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<document ID-DOI="10.5281/zenodo.272951" ID-GBIF-Dataset="e2c88f4c-3ac2-45f9-95e4-99b75561a081" ID-ISSN="1175-5326" ID-Zenodo-Dep="272951" ID-ZooBank="6D68A019-6F63-4AA4-A8B3-92D351F1F69B" checkinTime="1484409208030" checkinUser="plazi" docAuthor="Van, Rob W. M." docDate="2017" docId="03A8001077EAFF13FF14A53893DCFC52" docLanguage="en" docName="zootaxa.4217.1.1.pdf.imf" docOrigin="Zootaxa 1" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5ce3343f" docStyleName="zootaxa.2013.monograph" docTitle="Suberites crispolobatus Van, 2017, sp. nov." docType="treatment" docVersion="19" lastPageNumber="192" masterDocId="FF9178687757FFACFF83A36B907DFF9A" masterDocTitle="Sponges of the Guyana Shelf" masterLastPageNumber="225" masterPageNumber="1" pageNumber="190" updateTime="1645784257244" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Sponges of the Guyana Shelf</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Van, Rob W. M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>225</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.272951</mods:identifier>
<mods:identifier type="GBIF-Dataset">e2c88f4c-3ac2-45f9-95e4-99b75561a081</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">272951</mods:identifier>
<mods:identifier type="ZooBank">6D68A019-6F63-4AA4-A8B3-92D351F1F69B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5698760" ID-GBIF-Taxon="127419796" ID-Zenodo-Dep="5698760" LSID="urn:lsid:plazi:treatment:03A8001077EAFF13FF14A53893DCFC52" httpUri="http://treatment.plazi.org/id/03A8001077EAFF13FF14A53893DCFC52" lastPageId="191" lastPageNumber="192" pageId="189" pageNumber="190">
<subSubSection pageId="189" pageNumber="190" type="nomenclature">
<paragraph blockId="189.[151,532,1619,1678]" box="[151,532,1619,1645]" pageId="189" pageNumber="190">
<heading bold="true" box="[151,532,1619,1645]" fontSize="11" level="1" pageId="189" pageNumber="190" reason="1">
<emphasis bold="true" box="[151,532,1619,1645]" pageId="189" pageNumber="190">
<taxonomicName box="[151,431,1619,1645]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="189" pageNumber="190" phylum="Porifera" rank="species" species="crispolobatus" status="sp. nov.">
<emphasis bold="true" box="[151,431,1619,1645]" italics="true" pageId="189" pageNumber="190">Suberites crispolobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[439,532,1619,1645]" pageId="189" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="189.[151,532,1619,1678]" box="[151,323,1653,1678]" pageId="189" pageNumber="190">
<figureCitation box="[151,283,1653,1678]" captionStart="FIGURE 120" captionStartId="190.[151,250,1873,1895]" captionTargetBox="[151,1436,409,1852]" captionTargetId="figure@190.[151,1436,409,1852]" captionTargetPageId="190" captionText="FIGURE 120. Suberites crispolobatus sp. nov., a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a 1, habitus of paratypes RMNH Por. 10548, a 2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b – c, SEM images of the spicules, b, large tylostyle, b 1, details of b, c, small tylostyle, c 1, details of c." httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="189" pageNumber="190">Figures 120</figureCitation>
a–c
</paragraph>
</subSubSection>
<subSubSection lastPageId="190" lastPageNumber="191" pageId="189" pageNumber="190" type="materials_examined">
<paragraph blockId="189.[151,1437,1725,2038]" pageId="189" pageNumber="190">
<materialsCitation ID-GBIF-Occurrence="1424726535" collectingDate="1966-05-06" collectionCode="RMNH" collectorName="Snellius O. C. P. S." country="Suriname" elevation="59" latitude="7.0033" location="Suriname" longLatPrecision="7" longitude="-56.4417" pageId="189" pageNumber="190" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[151,385,1725,1750]" pageId="189" pageNumber="190">Material examined.</emphasis>
<typeStatus box="[393,497,1725,1750]" pageId="189" pageNumber="190">Holotype</typeStatus>
<collectionCode box="[505,589,1726,1749]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="189" pageNumber="190">RMNH</collectionCode>
Por. 9863,
<collectingCountry box="[720,830,1725,1750]" name="Suriname" pageId="189" pageNumber="190">Suriname</collectingCountry>
, ‘
<collectorName box="[846,1049,1725,1750]" pageId="189" pageNumber="190">Snellius O.C.P.S.</collectorName>
’
<collectingCountry box="[1062,1149,1726,1750]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
Shelf Expedition, station F40,
<geoCoordinate box="[208,317,1762,1786]" direction="north" orientation="latitude" pageId="189" pageNumber="190" precision="5" value="7.0033">7.0033°N</geoCoordinate>
<geoCoordinate box="[324,452,1761,1786]" direction="west" orientation="longitude" pageId="189" pageNumber="190" precision="5" value="-56.4417">56.4417°W</geoCoordinate>
, depth
<quantity box="[533,592,1761,1786]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="189" pageNumber="190" unit="m" value="59.0">
<elevation box="[533,592,1761,1786]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="189" pageNumber="190" unit="m" value="59.0">59 m</elevation>
</quantity>
, bottom sand,
<date box="[756,894,1761,1786]" pageId="189" pageNumber="190" value="1966-05-06">
<collectingDate box="[756,894,1761,1786]" pageId="189" pageNumber="190" value="1966-05-06">6 May 1966</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="189.[151,1437,1725,2038]" lastBlockId="190.[151,1436,151,357]" lastPageId="190" lastPageNumber="191" pageId="189" pageNumber="190">
<materialsCitation ID-GBIF-Occurrence="1424726522" collectingDate="1970-09-04" collectionCode="RMNH" country="Guyana" elevation="85" latitude="7.9" location="Guyana" longLatPrecision="5555" longitude="-57.5167" pageId="189" pageNumber="190" specimenCount="1" typeStatus="paratype">
<typeStatus box="[199,308,1798,1822]" pageId="189" pageNumber="190">Paratypes</typeStatus>
<collectionCode box="[319,403,1798,1821]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="189" pageNumber="190">RMNH</collectionCode>
Por. 9738,
<collectingCountry box="[544,634,1798,1822]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
, ‘Luymes’
<collectingCountry box="[769,856,1798,1822]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
Shelf Expedition, station 103,
<geoCoordinate box="[1226,1292,1798,1822]" direction="north" orientation="latitude" pageId="189" pageNumber="190" precision="5555" value="7.9">7.9°N</geoCoordinate>
<geoCoordinate box="[1303,1432,1797,1822]" direction="west" orientation="longitude" pageId="189" pageNumber="190" precision="5" value="-57.5167">57.5167°W</geoCoordinate>
, depth
<quantity box="[221,280,1833,1858]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="189" pageNumber="190" unit="m" value="85.0">
<elevation box="[221,280,1833,1858]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="189" pageNumber="190" unit="m" value="85.0">85 m</elevation>
</quantity>
, triangular dredge,
<collectingDate box="[499,703,1833,1858]" pageId="189" pageNumber="190" value="1970-09-04">
<date box="[499,703,1833,1858]" pageId="189" pageNumber="190" value="1970-09-04">4 September 1970</date>
</collectingDate>
(
<specimenCount box="[722,852,1833,1858]" pageId="189" pageNumber="190" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1424726547" collectingDate="1969-04-24" collectionCode="RMNH" collectorName="Por. &amp; Luymes O. C. P. S." country="Suriname" elevation="81" latitude="7.21" location="Suriname" longLatPrecision="555" longitude="-54.8617" pageId="189" pageNumber="190" specimenCount="1" typeStatus="paratype">
<collectionCode box="[871,955,1834,1857]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="189" pageNumber="190">RMNH</collectionCode>
<collectorName box="[963,1008,1834,1857]" pageId="189" pageNumber="190">Por.</collectorName>
9775,
<collectingCountry box="[1085,1195,1833,1858]" name="Suriname" pageId="189" pageNumber="190">Suriname</collectingCountry>
, ‘
<collectorName box="[1211,1408,1834,1858]" pageId="189" pageNumber="190">Luymes O.C.P.S.</collectorName>
II’
<collectingCountry box="[151,238,1870,1894]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
Shelf Expedition, station I115,
<geoCoordinate box="[621,701,1870,1894]" direction="north" orientation="latitude" pageId="189" pageNumber="190" precision="555" value="7.21">7.21°N</geoCoordinate>
<geoCoordinate box="[714,842,1869,1894]" direction="west" orientation="longitude" pageId="189" pageNumber="190" precision="5" value="-54.8617">54.8617°W</geoCoordinate>
, depth
<quantity box="[934,998,1870,1894]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="189" pageNumber="190" unit="m" value="81.0">
<elevation box="[934,998,1870,1894]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="189" pageNumber="190" unit="m" value="81.0">81 m</elevation>
</quantity>
, triangular dredge,
<date box="[1233,1400,1869,1894]" pageId="189" pageNumber="190" value="1969-04-24">
<collectingDate box="[1233,1400,1869,1894]" pageId="189" pageNumber="190" value="1969-04-24">24 April 1969</collectingDate>
</date>
(
<specimenCount pageId="189" pageNumber="190" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1424726541" collectingDate="1970-08-31" collectionCode="RMNH" country="Guyana" elevation="51" latitude="7.4167" location="Guyana" longLatPrecision="7" longitude="-57.1333" pageId="189" pageNumber="190" specimenCount="1" typeStatus="paratype">
<collectionCode box="[285,369,1906,1929]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="189" pageNumber="190">RMNH</collectionCode>
Por. 9814,
<collectingCountry box="[512,602,1906,1930]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
, ‘Luymes’
<collectingCountry box="[738,825,1906,1930]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
Shelf Expedition, station 68,
<geoCoordinate box="[1183,1292,1905,1930]" direction="north" orientation="latitude" pageId="189" pageNumber="190" precision="5" value="7.4167">7.4167°N</geoCoordinate>
<geoCoordinate box="[1303,1432,1905,1930]" direction="west" orientation="longitude" pageId="189" pageNumber="190" precision="5" value="-57.1333">57.1333°W</geoCoordinate>
, depth
<quantity box="[220,278,1941,1966]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="189" pageNumber="190" unit="m" value="51.0">
<elevation box="[220,278,1941,1966]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="189" pageNumber="190" unit="m" value="51.0">51 m</elevation>
</quantity>
, muddy sand bottom,
<date box="[526,707,1942,1966]" pageId="189" pageNumber="190" value="1970-08-31">
<collectingDate box="[526,707,1942,1966]" pageId="189" pageNumber="190" value="1970-08-31">31 August 1970</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1424726546" collectingDate="1970-09-02" collectionCode="RMNH" collectorName="Por." country="Guyana" elevation="69" latitude="7.65" location="Guyana" longLatPrecision="781" longitude="-57.25" pageId="189" pageNumber="190" specimenCount="1" typeStatus="paratype">
<collectionCode box="[719,803,1942,1965]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="189" pageNumber="190">RMNH</collectionCode>
<collectorName box="[810,854,1942,1965]" pageId="189" pageNumber="190">Por.</collectorName>
9840,
<collectingCountry box="[930,1020,1942,1966]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
, ‘Luymes’
<collectingCountry box="[1147,1234,1942,1966]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
Shelf Expedition, station 85,
<geoCoordinate box="[280,360,1977,2002]" direction="north" orientation="latitude" pageId="189" pageNumber="190" precision="555" value="7.65">7.65°N</geoCoordinate>
<geoCoordinate box="[370,471,1977,2002]" direction="west" orientation="longitude" pageId="189" pageNumber="190" precision="555" value="-57.25">57.25°W</geoCoordinate>
, depth
<quantity box="[557,618,1977,2002]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="189" pageNumber="190" unit="m" value="69.0">
<elevation box="[557,618,1977,2002]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="189" pageNumber="190" unit="m" value="69.0">69 m</elevation>
</quantity>
, bottom sand and shells,
<date box="[914,1123,1977,2002]" pageId="189" pageNumber="190" value="1970-09-02">
<collectingDate box="[914,1123,1977,2002]" pageId="189" pageNumber="190" value="1970-09-02">2 September 1970</collectingDate>
</date>
(
<specimenCount box="[1144,1277,1977,2002]" pageId="189" pageNumber="190" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1424726544" collectingDate="1966-05-10" collectionCode="RMNH" collectorName="Por. &amp; Snellius O. C. P. S. &amp; Agassiz" country="Suriname" elevation="68" lastPageId="190" lastPageNumber="191" latitude="7.26" location="Suriname" longLatPrecision="555" longitude="-56.6667" pageId="189" pageNumber="190" specimenCount="4" typeStatus="paratype">
<collectionCode box="[1298,1382,1978,2001]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="189" pageNumber="190">RMNH</collectionCode>
<collectorName box="[1392,1437,1978,2001]" pageId="189" pageNumber="190">Por.</collectorName>
9879, 9964,
<collectingCountry box="[291,401,2013,2038]" name="Suriname" pageId="189" pageNumber="190">Suriname</collectingCountry>
, ‘
<collectorName box="[416,617,2013,2038]" pageId="189" pageNumber="190">Snellius O.C.P.S.</collectorName>
’
<collectingCountry box="[629,716,2014,2038]" name="Guyana" pageId="189" pageNumber="190">Guyana</collectingCountry>
Shelf Expedition, station G56,
<geoCoordinate box="[1070,1150,2013,2038]" direction="north" orientation="latitude" pageId="189" pageNumber="190" precision="555" value="7.26">7.26°N</geoCoordinate>
<geoCoordinate box="[1157,1286,2013,2038]" direction="west" orientation="longitude" pageId="189" pageNumber="190" precision="5" value="-56.6667">56.6667°W</geoCoordinate>
, depth
<quantity lastPageId="190" lastPageNumber="191" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="6.8" metricValueMin="6.7" pageId="189" pageNumber="190" unit="m" value="67.5" valueMax="68.0" valueMin="67.0">
<elevation lastPageId="190" lastPageNumber="191" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="6.8" metricValueMin="6.7" pageId="189" pageNumber="190" unit="m" value="67.5" valueMax="68.0" valueMin="67.0">67–68 m</elevation>
</quantity>
,
<collectorName box="[191,280,151,176]" pageId="190" pageNumber="191">Agassiz</collectorName>
trawl,
<date box="[368,526,151,176]" pageId="190" pageNumber="191" value="1966-05-10">
<collectingDate box="[368,526,151,176]" pageId="190" pageNumber="191" value="1966-05-10">10 May 1966</collectingDate>
</date>
(
<specimenCount box="[547,695,151,176]" pageId="190" pageNumber="191" type="generic">4 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1424726543" collectingDate="1966-05-11" collectionCode="RMNH" collectorName="Por. &amp; Snellius O. C. P. S." country="Guyana" elevation="68" latitude="7.4233" location="Guyana" longLatPrecision="7" longitude="-56.9067" pageId="190" pageNumber="191" specimenCount="1" typeStatus="paratype">
<collectionCode box="[719,803,152,175]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="190" pageNumber="191">RMNH</collectionCode>
<collectorName box="[815,859,152,175]" pageId="190" pageNumber="191">Por.</collectorName>
9891,
<collectingCountry box="[945,1035,152,176]" name="Guyana" pageId="190" pageNumber="191">Guyana</collectingCountry>
, ‘
<collectorName box="[1055,1261,151,176]" pageId="190" pageNumber="191">Snellius O.C.P.S.</collectorName>
’
<collectingCountry box="[1278,1365,152,176]" name="Guyana" pageId="190" pageNumber="191">Guyana</collectingCountry>
Shelf Expedition, station H58,
<geoCoordinate box="[437,546,188,212]" direction="north" orientation="latitude" pageId="190" pageNumber="191" precision="5" value="7.4233">7.4233°N</geoCoordinate>
<geoCoordinate box="[555,683,188,213]" direction="west" orientation="longitude" pageId="190" pageNumber="191" precision="5" value="-56.9067">56.9067°W</geoCoordinate>
, depth
<quantity box="[768,871,188,213]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="6.9" metricValueMin="6.6" pageId="190" pageNumber="191" unit="m" value="67.5" valueMax="69.0" valueMin="66.0">
<elevation box="[768,871,188,213]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="6.9" metricValueMin="6.6" pageId="190" pageNumber="191" unit="m" value="67.5" valueMax="69.0" valueMin="66.0">66–69 m</elevation>
</quantity>
, bottom coarse sand,
<date box="[1120,1272,188,213]" pageId="190" pageNumber="191" value="1966-05-11">
<collectingDate box="[1120,1272,188,213]" pageId="190" pageNumber="191" value="1966-05-11">11 May 1966</collectingDate>
</date>
(
<specimenCount box="[1292,1424,188,213]" pageId="190" pageNumber="191" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1424726512" collectingDate="1966-05-06" collectionCode="RMNH" collectorName="Por. &amp; Snellius O. C. P. S." country="Suriname" elevation="59" latitude="7.0033" location="Suriname" longLatPrecision="7" longitude="-56.4417" pageId="190" pageNumber="191" specimenCount="23" typeStatus="paratype">
<collectionCode box="[151,235,224,247]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="190" pageNumber="191">RMNH</collectionCode>
<collectorName box="[242,287,224,247]" pageId="190" pageNumber="191">Por.</collectorName>
9899, 9960, 9985, 10503, 10548,
<collectingCountry box="[670,780,223,248]" name="Suriname" pageId="190" pageNumber="191">Suriname</collectingCountry>
, ‘
<collectorName box="[795,996,223,248]" pageId="190" pageNumber="191">Snellius O.C.P.S.</collectorName>
’
<collectingCountry box="[1008,1095,224,248]" name="Guyana" pageId="190" pageNumber="191">Guyana</collectingCountry>
Shelf Expedition, station F40,
<geoCoordinate box="[151,260,260,284]" direction="north" orientation="latitude" pageId="190" pageNumber="191" precision="5" value="7.0033">7.0033°N</geoCoordinate>
<geoCoordinate box="[273,402,260,285]" direction="west" orientation="longitude" pageId="190" pageNumber="191" precision="5" value="-56.4417">56.4417°W</geoCoordinate>
, depth
<quantity box="[496,562,260,285]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="190" pageNumber="191" unit="m" value="59.0">
<elevation box="[496,562,260,285]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="190" pageNumber="191" unit="m" value="59.0">59 m</elevation>
</quantity>
, bottom sand,
<date box="[747,896,260,285]" pageId="190" pageNumber="191" value="1966-05-06">
<collectingDate box="[747,896,260,285]" pageId="190" pageNumber="191" value="1966-05-06">6 May 1966</collectingDate>
</date>
(
<specimenCount box="[918,1083,260,285]" pageId="190" pageNumber="191" type="generic">23 specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="1424726526" collectingDate="1970-08-31" collectionCode="RMNH" collectorName="Por." country="Guyana" elevation="71" latitude="7.5833" location="Guyana" longLatPrecision="7" longitude="-57.0667" pageId="190" pageNumber="191" specimenCount="1" typeStatus="paratype">
<collectionCode box="[1109,1193,261,284]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="190" pageNumber="191">RMNH</collectionCode>
<collectorName box="[1207,1252,261,284]" pageId="190" pageNumber="191">Por.</collectorName>
9932,
<collectingCountry box="[1342,1432,260,284]" name="Guyana" pageId="190" pageNumber="191">Guyana</collectingCountry>
, ‘Luymes’
<collectingCountry box="[267,354,296,320]" name="Guyana" pageId="190" pageNumber="191">Guyana</collectingCountry>
Shelf Expedition, station 63,
<geoCoordinate box="[691,800,295,320]" direction="north" orientation="latitude" pageId="190" pageNumber="191" precision="5" value="7.5833">7.5833°N</geoCoordinate>
<geoCoordinate box="[807,936,295,320]" direction="west" orientation="longitude" pageId="190" pageNumber="191" precision="5" value="-57.0667">57.0667°W</geoCoordinate>
, depth
<quantity box="[1017,1077,296,320]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="190" pageNumber="191" unit="m" value="71.0">
<elevation box="[1017,1077,296,320]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="190" pageNumber="191" unit="m" value="71.0">71 m</elevation>
</quantity>
, sandy bottom,
<date box="[1256,1436,296,320]" pageId="190" pageNumber="191" value="1970-08-31">
<collectingDate box="[1256,1436,296,320]" pageId="190" pageNumber="191" value="1970-08-31">31 August 1970</collectingDate>
</date>
(
<specimenCount box="[162,293,332,357]" pageId="190" pageNumber="191" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="190" pageNumber="191" targetBox="[151,1436,409,1852]" targetPageId="190">
<paragraph blockId="190.[151,1436,1873,1956]" pageId="190" pageNumber="191">
<emphasis bold="true" box="[151,303,1873,1895]" pageId="190" pageNumber="191">FIGURE 120.</emphasis>
<taxonomicName box="[311,551,1873,1895]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="190" pageNumber="191" phylum="Porifera" rank="species" species="crispolobatus" status="sp. nov.">
<emphasis box="[311,551,1873,1895]" italics="true" pageId="190" pageNumber="191">
Suberites
<taxonomicName box="[413,551,1873,1895]" pageId="190" pageNumber="191" rank="species" species="crispolobatus" status="sp. nov.">crispolobatus</taxonomicName>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[560,643,1873,1894]" pageId="190" pageNumber="191" rank="species">
<emphasis bold="true" box="[560,643,1873,1894]" pageId="190" pageNumber="191">sp. nov.</emphasis>
</taxonomicNameLabel>
, a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a1, habitus of paratypes RMNH Por. 10548, a2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b–c, SEM images of the spicules, b, large tylostyle, b1, details of b, c, small tylostyle, c1, details of c.
</paragraph>
</caption>
<subSubSection pageId="191" pageNumber="192" type="description">
<paragraph blockId="191.[151,1437,151,968]" pageId="191" pageNumber="192">
<emphasis bold="true" box="[199,345,151,176]" pageId="191" pageNumber="192">Description.</emphasis>
Erect rounded branches, often with club-shaped endings, and with small side lobes along their length; branches usually divided, but only a few times. The
<typeStatus box="[825,923,188,213]" pageId="191" pageNumber="192">holotype</typeStatus>
<collectionCode box="[930,1014,189,212]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="191" pageNumber="192">RMNH</collectionCode>
Por. 9863 consists of a curled branch (
<figureCitation box="[159,260,223,248]" captionStart="FIGURE 120" captionStartId="190.[151,250,1873,1895]" captionTargetBox="[151,1436,409,1852]" captionTargetId="figure@190.[151,1436,409,1852]" captionTargetPageId="190" captionText="FIGURE 120. Suberites crispolobatus sp. nov., a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a 1, habitus of paratypes RMNH Por. 10548, a 2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b – c, SEM images of the spicules, b, large tylostyle, b 1, details of b, c, small tylostyle, c 1, details of c." httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="191" pageNumber="192">Fig. 120</figureCitation>
a), similar to other such branches of e.g.
<typeStatus box="[739,835,224,248]" pageId="191" pageNumber="192">paratype</typeStatus>
<collectionCode box="[846,930,224,247]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="191" pageNumber="192">RMNH</collectionCode>
Por. 10548 (
<figureCitation box="[1084,1186,223,248]" captionStart="FIGURE 120" captionStartId="190.[151,250,1873,1895]" captionTargetBox="[151,1436,409,1852]" captionTargetId="figure@190.[151,1436,409,1852]" captionTargetPageId="190" captionText="FIGURE 120. Suberites crispolobatus sp. nov., a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a 1, habitus of paratypes RMNH Por. 10548, a 2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b – c, SEM images of the spicules, b, large tylostyle, b 1, details of b, c, small tylostyle, c 1, details of c." httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="191" pageNumber="192">Fig. 120</figureCitation>
a1), but there are also wedge-shaped lobes, e.g.
<typeStatus box="[457,564,261,285]" pageId="191" pageNumber="192">paratypes</typeStatus>
<collectionCode box="[577,661,261,284]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="191" pageNumber="192">RMNH</collectionCode>
Por. 9985 (
<figureCitation box="[809,912,260,285]" captionStart="FIGURE 120" captionStartId="190.[151,250,1873,1895]" captionTargetBox="[151,1436,409,1852]" captionTargetId="figure@190.[151,1436,409,1852]" captionTargetPageId="190" captionText="FIGURE 120. Suberites crispolobatus sp. nov., a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a 1, habitus of paratypes RMNH Por. 10548, a 2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b – c, SEM images of the spicules, b, large tylostyle, b 1, details of b, c, small tylostyle, c 1, details of c." httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="191" pageNumber="192">Fig. 120</figureCitation>
a2). Surface smooth but with irregular folds (possibly deflated as an after-collection artifact, as is often the case in larger
<taxonomicName box="[1002,1106,295,320]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="191" pageNumber="192" phylum="Porifera" rank="genus">
<emphasis box="[1002,1106,295,320]" italics="true" pageId="191" pageNumber="192">Suberites</emphasis>
</taxonomicName>
species. There does not seem to be a massive basal body, individual branches have a pointed or thinner stalk-like basis. Sizes quite variable, up to
<quantity box="[151,220,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="191" pageNumber="192" unit="cm" value="25.0">25 cm</quantity>
long, up to
<quantity box="[357,441,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="191" pageNumber="192" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
in diameter.
<collectingRegion box="[589,623,368,391]" country="Haiti" name="Nord" pageId="191" pageNumber="192">No</collectingRegion>
apparent oscules but these are likely contracted. Color beige in alcohol. Consistency firm.
</paragraph>
<paragraph blockId="191.[151,1437,151,968]" pageId="191" pageNumber="192">
<emphasis bold="true" box="[199,309,439,464]" pageId="191" pageNumber="192">Skeleton.</emphasis>
Typically
<taxonomicName box="[427,531,439,464]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="191" pageNumber="192" phylum="Porifera" rank="genus">
<emphasis box="[427,531,439,464]" italics="true" pageId="191" pageNumber="192">Suberites</emphasis>
</taxonomicName>
-like, with bouquets of smaller tylostyles at the surface and more or less confused choanosomal skeleton. The center of the longer branches is more organized with tylostyles aligned lengthwise.
</paragraph>
<paragraph blockId="191.[151,1437,151,968]" box="[199,652,511,536]" pageId="191" pageNumber="192">
<emphasis bold="true" box="[199,304,511,536]" pageId="191" pageNumber="192">Spicules.</emphasis>
(
<figureCitation box="[319,417,511,536]" captionStart="FIGURE 120" captionStartId="190.[151,250,1873,1895]" captionTargetBox="[151,1436,409,1852]" captionTargetId="figure@190.[151,1436,409,1852]" captionTargetPageId="190" captionText="FIGURE 120. Suberites crispolobatus sp. nov., a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a 1, habitus of paratypes RMNH Por. 10548, a 2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b – c, SEM images of the spicules, b, large tylostyle, b 1, details of b, c, small tylostyle, c 1, details of c." httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="191" pageNumber="192">Figs 120</figureCitation>
b–c) Tylostyles only.
</paragraph>
<paragraph blockId="191.[151,1437,151,968]" pageId="191" pageNumber="192">
Tylostyles with prominent rounded heads, without obvious lobes except for thin growth stages which may have tuberculate heads; very variable in size, divisible into two size classes with some overlap, (1) larger (
<figureCitation box="[1272,1376,583,608]" captionStart="FIGURE 120" captionStartId="190.[151,250,1873,1895]" captionTargetBox="[151,1436,409,1852]" captionTargetId="figure@190.[151,1436,409,1852]" captionTargetPageId="190" captionText="FIGURE 120. Suberites crispolobatus sp. nov., a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a 1, habitus of paratypes RMNH Por. 10548, a 2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b – c, SEM images of the spicules, b, large tylostyle, b 1, details of b, c, small tylostyle, c 1, details of c." httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="191" pageNumber="192">Figs 120</figureCitation>
b,b1), 456–
<emphasis box="[207,249,620,644]" italics="true" pageId="191" pageNumber="192">724</emphasis>
–858
<date box="[312,346,620,644]" pageId="191" pageNumber="192">x 9</date>
–
<emphasis box="[361,406,620,644]" italics="true" pageId="191" pageNumber="192">11.4</emphasis>
–14 µm, and (2) smaller (
<figureCitation box="[693,798,620,645]" captionStart="FIGURE 120" captionStartId="190.[151,250,1873,1895]" captionTargetBox="[151,1436,409,1852]" captionTargetId="figure@190.[151,1436,409,1852]" captionTargetPageId="190" captionText="FIGURE 120. Suberites crispolobatus sp. nov., a, habitus of holotype RMNH Por. 9863 (scale bar = 1 cm), a 1, habitus of paratypes RMNH Por. 10548, a 2, habitus of two wedge-shaped paratypes RMNH Por. 9985 (scale bar = 1 cm), b – c, SEM images of the spicules, b, large tylostyle, b 1, details of b, c, small tylostyle, c 1, details of c." httpUri="https://zenodo.org/record/273070/files/figure.png" pageId="191" pageNumber="192">Figs 120</figureCitation>
c,c1), 186–
<emphasis box="[917,959,620,645]" italics="true" pageId="191" pageNumber="192">266</emphasis>
–396
<date box="[1022,1055,620,644]" pageId="191" pageNumber="192">x 3</date>
–
<emphasis box="[1071,1106,620,644]" italics="true" pageId="191" pageNumber="192">5.9</emphasis>
–10 µm.
</paragraph>
</subSubSection>
<subSubSection box="[199,1119,655,680]" pageId="191" pageNumber="192" type="distribution">
<paragraph blockId="191.[151,1437,151,968]" box="[199,1119,655,680]" pageId="191" pageNumber="192">
<emphasis bold="true" box="[199,498,655,680]" pageId="191" pageNumber="192">Distribution and ecology.</emphasis>
<collectingCountry box="[505,592,656,680]" name="Guyana" pageId="191" pageNumber="192">Guyana</collectingCountry>
Shelf, muddy sandy bottom at
<quantity box="[943,1042,655,680]" metricMagnitude="1" metricUnit="m" metricValue="6.8" metricValueMax="8.5" metricValueMin="5.1" pageId="191" pageNumber="192" unit="m" value="68.0" valueMax="85.0" valueMin="51.0">51–85 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection pageId="191" pageNumber="192" type="etymology">
<paragraph blockId="191.[151,1437,151,968]" pageId="191" pageNumber="192">
<emphasis bold="true" box="[199,334,692,717]" pageId="191" pageNumber="192">Etymology.</emphasis>
Crispus (L.) = curly,
<taxonomicName box="[574,655,692,717]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="191" pageNumber="192" phylum="Porifera" rank="species" species="lobatus">lobatus</taxonomicName>
(L.) = having lobes, referring to the curled branches with lobes found in this species, at least in the preserved specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="191" pageNumber="192" type="discussion">
<paragraph blockId="191.[151,1437,151,968]" pageId="191" pageNumber="192">
<emphasis bold="true" box="[199,315,764,789]" pageId="191" pageNumber="192">Remarks.</emphasis>
At first I assumed these specimens could be excessively ramose
<taxonomicName authority="Duchassaing &amp; Michelotti, 1864" authorityName="Duchassaing &amp; Michelotti" authorityYear="1864" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="191" pageNumber="192" phylum="Porifera" rank="species" species="aurantiacus">
<emphasis box="[1179,1436,764,789]" italics="true" pageId="191" pageNumber="192">Suberites aurantiacus</emphasis>
(Duchassaing &amp; Michelotti, 1864)
</taxonomicName>
(p. 99, as
<taxonomicName box="[652,733,801,824]" class="Demospongiae" family="Suberitidae" genus="Terpios" kingdom="Animalia" order="Hadromerida" pageId="191" pageNumber="192" phylum="Porifera" rank="genus">
<emphasis box="[652,733,801,824]" italics="true" pageId="191" pageNumber="192">Terpios</emphasis>
</taxonomicName>
) or a similar species, like
<taxonomicName authority="Wilson, 1902" authorityName="Wilson" authorityYear="1902" box="[1029,1399,799,824]" class="Demospongiae" family="Suberitidae" genus="Suberites" kingdom="Animalia" order="Hadromerida" pageId="191" pageNumber="192" phylum="Porifera" rank="species" species="lobatus">
<emphasis box="[1029,1223,799,824]" italics="true" pageId="191" pageNumber="192">Suberites lobatus</emphasis>
(
<bibRefCitation author="Wilson" box="[1238,1390,799,824]" pageId="191" pageNumber="192" refString="Wilson, H. V. (1902) [1900] The sponges collected in Porto Rico in 1899 by the U. S. Fish Commission Steamer Fish Hawk. Bulletin of the United States Fish Commission, 2, 375 - 411." type="journal article" year="1902">Wilson, 1902</bibRefCitation>
)
</taxonomicName>
(p. 399, as
<taxonomicName box="[238,424,836,861]" class="Demospongiae" family="Axinellidae" genus="Phakellia" kingdom="Animalia" order="Halichondrida" pageId="191" pageNumber="192" phylum="Porifera" rank="species" species="lobata">
<emphasis box="[238,424,836,861]" italics="true" pageId="191" pageNumber="192">Phakellia lobata</emphasis>
</taxonomicName>
, slides of the
<typeStatus box="[582,680,836,861]" pageId="191" pageNumber="192">holotype</typeStatus>
<collectionCode box="[687,768,836,860]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="191" pageNumber="192">USNM</collectionCode>
7684 examined), because spicule sizes are more or less the same of these two species. However, Dr Klaus Rützler (
<emphasis box="[792,896,871,896]" italics="true" pageId="191" pageNumber="192">in litteris</emphasis>
) convinced me that these have a massive base, and do not form such excessive long branches (see also Rützler &amp;
<collectingRegion box="[945,1012,908,933]" country="Bermuda" name="Smith's" pageId="191" pageNumber="192">Smith</collectingRegion>
1993). Possibly, some deeper water records of these two species may conform to the present new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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