A4DB8" box="[113,255,1644,1668]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="sardinayacu">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC072949A4DB8" bold="true" box="[113,255,1644,1668]" italics="true" pageId="21" pageNumber="22">P. sardinayacu</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C07396364DB8" box="[489,595,1644,1668]" gridcol="1" gridrow="10" pageId="21" pageNumber="22">Female (2)</td>
|
||
<td id="769D5DAFFFA70037C222C07397ED4DB8" box="[782,904,1644,1668]" gridcol="2" gridrow="10" pageId="21" pageNumber="22">14.2</td>
|
||
<td id="769D5DAFFFA70037C568C07390F94DB8" box="[1092,1180,1644,1668]" gridcol="3" gridrow="10" pageId="21" pageNumber="22">14.1</td>
|
||
<td id="769D5DAFFFA70037C474C07391D64DB8" box="[1368,1459,1644,1668]" gridcol="4" gridrow="10" pageId="21" pageNumber="22">14.3</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC09191D64D9A" box="[113,1459,1678,1702]" gridrow="11" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC09195484D9A" box="[113,301,1678,1702]" gridcol="0" gridrow="11" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC09094BD4D9A" box="[113,216,1678,1702]" family="Strabomantidae" genus="Urkuphryne" pageId="21" pageNumber="22" rank="species" species="merinoi">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC09094BD4D9A" bold="true" box="[113,216,1678,1702]" italics="true" pageId="21" pageNumber="22">U. merinoi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C09196364D9A" box="[489,595,1678,1702]" gridcol="1" gridrow="11" pageId="21" pageNumber="22">Female (5)</td>
|
||
<td id="769D5DAFFFA70037C222C09197ED4D9A" box="[782,904,1678,1702]" gridcol="2" gridrow="11" pageId="21" pageNumber="22">19.7 (0.3)</td>
|
||
<td id="769D5DAFFFA70037C568C09190F94D9A" box="[1092,1180,1678,1702]" gridcol="3" gridrow="11" pageId="21" pageNumber="22">19.4</td>
|
||
<td id="769D5DAFFFA70037C474C09191D64D9A" box="[1368,1459,1678,1702]" gridcol="4" gridrow="11" pageId="21" pageNumber="22">20.1</td>
|
||
</tr>
|
||
<tr id="354C34D3FFA70037C15DC0AF91D64DF4" box="[113,1459,1712,1736]" gridrow="12" pageId="21" pageNumber="22">
|
||
<th id="769D5DAFFFA70037C15DC0AF95484DF4" box="[113,301,1712,1736]" gridcol="0" gridrow="12" pageId="21" pageNumber="22">
|
||
<taxonomicName id="4C7C4D12FFA7FFDDC15DC0AF94BD4DF4" box="[113,216,1712,1736]" family="Strabomantidae" genus="Urkuphryne" pageId="21" pageNumber="22" rank="species" species="merinoi">
|
||
<emphasis id="B908EA83FFA7FFDDC15DC0AF94BD4DF4" bold="true" box="[113,216,1712,1736]" italics="true" pageId="21" pageNumber="22">U. merinoi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="769D5DAFFFA70037C0C5C0AF96364DF4" box="[489,595,1712,1736]" gridcol="1" gridrow="12" pageId="21" pageNumber="22">Male (3)</td>
|
||
<td id="769D5DAFFFA70037C222C0AF97ED4DF4" box="[782,904,1712,1736]" gridcol="2" gridrow="12" pageId="21" pageNumber="22">17.2 (1.8)</td>
|
||
<td id="769D5DAFFFA70037C568C0AF90F94DF4" box="[1092,1180,1712,1736]" gridcol="3" gridrow="12" pageId="21" pageNumber="22">15.2</td>
|
||
