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<document id="55174739579A06CA54D046532FE979A6" ID="10.11646/zootaxa.4258.3.1SLASH10916" ID-CLB-Dataset="33146" ID-DOI="10.11646/zootaxa.4258.3.1" ID-GBIF-Dataset="97942778-cef4-4cf6-b4e4-c077e633daa8" ID-ISSN="1175-5326" ID-Zenodo-Dep="569729" ID-ZooBank="DE89E162-B255-474C-A824-20DFA551DBBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493363817835" checkinUser="plazi" docAuthor="Penz, Carla M., Casagrande, Mirna M., Devries, Phil &amp; Simonsen, Thomas J." docDate="2017" docId="03BB87F4FFA4FF83E1FF2737B843FE74" docLanguage="en" docName="zootaxa.4258.3.1SLASH10916.pdf" docOrigin="Zootaxa 4258 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bia actorion subsp. occulta Casagrande &amp; Penz, NEW SSP." docType="treatment" docVersion="7" lastPageNumber="211" masterDocId="FF82FF8CFFACFF89E1682278BA7DFFFC" masterDocTitle="Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)" masterLastPageNumber="237" masterPageNumber="201" pageNumber="209" updateTime="1698439134094" updateUser="ExternalLinkService">
<mods:mods id="28D784C281ACF10BE1BBA4056E267580" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="83AA73E66C872A5E5690FDB879B8937E">
<mods:title id="478DF5AC64F72C22889C8751839CCA0F">Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)</mods:title>
</mods:titleInfo>
<mods:name id="E3225394D0C6830CA4AE09B5D0302537" type="personal">
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<mods:namePart id="E5B7D644641AEF79BF6D3E1B1FB3D8EA">Penz, Carla M.</mods:namePart>
</mods:name>
<mods:name id="3EABC3A3C8CC855FD4515C5814653610" type="personal">
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<mods:namePart id="19F6421DEE50F1559B3A47580594BD4C">Casagrande, Mirna M.</mods:namePart>
</mods:name>
<mods:name id="A1F52C0F1EC557AFBD3B28B33A423D16" type="personal">
<mods:role id="60E24BB218266C4F6FA6CFA3C31B40B3">
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<mods:namePart id="9382D2118582CDA3E298C8F3A7419250">Devries, Phil</mods:namePart>
</mods:name>
<mods:name id="9BA622009A580CBF9B9C8B6D6978A3BF" type="personal">
<mods:role id="98FD0015425960EA7782D8260059D9B7">
<mods:roleTerm id="E97B539ED23CF76AD29EC1DA358CD6CF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC50264FE77DCF3F6D858F4E2029C7F7">Simonsen, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CCB3872E47FC143F796A9604521127D0">text</mods:typeOfResource>
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<mods:titleInfo id="12FE3B181CA6129A3B208C2E2D8C97A4">
<mods:title id="63C9501B94F837D1993509AAEA42CEA2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="875BA2E51E37E3EB8A6DAE74A3D2BC12">
<mods:date id="BF5DEE2B3513DFD9349BA7252FD877C5">2017</mods:date>
<mods:detail id="9664D83ADFCBC32B1A99ED5E36A11D4A" type="volume">
<mods:number id="DA74E8A501937CA50CE9C38ECC31078B">4258</mods:number>
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<mods:detail id="CF1288106BE3359EF9590BE8DE9E4E0F" type="issue">
<mods:number id="4BE2812A591BC73C6BCC184D5729D2A4">3</mods:number>
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<mods:classification id="89969B8FD296A8330D17C10A41B9538B">journal article</mods:classification>
<mods:identifier id="BF4B58C4578EB83E3A45291CB7348E41" type="CLB-Dataset">33146</mods:identifier>
<mods:identifier id="1A66B5DA644D9294AF7EF4FA9549CF21" type="DOI">10.11646/zootaxa.4258.3.1</mods:identifier>
<mods:identifier id="303A73CF771E22DC53A23F5570BCEC83" type="GBIF-Dataset">97942778-cef4-4cf6-b4e4-c077e633daa8</mods:identifier>
<mods:identifier id="AE93DC4A9E89A22EB47E2A1970C00DBB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AEF94601F906E2E9D7EAB58F0E0FCD31" type="Zenodo-Dep">569729</mods:identifier>
<mods:identifier id="31AF0775EE53DA2AF8792BBA064B0ECF" type="ZooBank">DE89E162-B255-474C-A824-20DFA551DBBD</mods:identifier>
</mods:mods>
<treatment id="03BB87F4FFA4FF83E1FF2737B843FE74" ID-DOI="http://doi.org/10.5281/zenodo.6017375" ID-GBIF-Taxon="128810412" ID-Zenodo-Dep="6017375" LSID="urn:lsid:plazi:treatment:03BB87F4FFA4FF83E1FF2737B843FE74" httpUri="http://treatment.plazi.org/id/03BB87F4FFA4FF83E1FF2737B843FE74" lastPageId="10" lastPageNumber="211" pageId="8" pageNumber="209">
<subSubSection id="C3086569FFA4FF81E1FF2737BB0EFA76" pageId="8" pageNumber="209" type="nomenclature">
<paragraph id="8BAD36E2FFA4FF81E1FF2737B95DFA95" blockId="8.[151,800,1359,1418]" box="[151,800,1359,1385]" pageId="8" pageNumber="209">
<heading id="D0E5818EFFA4FF81E1FF2737B95DFA95" bold="true" box="[151,800,1359,1385]" fontSize="11" level="1" pageId="8" pageNumber="209" reason="1">
<emphasis id="B966EAF0FFA4FF81E1FF2737B95DFA95" bold="true" box="[151,800,1359,1385]" pageId="8" pageNumber="209">
<taxonomicName id="4C124D61FFA4FF81E1FF2737B8F3FA95" authority="Casagrande &amp; Penz" authorityName="Casagrande &amp; Penz" box="[151,654,1359,1385]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="subSpecies" species="actorion" status="subsp. nov." subSpecies="occulta">
<emphasis id="B966EAF0FFA4FF81E1FF2737BBF1FA95" bold="true" box="[151,396,1359,1385]" italics="true" pageId="8" pageNumber="209">Bia actorion occulta</emphasis>
Casagrande &amp; Penz
</taxonomicName>
,
<taxonomicNameLabel id="A255578BFFA4FF81E3F32737B95DFA95" box="[667,800,1359,1385]" pageId="8" pageNumber="209" rank="subSpecies">NEW SSP.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD36E2FFA4FF81E1FF2709BB0EFA76" blockId="8.[151,800,1359,1418]" box="[151,371,1393,1418]" pageId="8" pageNumber="209">
(
<figureCitation id="13292A67FFA4FF81E1F72709BA9EFA76" box="[159,227,1393,1418]" captionStart="FIGURE 3" captionStartId="9.[151,250,1853,1875]" captionTargetBox="[247,1326,215,1779]" captionTargetId="figure@9.[205,1373,193,1811]" captionTargetPageId="9" captionText="FIGURE 3. Adults in dorsal and ventral views. Taxon names appear below each set. Bia actorion actorion: a, male, Guyana, Kartabo (CMNH); b, female, Guyana, Two Hat Mountain, E Kanukus S Rupununi (USNM). Bia actorion ecuatoria DeVries &amp; Penz, new ssp.: c, male holotype, Ecuador, Sucumbios, Garza Cocha (PJD); d, female paratype, Ecuador, Sucumbios, Garza Cocha (PJD). Bia actorion occulta Casagrande &amp; Penz, new ssp.: e, male holotype, Brazil, Pará, Acará (DZUP); f, female paratype, Brazil, Pará, Sto Antonio do Tauá (DZUP). Intermediate between actorion and decaerulea: g, male, Suriname (RMNH); h, male, Suriname, Coronie (RMNH)." httpUri="https://zenodo.org/record/569732/files/figure.png" pageId="8" pageNumber="209">Figs 3</figureCitation>
e–f, 10c, 12)
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFA4FF81E1FF27CCB858F8E7" pageId="8" pageNumber="209" type="description">
<paragraph id="8BAD36E2FFA4FF81E1FF27CCB858F8E7" blockId="8.[151,1436,1460,2036]" pageId="8" pageNumber="209">
<emphasis id="B966EAF0FFA4FF81E1FF27CCBBD5FA31" bold="true" box="[151,424,1460,1485]" pageId="8" pageNumber="209">Diagnostic description.</emphasis>
Defined by the following combination of characters: (1) MF DFW white apical ocelli medium-small. (2) MF DFW orange band moderately wide and in M orange scales not conspicuously extended along veins. (3) M DFW iridescent band from anal margin to usually more than half of the CuA2 cell, with a scatter of scales reaching CuA2 (one specimen differed, where the band was ca. half of the cell). F DFW blue iridescence well developed but variable; usually extended across the entire DC, diffuse anteriorly; usually visible at the CuA1-CuA2 intersection; below the discal cell, the iridescent area is wide and it spreads towards the tornus. F seems indistinguishable from
<taxonomicName id="4C124D61FFA4FF81E0F124EEB826F951" box="[409,603,1686,1709]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="subSpecies" species="actorion" subSpecies="actorion">
<emphasis id="B966EAF0FFA4FF81E0F124EEB826F951" box="[409,603,1686,1709]" italics="true" pageId="8" pageNumber="209">actorion actorion</emphasis>
</taxonomicName>
. (4) M DFW androconial organ on the CuA-CuA1-CuA2 intersection dark brown, matching scale color of surrounding area. (5) DHW discal androconial pad chocolate-brown, comparable in color to the associated hairpencil. (6) M DHW discal hairpencil dark brown. (7) F VFW ripple pattern of the postmedial area similar to that of M.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFA4FF81E1AF255FB911F898" pageId="8" pageNumber="209" type="etymology">
<paragraph id="8BAD36E2FFA4FF81E1AF255FB911F898" blockId="8.[151,1436,1460,2036]" pageId="8" pageNumber="209">
<emphasis id="B966EAF0FFA4FF81E1AF255FBB36F8BC" bold="true" box="[199,331,1831,1856]" pageId="8" pageNumber="209">Etymology.</emphasis>
The name
<emphasis id="B966EAF0FFA4FF81E0AE2550B86AF8BC" box="[454,535,1832,1856]" italics="true" pageId="8" pageNumber="209">occulta</emphasis>
originates from the Latin word for hidden, alluding to the fact that this subspecies occupies the southeastern edge of the species’ geographical range.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFA4FF81E1AF2517BB6DF80F" pageId="8" pageNumber="209" type="materials_examined">
<paragraph id="8BAD36E2FFA4FF81E1AF2517BB6DF80F" blockId="8.[151,1436,1460,2036]" pageId="8" pageNumber="209">
<emphasis id="B966EAF0FFA4FF81E1AF2517BB0AF874" bold="true" box="[199,375,1903,1928]" pageId="8" pageNumber="209">Type material.</emphasis>
Holotype M (
<figureCitation id="13292A67FFA4FF81E34E2517B810F874" box="[550,621,1903,1928]" captionStart="FIGURE 3" captionStartId="9.[151,250,1853,1875]" captionTargetBox="[247,1326,215,1779]" captionTargetId="figure@9.[205,1373,193,1811]" captionTargetPageId="9" captionText="FIGURE 3. Adults in dorsal and ventral views. Taxon names appear below each set. Bia actorion actorion: a, male, Guyana, Kartabo (CMNH); b, female, Guyana, Two Hat Mountain, E Kanukus S Rupununi (USNM). Bia actorion ecuatoria DeVries &amp; Penz, new ssp.: c, male holotype, Ecuador, Sucumbios, Garza Cocha (PJD); d, female paratype, Ecuador, Sucumbios, Garza Cocha (PJD). Bia actorion occulta Casagrande &amp; Penz, new ssp.: e, male holotype, Brazil, Pará, Acará (DZUP); f, female paratype, Brazil, Pará, Sto Antonio do Tauá (DZUP). Intermediate between actorion and decaerulea: g, male, Suriname (RMNH); h, male, Suriname, Coronie (RMNH)." httpUri="https://zenodo.org/record/569732/files/figure.png" pageId="8" pageNumber="209">Fig. 3</figureCitation>
e), deposited in the DZUP collection, four labels separated by // and transcribed verbatim:
<materialsCitation id="3B7A3CBFFFA4FF81E0E125ECB950F82C" ID-GBIF-Occurrence="1499315711" collectingDate="2006-07-23" collectorName="P. Jauffret" location="Alca Viaria Km" municipality="Acara" pageId="8" pageNumber="209" specimenCount="2" stateProvince="Para" typeStatus="holotype">
<typeStatus id="54A98840FFA4FF81E0E125ECB850F850" box="[393,557,1940,1964]" pageId="8" pageNumber="209" type="holotype">HOLOTYPUS</typeStatus>
//
<collectingRegion id="49D6F800FFA4FF81E32425ECB802F850" box="[588,639,1940,1964]" country="Brazil" name="Para" pageId="8" pageNumber="209">Pará</collectingRegion>
,
<collectingMunicipality id="6BC9AC98FFA4FF81E3E425ECB8ADF850" box="[652,720,1940,1964]" pageId="8" pageNumber="209">Acará</collectingMunicipality>
,
<location id="8ECD6039FFA4FF81E3B425EBB9F1F857" LSID="urn:lsid:plazi:treatment:03BB87F4FFA4FF83E1FF2737B843FE74:8ECD6039FFA4FF81E3B425EBB9F1F857" box="[732,908,1939,1964]" municipality="Acara" name="Alca Viaria Km" pageId="8" pageNumber="209" stateProvince="Para">Alça Viária Km</location>
20,
<collectingDate id="EFE8E9CAFFA4FF81E2D725EBBE31F850" box="[959,1100,1939,1964]" pageId="8" pageNumber="209" value="2006-07-23">
<date id="FFAC1022FFA4FF81E2D725EBBE31F850" box="[959,1100,1939,1964]" pageId="8" pageNumber="209" value="2006-07-23">23-VII-2006</date>
</collectingDate>
,
<collectorName id="26E75334FFA4FF81E53025EBBEC0F850" box="[1112,1213,1939,1964]" pageId="8" pageNumber="209">P.Jauffret</collectorName>
leg. // DZ 22.447 //
<typeStatus id="54A98840FFA4FF81E1FF25CFBB74F82C" box="[151,265,1975,2000]" pageId="8" pageNumber="209" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C124D61FFA4FF81E07B25C0B950F82C" authority="Casagrande &amp; Penz, 2017" authorityName="Casagrande &amp; Penz" authorityYear="2017" box="[275,813,1975,2000]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="subSpecies" species="actorion" subSpecies="occulta">
<emphasis id="B966EAF0FFA4FF81E07B25C0BB84F82C" box="[275,505,1976,2000]" italics="true" pageId="8" pageNumber="209">Bia actorion occulta</emphasis>
Casagrande &amp; Penz, 2017
</taxonomicName>
</materialsCitation>
. Paratypes are listed in Appendix, and
<figureCitation id="13292A67FFA4FF81E58425CFBF50F82C" box="[1260,1325,1975,2000]" captionStart="FIGURE 3" captionStartId="9.[151,250,1853,1875]" captionTargetBox="[247,1326,215,1779]" captionTargetId="figure@9.[205,1373,193,1811]" captionTargetPageId="9" captionText="FIGURE 3. Adults in dorsal and ventral views. Taxon names appear below each set. Bia actorion actorion: a, male, Guyana, Kartabo (CMNH); b, female, Guyana, Two Hat Mountain, E Kanukus S Rupununi (USNM). Bia actorion ecuatoria DeVries &amp; Penz, new ssp.: c, male holotype, Ecuador, Sucumbios, Garza Cocha (PJD); d, female paratype, Ecuador, Sucumbios, Garza Cocha (PJD). Bia actorion occulta Casagrande &amp; Penz, new ssp.: e, male holotype, Brazil, Pará, Acará (DZUP); f, female paratype, Brazil, Pará, Sto Antonio do Tauá (DZUP). Intermediate between actorion and decaerulea: g, male, Suriname (RMNH); h, male, Suriname, Coronie (RMNH)." httpUri="https://zenodo.org/record/569732/files/figure.png" pageId="8" pageNumber="209">Fig. 3</figureCitation>
f shows a paratype F.
</paragraph>
</subSubSection>
<caption id="DF6D666AFFA5FF80E1FF2545B8FEF811" httpUri="https://zenodo.org/record/569732/files/figure.png" pageId="9" pageNumber="210" targetBox="[247,1326,215,1779]" targetPageId="9">
<paragraph id="8BAD36E2FFA5FF80E1FF2545B8FEF811" blockId="9.[151,1436,1853,2029]" pageId="9" pageNumber="210">
<emphasis id="B966EAF0FFA5FF80E1FF2545BB6EF8AF" bold="true" box="[151,275,1853,1875]" pageId="9" pageNumber="210">FIGURE 3.</emphasis>
Adults in dorsal and ventral views. Taxon names appear below each set.
<taxonomicName id="4C124D61FFA5FF80E56B2547BE9DF8A8" box="[1027,1248,1855,1876]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="subSpecies" species="actorion" subSpecies="actorion">
<emphasis id="B966EAF0FFA5FF80E56B2547BE9DF8A8" box="[1027,1248,1855,1876]" italics="true" pageId="9" pageNumber="210">Bia actorion actorion</emphasis>
</taxonomicName>
:
<emphasis id="B966EAF0FFA5FF80E5862545BE87F8AF" bold="true" box="[1262,1274,1853,1875]" pageId="9" pageNumber="210">a</emphasis>
, male, Guyana, Kartabo (CMNH);
<emphasis id="B966EAF0FFA5FF80E0302524BB1BF88E" bold="true" box="[344,358,1884,1906]" pageId="9" pageNumber="210">b</emphasis>
, female, Guyana, Two Hat Mountain, E Kanukus S Rupununi (USNM).
<taxonomicName id="4C124D61FFA5FF80E52C2525BAB6F86D" authority="DeVries &amp; Penz" authorityName="DeVries &amp; Penz" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="subSpecies" species="actorion" status="subsp. nov." subSpecies="ecuatoria">
<emphasis id="B966EAF0FFA5FF80E52C2525BF54F88E" box="[1092,1321,1885,1906]" italics="true" pageId="9" pageNumber="210">Bia actorion ecuatoria</emphasis>
DeVries &amp; Penz
</taxonomicName>
,
<taxonomicNameLabel id="A255578BFFA5FF80E1BE2504BB4EF86D" box="[214,307,1916,1937]" pageId="9" pageNumber="210" rank="subSpecies">new ssp.</taxonomicNameLabel>
:
<emphasis id="B966EAF0FFA5FF80E0562503BB34F86D" bold="true" box="[318,329,1915,1937]" pageId="9" pageNumber="210">c</emphasis>
, male holotype, Ecuador, Sucumbios, Garza Cocha (PJD);
<emphasis id="B966EAF0FFA5FF80E2D82503B9C3F86D" bold="true" box="[944,958,1915,1937]" pageId="9" pageNumber="210">d</emphasis>
, female paratype, Ecuador, Sucumbios, Garza Cocha (PJD).
<taxonomicName id="4C124D61FFA5FF80E04325E3B8A4F84C" authority="Casagrande &amp; Penz" authorityName="Casagrande &amp; Penz" box="[299,729,1946,1968]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="subSpecies" species="actorion" status="subsp. nov." subSpecies="occulta">
<emphasis id="B966EAF0FFA5FF80E04325E3B87CF84C" box="[299,513,1946,1968]" italics="true" pageId="9" pageNumber="210">Bia actorion occulta</emphasis>
Casagrande &amp; Penz
</taxonomicName>
,
<taxonomicNameLabel id="A255578BFFA5FF80E38E25E3B939F84C" box="[742,836,1947,1968]" pageId="9" pageNumber="210" rank="subSpecies">new ssp.</taxonomicNameLabel>
:
<emphasis id="B966EAF0FFA5FF80E23825E2B926F84C" bold="true" box="[848,859,1946,1968]" pageId="9" pageNumber="210">e</emphasis>
, male holotype, Brazil, Pará, Acará (DZUP);
<emphasis id="B966EAF0FFA5FF80E45725E2BF35F84C" bold="true" box="[1343,1352,1946,1968]" pageId="9" pageNumber="210">f</emphasis>
, female paratype, Brazil, Pará, Sto Antonio do Tauá (DZUP). Intermediate between
<taxonomicName id="4C124D61FFA5FF80E2AF25C1BE60F832" box="[967,1053,1977,1998]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="actorion">
<emphasis id="B966EAF0FFA5FF80E2AF25C1BE60F832" box="[967,1053,1977,1998]" italics="true" pageId="9" pageNumber="210">actorion</emphasis>
</taxonomicName>
and
<taxonomicName id="4C124D61FFA5FF80E53F25C0BEB5F832" box="[1111,1224,1976,1998]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="decaerulea">
<emphasis id="B966EAF0FFA5FF80E53F25C0BEB5F832" box="[1111,1224,1976,1998]" italics="true" pageId="9" pageNumber="210">decaerulea</emphasis>
</taxonomicName>
:
<emphasis id="B966EAF0FFA5FF80E5B225C0BE9AF832" bold="true" box="[1242,1255,1976,1998]" pageId="9" pageNumber="210">g</emphasis>
, male, Suriname (RMNH);
<emphasis id="B966EAF0FFA5FF80E06925AFBB72F811" bold="true" box="[257,271,2007,2029]" pageId="9" pageNumber="210">h</emphasis>
, male, Suriname, Coronie (RMNH).
</paragraph>
</caption>
<subSubSection id="C3086569FFA6FF83E1AF22EFB9ECFF4C" box="[199,913,151,176]" pageId="10" pageNumber="211" type="distribution">
<paragraph id="8BAD36E2FFA6FF83E1AF22EFB9ECFF4C" blockId="10.[151,1436,151,392]" box="[199,913,151,176]" pageId="10" pageNumber="211">
<emphasis id="B966EAF0FFA6FF83E1AF22EFB8F2FF4C" bold="true" box="[199,655,151,176]" pageId="10" pageNumber="211">Distribution and examined specimens.</emphasis>
<figureCitation id="13292A67FFA6FF83E3FD22EFB898FF4C" box="[661,741,151,176]" captionStart="FIGURE 12" captionStartId="26.[151,250,1021,1043]" captionTargetBox="[154,1414,195,985]" captionTargetId="figure@26.[151,1435,193,1000]" captionTargetPageId="26" captionText="FIGURE 12. Map showing collection localities of Bia species and subspecies (see Appendix). Some close localities are represented by one single dot. Dubious label data, localities that could not be accurately identified or were incomplete (e. g., Brazil, Pará) were not placed on map." httpUri="https://zenodo.org/record/569742/files/figure.png" pageId="10" pageNumber="211">Fig. 12</figureCitation>
and Appendix.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFA6FF83E1AF22C4B843FE74" pageId="10" pageNumber="211" type="discussion">
<paragraph id="8BAD36E2FFA6FF83E1AF22C4B843FE74" blockId="10.[151,1436,151,392]" pageId="10" pageNumber="211">
<emphasis id="B966EAF0FFA6FF83E1AF22C4BB46FF29" bold="true" box="[199,315,188,213]" pageId="10" pageNumber="211">Remarks.</emphasis>
Male specimens from
<collectingCountry id="F3057672FFA6FF83E35622C4B8F8FF29" box="[574,645,188,213]" name="Brazil" pageId="10" pageNumber="211">Brazil</collectingCountry>
,
<collectingRegion id="49D6F800FFA6FF83E3F822C4B8B8FF28" box="[656,709,188,212]" country="Brazil" name="Para" pageId="10" pageNumber="211">Pará</collectingRegion>
, Caxiuanã have a much reduced male dorsal forewing iridescent band. This band is absent in two specimens from
<collectingRegion id="49D6F800FFA6FF83E38E2298B966FF04" box="[742,795,224,248]" country="Brazil" name="Para" pageId="10" pageNumber="211">Pará</collectingRegion>
, Carajás and
<collectingRegion id="49D6F800FFA6FF83E2D522A7BE4FFF04" box="[957,1074,223,248]" country="Brazil" name="Maranhao" pageId="10" pageNumber="211">Maranhão</collectingRegion>
, Imperatriz. Inasmuch as these specimens markedly differ from those of other localities, we tentatively assign them to
<taxonomicName id="4C124D61FFA6FF83E5C6237DBF11FEE0" box="[1198,1388,260,284]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="subSpecies" species="actorion" subSpecies="occulta">
<emphasis id="B966EAF0FFA6FF83E5C6237DBF11FEE0" box="[1198,1388,260,284]" italics="true" pageId="10" pageNumber="211">actorion occulta</emphasis>
</taxonomicName>
but recognize that this should be re-visited when additional specimens become available. It would be of interest to confirm the existence of a north-south cline within
<taxonomicName id="4C124D61FFA6FF83E3822335B9D9FE98" box="[746,932,332,356]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="subSpecies" species="actorion" subSpecies="occulta">
<emphasis id="B966EAF0FFA6FF83E3822335B9D9FE98" box="[746,932,332,356]" italics="true" pageId="10" pageNumber="211">actorion occulta</emphasis>
</taxonomicName>
with regard to the size and presence of the male dorsal forewing iridescent band.
</paragraph>
</subSubSection>
</treatment>
</document>
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