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<document id="F22D985373D435BCB8BB438C527CEB5E" ID-DOI="10.1093/zoolinnean/zlae162" ID-ISSN="0024-4082" ID-Zenodo-Dep="14832689" ID-ZooBank="7-01-2184" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1738158242914" checkinUser="plazi" docAuthor="Ortega, Jhael A., Cisneros-Heredia, Diego F., Camper, Jeffrey D., Romero-Carvajal, Andrés, Negrete, Leonardo &amp; Ron, Santiago R." docDate="2025" docId="03D58787FF92FFEBC2F4C011965B4CF8" docLanguage="en" docName="zlae162.pdf" docOrigin="Zoological Journal of the Linnean Society 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae162" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Phyllonastes plateadensis Ortega, Cisneros-Heredia, Camper, Romero-Carvajal, Negrete &amp; Ron, 2025, sp.nov." docType="treatment" docVersion="4" lastPageNumber="36" masterDocId="FFECFFFFFFB2FFC8C12CC61F94654B3C" masterDocTitle="Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography" masterLastPageNumber="60" masterPageNumber="1" pageNumber="33" updateTime="1738929907134" updateUser="ExternalLinkService">
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<mods:titleInfo id="96A5F824C5F8D769927979948ADEC38B">
<mods:title id="FEE2278930CCB0476C5A527C848ABFCF">Systematics of minute strabomantid frogs allocated to the genus Noblella (Amphibia: Anura) with description of a new genus, seven new species, and insights into historical biogeography</mods:title>
</mods:titleInfo>
<mods:name id="81C86D9AB56C194EDC2AD4895A60B7AA" type="personal">
<mods:role id="2375345AA0B0E32897F943208A4E014A">
<mods:roleTerm id="C0F2C7228F152733E3CF3B56C23BAE78">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CDDC531855D5BD3DDC7BAF10F9E46E96">Ortega, Jhael A.</mods:namePart>
<mods:affiliation id="43FCB945D512E228601292A35B95A7C2">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
</mods:name>
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<mods:role id="A54E826A1F67667B32DB55CCD77310B8">
<mods:roleTerm id="A47CEE6BA7565B03FF776C6F8DE55DC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="60CB36D5B99D3D9433F5985D96D61E6F">Cisneros-Heredia, Diego F.</mods:namePart>
<mods:affiliation id="C6B6F374F1124D2F57C2740C2ED68DFD">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Museo de Zoología / Laboratorio de Zoología Terrestre, Quito, Ecuador &amp; Instituto Nacional de Biodiversidad INABIO, División de Herpetología, Quito, Ecuador</mods:affiliation>
</mods:name>
<mods:name id="D68C4DEA2250FCB430C89909D015FF7C" type="personal">
<mods:role id="C77EAC1AB33281445745238FAA8F6E18">
<mods:roleTerm id="B67049B8D1B551978E274F31F664F5BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F96031E03FB410586C6998808278EE33">Camper, Jeffrey D.</mods:namePart>
<mods:affiliation id="B18591C1EC16BDFD392F9BE95897A938">Department of Biology, Francis Marion University, Florence, SC 29506, USA</mods:affiliation>
</mods:name>
<mods:name id="9F83ADAB3D7178AA998DFE3B6FB7A249" type="personal">
<mods:role id="8E3CDA12C5398312D44307802C0F4536">
<mods:roleTerm id="E230BA5E0DE7C565FCE33D21DC204EB1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC76AF9159CB7DC24D10348D78E63063">Romero-Carvajal, Andrés</mods:namePart>
<mods:affiliation id="29A9D125736E5FBC4FE50BC2AED2D042">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
</mods:name>
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<mods:role id="1C51B89B7CEEAA726B60EB14C18F911A">
<mods:roleTerm id="3B57C8D4EE3D9A5F12CA11C555EE0F3A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD2D2FA1FBFDC8C11D0BD6A993C0FB05">Negrete, Leonardo</mods:namePart>
<mods:affiliation id="F1060F5E8F9C4011FCFE773F960E02FD">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<mods:role id="39341629ABD701C25B17F58680238482">
<mods:roleTerm id="A5FC4CDC6495F0A2271DBB5E7A93D5B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7B74DA5E70FE4D9898FF776582DCA46">Ron, Santiago R.</mods:namePart>
<mods:affiliation id="1E2CC4C89C54B4ABA0228EDED97A9138">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
<mods:nameIdentifier id="468A4FAC2B922F0CAB09105C4F0C8949" type="email">santiago.r.ron@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="145377CB6B7E0ECC26D67FEA725AEF99">text</mods:typeOfResource>
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<mods:title id="AC9FA64570DEA1344AFDFDDCC9CFCFAB">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="E157DC2AA6F43E159BC3ACA4ECEE091B">
<mods:date id="B0462FAAD7627662DE3D32C097FB5DC2">2025</mods:date>
<mods:detail id="86A45ABFFD95BD17A52C963B4A13FFC3" type="pubDate">
<mods:number id="D5DDCDF7FA884AA85A9AE2139CB64A91">2025-01-13</mods:number>
</mods:detail>
<mods:detail id="A8AFC512095A7E96A58525E01BB511CA" type="volume">
<mods:number id="34503046975A0029B1E6538AA4E351F6">203</mods:number>
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<mods:number id="AB2C993890A979F269CC27FC9BBEBE72">1</mods:number>
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<mods:start id="437772C4CCD60EDC46DE57263DA70D2C">1</mods:start>
<mods:end id="B6BC05EF4F34AD3AAA220BE833FE6642">60</mods:end>
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<mods:location id="A64BA53F4F8BBB5416B860F0521EDB1A">
<mods:url id="4E1AD405FD1D6869B0A8BC5401172DBE">https://doi.org/10.1093/zoolinnean/zlae162</mods:url>
</mods:location>
<mods:classification id="C6580AAD7465930886379B36A1FCAC4E">journal article</mods:classification>
<mods:identifier id="2CCE87CE1F59DB0EABB9604964A3E1C0" type="DOI">10.1093/zoolinnean/zlae162</mods:identifier>
<mods:identifier id="192A97576F98337030BDC5171ED35E66" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="BC703226FB60FB4DBA9D6372917537F7" type="Zenodo-Dep">14832689</mods:identifier>
<mods:identifier id="FD4E10A37985384A559848C5F088571F" type="ZooBank">7-01-2184</mods:identifier>
</mods:mods>
<treatment id="03D58787FF92FFEBC2F4C011965B4CF8" ID-DOI="http://doi.org/10.5281/zenodo.14832719" ID-Zenodo-Dep="14832719" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8" httpUri="http://treatment.plazi.org/id/03D58787FF92FFEBC2F4C011965B4CF8" lastPageId="35" lastPageNumber="36" pageId="32" pageNumber="33">
<subSubSection id="C366651AFF92FFE8C2F4C01191464D15" box="[984,1315,1550,1577]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BC33691FF92FFE8C2F4C01191464D15" blockId="32.[825,1474,1550,1645]" box="[984,1315,1550,1577]" pageId="32" pageNumber="33">
<emphasis id="B908EA83FF92FFE8C2F4C01191464D15" bold="true" box="[984,1315,1550,1577]" pageId="32" pageNumber="33">
<taxonomicName id="4C7C4D12FF92FFE8C2F4C01190B64D15" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[984,1235,1550,1577]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="plateadensis" status="sp. nov.">
<emphasis id="B908EA83FF92FFE8C2F4C01190B64D15" bold="true" box="[984,1235,1550,1577]" italics="true" pageId="32" pageNumber="33">Phyllonastes plateadensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57F8FF92FFE8C5FBC01091464D15" box="[1239,1315,1551,1577]" pageId="32" pageNumber="33" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C366651AFF92FFE8C215C02997B94D51" pageId="32" pageNumber="33" type="description">
<paragraph id="8BC33691FF92FFE8C215C02997B94D51" blockId="32.[825,1474,1550,1645]" pageId="32" pageNumber="33">
LSID:
<uri id="FFED3A93FF92FFE8C251C02A97B94D51" pageId="32" pageNumber="33">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C366651AFF92FFE9C215C091955A4A19" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BC33691FF92FFE8C215C09191324C5E" blockId="32.[822,1475,1678,1894]" pageId="32" pageNumber="33">
<materialsCitation id="3B143CCCFF92FFE8C215C09191324C5E" collectingDate="2016-09-23" collectionCode="QCAZ" collectorName="Diego Almeida &amp; Kunam Nusirquia &amp; Fernando Ayala &amp; Javier Pinto &amp; Alex Achig" elevation="2055" latitude="-4.6027" location="ZamoraChinchipe Province" longLatPrecision="7" longitude="-78.831" municipality="Cerro Plateado Biological Reserve" pageId="32" pageNumber="33" specimenCode="QCAZ 65015" specimenCount="1" typeStatus="holotype">
<emphasis id="B908EA83FF92FFE8C215C09191314D9A" box="[825,1364,1678,1702]" italics="true" pageId="32" pageNumber="33">
<typeStatus id="54C78833FF92FFE8C215C09197F74D9A" box="[825,914,1678,1702]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
(
<figureCitation id="13472A14FF92FFE8C285C09097904D9A" box="[937,1013,1678,1702]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="32" pageNumber="33">Figs 20</figureCitation>
,
<figureCitation id="13472A14FF92FFE8C52BC09190444D9A" box="[1031,1057,1678,1702]" captionStart="Figure 21" captionStartId="35.[113,178,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-421@35.[306,1266,144,1064]" captionTargetPageId="35" captionText="Figure 21. Palmar (A) and plantar (B) surfaces of Phyllonastes plateadensis. Photographs of right hand and right foot of the holotype QCAZ 65015." figureDoi="http://doi.org/10.5281/zenodo.14832748" httpUri="https://zenodo.org/record/14832748/files/figure.png" pageId="32" pageNumber="33">21</figureCitation>
,
<figureCitation id="13472A14FF92FFE8C51EC091903E4D9A" box="[1074,1115,1678,1702]" captionStart="Figure 22" captionStartId="36.[129,194,1772,1796]" captionTargetBox="[251,1360,158,1734]" captionTargetId="figure-100@36.[226,1376,144,1744]" captionTargetPageId="36" captionText="Figure 22. Skulls of Phyllonastes species.A, Phyllonastes myrmecoides QCAZ 53583. B, Phyllonastes plateadensis QCAZ 65015. C, Phyllonastes sardinayacu, QCAZ 58821." figureDoi="http://doi.org/10.5281/zenodo.14832750" httpUri="https://zenodo.org/record/14832750/files/figure.png" pageId="32" pageNumber="33">22B</figureCitation>
,
<figureCitation id="13472A14FF92FFE8C540C09190F24D9A" box="[1132,1175,1678,1702]" captionStart="Figure 23" captionStartId="37.[112,177,1566,1590]" captionTargetBox="[148,1425,146,1537]" captionTargetId="figure-197@37.[146,1426,144,1539]" captionTargetPageId="37" captionText="Figure 23. Skeletons of Phyllonastes species. A, Phyllonastes plateadensis QCAZ 65015, SVL = 15.70 mm. B, Phyllonastes sardinayacu QCAZ 58821, SVL = 14.32 mm. C, Phyllonastes personinus QCAZ 58818, SVL = 16.91 mm." figureDoi="http://doi.org/10.5281/zenodo.14832752" httpUri="https://zenodo.org/record/14832752/files/figure.png" pageId="32" pageNumber="33">23A</figureCitation>
):
<specimenCode id="DBDA9EEAFF92FFE8C5EEC09191314D9A" box="[1218,1364,1678,1702]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="32" pageNumber="33" type="Museum">QCAZ 65015</specimenCode>
</emphasis>
(field no. SC-PUCE56489)
<specimenCount id="9D7AFD18FF92FFE8C2D9C0B290984DF9" box="[1013,1277,1709,1733]" pageId="32" pageNumber="33">adultfemalefromEcuador</specimenCount>
,
<location id="8EA3604AFF92FFE8C42FC0B297F74DD9" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF92FFE8C42FC0B297F74DD9" latitude="-4.6027" longLatPrecision="7" longitude="-78.831" municipality="Cerro Plateado Biological Reserve" name="ZamoraChinchipe Province" pageId="32" pageNumber="33">ZamoraChinchipe Province</location>
,
<location id="8EA3604AFF92FFE8C28DC0D290294DD9" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF92FFE8C28DC0D290294DD9" box="[929,1100,1741,1765]" latitude="-4.6027" longLatPrecision="7" longitude="-78.831" municipality="Cerro Plateado Biological Reserve" name="Canton Palanda" pageId="32" pageNumber="33">Canton Palanda</location>
,
<location id="8EA3604AFF92FFE8C576C0D291774DD9" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF92FFE8C576C0D291774DD9" box="[1114,1298,1741,1765]" latitude="-4.6027" longLatPrecision="7" longitude="-78.831" municipality="Cerro Plateado Biological Reserve" name="La Canela Parish" pageId="32" pageNumber="33">La Canela Parish</location>
,
<collectingMunicipality id="6BA7ACEBFF92FFE8C40CC0D2979D4C38" pageId="32" pageNumber="33">Cerro Plateado Biological Reserve</collectingMunicipality>
,
<location id="8EA3604AFF92FFE8C52EC0F390994C38" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF92FFE8C52EC0F390994C38" box="[1026,1276,1772,1796]" latitude="-4.6027" longLatPrecision="7" longitude="-78.831" municipality="Cerro Plateado Biological Reserve" name="surroundings of Camp 3" pageId="32" pageNumber="33">surroundings of Camp 3</location>
,
<location id="8EA3604AFF92FFE8C42BC0F397CF4C18" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF92FFE8C42BC0F397CF4C18" latitude="-4.6027" longLatPrecision="7" longitude="-78.831" municipality="Cerro Plateado Biological Reserve" name="trail from Mirador to Camp 3" pageId="32" pageNumber="33">trail from Mirador to Camp 3</location>
(
<geoCoordinate id="EE485056FF92FFE8C296C114907E4C1F" box="[954,1051,1803,1827]" degrees="4.6027" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="5" value="-4.6027">4.6027°S</geoCoordinate>
,
<geoCoordinate id="EE485056FF92FFE8C50AC11390C54C18" box="[1062,1184,1804,1828]" degrees="78.8310" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5" value="-78.831">78.8310°W</geoCoordinate>
),
<elevation id="0051D1A2FF92FFE8C59FC11491524C1F" box="[1203,1335,1803,1827]" metricMagnitude="3" metricUnit="m" metricValue="2.055" pageId="32" pageNumber="33" unit="m" value="2055.0">
<quantity id="4C849B74FF92FFE8C59FC11491614C1F" box="[1203,1284,1803,1827]" metricMagnitude="3" metricUnit="m" metricValue="2.055" pageId="32" pageNumber="33" unit="m" value="2055.0">2055 m</quantity>
a.s.l.
</elevation>
Collected by
<collectorName id="26895347FF92FFE8C215C13497B04C7F" box="[825,981,1835,1859]" pageId="32" pageNumber="33">Diego Almeida</collectorName>
,
<collectorName id="26895347FF92FFE8C2F1C13490F34C7F" box="[989,1174,1835,1859]" pageId="32" pageNumber="33">Kunam Nusirquia</collectorName>
,
<collectorName id="26895347FF92FFE8C5B1C134915A4C7F" box="[1181,1343,1835,1859]" pageId="32" pageNumber="33">Fernando Ayala</collectorName>
,
<collectorName id="26895347FF92FFE8C464C13491DA4C7F" box="[1352,1471,1835,1859]" pageId="32" pageNumber="33">Javier Pinto</collectorName>
,
<collectorName id="26895347FF92FFE8C215C15597CE4C5A" box="[825,939,1866,1894]" pageId="32" pageNumber="33">Alex Áchig</collectorName>
, and Malki Bustos on
<date id="FFC21051FF92FFE8C5A5C15591314C5E" box="[1161,1364,1866,1890]" pageId="32" pageNumber="33" value="2016-09-23">
<collectingDate id="EF86E9B9FF92FFE8C5A5C15591314C5E" box="[1161,1364,1866,1890]" pageId="32" pageNumber="33" value="2016-09-23">23 September 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC33691FF92FFE9C215C19B955A4A19" blockId="32.[824,1474,1923,1979]" lastBlockId="33.[113,764,144,293]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<materialsCitation id="3B143CCCFF92FFE9C215C19B95594A19" collectingDate="2016-09-26" collectionCode="QCAZ" collectorName="Diego Almeida &amp; Kunam Nusirquia &amp; Javier Pinto &amp; Alex Achig &amp; Malki Bustos" country="Ecuador" elevation="1980" lastPageId="33" lastPageNumber="34" latitude="-4.5993" location="Canton Palanda" longLatPrecision="7" longitude="-78.835" municipality="Biological Reserve" pageId="32" pageNumber="33" specimenCode="QCAZ 65016" specimenCount="1" stateProvince="Zamora Chinchipe Province" typeStatus="paratype">
<emphasis id="B908EA83FF92FFE8C215C19B90264CA7" box="[825,1091,1924,1948]" italics="true" pageId="32" pageNumber="33">
<typeStatus id="54C78833FF92FFE8C215C19B97FD4CA7" box="[825,920,1924,1947]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
(N = 1;
<figureCitation id="13472A14FF92FFE8C2DCC19B90554CA0" box="[1008,1072,1924,1948]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="32" pageNumber="33">Fig. 20</figureCitation>
):
</emphasis>
<specimenCode id="DBDA9EEAFF92FFE8C566C19B90BD4CA7" box="[1098,1240,1923,1948]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="32" pageNumber="33" type="Museum">QCAZ 65016</specimenCode>
<specimenCount id="9D7AFD18FF92FFE8C5F1C19C914B4CA7" box="[1245,1326,1923,1947]" pageId="32" pageNumber="33" type="juvenile">juvenile</specimenCount>
from
<collectingCountry id="F36B7601FF92FFE8C44BC19C91DB4CA7" box="[1383,1470,1923,1947]" name="Ecuador" pageId="32" pageNumber="33">Ecuador</collectingCountry>
,
<collectingRegion id="49B8F873FF92FFE8C214C1BC903F4C87" box="[824,1114,1955,1979]" country="Ecuador" name="Zamora-Chinchipe" pageId="32" pageNumber="33">Zamora Chinchipe Province</collectingRegion>
,
<location id="8EA3604AFF92FFE8C54FC1BC916D4C87" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF92FFE8C54FC1BC916D4C87" box="[1123,1288,1955,1979]" country="Ecuador" latitude="-4.5993" longLatPrecision="7" longitude="-78.835" municipality="Biological Reserve" name="Canton Palanda" pageId="32" pageNumber="33" stateProvince="Zamora Chinchipe Province">Canton Palanda</location>
,
<location id="8EA3604AFF92FFE8C43CC1BC91DB4C87" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF92FFE8C43CC1BC91DB4C87" box="[1296,1470,1955,1979]" country="Ecuador" latitude="-4.5993" longLatPrecision="7" longitude="-78.835" municipality="Biological Reserve" name="La Canela Parish" pageId="32" pageNumber="33" stateProvince="Zamora Chinchipe Province">La Canela Parish</location>
,
<location id="8EA3604AFF93FFE9C15DC68F95BB4B94" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF93FFE9C15DC68F95BB4B94" box="[113,478,144,168]" country="Ecuador" latitude="-4.5993" longLatPrecision="7" longitude="-78.835" municipality="Biological Reserve" name="Biological Reserve Cerro Plateado" pageId="33" pageNumber="34" stateProvince="Zamora Chinchipe Province">Biological Reserve Cerro Plateado</location>
,
<location id="8EA3604AFF93FFE9C0C1C68F96934B94" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF93FFE9C0C1C68F96934B94" box="[493,758,144,168]" country="Ecuador" latitude="-4.5993" longLatPrecision="7" longitude="-78.835" municipality="Biological Reserve" name="surroundings of Camp 3" pageId="33" pageNumber="34" stateProvince="Zamora Chinchipe Province">surroundings of Camp 3</location>
,
<location id="8EA3604AFF93FFE9C15DC6B096644BF4" LSID="urn:lsid:plazi:treatment:03D58787FF92FFEBC2F4C011965B4CF8:8EA3604AFF93FFE9C15DC6B096644BF4" box="[113,513,175,200]" country="Ecuador" latitude="-4.5993" longLatPrecision="7" longitude="-78.835" municipality="Biological Reserve" name="trail of the upper section of Rio Negro" pageId="33" pageNumber="34" stateProvince="Zamora Chinchipe Province">trail of the upper section of Río Negro</location>
(
<geoCoordinate id="EE485056FF93FFE9C323C6B096154BFB" box="[527,624,175,199]" degrees="4.5993" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="-4.5993">4.5993°S</geoCoordinate>
,
<geoCoordinate id="EE485056FF93FFE9C357C6B096964BFB" box="[635,755,175,199]" degrees="78.8350" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="5" value="-78.835">78.8350°W</geoCoordinate>
)
<elevation id="0051D1A2FF93FFE9C15FC6D0949C4BDB" box="[115,249,207,231]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="33" pageNumber="34" unit="m" value="1980.0">
<quantity id="4C849B74FF93FFE9C15FC6D094A04BDA" box="[115,197,207,231]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="33" pageNumber="34" unit="m" value="1980.0">1980 m</quantity>
a.s.l.
</elevation>
Collected by
<collectorName id="26895347FF93FFE9C0BCC6D096554BDB" box="[400,560,207,231]" pageId="33" pageNumber="34">Diego Almeida</collectorName>
,
<collectorName id="26895347FF93FFE9C310C6D0969C4BDB" box="[572,761,207,231]" pageId="33" pageNumber="34">Kunam Nusirquia</collectorName>
, Fernando Ayala,
<collectorName id="26895347FF93FFE9C00EC6F195F94A3A" box="[290,412,238,262]" pageId="33" pageNumber="34">Javier Pinto</collectorName>
,
<collectorName id="26895347FF93FFE9C084C6F1967C4A36" box="[424,537,238,266]" pageId="33" pageNumber="34">Alex Áchig</collectorName>
and
<collectorName id="26895347FF93FFE9C363C6F196B24A39" box="[591,727,238,262]" pageId="33" pageNumber="34">Malki Bustos</collectorName>
on
<date id="FFC21051FF93FFE9C15DC71295594A19" box="[113,316,269,293]" pageId="33" pageNumber="34" value="2016-09-26">
<collectingDate id="EF86E9B9FF93FFE9C15DC71295594A19" box="[113,316,269,293]" pageId="33" pageNumber="34" value="2016-09-26">26 September 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C366651AFF93FFE9C15DC75894AA4A8B" pageId="33" pageNumber="34" type="vernacular_names">
<paragraph id="8BC33691FF93FFE9C15DC75896854A63" blockId="33.[113,736,327,351]" box="[113,736,327,351]" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C15DC75895D64A62" box="[113,435,327,351]" italics="true" pageId="33" pageNumber="34">Proposed standard English name:</emphasis>
Cerro Plateado leaflitter frog.
</paragraph>
<paragraph id="8BC33691FF93FFE9C15DC79F94AA4A8B" blockId="33.[113,762,384,439]" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C15DC79F95A74AA4" box="[113,450,384,408]" italics="true" pageId="33" pageNumber="34">Proposed standard Spanish name:</emphasis>
Cutín de Hojarasca de Cerro Plateado.
</paragraph>
</subSubSection>
<subSubSection id="C366651AFF93FFE9C15DC7C695574EA1" pageId="33" pageNumber="34" type="description">
<paragraph id="8BC33691FF93FFE9C15DC7C695574EA1" blockId="33.[112,764,473,1437]" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C15DC7C694B64ACD" box="[113,211,473,497]" italics="true" pageId="33" pageNumber="34">Definition</emphasis>
(
<figureCitation id="13472A14FF93FFE9C1CBC7C695564ACD" box="[231,307,473,497]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="33" pageNumber="34">Figs 20</figureCitation>
,
<figureCitation id="13472A14FF93FFE9C06DC7C6953E4ACD" box="[321,347,473,497]" captionStart="Figure 21" captionStartId="35.[113,178,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-421@35.[306,1266,144,1064]" captionTargetPageId="35" captionText="Figure 21. Palmar (A) and plantar (B) surfaces of Phyllonastes plateadensis. Photographs of right hand and right foot of the holotype QCAZ 65015." figureDoi="http://doi.org/10.5281/zenodo.14832748" httpUri="https://zenodo.org/record/14832748/files/figure.png" pageId="33" pageNumber="34">21</figureCitation>
,
<figureCitation id="13472A14FF93FFE9C045C7C695F64ACD" box="[361,403,473,497]" captionStart="Figure 22" captionStartId="36.[129,194,1772,1796]" captionTargetBox="[251,1360,158,1734]" captionTargetId="figure-100@36.[226,1376,144,1744]" captionTargetPageId="36" captionText="Figure 22. Skulls of Phyllonastes species.A, Phyllonastes myrmecoides QCAZ 53583. B, Phyllonastes plateadensis QCAZ 65015. C, Phyllonastes sardinayacu, QCAZ 58821." figureDoi="http://doi.org/10.5281/zenodo.14832750" httpUri="https://zenodo.org/record/14832750/files/figure.png" pageId="33" pageNumber="34">22B</figureCitation>
,
<figureCitation id="13472A14FF93FFE9C08CC7C695AE4ACD" box="[416,459,473,497]" captionStart="Figure 23" captionStartId="37.[112,177,1566,1590]" captionTargetBox="[148,1425,146,1537]" captionTargetId="figure-197@37.[146,1426,144,1539]" captionTargetPageId="37" captionText="Figure 23. Skeletons of Phyllonastes species. A, Phyllonastes plateadensis QCAZ 65015, SVL = 15.70 mm. B, Phyllonastes sardinayacu QCAZ 58821, SVL = 14.32 mm. C, Phyllonastes personinus QCAZ 58818, SVL = 16.91 mm." figureDoi="http://doi.org/10.5281/zenodo.14832752" httpUri="https://zenodo.org/record/14832752/files/figure.png" pageId="33" pageNumber="34">23A</figureCitation>
;
<tableCitation id="C6FE032AFF93FFE9C0F0C7C696564ACD" box="[476,563,473,497]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="33" pageNumber="34" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
and
<tableCitation id="C6FE032AFF93FFE9C346C7C696124ACD" box="[618,631,473,497]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="33" pageNumber="34" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
<emphasis id="B908EA83FF93FFE9C35BC7C696EF4ACC" box="[631,650,473,496]" italics="true" pageId="33" pageNumber="34">):</emphasis>
We assign the new species to the genus
<taxonomicName id="4C7C4D12FF93FFE9C0BBC7E79675492C" authorityName="Heyer" authorityYear="1977" box="[407,528,504,528]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B908EA83FF93FFE9C0BBC7E79675492C" box="[407,528,504,528]" italics="true" pageId="33" pageNumber="34">Phyllonastes</emphasis>
</taxonomicName>
based on its phylogenetic relationships. The new species is characterized by: (1) skin on dorsum and flanks finely shagreen, dorsolateral folds absent, ventral surfaces weakly areolate, discoidal fold present; (2) tympanic membrane not differentiated, tympanic annulus present but inconspicuous, supratympanic fold and postrictal tubercles absent; (3) snout broadly rounded in dorsal view and rounded and lateral view; (4) upper eyelid without tubercles, cranial crests absent; (5) vomerine teeth absent; (6) vocal slits and nuptial pads unknown; (7) fingers not expanded distally, finger tips subacuminate, without papillae, finger I shorter than finger II, supernumerary tubercles present; (8) fingers bearing narrow lateral fringes, ill-defined pads and circumferential grooves; (9) distal phalanges blunt or T-shaped, phalangeal formula of hand 2-2-3-3; (10) ulnar tubercles absent; (11) knees and heels without tubercles, outer edge of tarsus without tubercles, inner edge of tarsus bearing one rounded tubercle; (12) inner metatarsal tubercle small, rounded in ventral view and very low in lateral view, smaller than rounded outer metatarsal tubercle; (13) toes slightly expanded distally, with acuminate toe tips and lacking papillae, supernumerary tubercles ill defined, narrow lateral fringes on toes, toe basal webbing absent, all toes bearing ill-defined circumferential grooves, toe V shorter than Toe III; (14) in life, dorsal surfaces light brown with scattered white dots; circular dark brown sacral lateral marks delineated by a paler line, cloacal region cream surrounded by a dark brown stripe, facial mask dark brown extending from tip of snout to groins, flanks brown, ventral surfaces brown with white spots; and (15) SVL in adult males unknown, adult female
<quantity id="4C849B74FF93FFE9C3BFC37A969F4E41" box="[659,762,1381,1405]" metricMagnitude="-2" metricUnit="m" metricValue="1.935" pageId="33" pageNumber="34" unit="mm" value="19.35">19.35 mm</quantity>
(
<emphasis id="B908EA83FF93FFE9C151C39A94F54EA0" box="[125,144,1413,1436]" italics="true" pageId="33" pageNumber="34">N</emphasis>
= 1) (
<tableCitation id="C6FE032AFF93FFE9C1FAC39A95444EA1" box="[214,289,1413,1437]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="33" pageNumber="34" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366651AFF93FFE9C15DC3D2954B4C67" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BC33691FF93FFE9C15DC3D2954B4C67" blockId="33.[113,764,1484,1884]" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C15DC3D295B14ED8" box="[113,468,1484,1508]" italics="true" pageId="33" pageNumber="34">
Diagnosis:
<taxonomicName id="4C7C4D12FF93FFE9C1CEC3D395B14ED8" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[226,468,1484,1508]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="plateadensis">Phyllonastes plateadensis</taxonomicName>
</emphasis>
resembles its closest species,
<taxonomicName id="4C7C4D12FF93FFE9C15DC3F394BA4D3F" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[113,223,1515,1539]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="dicaprioi">
<emphasis id="B908EA83FF93FFE9C15DC3F394BA4D3F" box="[113,223,1515,1539]" italics="true" pageId="33" pageNumber="34">P. dicaprioi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7C4D12FF93FFE9C038C3F395004D3F" baseAuthorityName="Lynch" baseAuthorityYear="1986" box="[276,357,1515,1539]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FF93FFE9C038C3F395004D3F" box="[276,357,1515,1539]" italics="true" pageId="33" pageNumber="34">P. heyeri</emphasis>
</taxonomicName>
, by having the snout rounded in lateral view, a tympanic annulus, supernumerary palmar tubercles, three phalanges in finger IV, and by the absence of papillae in fingers and toes. It differs from both by lacking a supratympanic fold (present in
<taxonomicName id="4C7C4D12FF93FFE9C03EC07695184DBD" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[274,381,1641,1665]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="dicaprioi">
<emphasis id="B908EA83FF93FFE9C03EC07695184DBD" box="[274,381,1641,1665]" italics="true" pageId="33" pageNumber="34">P. dicaprioi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7C4D12FF93FFE9C080C07695994DBD" baseAuthorityName="Lynch" baseAuthorityYear="1986" box="[428,508,1641,1665]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FF93FFE9C080C07695994DBD" box="[428,508,1641,1665]" italics="true" pageId="33" pageNumber="34">P. heyeri</emphasis>
</taxonomicName>
) and by having a broadly rounded snout in dorsal view (snout rounded in
<taxonomicName id="4C7C4D12FF93FFE9C3A5C096969F4D9C" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[649,762,1672,1696]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="dicaprioi">
<emphasis id="B908EA83FF93FFE9C3A5C096969F4D9C" box="[649,762,1672,1696]" italics="true" pageId="33" pageNumber="34">P. dicaprioi</emphasis>
</taxonomicName>
and subacuminate in
<taxonomicName id="4C7C4D12FF93FFE9C07AC0B795CE4D83" baseAuthorityName="Lynch" baseAuthorityYear="1986" box="[342,427,1703,1727]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FF93FFE9C07AC0B795CE4D83" box="[342,427,1703,1727]" italics="true" pageId="33" pageNumber="34">P. heyeri</emphasis>
</taxonomicName>
), rounded inner tarsal tubercle (inner tarsal tubercle conical in
<taxonomicName id="4C7C4D12FF93FFE9C0EEC0D896544DE3" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[450,561,1735,1759]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="dicaprioi">
<emphasis id="B908EA83FF93FFE9C0EEC0D896544DE3" box="[450,561,1735,1759]" italics="true" pageId="33" pageNumber="34">P. dicaprioi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7C4D12FF93FFE9C344C0D896DE4DE3" baseAuthorityName="Lynch" baseAuthorityYear="1986" box="[616,699,1735,1759]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FF93FFE9C344C0D896DE4DE3" box="[616,699,1735,1759]" italics="true" pageId="33" pageNumber="34">P. heyeri</emphasis>
</taxonomicName>
), and circumferential grooves of fingers (fingers without circumferential grooves in
<taxonomicName id="4C7C4D12FF93FFE9C025C11995134C21" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[265,374,1797,1821]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="dicaprioi">
<emphasis id="B908EA83FF93FFE9C025C11995134C21" box="[265,374,1797,1821]" italics="true" pageId="33" pageNumber="34">P. dicaprioi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7C4D12FF93FFE9C085C119959C4C21" baseAuthorityName="Lynch" baseAuthorityYear="1986" box="[425,505,1797,1821]" class="Amphibia" family="Craugastoridae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="heyeri">