<td id="769D5DAFFFA70037C474C0AF91D64DF4" box="[1368,1459,1712,1736]" gridcol="4" gridrow="12" pageId="21" pageNumber="22">18.7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA7FFDEC269C113948E4AFE" blockId="21.[810,1459,1804,1953]" lastBlockId="22.[128,779,144,450]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">Hindlimbs robust; knee and heel without tubercles; tarsal folds (inner and outer) absent, outer tarsal tubercles absent; inner tarsal tubercle present, prominent and round; inner metatarsal tubercle present, elongate in ventral view and flat in lateral profile; well-defined, prominent and round outer metatarsal tubercle; plantar surface with several rounded and conspicuous supernumerary tubercles; round subarticular tubercles ill defined, more conspicuous at the base of toes; toes slightly expanded distally, toes with narrow lateral fringes; basal webbing between toes; discs on toes ill defined, narrow, with acuminated tip, without papilla; ill-defined circumferential grooves; relative lengths of toes I <II <V <III <IV; toe III longer than toe V (toe III reaches the distal border of the second subarticular tubercle of toe IV; Toe V almost reaches half of the second tubercle of toe IV).</paragraph>
|
||
<paragraph id="8BC33691FFA4FFDEC1ADC7FC96914F33" blockId="22.[128,779,483,1040]" pageId="22" pageNumber="23">
|
||
<emphasis id="B908EA83FFA4FFDEC1ADC7FC976C4AC7" box="[129,777,483,507]" italics="true" pageId="22" pageNumber="23">
|
||
Colour of
|
||
<typeStatus id="54C78833FFA4FFDEC1CEC7FC95514AC7" box="[226,308,483,507]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
in life (based on digital photographs) (
|
||
<figureCitation id="13472A14FFA4FFDEC398C7FB96924AC7" box="[692,759,483,507]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Dorsal surface light brown, with darker, scattered, and low tubercles and a darker hourglass-shaped blotch extending from the interorbital to the sacral region; circular dark brown sacral lateral patches well defined. Facial mask dark brown extending from the tip of snout to the midflank and delineated along its upper edge by a light brown line. Lips dark brown with irregular white flecks. Cloacal region dark brown surrounded by a paler stripe. Flanks light brown posteriorly and with the extension of the facial mask anteriorly. Upper arm dorsal surfaces yellowish brown, elbow and forearm light brown, becoming darker towards the hand. Dorsal surfaces of thighs and proximal half of shanks light brown; distal half of shanks and tarsus yellowish brown; dorsal surface of tarsus dark brown. Ventral surfaces of throat, chest, forearms, belly, and thighs dark brown (almost black) bearing irregular white flecks in greater quantity on the belly. Ventral surfaces of upper arms and distal half of shanks reddish brown. Distal half of shanks and heels bearing yellowish brown spots.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA4FFDEC1ADC22E963E4E78" blockId="22.[129,780,1073,1348]" pageId="22" pageNumber="23">
|
||
<emphasis id="B908EA83FFA4FFDEC1ADC22E96554F75" box="[129,560,1073,1097]" italics="true" pageId="22" pageNumber="23">
|
||
Colour of
|
||
<typeStatus id="54C78833FFA4FFDEC1C8C22E95534F75" box="[228,310,1073,1097]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
in preservative (
|
||
<figureCitation id="13472A14FFA4FFDEC0F6C22D967B4F75" box="[474,542,1073,1097]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Dorsal surface from the scapular region to the tip of snout dark brown, bearing cream blotches that become smaller and less abundant towards the snout. Dorsum cream from the scapular region to the vent. Dorsal surfaces of forearms, hands, thighs, and feet dark brown. Dorsal upper arms and shanks cream. Ventral surfaces dark brown; upper arms and shanks cream. Belly and ventral surfaces of thighs bearing cream irregular blotches. Cloacal region dark brown with a medial cream longitudinal stripe.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FFA4FFDEC1ADC37995D24DE9" blockId="22.[128,779,1381,1750]" pageId="22" pageNumber="23">
|
||
<emphasis id="B908EA83FFA4FFDEC1ADC37995214E41" box="[129,324,1381,1405]" italics="true" pageId="22" pageNumber="23">
|
||
Variation (
|
||
<figureCitation id="13472A14FFA4FFDEC1C2C37995574E41" box="[238,306,1381,1405]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
):
|
||
</emphasis>
|
||
In this section, traits refer to preserved individuals unless otherwise mentioned. Tympanic annulus can be differentiated beneath skin only anteriorly (e.g. QCAZ 66087) or be inconspicuous (i.e. QCAZ 66088–89); dorsal skin might be smooth (e.g. QCAZ 66087) or finely shagreen (e.g. QCAZ 66086); outer palmar tubercle can be elongate (e.g. QCAZ 66087). Snout is subacuminate in the only subadult male (QCAZ 66088). Dorsum might be completely dark brown (e.g. QCAZ 66087), dark brown bearing some cream blotches (e.g. QCAZ 66086), or cream with a dark brown snout (QCAZ 66088). Colour variation is shown in
|
||
<figureCitation id="13472A14FFA4FFDEC328C08096004D8B" box="[516,613,1695,1719]" captionStart="Figure 10" captionStartId="20.[129,194,1278,1302]" captionTargetBox="[166,1436,151,1242]" captionTargetId="figure-312@20.[161,1441,144,1250]" captionTargetPageId="20" captionText="Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects." figureDoi="http://doi.org/10.5281/zenodo.14832715" httpUri="https://zenodo.org/record/14832715/files/figure.png" pageId="22" pageNumber="23">Figure 10</figureCitation>
|
||
. Morphometric variation is detailed in
|
||
<tableCitation id="C6FE032AFFA4FFDEC04AC0A195D44DEA" box="[358,433,1726,1750]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="22" pageNumber="23" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA4FFDEC1ADC0E8917E4A79" pageId="22" pageNumber="23" type="multiple">
|
||
<paragraph id="8BC33691FFA4FFDEC1ADC0E8917E4A79" blockId="22.[129,778,1783,1964]" lastBlockId="22.[825,1476,144,325]" pageId="22" pageNumber="23">
|
||
<emphasis id="B908EA83FFA4FFDEC1ADC0E895C14C13" italics="true" pageId="22" pageNumber="23">
|
||
Distribution, natural history, and conservation status (
|
||
<figureCitation id="13472A14FFA4FFDEC396C0E796924C33" box="[698,759,1783,1807]" captionStart="Figure 3" captionStartId="9.[113,175,1807,1831]" captionTargetBox="[149,1423,152,1776]" captionTargetId="figure-7@9.[146,1426,149,1779]" captionTargetPageId="9" captionText="Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed." figureDoi="http://doi.org/10.5281/zenodo.14832695" httpUri="https://zenodo.org/record/14832695/files/figure.png" pageId="22" pageNumber="23">Fig. 3</figureCitation>
|
||
):
|
||
<taxonomicName id="4C7C4D12FFA4FFDEC1ADC10895C14C13" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[129,420,1815,1839]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cerrogolondrinas">Phyllonastes cerrogolondrinas</taxonomicName>
|
||
</emphasis>
|
||
is known from the surroundings of Cerro Golondrinas Protective Forest from
|
||
<quantity id="4C849B74FFA4FFDEC373C129976F4C71" box="[607,778,1846,1870]" metricMagnitude="3" metricUnit="m" metricValue="2.5115" metricValueMax="2.554" metricValueMin="2.469" pageId="22" pageNumber="23" unit="m" value="2511.5" valueMax="2554.0" valueMin="2469.0">2469 to 2554 m</quantity>
|
||
a.s.l. All individuals were collected in Eastern Montane Forest (
|
||
<quantity id="4C849B74FFA4FFDEC1A4C16A95794CB0" box="[136,284,1909,1933]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="3.6" metricValueMin="1.3" pageId="22" pageNumber="23" unit="m" value="2450.0" valueMax="3600.0" valueMin="1300.0">1300–3600 m</quantity>
|
||
a.s.l.) at night, between 20:00 and 22:00 h. They were found on leaf litter near streams. Collections were made in secondary
|
||
<emphasis id="B908EA83FFA4FFDEC285C68E90744B94" box="[937,1041,144,168]" italics="true" pageId="22" pageNumber="23">terra firme</emphasis>
|
||
forest. Because of the lack of information on population size and geographical range, we suggest assigning
|
||
<taxonomicName id="4C7C4D12FFA4FFDEC216C6D0979D4BDB" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[826,1016,207,231]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA4FFDEC216C6D0979D4BDB" box="[826,1016,207,231]" italics="true" pageId="22" pageNumber="23">P. cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
to the Data Deficient Red List Category (based on
|
||
<bibRefCitation id="EFED4B60FFA4FFDEC282C6F191D54A3A" author="IUCN Standards and Petitions Committee & Using the IUCN Red List Categories" box="[942,1456,238,262]" pageId="22" pageNumber="23" refId="ref39535" refString="IUCN Standards and Petitions Committee. Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee, 2023. https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (25 January 2024, date last accessed)." type="url" year="2023">IUCN Standards and Petitions Committee 2023</bibRefCitation>
|
||
); nevertheless, its presence in secondary forest suggests adaptability to anthropically modified environments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFFA4FFDEC215C77990904829" pageId="22" pageNumber="23" type="etymology">
|
||
<paragraph id="8BC33691FFA4FFDEC215C77990904829" blockId="22.[823,1476,358,789]" pageId="22" pageNumber="23">
|
||
<emphasis id="B908EA83FFA4FFDEC215C77997CD4A42" box="[825,936,358,382]" italics="true" pageId="22" pageNumber="23">Etymology:</emphasis>
|
||
The specific epithet is a toponym in apposition, and it refers to the
|
||
<typeStatus id="54C78833FFA4FFDEC2F9C79990664AA2" box="[981,1027,390,414]" pageId="22" pageNumber="23">type</typeStatus>
|
||
locality. The Cerro Golondrinas Protective Forestis 1500 haandislocatedwithintheChocóBiogeographical Region in the province of
|
||
<collectingRegion id="49B8F873FFA4FFDEC577C7DB90C44AE0" box="[1115,1185,452,476]" country="Ecuador" name="Carchi" pageId="22" pageNumber="23">Carchi</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36B7601FFA4FFDEC59CC7DB916D4AE0" box="[1200,1288,452,476]" name="Ecuador" pageId="22" pageNumber="23">Ecuador</collectingCountry>
|
||
. It is part of the Biological Corridor Chiles–Mataje. The Cerro Golondrinas Protective Forest is an important habitat for flora and fauna, with unique characteristics of richness and endemism. The forest is protected by the NGO Ecominga.
|
||
<taxonomicName id="4C7C4D12FFA4FFDEC589C45E91A14965" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1189,1476,577,601]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="cerrogolondrinas">
|
||
<emphasis id="B908EA83FFA4FFDEC589C45E91A14965" box="[1189,1476,577,601]" italics="true" pageId="22" pageNumber="23">Phyllonastes cerrogolondrinas</emphasis>
|
||
</taxonomicName>
|
||
is also a posthumous tribute to Jaime Levi, a naturalist who promoted the creation of the Cerro Golondrinas Protective Forest. This species name was chosen with the inhabitants of the communities of La Plata, Morán, and Golondrinas (
|
||
<collectingRegion id="49B8F873FFA4FFDEC43DC4A091DE49EB" box="[1297,1467,703,727]" country="Ecuador" name="Carchi" pageId="22" pageNumber="23">Carchi Province</collectingRegion>
|
||
) in an outreach effort to empower local communities in favour of environmental education and conservation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |