treatments-xml/data/03/D5/87/03D58784817AFFB2FDD1FB72FBC8FAD4.xml

<document id="FB24E45451CC0E264C24C63EE50A27CC" ID-CLB-Dataset="22393" ID-DOI="10.5852/ejt.2018.438" ID-GBIF-Dataset="34ad2255-e191-4870-83e8-74819ca0fe06" ID-Zenodo-Dep="3785478" ID-ZooBank="00B57E96-225C-4646-9CB8-B3514C4EC7E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588622038275" checkinUser="valdenar" docAuthor="Jia, Fenglong &amp; Tang, Yu-dan" docDate="2018" docId="03D58784817AFFB2FDD1FB72FBC8FAD4" docLanguage="en" docName="ejt-438_jia_tang.pdf.imf" docOrigin="European Journal of Taxonomy 438" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helochares fuliginosus d'Orchymont 1932" docType="treatment" docVersion="6" lastPageNumber="9" masterDocId="FFECFFFC817FFFBAFFC7FF81FFF9FFFF" masterDocTitle="A revision of the Chinese Helochares (s. str.) Mulsant, 1844 (Coleoptera, Hydrophilidae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="6" updateTime="1698838436565" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C71957A28CFD847C37651999FC8A8495">A revision of the Chinese Helochares (s. str.) Mulsant, 1844 (Coleoptera, Hydrophilidae)</mods:title>
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<mods:namePart id="49F75876013EA8912B1A9325ED98198B">Jia, Fenglong</mods:namePart>
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<mods:namePart id="D76F3FF875AB73CB113ECF22E130EA95">Tang, Yu-dan</mods:namePart>
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<mods:affiliation id="7AD8884FF43D5C3C56D63A6DB19CE74E">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China.</mods:affiliation>
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<treatment id="03D58784817AFFB2FDD1FB72FBC8FAD4" ID-DOI="http://doi.org/10.5281/zenodo.3794613" ID-GBIF-Taxon="163943953" ID-Zenodo-Dep="3794613" LSID="urn:lsid:plazi:treatment:03D58784817AFFB2FDD1FB72FBC8FAD4" httpUri="http://treatment.plazi.org/id/03D58784817AFFB2FDD1FB72FBC8FAD4" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
<subSubSection id="C3666519817AFFBFFDD1FB72FBE7FAF2" box="[534,1054,1267,1294]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BC33692817AFFBFFDD1FB72FBE7FAF2" blockId="5.[534,1054,1267,1328]" box="[534,1054,1267,1294]" pageId="5" pageNumber="6">
<heading id="D08B81FE817AFFBFFDD1FB72FBE7FAF2" box="[534,1054,1267,1294]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="4C7C4D11817AFFBFFDD1FB72FBE7FAF2" ID-CoL="8Y496" authority="d'Orchymont, 1932" authorityName="d'Orchymont" authorityYear="1932" box="[534,1054,1267,1294]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B908EA80817AFFBFFDD1FB72FCD6FAF1" bold="true" box="[534,815,1267,1294]" italics="true" pageId="5" pageNumber="6">Helochares fuliginosus</emphasis>
<bibRefCitation id="EFED4B63817AFFBFFCF1FB72FBE7FAF2" author="d'Orchymont A." box="[822,1054,1267,1293]" pageId="5" pageNumber="6" pagination="623 - 714" refId="ref16315" refString="d'Orchymont A. 1932. Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement IX (Tropische Binnengewasser II): 623 - 714." type="book chapter" year="1932">d’Orchymont, 1932</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="C3666519817AFFBFFD5AFA97FC6FFACF" box="[669,918,1302,1328]" pageId="5" pageNumber="6" type="description">
<paragraph id="8BC33692817AFFBFFD5AFA97FC6FFACF" blockId="5.[534,1054,1267,1328]" box="[669,918,1302,1328]" pageId="5" pageNumber="6">
<figureCitation id="13472A17817AFFBFFD5AFA97FCEDFACF" box="[669,788,1302,1328]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="5" pageNumber="6">Figs 9–17</figureCitation>
, 
<figureCitation id="13472A17817AFFBFFCE6FA97FC6FFACF" box="[801,918,1302,1328]" captionStart-0="Figs 41–46. – 41–42" captionStart-1="Figs 47–55" captionStartId-0="20.[189,241,1930,1956]" captionStartId-1="22.[189,241,1894,1920]" captionTargetBox-0="[235,1344,268,1888]" captionTargetBox-1="[280,1297,268,1859]" captionTargetId-0="figure@20.[220,1367,254,1900]" captionTargetId-1="figure@22.[275,1312,253,1869]" captionTargetPageId-0="20" captionTargetPageId-1="22" captionText-0="Figs 41–46. – 41–42. Helochares pallens (MacLeay, 1825). Elytra. 41. Dorsal. 42. Ventral. 43, 45. Paratype of H. vitalisi d’Orchymont, 1919. 43. Data of labels. 45. Habitus, dorsal. 44, 46. Paratype of H. fuliginosus d’Orchymont, 1932. 44. Data of labels. 46. Habitus, dorsal." captionText-1="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi-0="http://doi.org/10.5281/zenodo.3785492" figureDoi-1="http://doi.org/10.5281/zenodo.3785494" httpUri-0="https://zenodo.org/record/3785492/files/figure.png" httpUri-1="https://zenodo.org/record/3785494/files/figure.png" pageId="5" pageNumber="6">44, 46–47</figureCitation>
</paragraph>
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<subSubSection id="C3666519817AFFBFFF7AFAD9FCACFA8D" box="[189,853,1368,1394]" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BC33692817AFFBFFF7AFAD9FCACFA8D" blockId="5.[189,853,1368,1394]" box="[189,853,1368,1394]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB6C11BC817AFFBFFF7AFAD9FCACFA8D" box="[189,853,1368,1394]" pageId="5" pageNumber="6">
<treatmentCitation id="0ADD1083817AFFBFFF7AFAD9FCA8FA8D" author="d'Orchymont A." box="[189,849,1368,1394]" page="689" pageId="5" pageNumber="6" year="1932">
<taxonomicName id="4C7C4D11817AFFBFFF7AFAD9FCA8FA8D" ID-CoL="8Y496" authority="d'Orchymont, 1932: 689" authorityName="d'Orchymont" authorityPageNumber="689" authorityYear="1932" box="[189,849,1368,1394]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B908EA80817AFFBFFF7AFAD9FEBAFA8D" box="[189,323,1368,1394]" italics="true" pageId="5" pageNumber="6">Helochares</emphasis>
(s. str.) 
<emphasis id="B908EA80817AFFBFFE65FAD9FDDAFA8D" box="[418,547,1368,1394]" italics="true" pageId="5" pageNumber="6">fuliginosus</emphasis>
<bibRefCitation id="EFED4B63817AFFBFFDECFAD9FCA8FA8D" author="d'Orchymont A." box="[555,849,1368,1394]" pageId="5" pageNumber="6" pagination="623 - 714" refId="ref16315" refString="d'Orchymont A. 1932. Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement IX (Tropische Binnengewasser II): 623 - 714." type="book chapter" year="1932">d’Orchymont, 1932: 689</bibRefCitation>
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.
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<subSubSection id="C3666519817AFFBFFF7AFA18FBE9F8E0" pageId="5" pageNumber="6" type="diagnosis">
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<heading id="D08B81FE817AFFBFFF7AFA18FECFFA4C" bold="true" box="[189,310,1433,1459]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B908EA80817AFFBFFF7AFA18FECFFA4C" bold="true" box="[189,310,1433,1459]" pageId="5" pageNumber="6">Diagnosis</emphasis>
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</paragraph>
<paragraph id="8BC33692817AFFBFFF7AFA49FBE9F8E0" blockId="5.[189,1399,1480,1823]" pageId="5" pageNumber="6">
Length 
<quantity id="4C849B77817AFFBFFEDCFA49FE56FA1D" box="[283,431,1480,1506]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="5" pageNumber="6" unit="mm" value="3.95" valueMax="4.3" valueMin="3.6">3.6–4.3 mm</quantity>
. Dorsum with dense and fine punctures that are finer than in 
<taxonomicName id="4C7C4D11817AFFBFFB61FA48FAF7FA1D" authority="Jia &amp; Tang, 2018" authorityName="Jia &amp; Tang" authorityYear="2018" box="[1190,1294,1481,1506]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="songi" status="sp. nov.">
<emphasis id="B908EA80817AFFBFFB61FA48FAF7FA1D" box="[1190,1294,1481,1506]" italics="true" pageId="5" pageNumber="6">H. songi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57FB817AFFBFFADFFA49FA8EFA1D" box="[1304,1399,1480,1506]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
Clypeus somewhat impressed in front of eyes (
<figureCitation id="13472A17817AFFBFFD2BFA6AFC8EF9FA" box="[748,887,1515,1541]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="5" pageNumber="6">Figs 12, 14</figureCitation>
). Maxillary palps ca 1.2× as long as width of head, each palpomere uniformly yellowish brown (
<figureCitation id="13472A17817AFFBFFCA7F98FFC09F9D7" box="[864,1008,1550,1576]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="5" pageNumber="6">Figs 13–14</figureCitation>
). Pedicel subequal in length to antennomeres 3–6 combined. Submentum with a small glabrous area medially and with a few coarse punctures laterally (
<figureCitation id="13472A17817AFFBFFE6BF9D5FDFFF990" box="[428,518,1620,1647]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
). Elytra almost parallel from base to posterior third, somewhat explanate laterally, with distinct systematic punctures (
<figureCitation id="13472A17817AFFBFFD0EF9F9FCE8F96D" box="[713,785,1656,1682]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
). Mesoventrite with a low transverse ridge medially and a low longitudinal carina (
<figureCitation id="13472A17817AFFBFFDEBF91AFD7DF949" box="[556,644,1691,1718]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="5" pageNumber="6">Fig. 15</figureCitation>
). Metaventrite without glabrous area medially. Femora densely pubescent except apical eighth (
<figureCitation id="13472A17817AFFBFFDFAF93FFD30F927" box="[573,713,1726,1752]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="5" pageNumber="6">Figs 10, 17</figureCitation>
). Length of parameres ca 6.5× as long as width of apex, subparallel, slightly expanded subapically and round apically, apex without inwardly directed tooth; median lobe expanded subapically, abruptly narrowed apically (
<figureCitation id="13472A17817AFFBFFC6CF885FBFAF8E0" box="[939,1027,1796,1823]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="5" pageNumber="6">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3666519817AFFBCFF7AF8C7FC79FBFC" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BC33692817AFFBFFF7AF8C7FE53F89F" blockId="5.[189,426,1862,1888]" box="[189,426,1862,1888]" pageId="5" pageNumber="6">
<heading id="D08B81FE817AFFBFFF7AF8C7FE53F89F" bold="true" box="[189,426,1862,1888]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B908EA80817AFFBFFF7AF8C7FE53F89F" bold="true" box="[189,426,1862,1888]" pageId="5" pageNumber="6">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B143CCF817AFFBFFF2DF8F4FE3BF82A" ID-GBIF-Occurrence="2610058765" pageId="5" pageNumber="6" typeStatus="paratype">
<paragraph id="8BC33692817AFFBFFF2DF8F4FEA5F870" blockId="5.[189,1399,1909,2006]" box="[234,348,1909,1935]" pageId="5" pageNumber="6">
<emphasis id="B908EA80817AFFBFFF2DF8F4FEA5F870" bold="true" box="[234,348,1909,1935]" pageId="5" pageNumber="6">
<typeStatus id="54C78830817AFFBFFF2DF8F4FEA5F870" box="[234,348,1909,1935]" pageId="5" pageNumber="6" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC33692817AFFBFFF7AF819FE3BF82A" blockId="5.[189,1399,1909,2006]" pageId="5" pageNumber="6">
<collectingRegion id="49B8F870817AFFBFFF7AF819FEFCF84D" box="[189,261,1944,1970]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="6">JAVA</collectingRegion>
: 1 ♂, Buitenzorg, “Bot. Gart. Quellmorast, 13. 1928,” Exp. Thienemann leg., A. d’Orchymont det. (handwritten) (
<collectionCode id="ED6DAE57817AFFBFFEABF83AFE4CF82A" box="[364,437,1979,2005]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="5" pageNumber="6">IRSN</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BC336928179FFBCFF2DFE8AFE5FFEDA" blockId="6.[189,1399,267,364]" box="[234,422,267,293]" pageId="6" pageNumber="7">
<emphasis id="B908EA808179FFBCFF2DFE8AFE5FFEDA" bold="true" box="[234,422,267,293]" pageId="6" pageNumber="7">Other material</emphasis>
</paragraph>
<paragraph id="8BC336928179FFBCFF7AFEAFFC7CFE94" blockId="6.[189,1399,267,364]" pageId="6" pageNumber="7">
<collectingCountry id="F36B76028179FFBCFF7AFEAFFEF7FEB7" box="[189,270,302,328]" name="Laos" pageId="6" pageNumber="7">LAOS</collectingCountry>
: 
<specimenCount id="9D7AFD1B8179FFBCFEDCFEAFFEB3FEB6" box="[283,330,302,329]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
, 
<collectingRegion id="49B8F8708179FFBCFE9CFEAEFDBDFEB7" box="[347,580,302,329]" country="Laos" name="Houaphan" pageId="6" pageNumber="7">Hua Phan Province</collectingRegion>
, Ban Saluei Phu Phan Mt. Env., 
<geoCoordinate id="EE4850558179FFBCFC14FEADFBC6FEB6" box="[979,1087,300,329]" degrees="20" direction="north" minutes="13" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="20.216667">20º13′ N</geoCoordinate>
, 
<geoCoordinate id="EE4850558179FFBCFB8AFEADFB3AFEB6" box="[1101,1219,300,329]" degrees="103" direction="east" minutes="59" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="103.98333">103º59′ E</geoCoordinate>
, 
<quantity id="4C849B778179FFBCFB15FEAFFA8EFEB6" box="[1234,1399,302,329]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="6" pageNumber="7" unit="m" value="1650.0" valueMax="2000.0" valueMin="1300.0">1300–2000 m</quantity>
a.s.l., 
<date id="FFC210528179FFBCFEC5FED0FE42FE94" box="[258,443,337,363]" pageId="6" pageNumber="7" value="2004-04-06" valueMax="2004-04-18" valueMin="2004-04-06">6–18 Apr. 2004</date>
, J. Bezdek leg., Hebauer det. (
<collectionCode id="ED6DAE578179FFBCFCE6FED0FC83FE94" box="[801,890,337,363]" country="Czech Republic" name="National Museum Prague" pageId="6" pageNumber="7">NMPC</collectionCode>
).
</paragraph>
<paragraph id="8BC336928179FFBCFF7AFE12FC79FBFC" blockId="6.[189,1399,401,1027]" pageId="6" pageNumber="7">
CHINA: 
<emphasis id="B908EA808179FFBCFEE9FE13FE38FE53" bold="true" box="[302,449,402,428]" pageId="6" pageNumber="7">Guangdong</emphasis>
: 6 ♂♂, 
<specimenCount id="9D7AFD1B8179FFBCFDEAFE12FD8AFE52" box="[557,627,403,429]" pageId="6" pageNumber="7" type="female">9 ♀♀</specimenCount>
, Zhaoqing, Heishiding, 
<date id="FFC210528179FFBCFC5AFE12FBA1FE52" box="[925,1112,403,429]" pageId="6" pageNumber="7" value="2013-05-04" valueMax="2013-05-06" valueMin="2013-05-04">4–6 May. 2013</date>
, Fenglong Jia, Yue Jia, Bingjie Chen, Renchao Lin and Weilin Xu leg.; 
<specimenCount id="9D7AFD1B8179FFBCFD2CFE37FCB7FE2F" box="[747,846,438,464]" pageId="6" pageNumber="7" type="generic">74 spec.</specimenCount>
, Fengkai, Heishiding, 
<geoCoordinate id="EE4850558179FFBCFB93FE35FB46FE2F" box="[1108,1215,436,464]" degrees="23" direction="north" minutes="31" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="23.516666">23°31′ N</geoCoordinate>
, 
<geoCoordinate id="EE4850558179FFBCFB0CFE35FAC6FE2F" box="[1227,1343,436,464]" degrees="111" direction="east" minutes="52" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="111.86667">111°52′ E</geoCoordinate>
, 
<date id="FFC210528179FFBCFA8DFE37FEAFFE0C" pageId="6" pageNumber="7" value="2014-04-20" valueMax="2014-04-22" valueMin="2014-04-20">20– 22 Apr. 2014</date>
, Fenglong Jia, Renchao Lin and Yudan Tang leg.; 
<specimenCount id="9D7AFD1B8179FFBCFC52FE58FC11FE0B" box="[917,1000,473,500]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
, Fengkai, Heishiding, in a shallow puddle with plenty of leaf litter, 
<date id="FFC210528179FFBCFD8FFE7DFD09FDE9" box="[584,752,508,534]" pageId="6" pageNumber="7" value="2010-08-13">13 Aug. 2010</date>
, Fenglong Jia leg.; 
<specimenCount id="9D7AFD1B8179FFBCFC18FE7DFBCFFDE8" box="[991,1078,508,535]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
, Gaoming, Yangmei town, 
<date id="FFC210528179FFBCFF7AFD9EFE71FDC6" box="[189,392,543,570]" pageId="6" pageNumber="7" value="2006-04-23" valueMax="2006-04-26" valueMin="2006-04-23">23–26 Apr. 2006</date>
, Fenglong Jia leg.; 
<specimenCount id="9D7AFD1B8179FFBCFDB3FDA1FD21FDC5" box="[628,728,544,570]" pageId="6" pageNumber="7" type="generic">37 spec.</specimenCount>
, Zhuhai, Sun Yat-sen Univerisity, near mountain behind campus, 
<date id="FFC210528179FFBCFEE2FDC3FE3FFDA2" box="[293,454,578,605]" pageId="6" pageNumber="7" value="2011-07-05" valueMax="2011-07-08" valueMin="2011-07-05">5–8 Jul. 2011</date>
, Fenglong Jia leg.; 
<specimenCount id="9D7AFD1B8179FFBCFD6BFDC2FCF7FDA2" box="[684,782,579,605]" pageId="6" pageNumber="7" type="generic">46 spec.</specimenCount>
, Shenzhen, Maluanshan, 
<date id="FFC210528179FFBCFBFDFDC2FB2DFDA2" box="[1082,1236,579,605]" pageId="6" pageNumber="7" value="2014-06-13">13 Jun. 2014</date>
, Fenglong Jia and Weicai Xie leg.; 
<specimenCount id="9D7AFD1B8179FFBCFE7FFDE7FDE5FD7F" box="[440,540,614,640]" pageId="6" pageNumber="7" type="generic">10 spec.</specimenCount>
, Shenzhen, Wutongshan, Hengpailing, 
<date id="FFC210528179FFBCFC28FDE7FB6CFD7F" box="[1007,1173,614,641]" pageId="6" pageNumber="7" value="2011-05-15">15 May. 2011</date>
, Fenglong Jia leg.; 
<specimenCount id="9D7AFD1B8179FFBCFF7AFD08FEE8FD5B" box="[189,273,649,676]" pageId="6" pageNumber="7" type="generic">2 spec.</specimenCount>
, Shenzhen, Paiyashan, 
<date id="FFC210528179FFBCFDE6FD08FD47FD5C" box="[545,702,649,676]" pageId="6" pageNumber="7" value="2011-05-17">17 May 2011</date>
, Fenglong Jia leg.; 
<specimenCount id="9D7AFD1B8179FFBCFC66FD08FC0DFD5B" box="[929,1012,649,676]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
, Shenzhen, Qiniangshan, 
<date id="FFC210528179FFBCFAE5FD08FF02FD39" pageId="6" pageNumber="7" value="2010-06-17">17 Jun. 2010</date>
, Fenglong Jia leg.; 
<specimenCount id="9D7AFD1B8179FFBCFE1BFD2DFDC9FD38" box="[476,560,684,711]" pageId="6" pageNumber="7" type="generic">8 spec.</specimenCount>
, Danxiashan, 
<date id="FFC210528179FFBCFD13FD2DFCA7FD39" box="[724,862,684,711]" pageId="6" pageNumber="7" value="2012-06-08">8 Jun. 2012</date>
, light trap, Fenglong Jia leg.; 3 unsexed spec., Danxiashan, 
<date id="FFC210528179FFBCFE92FD51FE13FD15" box="[341,490,720,746]" pageId="6" pageNumber="7" value="2008-05-02">2 May. 2008</date>
, Fenglong and Keqing Song leg.; 
<specimenCount id="9D7AFD1B8179FFBCFCB2FD4EFC30FD15" box="[885,969,719,746]" pageId="6" pageNumber="7" type="generic">6 spec.</specimenCount>
, Luogang, 
<date id="FFC210528179FFBCFB8DFD51FB1AFD15" box="[1098,1251,719,746]" pageId="6" pageNumber="7" value="1958-04-20">20 Apr. 1958</date>
, Zhelong Pu leg. – 
<emphasis id="B908EA808179FFBCFEC2FD72FE6DFCF2" bold="true" box="[261,404,755,781]" pageId="6" pageNumber="7">
<collectingCountry id="F36B76028179FFBCFEC2FD72FE6DFCF2" box="[261,404,755,781]" name="Hong Kong" pageId="6" pageNumber="7">Hong Kong</collectingCountry>
</emphasis>
: 2 ♂♂, 
<specimenCount id="9D7AFD1B8179FFBCFE34FD72FDE6FCF2" box="[499,543,755,781]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
, Shuikou, 
<date id="FFC210528179FFBCFD5AFD72FCCEFCF2" box="[669,823,755,781]" pageId="6" pageNumber="7" value="2013-04-27">27 Apr. 2013</date>
, Fenglong Jia, Yingming Lee and Eric Chen leg.; 
<specimenCount id="9D7AFD1B8179FFBCFF7AFC97FEEAFCCF" box="[189,275,790,816]" pageId="6" pageNumber="7" type="generic">4 spec.</specimenCount>
, Dabujiao, 
<date id="FFC210528179FFBCFE5AFC97FDC5FCCF" box="[413,572,790,816]" pageId="6" pageNumber="7" value="2013-04-27">27 Apr. 2013</date>
, Fenglong Jia, Yingming Lee and Eric Chen leg.; 
<specimenCount id="9D7AFD1B8179FFBCFB50FC97FB17FCCF" box="[1175,1262,790,816]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
, Fengkeng, 
<geoCoordinate id="EE4850558179FFBCFF7AFCB6FEA5FCAB" box="[189,348,823,852]" degrees="22" direction="north" minutes="31.695" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="22.52825">22°31.695′ N</geoCoordinate>
, 
<geoCoordinate id="EE4850558179FFBCFEA0FCB6FDF6FCAB" box="[359,527,823,852]" degrees="114" direction="east" minutes="13.450" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="114.22417">114°13.450′ E</geoCoordinate>
, 
<quantity id="4C849B778179FFBCFDDDFCB8FDACFCAB" box="[538,597,825,852]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="m" value="13.0">13 m</quantity>
a.s.l., 
<date id="FFC210528179FFBCFD67FCB8FCD3FCAC" box="[672,810,825,852]" pageId="6" pageNumber="7" value="2014-06-09">9 Jun. 2014</date>
, Fenglong Jia, Weicai Xie and Jiahuang Chen leg.; 
<specimenCount id="9D7AFD1B8179FFBCFF7AFCDDFEEAFC88" box="[189,275,860,887]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
, Rongshuao, 
<geoCoordinate id="EE4850558179FFBCFE73FCDBFDACFC88" box="[436,597,858,887]" degrees="22" direction="north" minutes="32.646" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="22.5441">22°32.646′ N</geoCoordinate>
, 
<geoCoordinate id="EE4850558179FFBCFDA5FCDBFCF4FC88" box="[610,781,858,887]" degrees="114" direction="east" minutes="14.926" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="114.248764">114°14.926′ E</geoCoordinate>
, 
<quantity id="4C849B778179FFBCFCDCFCDDFCA1FC88" box="[795,856,860,887]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="10.0">10 m</quantity>
a.s.l., Fenglong Jia, Weicai Xie and Jiahuang Chen leg. – 
<emphasis id="B908EA808179FFBCFE8DFCFEFE64FC66" bold="true" box="[330,413,895,921]" pageId="6" pageNumber="7">Fujian</emphasis>
: 
<specimenCount id="9D7AFD1B8179FFBCFE6AFC01FDF8FC65" box="[429,513,896,922]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
, Nanjing, Hexi town pond, 
<date id="FFC210528179FFBCFC83FC01FC2FFC65" box="[836,982,895,922]" pageId="6" pageNumber="7" value="2013-07-13">13 Jul. 2013</date>
, Fenglong Jia leg. – 
<emphasis id="B908EA808179FFBCFB0FFCFEFACDFC66" bold="true" box="[1224,1332,895,921]" pageId="6" pageNumber="7">Guangxi</emphasis>
: 1 ♂, Shiwandashan, 
<date id="FFC210528179FFBCFEB0FC22FDC8FC42" box="[375,561,931,957]" pageId="6" pageNumber="7" value="2013-04-08" valueMax="2013-04-12" valueMin="2013-04-08">8–12 Apr. 2013</date>
, Haidong Chen leg.; 
<specimenCount id="9D7AFD1B8179FFBCFCF7FC22FC7CFC42" box="[816,901,931,957]" pageId="6" pageNumber="7" type="generic">1 spec.</specimenCount>
, Shiwandashan, Forest Park, 
<quantity id="4C849B778179FFBCFB23FC22FAC9FC42" box="[1252,1328,931,957]" metricMagnitude="2" metricUnit="m" metricValue="2.3899999999999997" pageId="6" pageNumber="7" unit="m" value="239.0">239 m</quantity>
a.s.l., edge of upper river, 
<date id="FFC210528179FFBCFE72FC47FDB4FC1F" box="[437,589,966,992]" pageId="6" pageNumber="7" value="2011-07-19">19 Jul. 2011</date>
, Keqing Song leg. – 
<emphasis id="B908EA808179FFBCFC97FC47FC5CFC1F" bold="true" box="[848,933,966,992]" pageId="6" pageNumber="7">
<collectingCountry id="F36B76028179FFBCFC97FC47FC5CFC1F" box="[848,933,966,992]" name="Macao" pageId="6" pageNumber="7">Macao</collectingCountry>
</emphasis>
: 
<specimenCount id="9D7AFD1B8179FFBCFC70FC47FBF7FC1F" box="[951,1038,966,992]" pageId="6" pageNumber="7" type="generic">3 spec.</specimenCount>
, Luhuan, Dieshitang, 
<date id="FFC210528179FFBCFADFFC47FF02FBFC" pageId="6" pageNumber="7" value="2014-03-27">27 Mar. 2014</date>
, Fenglong Jia leg. [All data transcribed from Chinese.]
</paragraph>
</subSubSection>
<subSubSection id="C36665198179FFB2FF7AFBABFCF8FD1A" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8BC336928179FFBCFF7AFBABFE92FBBB" blockId="6.[189,363,1066,1092]" box="[189,363,1066,1092]" pageId="6" pageNumber="7">
<heading id="D08B81FE8179FFBCFF7AFBABFE92FBBB" bold="true" box="[189,363,1066,1092]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B908EA808179FFBCFF7AFBABFE92FBBB" bold="true" box="[189,363,1066,1092]" pageId="6" pageNumber="7">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BC336928179FFBCFF7AFBDBFD2EFAFF" blockId="6.[189,1398,1113,1280]" pageId="6" pageNumber="7">
FORM AND COLOUR. Length 
<quantity id="4C849B778179FFBCFDCDFBD8FD66FB8B" box="[522,671,1113,1140]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="6" pageNumber="7" unit="mm" value="3.95" valueMax="4.3" valueMin="3.6">3.6–4.3 mm</quantity>
, width 
<quantity id="4C849B778179FFBCFD38FBD8FC6CFB8B" box="[767,917,1113,1140]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="6" pageNumber="7" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
, oval, moderately convex. Head black, yellowish brown laterally (
<figureCitation id="13472A178179FFBCFE3DFBFDFD7AFB69" box="[506,643,1148,1174]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Figs 12–14</figureCitation>
); pronotum and elytra dark brown to black, both with yellowish or reddish brown margin laterally (
<figureCitation id="13472A178179FFBCFDBFFB1EFD2DFB45" box="[632,724,1183,1210]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
); maxillary palps uniform yellow or yellowish brown (
<figureCitation id="13472A178179FFBCFF02FB42FEB4FB22" box="[197,333,1219,1245]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Figs 13–14</figureCitation>
); labial palps yellow, antenna yellowish brown. Ventral side dark brown to black (
<figureCitation id="13472A178179FFBCFAD7FB42FA9EFB22" box="[1296,1383,1219,1245]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
); legs dark with yellowish tarsomeres (
<figureCitation id="13472A178179FFBCFDB4FB67FD30FAFF" box="[627,713,1254,1280]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BC336928179FFBCFF7AFAA9FB5EF917" blockId="6.[189,1399,1319,1768]" pageId="6" pageNumber="7">
HEAD. Labrum with fine and dense punctures, smooth between punctures, anterior margin broadly emarginate. Clypeus broad, anterior margin clearly broadly emarginate, moderately expanded in front of eyes, lateral margin somewhat rounded, clearly impressed in front of eyes (
<figureCitation id="13472A178179FFBCFBA5FAEFFB14FA77" box="[1122,1261,1390,1416]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Figs 12–14</figureCitation>
). Punctures on clypeus coarser and sparser than those on labrum, smooth between punctures, systematic punctures almost of same size as ground punctures (
<figureCitation id="13472A178179FFBCFD60FA35FD06FA31" box="[679,767,1460,1486]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
). Eyes of moderate size, clearly emarginate anteriorly (
<figureCitation id="13472A178179FFBCFF02FA56FEA9FA0E" box="[197,336,1495,1521]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Figs 12–14</figureCitation>
), separated by ca 4.0× the width of one eye. Ground punctures on frons similar to those on clypeus, with clear systematic punctures in front of eyes. Maxillary palps long and slender, ca 1.2× as wide as head, second and third palpomeres curved inwards, apical palpomere almost as long as penultimate, asymmetrical (
<figureCitation id="13472A178179FFBCFDC1F9C0FDA7F9A3" box="[518,606,1601,1628]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 16</figureCitation>
). Antennae with 9 antennomeres, scape ca 1.5× as long as pedicel, pedicel subequal in length to antennomeres 3–6 combined, club loosely with dense pubescence. 
<collectingRegion id="49B8F8708179FFBCFAD4F9E4FA8FF980" box="[1299,1398,1637,1663]" country="Myanmar" name="Chin" pageId="6" pageNumber="7">Mentum</collectingRegion>
ca as wide as long, subquadrate, with strong v-shaped impression anteromedially, with coarse punctures and wrinkles posteriorly (
<figureCitation id="13472A178179FFBCFE32F92BFDA9F93A" box="[501,592,1706,1733]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 16</figureCitation>
). Submentum with a small glabrous area medially and with a few coarse punctures laterally. Maxilla with coarse punctures, smooth between punctures.
</paragraph>
<paragraph id="8BC336928179FFB2FF7AF891FE85FDE3" blockId="6.[189,1399,1807,2010]" lastBlockId="8.[189,1399,267,540]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
THORAX. Pronotum ca 2× as wide as long, widest posteriorly, posterolateral margins broadly round, anterior margin smooth, with a very fine transverse groove, lateral margin with stronger groove, posterior margin without such groove; posterior margin almost straight. Ground punctures on pronotum slightly coarser than on head, with distinct setiferous systematic punctures (
<figureCitation id="13472A178179FFBCFC32F8F8FBA9F86C" box="[1013,1104,1913,1939]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 13</figureCitation>
). Prosternum bulging in middle, not carinate, pubescent, with transverse groove (
<figureCitation id="13472A178179FFBCFCA3F81DFC44F848" box="[868,957,1948,1975]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="6" pageNumber="7">Fig. 16</figureCitation>
), protruding anteriorly. Mesoventrite with a low transverse ridge medially, which does not bear an elevated tooth or projection but with a tuft of long setae (
<figureCitation id="13472A178177FFB2FE7BFE8AFDEEFED9" box="[444,535,267,294]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="8" pageNumber="9">Fig. 15</figureCitation>
), behind ridge with a low longitudinal carina. Metaventrite pubescent, with somewhat convex middle portion, without glabrous area. Metepisterna ca 3.5× as long as wide, subparallel. Scutellum triangular, with similar punctures to elytra. Elytra with ground punctures similar to those on pronotum and with 3 distinct rows of systematic punctures (
<figureCitation id="13472A178177FFB2FBD4FEF4FB90FE70" box="[1043,1129,373,399]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
), without sutural stria; epipleura very broad anteriorly, reaching posterior margin of first visible abdominal sternite, with sparse, coarse punctures. Femora densely pubescent, with apical eighth glabrous (
<figureCitation id="13472A178177FFB2FBD8FE3AFB5FFE2A" box="[1055,1190,443,469]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="8" pageNumber="9">Figs 10, 17</figureCitation>
). Metatarsomeres with dense white hairs ventrally and some long swimming hairs dorsally. Claws of moderate size, rather strongly curved.
</paragraph>
<caption id="DF03661A8178FFBDFF7AF80BFDD0F814" ID-DOI="http://doi.org/10.5281/zenodo.3785484" ID-Zenodo-Dep="3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,241,1930,1956]" targetBox="[262,1323,269,1889]" targetPageId="7">
<paragraph id="8BC336928178FFBDFF7AF80BFDD0F814" blockId="7.[189,1399,1930,2027]" pageId="7" pageNumber="8">
<emphasis id="B908EA808178FFBDFF7AF80BFEC0F85B" bold="true" box="[189,313,1930,1956]" pageId="7" pageNumber="8">Figs 9–17.</emphasis>
<taxonomicName id="4C7C4D118178FFBDFEF9F80BFCC0F85B" authority="d'Orchymont, 1932" authorityName="d'Orchymont" authorityYear="1932" box="[318,825,1930,1956]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B908EA808178FFBDFEF9F80BFDB2F85B" box="[318,587,1930,1956]" italics="true" pageId="7" pageNumber="8">Helochares fuliginosus</emphasis>
<bibRefCitation id="EFED4B638178FFBDFD97F80BFCC0F85B" author="d'Orchymont A." box="[592,825,1930,1956]" pageId="7" pageNumber="8" pagination="623 - 714" refId="ref16315" refString="d'Orchymont A. 1932. Zur Kenntnis der Kolbenwasserkafer (Palpicornia) von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement IX (Tropische Binnengewasser II): 623 - 714." type="book chapter" year="1932">d’Orchymont, 1932</bibRefCitation>
</taxonomicName>
. 
<emphasis id="B908EA808178FFBDFC84F80BFC82F85B" bold="true" box="[835,891,1930,1956]" pageId="7" pageNumber="8">9–11</emphasis>
. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 
<emphasis id="B908EA808178FFBDFF7AF82CFEFEF838" bold="true" box="[189,263,1965,1991]" pageId="7" pageNumber="8">12–13</emphasis>
. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 
<emphasis id="B908EA808178FFBDFCA2F82CFC7BF838" bold="true" box="[869,898,1965,1991]" pageId="7" pageNumber="8">14</emphasis>
. Head, dorsal. 
<emphasis id="B908EA808178FFBDFBF1F82CFBAAF838" bold="true" box="[1078,1107,1965,1991]" pageId="7" pageNumber="8">15</emphasis>
. Mesoventrite. 
<emphasis id="B908EA808178FFBDFACBF82CFAD0F838" bold="true" box="[1292,1321,1965,1991]" pageId="7" pageNumber="8">16</emphasis>
. Head and prosternum. 
<emphasis id="B908EA808178FFBDFE44F850FE59F814" bold="true" box="[387,416,2001,2027]" pageId="7" pageNumber="8">17</emphasis>
. Abdomen.
</paragraph>
</caption>
<paragraph id="8BC336928177FFB2FF7AFDC2FAA2FDA2" blockId="8.[189,1371,579,606]" box="[189,1371,579,606]" pageId="8" pageNumber="9">
ABDOMEN. All visible abdominal ventrites with dense pubescence; first ventrite not carinate (
<figureCitation id="13472A178177FFB2FB31FDC2FAB4FDA2" box="[1270,1357,579,605]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BC336928177FFB2FF7AFD04FCF8FD1A" blockId="8.[189,1399,644,741]" pageId="8" pageNumber="9">
AEDEAGUS. Broad, paramere ca 6.5× as long as width of apex, subparallel, slightly expanded subapically and round apically, apex without small, inwardly directed tooth; median lobe somewhat expanded subapically, abruptly narrowed apically (
<figureCitation id="13472A178177FFB2FD5CFD4AFD0AFD1A" box="[667,755,715,741]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="8" pageNumber="9">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36665198177FFB2FF7AFC8DFE96FC1D" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BC336928177FFB2FF7AFC8DFE30FCD9" blockId="8.[189,457,780,806]" box="[189,457,780,806]" pageId="8" pageNumber="9">
<heading id="D08B81FE8177FFB2FF7AFC8DFE30FCD9" bold="true" box="[189,457,780,806]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B908EA808177FFB2FF7AFC8DFE30FCD9" bold="true" box="[189,457,780,806]" pageId="8" pageNumber="9">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC336928177FFB2FF7AFCBAFE96FC1D" blockId="8.[189,1399,827,994]" pageId="8" pageNumber="9">
This species is closest to 
<taxonomicName id="4C7C4D118177FFB2FE24FCBDFDBFFCAA" authority="Jia &amp; Tang, 2018" authorityName="Jia &amp; Tang" authorityYear="2018" box="[483,582,828,853]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="songi" status="sp. nov.">
<emphasis id="B908EA808177FFB2FE24FCBDFDBFFCAA" box="[483,582,828,853]" italics="true" pageId="8" pageNumber="9">H. songi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B57FB8177FFB2FD8AFCBDFD53FCA9" box="[589,682,828,854]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can easily be distinguished by its smaller size; clypeus yellowish brown laterally (
<figureCitation id="13472A178177FFB2FD9FFCDFFD25FC87" box="[600,732,862,888]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="8" pageNumber="9">Fig. 12–14</figureCitation>
); elytra with yellowish or reddish brown margin laterally (
<figureCitation id="13472A178177FFB2FF01FC00FEE5FC64" box="[198,284,897,923]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
); maxillary palps uniform yellow or yellowish brown (
<figureCitation id="13472A178177FFB2FC63FC00FBD4FC64" box="[932,1069,897,923]" captionStart="Figs 9–17" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[262,1323,269,1889]" captionTargetId="figure@7.[189,1398,250,1909]" captionTargetPageId="7" captionText="Figs 9–17. Helochares fuliginosus d’Orchymont, 1932. 9–11. Habitus. 9. Dorsal. 10. Ventral. 11. Lateral. 12–13. Head and pronotum. 12. Dorsal. 13. Dorsolateral. 14. Head, dorsal. 15. Mesoventrite. 16. Head and prosternum. 17. Abdomen." figureDoi="http://doi.org/10.5281/zenodo.3785484" httpUri="https://zenodo.org/record/3785484/files/figure.png" pageId="8" pageNumber="9">Figs 13–14</figureCitation>
); pronotum and elytra more finely punctate; parameres ca 6.5× as long as width of apex, apex without a small, inwardly directed tooth (
<figureCitation id="13472A178177FFB2FECEFC49FE98FC1D" box="[265,353,968,994]" captionStart="Figs 47–55" captionStartId="22.[189,241,1894,1920]" captionTargetBox="[280,1297,268,1859]" captionTargetId="figure@22.[275,1312,253,1869]" captionTargetPageId="22" captionText="Figs 47–55. Aedeagi in species of Helochares Mulsant, 1844. 47. H. fuliginosus d’Orchymont, 1932. 48–49. H. songi sp. nov. 50–51. H. atropiceus Régimbart, 1903. 50. From Guangxi. 51. From Hong Kong. 52. H. pallens (MacLeay, 1825). 53–55. H. obscurus (Müller, 1776). 53. From Xinjiang. 54–55. After Hansen (1987)." figureDoi="http://doi.org/10.5281/zenodo.3785494" httpUri="https://zenodo.org/record/3785494/files/figure.png" pageId="8" pageNumber="9">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36665198177FFB2FF7AFB88FC02FB67" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="8BC336928177FFB2FF7AFB88FEE5FBDC" blockId="8.[189,284,1033,1059]" box="[189,284,1033,1059]" pageId="8" pageNumber="9">
<heading id="D08B81FE8177FFB2FF7AFB88FEE5FBDC" bold="true" box="[189,284,1033,1059]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B908EA808177FFB2FF7AFB88FEE5FBDC" bold="true" box="[189,284,1033,1059]" pageId="8" pageNumber="9">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC336928177FFB2FF7AFBB9FC02FB67" blockId="8.[189,1399,1080,1177]" pageId="8" pageNumber="9">
This species occurs in small stagnant pools and on wet stones on ground with grass or leaf litter near rivers or streams, but never collected in habitats where 
<taxonomicName id="4C7C4D118177FFB2FC82FBDAFC4BFB8A" authorityName="Sharp" authorityYear="1882" box="[837,946,1115,1141]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B908EA808177FFB2FC82FBDAFC4BFB8A" box="[837,946,1115,1141]" italics="true" pageId="8" pageNumber="9">Oocyclus</emphasis>
</taxonomicName>
spp. occur. It is occasionally collected at light. The female carries the egg case under the abdominal ventrites.
</paragraph>
</subSubSection>
<subSubSection id="C36665198177FFB2FF7AFB3EFBC8FAD4" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BC336928177FFB2FF7AFB3EFEAFFB26" blockId="8.[189,342,1215,1241]" box="[189,342,1215,1241]" pageId="8" pageNumber="9">
<heading id="D08B81FE8177FFB2FF7AFB3EFEAFFB26" bold="true" box="[189,342,1215,1241]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B908EA808177FFB2FF7AFB3EFEAFFB26" bold="true" box="[189,342,1215,1241]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC336928177FFB2FF7AFB6FFBC8FAD4" blockId="8.[189,1398,1262,1324]" pageId="8" pageNumber="9">
<collectingCountry id="F36B76028177FFB2FF7AFB6FFEFAFAF7" box="[189,259,1262,1288]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
(
<collectingRegion id="49B8F8708177FFB2FEDFFB6FFE91FAF7" box="[280,360,1262,1288]" country="China" name="Fujian" pageId="8" pageNumber="9">Fujian</collectingRegion>
, 
<collectingRegion id="49B8F8708177FFB2FEBDFB6FFDFFFAF7" box="[378,518,1262,1288]" country="China" name="Guangdong" pageId="8" pageNumber="9">Guangdong</collectingRegion>
, 
<collectingRegion id="49B8F8708177FFB2FDDFFB6FFD79FAF7" box="[536,640,1262,1288]" country="China" name="Guangxi" pageId="8" pageNumber="9">Guangxi</collectingRegion>
, 
<collectingCountry id="F36B76028177FFB2FD56FB6EFCDDFAF6" box="[657,804,1263,1289]" name="Hong Kong" pageId="8" pageNumber="9">Hong Kong</collectingCountry>
, 
<collectingCountry id="F36B76028177FFB2FCF1FB6EFC73FAF6" box="[822,906,1263,1289]" name="Macao" pageId="8" pageNumber="9">Macau</collectingCountry>
), 
<collectingCountry id="F36B76028177FFB2FC63FB6FFBD9FAF7" box="[932,1056,1262,1288]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
, 
<collectingCountry id="F36B76028177FFB2FBF6FB6FFB5AFAF7" box="[1073,1187,1262,1288]" name="Indonesia" pageId="8" pageNumber="9">Indonesia</collectingCountry>
(
<collectingRegion id="49B8F8708177FFB2FB7EFB6FFB08FAF7" box="[1209,1265,1262,1288]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="9">Java</collectingRegion>
, 
<collectingRegion id="49B8F8708177FFB2FAC3FB6FFA93FAF7" box="[1284,1386,1262,1288]" country="Indonesia" name="Sumatera Utara" pageId="8" pageNumber="9">Sumatra</collectingRegion>
), 
<collectingCountry id="F36B76028177FFB2FF7AFA90FED0FAD4" box="[189,297,1297,1323]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
(peninsula) (
<bibRefCitation id="EFED4B638177FFB2FE04FA90FD9AFAD4" author="Hansen M." box="[451,611,1297,1323]" pageId="8" pageNumber="9" refId="ref15262" refString="Hansen M. 1999. World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Stenstrup, Denmark." type="book" year="1999">Hansen 1999</bibRefCitation>
). New record for 
<collectingCountry id="F36B76028177FFB2FCF5FA90FC81FAD4" box="[818,888,1297,1323]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
and 
<collectingCountry id="F36B76028177FFB2FC76FA90FBD7FAD4" box="[945,1070,1297,1323]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>