<emphasis id="B908EA83FF93FFE9C085C119959C4C21" box="[425,505,1797,1821]" italics="true" pageId="33" pageNumber="34">P. heyeri</emphasis>
</taxonomicName>
). For a comparison with other
<taxonomicName id="4C7C4D12FF93FFE9C181C13A95174C01" box="[173,370,1829,1853]" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C181C13A95414C01" box="[173,292,1829,1853]" italics="true" pageId="33" pageNumber="34">Phyllonastes</emphasis>
species
</taxonomicName>
that are more phylogenetically distant, see
<tableCitation id="C6FE032AFF93FFE9C1BBC15B948E4C60" box="[151,235,1860,1884]" captionStart="Table 2" captionStartId="11.[109,164,664,689]" captionTargetPageId="11" captionText="Table 2. General morphological characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined. Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al.(2007), De la Riva et al. (2008a, b, 2018), Hedges et al. (2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al. (2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFB9FFC3C141C48796BE4865" pageId="33" pageNumber="34" tableUuid="DF036619FFB9FFC3C141C48796BE4865">Tables 2</tableCitation>
and
<tableCitation id="C6FE032AFF93FFE9C037C15B954D4C60" box="[283,296,1860,1884]" captionStart="Table 3" captionStartId="18.[109,164,620,645]" captionTargetPageId="18" captionText="Table 3. General morphological characters of Phyllonastes and Urkuphryne species. Finger and toe characters of Phyllonastes and Urkuphryne species. Abbreviations: A, absent; Exp, expanded; I, indistinct; NExp, not expanded; P, present; SExp, slightly expanded; Und, undifferentiated; Ukn, unknown; Wdef, weakly defined.Character states are based on specimens deposited at Museo de Zoología of Pontificia Universidad Católica del Ecuador (QCAZ) and from the literature: Boulenger (1898), Miranda-Ribeiro (1920), Parker (1926, 1932), Miranda-Ribeiro (1937), Cochran (1955), Lutz (1958), Griffiths (1959), Bokermann (1966), Heyer (1969), Bokermann (1975), Lynch (1975, 1976, 1986), Heyer (1977), De la Riva (1992, 2007), Köhler (2000), Aguayo-Vedia and Harvey (2001), Caramaschi and Pombal (2006), Lehr (2006), Heinicke et al. (2007), De la Riva et al. (2008a, b, 2018), Hedges et al.(2008), Guayasamin and Terán-Valdez (2009), Lehr and Catenazzi (2009a, b, 2010), Harvey et al. (2013), De la Riva and Burrowes (2014), Catenazzi et al. (2015, 2020), Catenazzi and Ttito (2016, 2018, 2019), Mamani et al.(2017), Reyes-Puig et al. (2019, 2020); Santa Cruz et al. (2019), Dubois et al. (2021), Motta et al. (2021), Reyes-Puig et al. (2021)." httpUri="http://table.plazi.org/id/DF036619FFA0FFDAC141C47390344811" pageId="33" pageNumber="34" tableUuid="DF036619FFA0FFDAC141C47390344811">3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C366651AFF93FFEBC15DC16196204EA6" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="description">
<paragraph id="8BC33691FF93FFE9C15DC16190AB4A3A" blockId="33.[113,762,1917,1973]" lastBlockId="33.[809,1461,144,1359]" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C15DC16196EF4CA9" box="[113,650,1917,1941]" italics="true" pageId="33" pageNumber="34">
Description of the
<typeStatus id="54C78833FF93FFE9C001C162951A4CA9" box="[301,383,1917,1941]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
(
<figureCitation id="13472A14FF93FFE9C0B8C16195B84CA9" box="[404,477,1917,1941]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="33" pageNumber="34">Figs 20</figureCitation>
,
<figureCitation id="13472A14FF93FFE9C0C0C16296634CA9" box="[492,518,1917,1941]" captionStart="Figure 21" captionStartId="35.[113,178,1091,1115]" captionTargetBox="[306,1266,144,1064]" captionTargetId="figure-421@35.[306,1266,144,1064]" captionTargetPageId="35" captionText="Figure 21. Palmar (A) and plantar (B) surfaces of Phyllonastes plateadensis. Photographs of right hand and right foot of the holotype QCAZ 65015." figureDoi="http://doi.org/10.5281/zenodo.14832748" httpUri="https://zenodo.org/record/14832748/files/figure.png" pageId="33" pageNumber="34">21</figureCitation>
,
<figureCitation id="13472A14FF93FFE9C339C162965B4CA9" box="[533,574,1917,1941]" captionStart="Figure 22" captionStartId="36.[129,194,1772,1796]" captionTargetBox="[251,1360,158,1734]" captionTargetId="figure-100@36.[226,1376,144,1744]" captionTargetPageId="36" captionText="Figure 22. Skulls of Phyllonastes species.A, Phyllonastes myrmecoides QCAZ 53583. B, Phyllonastes plateadensis QCAZ 65015. C, Phyllonastes sardinayacu, QCAZ 58821." figureDoi="http://doi.org/10.5281/zenodo.14832750" httpUri="https://zenodo.org/record/14832750/files/figure.png" pageId="33" pageNumber="34">22B</figureCitation>
,
<figureCitation id="13472A14FF93FFE9C360C16296124CA9" box="[588,631,1917,1941]" captionStart="Figure 23" captionStartId="37.[112,177,1566,1590]" captionTargetBox="[148,1425,146,1537]" captionTargetId="figure-197@37.[146,1426,144,1539]" captionTargetPageId="37" captionText="Figure 23. Skeletons of Phyllonastes species. A, Phyllonastes plateadensis QCAZ 65015, SVL = 15.70 mm. B, Phyllonastes sardinayacu QCAZ 58821, SVL = 14.32 mm. C, Phyllonastes personinus QCAZ 58818, SVL = 16.91 mm." figureDoi="http://doi.org/10.5281/zenodo.14832752" httpUri="https://zenodo.org/record/14832752/files/figure.png" pageId="33" pageNumber="34">23A</figureCitation>
):
</emphasis>
Adult female (
<emphasis id="B908EA83FF93FFE9C196C18295294C89" box="[186,332,1949,1973]" italics="true" pageId="33" pageNumber="34">QCAZ 65015</emphasis>
). Measurements (in millimetres): SVL, 19.35; tibia length, 7.79; foot length, 8.66; head length, 5.74; head width, 7.05; eye diameter, 2.14; tympanum diameter, 1.18; interorbital distance, 1.93; upper eyelid width, 1.31; internarial distance, 1.79; eye–nostril distance, 1.51.
</paragraph>
<paragraph id="8BC33691FF93FFE9C269C7129154493D" blockId="33.[809,1461,144,1359]" pageId="33" pageNumber="34">Head wider than long, head wider than body; canthus rostralis straight in lateral view; loreal region slightly concave in dorsal view; cranial crests absent; upper eyelid lacking tubercles. Tympanic annulus inconspicuous but visible beneath skin; tympanic membrane not differentiated; supratympanic fold and postrictal tubercles absent. Vomerine teeth absent, vocal slits and nuptial pads absent. Snout broadly rounded in dorsal view and rounded in lateral view, without rostral papilla.</paragraph>
<paragraph id="8BC33691FF93FFE9C269C41790CE4845" blockId="33.[809,1461,144,1359]" pageId="33" pageNumber="34">Skin on dorsum and flanks finely shagreen; dorsolateral folds absent; ventral surfaces weakly areolate; skin in cloacal region areolate; discoidal fold present. Ulnar tubercles absent; palmar tubercles low, outer palmar tubercle circular, larger than elongate thenar tubercle; subarticular tubercles ill defined, more conspicuous at the base of fingers, round in ventral and lateral view; three supernumerary tubercles present at the base of fingers; narrow lateral dermal fringes on fingers, fingers not expanded distally; discs narrow, bearing round and inconspicuous pads with an ill-defined circumferential groove; fingertips subacuminate lacking papillae; relative length of fingers is I &lt;IV &lt;II &lt;III; phalangeal formula of hand is 2-2-3-3.</paragraph>
<paragraph id="8BC33691FF93FFE9C269C59F905E4E73" blockId="33.[809,1461,144,1359]" pageId="33" pageNumber="34">Hindlimbs robust; knees and heels without tubercles; tarsal folds (inner and outer) absent, outer tarsal tubercle absent; inner tarsal tubercle present, small and round; inner metatarsal tubercle small (ill defined in preservative), rounded in ventral and lateral views; well-defined and round outer metatarsal tubercle, bigger than the inner one; plantar surface with ill-defined supernumerary tubercles; round subarticular tubercles ill defined; toes with narrow dermal lateral fringes; webbing between toes absent; toes slightly expanded distally, discs on toes ill defined (ill defined in preservative), narrow with acuminated tip, papilla on tip not visible; ill-defined circumferential grooves; relative lengths of toes I &lt;II &lt;V &lt;III &lt;IV; toe III longer than toe V (toe III reaches the distal border of the second subarticular tubercle of toe IV; toe V almost reaches the proximal border of the second tubercle of toe IV).</paragraph>
<paragraph id="8BC33691FF93FFE9C206C36F905D4C23" blockId="33.[809,1460,1392,1823]" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C206C36F91D64EBB" box="[810,1459,1392,1416]" italics="true" pageId="33" pageNumber="34">
Colour of
<typeStatus id="54C78833FF93FFE9C2A0C36F97BB4EB4" box="[908,990,1392,1416]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
in life (based on digital photographs) (
<figureCitation id="13472A14FF93FFE9C472C36E91C44EB4" box="[1374,1441,1392,1416]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="33" pageNumber="34">Fig. 20</figureCitation>
):
</emphasis>
Dorsal surfaces light brown, with some scattered white dots; circular dark brown sacral lateral patches well defined, delineated by a paler line. Cloacal region cream, surrounded by a dark brown stripe. Facial mask dark brown extending from tip of snout to groins and delineated along its dorsal edge by a light orange line, more conspicuous rostrally. Flanks light brown, bearing the dark brown mask extension. Lips bearing small and white spots. Forearms with two dark brown blotches delineated by a paler line; the proximal blotch is bigger than the distal one. Heels bearing a dark brown blotch with a paler border; tarsus bearing dark brown blotches surrounded by a paler border dorsally. Ventral surfaces brown with white spots that become bigger towards the cloacal region.
</paragraph>
<paragraph id="8BC33691FF93FFEAC206C15E90EA4C6F" blockId="33.[810,1459,1857,1975]" lastBlockId="34.[825,1476,1820,1875]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="B908EA83FF93FFE9C206C15E90B34C64" box="[810,1238,1857,1881]" italics="true" pageId="33" pageNumber="34">
Colour of
<typeStatus id="54C78833FF93FFE9C2A0C15E97BB4C65" box="[908,990,1857,1881]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
in preservative (
<figureCitation id="13472A14FF93FFE9C5ADC15E90A14C65" box="[1153,1220,1857,1881]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="33" pageNumber="34">Fig. 20</figureCitation>
):
</emphasis>
Dorsal surfaces light brown, with dark brown sacral lateral circular blotches surrounded by a cream edge. Facial mask dark brown extending from tip of snout to near groins, limited on its upper side by a thin cream-coloured line. Ventral surfaces light brown, bearing ill-defined white spots that become bigger posteriorly. Posterior surfaces of thighs light brown. Cloacal region white, surrounded by a dark brown stripe, and bearing small and dispersed dark brown flecks. Forearms dorsally with two dark brown blotches delineated by a paler line; the proximal blotch is bigger and better defined than the distal one.
</paragraph>
<caption id="DF036619FF90FFEAC1ADC08395244DD0" ID-DOI="http://doi.org/10.5281/zenodo.14832746" ID-Zenodo-Dep="14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="34" pageNumber="35" startId="34.[129,194,1692,1716]" targetBox="[166,1429,150,1624]" targetPageId="34" targetType="figure">
<paragraph id="8BC33691FF90FFEAC1ADC08395244DD0" blockId="34.[129,1474,1692,1772]" pageId="34" pageNumber="35">
<emphasis id="B908EA83FF90FFEAC1ADC08394834D88" bold="true" box="[129,230,1692,1716]" pageId="34" pageNumber="35">Figure 20.</emphasis>
Live and preserved specimens of
<taxonomicName id="4C7C4D12FF90FFEAC30DC083977D4D88" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[545,792,1692,1716]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="plateadensis">
<emphasis id="B908EA83FF90FFEAC30DC083977D4D88" bold="true" box="[545,792,1692,1716]" italics="true" pageId="34" pageNumber="35">Phyllonastes plateadensis</emphasis>
</taxonomicName>
. Lateral, dorsal, and ventral views of: A, B, holotype
<emphasis id="B908EA83FF90FFEAC42AC08391E24D88" box="[1286,1415,1692,1716]" italics="true" pageId="34" pageNumber="35">QCAZ 65015</emphasis>
, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life. B, D, in preservative. Scale bar is given only for preserved adults.
</paragraph>
</caption>
<paragraph id="8BC33691FF90FFEBC215C16496204EA6" blockId="34.[825,1474,1915,1970]" lastBlockId="35.[112,763,1191,1435]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="B908EA83FF90FFEAC215C16497994CAE" box="[825,1020,1915,1939]" italics="true" pageId="34" pageNumber="35">
Variation (
<figureCitation id="13472A14FF90FFEAC28AC164978F4CAF" box="[934,1002,1915,1939]" captionStart="Figure 20" captionStartId="34.[129,194,1692,1716]" captionTargetBox="[166,1429,150,1624]" captionTargetId="figure-92@34.[161,1441,144,1664]" captionTargetPageId="34" captionText="Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults." figureDoi="http://doi.org/10.5281/zenodo.14832746" httpUri="https://zenodo.org/record/14832746/files/figure.png" pageId="34" pageNumber="35">Fig. 20</figureCitation>
):
</emphasis>
This section refers to living individuals unless otherwise mentioned. Juvenile QCAZ 65016 has more and better-defined dark brown circular marks on the tarsus. Facial mask can be ill defined and weakly contrasting, as in QCAZ 65016. Juvenile ventral coloration is darker than in adult, with a darker throat and white spots restricted to belly and thighs, as in QCAZ 65016. Tip of toes may be subacuminate, as in QCAZ 65016. Juvenile presents more inconspicuous palmar and plantar tubercles, round and lower than those in adults in lateral view. Morphometric variation is detailed in
<tableCitation id="C6FE032AFF91FFEBC0D9C39C965A4EA7" box="[501,575,1411,1435]" captionStart="Table 4" captionStartId="21.[113,168,1190,1214]" captionTargetPageId="21" captionText="Table 4. Snout–vent length (SVL; in millimetres) and SD (in parentheses) for adult females and males.of Phyllonastes and Urkuphryne new species. The number of individuals is given in parentheses after the sex.For each group, minimum (Min) and maximum (Max) values are also shown." httpUri="http://table.plazi.org/id/DF036619FFA7FFDDC15DC2B994D34FCA" pageId="35" pageNumber="36" tableUuid="DF036619FFA7FFDDC15DC2B994D34FCA">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF036619FF91FFEBC15DC25C94D64F4B" ID-DOI="http://doi.org/10.5281/zenodo.14832748" ID-Zenodo-Dep="14832748" httpUri="https://zenodo.org/record/14832748/files/figure.png" pageId="35" pageNumber="36" startId="35.[113,178,1091,1115]" targetBox="[306,1266,144,1064]" targetPageId="35" targetType="figure">
<paragraph id="8BC33691FF91FFEBC15DC25C94D64F4B" blockId="35.[113,1441,1091,1143]" pageId="35" pageNumber="36">
<emphasis id="B908EA83FF91FFEBC15DC25C94B04F67" bold="true" box="[113,213,1091,1115]" pageId="35" pageNumber="36">Figure 21.</emphasis>
Palmar (A) and plantar (B) surfaces of
<taxonomicName id="4C7C4D12FF91FFEBC364C25C975A4F67" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[584,831,1091,1115]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="plateadensis">
<emphasis id="B908EA83FF91FFEBC364C25C975A4F67" bold="true" box="[584,831,1091,1115]" italics="true" pageId="35" pageNumber="36">Phyllonastes plateadensis</emphasis>
</taxonomicName>
. Photographs of right hand and right foot of the holotype
<emphasis id="B908EA83FF91FFEBC44DC25B94C84F4B" italics="true" pageId="35" pageNumber="36">QCAZ 65015</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C366651AFF91FFEBC15DC3A395D94C50" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BC33691FF91FFEBC15DC3A395D94C50" blockId="35.[113,765,1468,1900]" pageId="35" pageNumber="36">
<emphasis id="B908EA83FF91FFEBC15DC3A395014ECF" italics="true" pageId="35" pageNumber="36">
Distribution, natural history, and conservation status (
<figureCitation id="13472A14FF91FFEBC387C3A2968D4EE8" box="[683,744,1468,1492]" captionStart="Figure 3" captionStartId="9.[113,175,1807,1831]" captionTargetBox="[149,1423,152,1776]" captionTargetId="figure-7@9.[146,1426,149,1779]" captionTargetPageId="9" captionText="Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed." figureDoi="http://doi.org/10.5281/zenodo.14832695" httpUri="https://zenodo.org/record/14832695/files/figure.png" pageId="35" pageNumber="36">Fig. 3</figureCitation>
):
<taxonomicName id="4C7C4D12FF91FFEBC15DC3C495014ECF" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[113,356,1499,1523]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="plateadensis">Phyllonastes plateadensis</taxonomicName>
</emphasis>
is known from the surroundings of the Biological Reserve Cerro Plateado from
<quantity id="4C849B74FF91FFEBC330C3E496AD4D2E" box="[540,712,1531,1555]" metricMagnitude="3" metricUnit="m" metricValue="2.0365" metricValueMax="2.093" metricValueMin="1.98" pageId="35" pageNumber="36" unit="m" value="2036.5" valueMax="2093.0" valueMin="1980.0">1980 to 2093 m</quantity>
a.s.l. Cerro Plateado is in Cordillera del Cóndor, a sub-Andean cordillera separate from the main Andean chain. Individuals were collected in primary Eastern Montane Forest, at night (22:00– 23:00 h), on mossy soils and on leaf litter, at the edge of creeks next to water pools. Cerro Plateado is a sandstone formation. Forests in sandstone at Cordillera del Cóndor are characterized by dwarf trees with a canopy of ~
<quantity id="4C849B74FF91FFEBC0E8C0C995894DD2" box="[452,492,1750,1774]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="36" unit="m" value="5.0">5 m</quantity>
(Schulenberg and Awbrey 1997). Because of the lack of information related to population size and geographical range, we assign
<taxonomicName id="4C7C4D12FF91FFEBC0D4C10996E64C11" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[504,643,1813,1837]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="plateadensis">
<emphasis id="B908EA83FF91FFEBC0D4C10996E64C11" box="[504,643,1813,1837]" italics="true" pageId="35" pageNumber="36">P. plateadensis</emphasis>
</taxonomicName>
to the Data Deficient IUCN Red List Category (based on
<bibRefCitation id="EFED4B60FF91FFEBC361C12B95CE4C50" author="IUCN Standards and Petitions Committee &amp; Using the IUCN Red List Categories" pageId="35" pageNumber="36" refId="ref39535" refString="IUCN Standards and Petitions Committee. Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee, 2023. https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (25 January 2024, date last accessed)." type="url" year="2023">IUCN Standards and Petitions Committee 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366651AFF91FFEBC15DC192965B4CF8" pageId="35" pageNumber="36" type="etymology">
<paragraph id="8BC33691FF91FFEBC15DC192965B4CF8" blockId="35.[113,762,1933,1989]" pageId="35" pageNumber="36">
<emphasis id="B908EA83FF91FFEBC15DC19294844C99" box="[113,225,1933,1957]" italics="true" pageId="35" pageNumber="36">Etymology:</emphasis>
The specific epithet
<taxonomicName id="4C7C4D12FF91FFEBC091C19296574C99" authorityName="Ortega &amp; Cisneros-Heredia &amp; Camper &amp; Romero-Carvajal &amp; Negrete &amp; Ron" authorityYear="2025" box="[445,562,1933,1957]" class="Amphibia" family="Craugastoridae" genus="Phyllonastes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="plateadensis">
<emphasis id="B908EA83FF91FFEBC091C19296574C99" box="[445,562,1933,1957]" italics="true" pageId="35" pageNumber="36">plateadensis</emphasis>
</taxonomicName>
is a toponym noun in apposition and it refers to the
<typeStatus id="54C78833FF91FFEBC090C1B2958F4CF9" box="[444,490,1965,1989]" pageId="35" pageNumber="36">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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