344 lines
43 KiB
XML
344 lines
43 KiB
XML
<document id="5D1863EEB2DA5B2B717A5D9FD9BA4966" ID-DOI="10.1093/zoolinnean/zlae162" ID-ISSN="0024-4082" ID-Zenodo-Dep="14832689" ID-ZooBank="7-01-2184" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1738158242914" checkinUser="plazi" docAuthor="Ortega, Jhael A., Cisneros-Heredia, Diego F., Camper, Jeffrey D., Romero-Carvajal, Andrés, Negrete, Leonardo & Ron, Santiago R." docDate="2025" docId="03D58787FF9AFFE2C52FC49990A44C84" docLanguage="en" docName="zlae162.pdf" docOrigin="Zoological Journal of the Linnean Society 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae162" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Urkuphryne Ortega, Cisneros-Heredia, Camper, Romero-Carvajal, Negrete & Ron, 2025, gen. nov." docType="treatment" docVersion="4" lastPageNumber="43" masterDocId="FFECFFFFFFB2FFC8C12CC61F94654B3C" masterDocTitle="Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography" masterLastPageNumber="60" masterPageNumber="1" pageNumber="41" updateTime="1738929907134" updateUser="ExternalLinkService">
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<mods:title id="4B570C5B0CC9AA162C6242F245E6B18E">Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography</mods:title>
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<mods:namePart id="CC1369B48D392CE5AFE5909272D97A0B">Ortega, Jhael A.</mods:namePart>
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<mods:affiliation id="6F3428D0D7F779091F18AF157930A152">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="B918762B3C9579D6B229F55FA14FABE3">Cisneros-Heredia, Diego F.</mods:namePart>
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<mods:affiliation id="C8607686598954F9E1681B15F1688754">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Museo de Zoología / Laboratorio de Zoología Terrestre, Quito, Ecuador & Instituto Nacional de Biodiversidad INABIO, División de Herpetología, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="499D2F2BFC4F74885214BE92FD5247A2">Camper, Jeffrey D.</mods:namePart>
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<mods:affiliation id="AF5E44DE970EBFC7AEC68213CE3783DA">Department of Biology, Francis Marion University, Florence, SC 29506, USA</mods:affiliation>
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<mods:namePart id="2E2EDCDD3ADD22EB59E7B822ADC4BF5E">Romero-Carvajal, Andrés</mods:namePart>
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<mods:affiliation id="FD1A06E5F06F4225DB8A13D56CAE34C8">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="D65D217980103F4BB9C8949F2291870C">Negrete, Leonardo</mods:namePart>
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<mods:affiliation id="68EEC68156F895A97A407264BAC0E2D6">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:namePart id="53AA12D4A81FE9C1FD4FC2DC403AA6C0">Ron, Santiago R.</mods:namePart>
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<mods:nameIdentifier id="49FD8FEDC0010CD6CB22DBBD7B221D24" type="email">santiago.r.ron@gmail.com</mods:nameIdentifier>
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<mods:title id="BF7B86B713535C6BDC827DCB7F88CAD8">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5777F854BA72D74F439EC85FAA8FA105">2025</mods:date>
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<treatment id="03D58787FF9AFFE2C52FC49990A44C84" ID-DOI="http://doi.org/10.5281/zenodo.14832733" ID-Zenodo-Dep="14832733" LSID="urn:lsid:plazi:treatment:03D58787FF9AFFE2C52FC49990A44C84" httpUri="http://treatment.plazi.org/id/03D58787FF9AFFE2C52FC49990A44C84" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
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<taxonomicName id="4C7C4D12FF9AFFE0C52FC49990EE499D" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1027,1163,646,673]" class="Amphibia" family="Strabomantidae" genus="Urkuphryne" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="40" pageNumber="41" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B908EA83FF9AFFE0C52FC49990EE499D" bold="true" box="[1027,1163,646,673]" italics="true" pageId="40" pageNumber="41">Urkuphryne</emphasis>
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<taxonomicNameLabel id="A23B57F8FF9AFFE0C5BDC498909D499D" box="[1169,1272,647,673]" pageId="40" pageNumber="41" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="C366651AFF9AFFE0C57BC4A997BA4828" pageId="40" pageNumber="41" type="description">
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<paragraph id="8BC33691FF9AFFE0C57BC4A990C049EC" blockId="40.[825,1474,694,788]" box="[1111,1189,694,720]" pageId="40" pageNumber="41">
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<emphasis id="B908EA83FF9AFFE0C57BC4A990C049EC" bold="true" box="[1111,1189,694,720]" pageId="40" pageNumber="41">
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(
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<figureCitation id="13472A14FF9AFFE0C54DC4A990FF49EC" box="[1121,1178,694,720]" captionStart="Figure 4" captionStartId="10.[129,193,1519,1543]" captionTargetBox="[164,1431,147,1488]" captionTargetId="figure-150@10.[161,1441,144,1491]" captionTargetPageId="10" captionText="Figure 4. Phylogenetic relationships of Phyllonastes + Urkuphrynegen. nov.(= Noblella northern clade). Maximum likelihood tree obtained for genes 16S, 12S, TYR and RAG1 from a matrix of 3846 bp for 170 terminals.Support values are on the corresponding branches:aLRT values before the slash and non-parametric bootstrap after the slash; triangles on the branches represent aLRT or bootstrap support values of 100%. For each specimen the voucher number is shown, or in its absence, the GenBank accession number.The outgroup is not shown.Black bars represent species recovered by contrasting morphological and genetic information. Red boxes represent species recovered by bPTP analysis. Abbreviations:Morpho. = Morphology; Consens.= Consensus.Photography of Bahius bilineatus by Mauro Teixeira Jr.Photography of Phyllonastes coloma by Luis A.Coloma.Photography of Phyllonastes mindo by Matthijs Hollanders.Photography of Phyllonastes myrmecoides by Mauricio Ortega. Photography of Phyllonastes naturetrekii by Juan Pablo Reyes.Photography of Phyllonastes worleyae by David Brito." figureDoi="http://doi.org/10.5281/zenodo.14832699" httpUri="https://zenodo.org/record/14832699/files/figure.png" pageId="40" pageNumber="41">Fig. 4</figureCitation>
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)
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</emphasis>
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<paragraph id="8BC33691FF9AFFE0C215C4C297BA4828" blockId="40.[825,1474,694,788]" pageId="40" pageNumber="41">
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LSID:
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urn:lsid:zoobank.org:act:
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<subSubSection id="C366651AFF9AFFE0C215C526903A4F07" pageId="40" pageNumber="41" type="multiple">
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<paragraph id="8BC33691FF9AFFE0C215C526902E486C" blockId="40.[825,1099,825,849]" box="[825,1099,825,849]" pageId="40" pageNumber="41">
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<emphasis id="B908EA83FF9AFFE0C215C526971E486D" box="[825,891,825,849]" italics="true" pageId="40" pageNumber="41">Order:</emphasis>
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<taxonomicName id="4C7C4D12FF9AFFE0C2ADC5269020486D" authority="Oppel, 1811" authorityName="Oppel" authorityYear="1811" box="[897,1093,825,849]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="40" pageNumber="41" phylum="Chordata" rank="order">
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Anura
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<bibRefCitation id="EFED4B60FF9AFFE0C2E9C5269020486D" author="Oppel M" box="[965,1093,825,849]" pageId="40" pageNumber="41" refId="ref40729" refString="Oppel M. Die Ordnung, Familien und Gattungen der Reptilien als Prodrom einer Naturgeschichte derselben. Munchen: Joseph Lindauer, 1811." type="book" year="1811">Oppel, 1811</bibRefCitation>
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</taxonomicName>
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.
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||
</paragraph>
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<paragraph id="8BC33691FF9AFFE0C215C56D914748B5" blockId="40.[825,1314,882,906]" box="[825,1314,882,906]" pageId="40" pageNumber="41">
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<emphasis id="B908EA83FF9AFFE0C215C56D97D348B6" box="[825,950,882,906]" italics="true" pageId="40" pageNumber="41">Superfamily:</emphasis>
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Brachycephaloidea
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||
<bibRefCitation id="EFED4B60FF9AFFE0C5ABC56D917948B6" author="Gunther ACLG" box="[1159,1308,882,906]" pageId="40" pageNumber="41" pagination="319 - 28" refId="ref38757" refString="Gunther ACLG. Neue Batrachier in der Sammlung des britischen Museums. Archiv fur Naturgeschichte 1858; 24: 319 - 28. https: // doi. org / 10.5962 / bhl. part. 5288" type="journal article" year="1858">Günther, 1858</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BC33691FF9AFFE0C215C5B4909348FF" blockId="40.[825,1270,939,963]" box="[825,1270,939,963]" pageId="40" pageNumber="41">
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<emphasis id="B908EA83FF9AFFE0C215C5B497E148FF" box="[825,900,939,963]" italics="true" pageId="40" pageNumber="41">Family:</emphasis>
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<taxonomicName id="4C7C4D12FF9AFFE0C2A5C5B4909548FF" authority="Hedges et al., 2008" authorityName="Hedges" authorityYear="2008" box="[905,1264,939,963]" family="Strabomantidae" pageId="40" pageNumber="41" rank="family">
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||
Strabomantidae
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||
<bibRefCitation id="EFED4B60FF9AFFE0C51DC5B4909548FF" author="Hedges SB & Duellman WE & Heinicke MP" box="[1073,1264,939,963]" pageId="40" pageNumber="41" pagination="1 - 182" refId="ref38978" refString="Hedges SB, Duellman WE, Heinicke MP. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 2008; 1737: 1 - 182." type="journal article" year="2008">
|
||
Hedges
|
||
<emphasis id="B908EA83FF9AFFE0C5AFC5B390D748FF" box="[1155,1202,939,963]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
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||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FF9AFFE0C215C5FA916D48C0" blockId="40.[825,1288,997,1021]" box="[825,1288,997,1021]" pageId="40" pageNumber="41">
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||
<emphasis id="B908EA83FF9AFFE0C215C5FA97C648C1" box="[825,931,997,1021]" italics="true" pageId="40" pageNumber="41">Subfamily:</emphasis>
|
||
<taxonomicName id="4C7C4D12FF9AFFE0C281C5FA916748C1" authority="Hedges et al., 2008" authorityName="Hedges" authorityYear="2008" box="[941,1282,997,1021]" class="Amphibia" family="Craugastoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="40" pageNumber="41" phylum="Chordata" rank="subFamily" subFamily="Holoadeninae">
|
||
Holoadeninae
|
||
<bibRefCitation id="EFED4B60FF9AFFE0C56FC5FA916748C1" author="Hedges SB & Duellman WE & Heinicke MP" box="[1091,1282,997,1021]" pageId="40" pageNumber="41" pagination="1 - 182" refId="ref38978" refString="Hedges SB, Duellman WE, Heinicke MP. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 2008; 1737: 1 - 182." type="journal article" year="2008">
|
||
Hedges
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||
<emphasis id="B908EA83FF9AFFE0C5B9C5FA90A148C1" box="[1173,1220,997,1021]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
</taxonomicName>
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||
.
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||
</paragraph>
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||
<paragraph id="8BC33691FF9AFFE0C215C23B903A4F07" blockId="40.[825,1119,1060,1084]" box="[825,1119,1060,1084]" pageId="40" pageNumber="41">
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<emphasis id="B908EA83FF9AFFE0C215C23B979A4F00" box="[825,1023,1060,1084]" italics="true" pageId="40" pageNumber="41">
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Genus:
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||
<taxonomicName id="4C7C4D12FF9AFFE0C2A4C23B979A4F00" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[904,1023,1060,1084]" family="Strabomantidae" genus="Urkuphryne" pageId="40" pageNumber="41" rank="genus" status="gen. nov.">Urkuphryne</taxonomicName>
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||
</emphasis>
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||
<taxonomicNameLabel id="A23B57F8FF9AFFE0C528C23B903A4F07" box="[1028,1119,1060,1084]" pageId="40" pageNumber="41" rank="genus">gen. nov.</taxonomicNameLabel>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C366651AFF9AFFE0C215C24191F94F49" box="[825,1436,1117,1141]" pageId="40" pageNumber="41" type="type_taxon">
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<paragraph id="8BC33691FF9AFFE0C215C24191F94F49" blockId="40.[825,1436,1117,1141]" box="[825,1436,1117,1141]" pageId="40" pageNumber="41">
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||
<emphasis id="B908EA83FF9AFFE0C215C24190E04F49" box="[825,1157,1117,1141]" italics="true" pageId="40" pageNumber="41">
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Type species:
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||
<taxonomicName id="4C7C4D12FF9AFFE0C293C24290E04F49" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[959,1157,1117,1141]" family="Strabomantidae" genus="Urkuphryne" pageId="40" pageNumber="41" rank="species" species="merinoi" status="sp. nov.">Urkuphryne merinoi</taxonomicName>
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||
</emphasis>
|
||
<taxonomicNameLabel id="A23B57F8FF9AFFE0C5A7C24190B34F49" box="[1163,1238,1118,1141]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(described below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFF9AFFE0C215C28890D54C3D" pageId="40" pageNumber="41" type="description">
|
||
<paragraph id="8BC33691FF9AFFE0C215C28890D54C3D" blockId="40.[824,1475,1175,1794]" pageId="40" pageNumber="41">
|
||
<emphasis id="B908EA83FF9AFFE0C215C28897FC4F93" box="[825,921,1175,1199]" italics="true" pageId="40" pageNumber="41">Definition</emphasis>
|
||
(
|
||
<tableCitation id="C6FE032AFF9AFFE0C283C288906D4F93" box="[943,1032,1175,1199]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="40" pageNumber="41" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="C6FE032AFF9AFFE0C56FC28890354F93" box="[1091,1104,1175,1199]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="40" pageNumber="41" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
|
||
<emphasis id="B908EA83FF9AFFE0C57CC28890064F92" box="[1104,1123,1175,1198]" italics="true" pageId="40" pageNumber="41">):</emphasis>
|
||
(1) Larger than its sister genus,
|
||
<taxonomicName id="4C7C4D12FF9AFFE0C215C2A997CA4FF2" authorityName="Heyer" authorityYear="1977" box="[825,943,1206,1230]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF9AFFE0C215C2A997CA4FF2" box="[825,943,1206,1230]" italics="true" pageId="40" pageNumber="41">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
(average adult female SVL =
|
||
<quantity id="4C849B74FF9AFFE0C5F7C2A9915C4FF1" box="[1243,1337,1206,1230]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="40" pageNumber="41" unit="mm" value="19.7">19.7 mm</quantity>
|
||
, adult male =
|
||
<quantity id="4C849B74FF9AFFE0C216C2CA97F04FD1" box="[826,917,1237,1261]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="40" pageNumber="41" unit="mm" value="17.2">17.2 mm</quantity>
|
||
,
|
||
<tableCitation id="C6FE032AFF9AFFE0C2B6C2CA97874FD1" box="[922,994,1237,1261]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="40" pageNumber="41" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
|
||
); (2) head wider than long, narrower than body, body robust; (3) columella, tympanic annulus and tympanic membrane present; (4) cranial crest absent; (5) finger I <finger II; (6) distal phalanges of fingers and toes rounded; (7) knees and heels without tubercles; (8) presence of inner tarsal fold; (9) fingers and toes lacking circumferential grooves and lateral dermal fringes; (10) toe V <toe III; (11) hand phalangeal formula 2-2- 3-3; foot phalangeal formula 2-2-3-4-3; (12) dark facial mask extending to the groins, triangular cloacal blotch present; (13) thin frontoparietals, longer than wide, big orbital fossae; (14) very prominent premaxillary and maxillary teeth present; (15) large nasal bones, almost in contact with maxilla; (16) zygomatic branch of squamosal shorter than otic branch; (17) large vomers with protruding vomerine teeth; (18) occipital condyles widely separated from each other; (19) prominent medial ridges present in all presacral and sacral vertebrae; (20) urostyle crest very prominent; (21) broadly expanded sacral diapophyses; (22) very large prepollex and prehallux present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFF9AFFE0C215C13C97D14C85" pageId="40" pageNumber="41" type="discussion">
|
||
<paragraph id="8BC33691FF9AFFE0C215C13C97D14C85" blockId="40.[825,1474,1827,1977]" pageId="40" pageNumber="41">
|
||
<emphasis id="B908EA83FF9AFFE0C215C13C97F24C07" box="[825,919,1827,1851]" italics="true" pageId="40" pageNumber="41">Remarks:</emphasis>
|
||
Autapomorphies for this genus are the reduction of the distal expansion of the phalanges of fingers and toes (
|
||
<figureCitation id="13472A14FF9AFFE0C45CC15C91D94C66" box="[1392,1468,1859,1883]" captionStart="Figure 7" captionStartId="14.[129,194,1836,1860]" captionTargetBox="[347,1255,145,1807]" captionTargetId="figure-62@14.[346,1257,144,1808]" captionTargetPageId="14" captionText="Figure 7. Ancestral trait reconstruction for discrete morphological characters for Phyllonastes and related genera. The five synapomorphic characters of Phyllonastes are shown. Stars indicate statistically significant support for the most likely character state." figureDoi="http://doi.org/10.5281/zenodo.14832707" httpUri="https://zenodo.org/record/14832707/files/figure.png" pageId="40" pageNumber="41">Figs 7D</figureCitation>
|
||
,
|
||
<figureCitation id="13472A14FF9AFFE0C215C17D97234C46" box="[825,838,1890,1914]" captionStart="Figure 8" captionStartId="16.[128,193,1174,1198]" captionTargetBox="[165,1437,148,1141]" captionTargetId="figure-444@16.[161,1441,144,1146]" captionTargetPageId="16" captionText="Figure 8. Ancestral trait reconstruction for the terminal expansion of distal phalanges for Phyllonastes and related genera. The characters analysed are the ratio between the terminal expansion and the length of the distal phalanx of the third finger and the fourth toe." figureDoi="http://doi.org/10.5281/zenodo.14832709" httpUri="https://zenodo.org/record/14832709/files/figure.png" pageId="40" pageNumber="41">8</figureCitation>
|
||
), the absence of supernumerary plantar tubercles (Supporting Information,
|
||
<figureCitation id="13472A14FF9AFFE0C2EBC19E90444CA5" box="[967,1057,1921,1945]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="40" pageNumber="41">Fig. S20</figureCitation>
|
||
), and protruding vomerine teeth (see ‘
|
||
<emphasis id="B908EA83FF9AFFE0C213C1BE97FB4C84" box="[831,926,1953,1976]" italics="true" pageId="40" pageNumber="41">Diagnosis</emphasis>
|
||
’).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF036619FF9BFFE1C15FC0DE948F4DC9" ID-DOI="http://doi.org/10.5281/zenodo.14832758" ID-Zenodo-Dep="14832758" httpUri="https://zenodo.org/record/14832758/files/figure.png" pageId="41" pageNumber="42" startId="41.[115,180,1729,1753]" targetBox="[149,1419,154,1695]" targetPageId="41" targetType="figure">
|
||
<paragraph id="8BC33691FF9BFFE1C15FC0DE948F4DC9" blockId="41.[113,1414,1729,1781]" pageId="41" pageNumber="42">
|
||
<emphasis id="B908EA83FF9BFFE1C15FC0DE94BD4DE5" bold="true" box="[115,216,1729,1753]" pageId="41" pageNumber="42">Figure 26.</emphasis>
|
||
Live and preserved specimens of
|
||
<taxonomicName id="4C7C4D12FF9BFFE1C33EC0DE96814DE4" authorityName="Harvey, Almendariz, Brito & Batallas" authorityYear="2013" box="[530,740,1729,1753]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="personinus">
|
||
<emphasis id="B908EA83FF9BFFE1C33EC0DE96814DE4" box="[530,740,1729,1753]" italics="true" pageId="41" pageNumber="42">Phyllonastes personinus</emphasis>
|
||
</taxonomicName>
|
||
. A, QCAZ 59437. B, QCAZ 71453. C, QCAZ 31118. A, B, in life. C, in preservative.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C366651AFF9BFFE2C15DC13A91534DFD" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="diagnosis">
|
||
<paragraph id="8BC33691FF9BFFE2C15DC13A95D44DFD" blockId="41.[113,764,1829,1978]" lastBlockId="42.[128,779,1423,1980]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
|
||
<emphasis id="B908EA83FF9BFFE1C15DC13A95384C01" box="[113,349,1829,1853]" italics="true" pageId="41" pageNumber="42">
|
||
Diagnosis:
|
||
<taxonomicName id="4C7C4D12FF9BFFE1C1CAC13A95384C01" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[230,349,1829,1853]" family="Strabomantidae" genus="Urkuphryne" pageId="41" pageNumber="42" rank="genus">Urkuphryne</taxonomicName>
|
||
</emphasis>
|
||
can be distinguished easily from its sister clade, the genus
|
||
<taxonomicName id="4C7C4D12FF9BFFE1C0A9C15B95994C60" authorityName="Heyer" authorityYear="1977" box="[389,508,1860,1884]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF9BFFE1C0A9C15B95994C60" box="[389,508,1860,1884]" italics="true" pageId="41" pageNumber="42">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
by (characteristics of
|
||
<taxonomicName id="4C7C4D12FF9BFFE1C15DC17C948D4C47" authorityName="Heyer" authorityYear="1977" box="[113,232,1891,1915]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF9BFFE1C15DC17C948D4C47" box="[113,232,1891,1915]" italics="true" pageId="41" pageNumber="42">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
shown in parentheses): (1) tips of fingers and toes subacuminate and rounded (tips pointed); (2) vomerine teeth present (vomerine teeth absent), and (3) larger size (
|
||
<tableCitation id="C6FE032AFF9BFFE1C3B3C1BD968F4C86" box="[671,746,1954,1978]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="41" pageNumber="42" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
|
||
). Osteologically,
|
||
<taxonomicName id="4C7C4D12FF9BFFE1C2EBC13A905B4C01" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[967,1086,1829,1853]" family="Strabomantidae" genus="Urkuphryne" pageId="41" pageNumber="42" rank="genus">
|
||
<emphasis id="B908EA83FF9BFFE1C2EBC13A905B4C01" box="[967,1086,1829,1853]" italics="true" pageId="41" pageNumber="42">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7C4D12FF9BFFE1C5EEC13A915C4C01" authorityName="Heyer" authorityYear="1977" box="[1218,1337,1829,1853]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF9BFFE1C5EEC13A915C4C01" box="[1218,1337,1829,1853]" italics="true" pageId="41" pageNumber="42">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
by having a shorter skull and a slightly broader maxilla, with a broader space between angulosplenials, thinner frontoparietals and larger orbital fossae (broader frontoparietals with smaller orbital fossae), bigger and more conspicuous premaxillary and maxillary teeth (premaxillary and maxillary teeth present but smaller), broader nasal bones almost in contact with the maxilla (nasal bones small, broadly separate from maxilla), vomers large and prominent, bearing protruding teeth (vomers reduced, broadly separated from each other, vomerine teeth absent), prominent medial ridges present on all presacral and sacral vertebrae (low medial ridges present only on presacral vertebrae), very prominent prepollex and prehallux (small prepollex and prehallux), and terminal phalanges rounded in fingers and toes (terminal phalanges expanded in fingers and toes).
|
||
</paragraph>
|
||
<caption id="DF036619FF98FFE2C1ADC35897EF4E63" ID-DOI="http://doi.org/10.5281/zenodo.14832760" ID-Zenodo-Dep="14832760" box="[129,906,1351,1375]" httpUri="https://zenodo.org/record/14832760/files/figure.png" pageId="42" pageNumber="43" startId="42.[129,194,1351,1375]" targetBox="[179,1428,174,1312]" targetPageId="42" targetType="figure">
|
||
<paragraph id="8BC33691FF98FFE2C1ADC35897EF4E63" blockId="42.[129,906,1351,1375]" box="[129,906,1351,1375]" pageId="42" pageNumber="43">
|
||
<emphasis id="B908EA83FF98FFE2C1ADC35894804E63" bold="true" box="[129,229,1351,1375]" pageId="42" pageNumber="43">Figure 27.</emphasis>
|
||
Skeletons of
|
||
<taxonomicName id="4C7C4D12FF98FFE2C04DC35896514E63" authorityName="Harvey, Almendariz, Brito & Batallas" authorityYear="2013" box="[353,564,1351,1375]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="personinus">
|
||
<emphasis id="B908EA83FF98FFE2C04DC35896514E63" box="[353,564,1351,1375]" italics="true" pageId="42" pageNumber="43">Phyllonastes personinus</emphasis>
|
||
</taxonomicName>
|
||
. A, QCAZ 58818. B, QCAZ 31115.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC33691FF98FFE2C1B0C0D797B64DBE" blockId="42.[128,779,1423,1980]" lastBlockId="42.[825,1475,1423,1666]" pageId="42" pageNumber="43">
|
||
<taxonomicName id="4C7C4D12FF98FFE2C1B0C0D795714DDC" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[156,276,1736,1760]" family="Strabomantidae" genus="Urkuphryne" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C1B0C0D795714DDC" box="[156,276,1736,1760]" italics="true" pageId="42" pageNumber="43">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7C4D12FF98FFE2C0B7C0D796634DDC" authorityName="Heyer" authorityYear="1969" box="[411,518,1736,1760]" class="Amphibia" family="Craugastoridae" genus="Barycholos" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C0B7C0D796634DDC" box="[411,518,1736,1760]" italics="true" pageId="42" pageNumber="43">Barycholos</emphasis>
|
||
</taxonomicName>
|
||
by the absence of dorsolateral folds (dorsolateral folds present), by the reduction of the distal expansion of the phalanges of fingers and toes, and by the arrangement of the vomerine teeth (small, oblique, broadly separated, and posteromedial to choanae in
|
||
<taxonomicName id="4C7C4D12FF98FFE2C335C15996F44C62" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[537,657,1862,1886]" family="Strabomantidae" genus="Urkuphryne" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C335C15996F44C62" box="[537,657,1862,1886]" italics="true" pageId="42" pageNumber="43">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
; and in two long arcuate series extending beyond the choana in
|
||
<taxonomicName id="4C7C4D12FF98FFE2C3A0C17A96924C41" authorityName="Heyer" authorityYear="1969" box="[652,759,1893,1917]" class="Amphibia" family="Craugastoridae" genus="Barycholos" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C3A0C17A96924C41" box="[652,759,1893,1917]" italics="true" pageId="42" pageNumber="43">Barycholos</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C7C4D12FF98FFE2C1ADC19B949C4CA0" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[129,249,1924,1948]" family="Strabomantidae" genus="Urkuphryne" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C1ADC19B949C4CA0" box="[129,249,1924,1948]" italics="true" pageId="42" pageNumber="43">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C7C4D12FF98FFE2C333C19B96064CA0" box="[543,611,1924,1948]" family="Strabomantidae" genus="Bahius" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C333C19B96064CA0" box="[543,611,1924,1948]" italics="true" pageId="42" pageNumber="43">Bahius</emphasis>
|
||
</taxonomicName>
|
||
by the absence of circumferential grooves of fingers and toes (both present in
|
||
<taxonomicName id="4C7C4D12FF98FFE2C27AC39097FF4E9B" box="[854,922,1423,1447]" family="Strabomantidae" genus="Bahius" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C27AC39097FF4E9B" box="[854,922,1423,1447]" italics="true" pageId="42" pageNumber="43">Bahius</emphasis>
|
||
</taxonomicName>
|
||
) and by the presence of an inconspicuous and thin inner tarsal fold (conspicuous, short, tubercle-like inner tarsal fold present in
|
||
<taxonomicName id="4C7C4D12FF98FFE2C2CAC3D1904F4EDA" box="[998,1066,1486,1510]" family="Strabomantidae" genus="Bahius" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C2CAC3D1904F4EDA" box="[998,1066,1486,1510]" italics="true" pageId="42" pageNumber="43">Bahius</emphasis>
|
||
</taxonomicName>
|
||
). The presence of a tympanic membrane, tympanic annulus, and inner tarsal tubercle differentiates
|
||
<taxonomicName id="4C7C4D12FF98FFE2C241C01397804D18" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[877,997,1548,1572]" family="Strabomantidae" genus="Urkuphryne" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C241C01397804D18" box="[877,997,1548,1572]" italics="true" pageId="42" pageNumber="43">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C7C4D12FF98FFE2C502C01390C94D18" authorityName="Griffiths" authorityYear="1959" box="[1070,1196,1548,1572]" class="Amphibia" family="Craugastoridae" genus="Euparkerella" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C502C01390C94D18" box="[1070,1196,1548,1572]" italics="true" pageId="42" pageNumber="43">Euparkerella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D12FF98FFE2C5C6C013912F4D18" authorityName="Miranda-Ribeiro" authorityYear="1920" box="[1258,1354,1548,1572]" class="Amphibia" family="Craugastoridae" genus="Holoaden" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C5C6C013912F4D18" box="[1258,1354,1548,1572]" italics="true" pageId="42" pageNumber="43">Holoaden</emphasis>
|
||
</taxonomicName>
|
||
(tympanic membrane, tympanic annulus, and tarsal tubercle absent), two genera sequentially sister to
|
||
<taxonomicName id="4C7C4D12FF98FFE2C573C05490B24D5F" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1119,1239,1611,1635]" family="Strabomantidae" genus="Urkuphryne" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C573C05490B24D5F" box="[1119,1239,1611,1635]" italics="true" pageId="42" pageNumber="43">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C7C4D12FF98FFE2C5D8C05491084D5F" authority="Baryc" authorityName="Heyer" authorityYear="1977" box="[1268,1389,1611,1635]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="42" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C5D8C05491084D5F" box="[1268,1389,1611,1635]" italics="true" pageId="42" pageNumber="43">Phyllonastes</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<emphasis id="B908EA83FF98FFE2C4A6C05397A84DBE" italics="true" pageId="42" pageNumber="43">
|
||
Baryc holos +
|
||
<taxonomicName id="4C7C4D12FF98FFE2C2A5C07597A84DBE" box="[905,973,1642,1666]" family="Strabomantidae" genus="Bahius" pageId="42" pageNumber="43" rank="genus">Bahius</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC33691FF98FFE2C215C0B591534DFD" blockId="42.[825,1334,1706,1730]" box="[825,1334,1706,1730]" pageId="42" pageNumber="43">
|
||
<emphasis id="B908EA83FF98FFE2C215C0B597EA4DFD" box="[825,911,1706,1729]" italics="true" pageId="42" pageNumber="43">Content:</emphasis>
|
||
One species,
|
||
<taxonomicName id="4C7C4D12FF98FFE2C533C0B590804DFD" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[1055,1253,1706,1730]" family="Strabomantidae" genus="Urkuphryne" pageId="42" pageNumber="43" rank="species" species="merinoi" status="sp. nov.">
|
||
<emphasis id="B908EA83FF98FFE2C533C0B590804DFD" box="[1055,1253,1706,1730]" italics="true" pageId="42" pageNumber="43">Urkuphryne merinoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B57F8FF98FFE2C5C6C0B591534DFD" box="[1258,1334,1706,1729]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFF98FFE2C215C0F6913D4C63" pageId="42" pageNumber="43" type="etymology">
|
||
<paragraph id="8BC33691FF98FFE2C215C0F6913D4C63" blockId="42.[825,1475,1769,1887]" pageId="42" pageNumber="43">
|
||
<emphasis id="B908EA83FF98FFE2C215C0F697CD4C3D" box="[825,936,1769,1793]" italics="true" pageId="42" pageNumber="43">Etymology:</emphasis>
|
||
The name refers to the
|
||
<typeStatus id="54C78833FF98FFE2C5B3C0F690A84C3D" box="[1183,1229,1769,1793]" pageId="42" pageNumber="43">type</typeStatus>
|
||
locality of the
|
||
<typeStatus id="54C78833FF98FFE2C448C0F691F74C3D" box="[1380,1426,1769,1793]" pageId="42" pageNumber="43">type</typeStatus>
|
||
species, ‘Cerro Golondrinas protective forest’ (Golondrinas hill).
|
||
<taxonomicName id="4C7C4D12FF98FFE2C215C13797D54C7C" authorityName="Ortega & Cisneros-Heredia & Camper & Romero-Carvajal & Negrete & Ron" authorityYear="2025" box="[825,944,1832,1856]" family="Strabomantidae" genus="Urkuphryne" pageId="42" pageNumber="43" rank="genus">
|
||
<emphasis id="B908EA83FF98FFE2C215C13797D54C7C" box="[825,944,1832,1856]" italics="true" pageId="42" pageNumber="43">Urkuphryne</emphasis>
|
||
</taxonomicName>
|
||
is a compound word:
|
||
<emphasis id="B908EA83FF98FFE2C5A3C13790A74C7C" box="[1167,1218,1832,1856]" italics="true" pageId="42" pageNumber="43">Urku</emphasis>
|
||
means ‘hill’ in the Native American Quichua language, and
|
||
<emphasis id="B908EA83FF98FFE2C5BDC15890B04C63" box="[1169,1237,1863,1887]" italics="true" pageId="42" pageNumber="43">phryne</emphasis>
|
||
means ‘frog’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366651AFF98FFE2C215C19F90A44C84" pageId="42" pageNumber="43" type="distribution">
|
||
<paragraph id="8BC33691FF98FFE2C215C19F90A44C84" blockId="42.[824,1474,1920,1976]" pageId="42" pageNumber="43">
|
||
<emphasis id="B908EA83FF98FFE2C215C19F97D24CA4" box="[825,951,1920,1944]" italics="true" pageId="42" pageNumber="43">Distribution:</emphasis>
|
||
Pacific Basin of Northern
|
||
<collectingCountry id="F36B7601FF98FFE2C593C19F91734CA4" box="[1215,1302,1920,1944]" name="Ecuador" pageId="42" pageNumber="43">Ecuador</collectingCountry>
|
||
, western Andean slopes at elevations
|
||
<quantity id="4C849B74FF98FFE2C52CC18090F54C8B" box="[1024,1168,1951,1975]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="42" pageNumber="43" unit="m" value="2650.0" valueMax="2800.0" valueMin="2500.0">2500–2800 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |