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treatments-xml/data/03/A5/87/03A587E7A873FFF60CBA16E8FC4A93A2.xml
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<document id="3FFA97044FAB17C7CADD7989A03A9B10" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A873FFF60CBA16E8FC4A93A2" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus formosus Perkins 2019, new species" docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="17" updateTime="1698775642300" updateUser="ExternalLinkService">
<mods:mods id="25898C0AC79EEF0F5635917EF8ABE18B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F29CFFE98B58C834F7A6096B13FDA38E">
<mods:title id="8D279AD0A5400198876B5997ABC3EEE3">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<mods:namePart id="3567CA27EC742335FB8C8FA633904609">Perkins, Philip D.</mods:namePart>
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<mods:title id="A6E78B37B472AF0F701FD588CB9360C1">Zootaxa</mods:title>
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<mods:part id="2EBCD50278C15F29B4DB2824D5450D8F">
<mods:date id="ED0222B481078C2A1076C0600F7FE429">2019</mods:date>
<mods:detail id="D3D4CF4497E207D8AD8838D67A841C54" type="pubDate">
<mods:number id="B9F0B7AC2166DF8063936628B2499015">2019-12-12</mods:number>
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<mods:classification id="E17DE7556BE081BC0FA01EABB923047E">journal article</mods:classification>
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<treatment id="03A587E7A873FFF60CBA16E8FC4A93A2" ID-DOI="http://doi.org/10.5281/zenodo.5941789" ID-GBIF-Taxon="160798404" ID-Zenodo-Dep="5941789" LSID="urn:lsid:plazi:treatment:03A587E7A873FFF60CBA16E8FC4A93A2" httpUri="http://treatment.plazi.org/id/03A587E7A873FFF60CBA16E8FC4A93A2" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
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<emphasis id="B978EAE3A873FFF40CBA16E8FDCD9693" bold="true" box="[151,556,1196,1223]" pageId="16" pageNumber="17">
<taxonomicName id="4C0C4D72A873FFF40CBA16E8FE759693" ID-CoL="8Y4Q7" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[151,404,1196,1223]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="formosus" status="sp. nov.">
<emphasis id="B978EAE3A873FFF40CBA16E8FE759693" bold="true" box="[151,404,1196,1223]" italics="true" pageId="16" pageNumber="17">Hydrochus formosus</emphasis>
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<figureCitation id="13372A74A873FFF40CBA1694FF0496BE" box="[151,229,1232,1259]" captionStart="FIGURE 2" captionStartId="14.[151,250,2009,2034]" captionTargetBox="[199,1400,435,1940]" captionTargetId="figure@14.[151,1435,417,1984]" captionTargetPageId="14" captionText="FIGURE 2. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3587185" httpUri="https://zenodo.org/record/3587185/files/figure.png" pageId="16" pageNumber="17">Figs. 2</figureCitation>
,
<figureCitation id="13372A74A873FFF40CDC1695FEEC96BE" box="[241,269,1233,1259]" captionStart="FIGURE 12" captionStartId="28.[151,250,1701,1726]" captionTargetBox="[171,1431,215,1649]" captionTargetId="figure@28.[151,1435,181,1676]" captionTargetPageId="28" captionText="FIGURE 12. Hydrochus formosus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587205" httpUri="https://zenodo.org/record/3587205/files/figure.png" pageId="16" pageNumber="17">12</figureCitation>
</paragraph>
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<emphasis id="B978EAE3A873FFF40CBA175DFEA89766" bold="true" box="[151,329,1304,1331]" pageId="16" pageNumber="17">Type Material.</emphasis>
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<typeStatus id="54B78853A873FFF40D7C175DFE5F9766" box="[337,446,1305,1331]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
(male):
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“
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<collectingCountry id="F31B7661A873FFF40E1C175DFD489766" box="[561,681,1305,1331]" name="Venezuela" pageId="16" pageNumber="17">Venezuela</collectingCountry>
</emphasis>
:
<collectingRegion id="49C8F813A873FFF40E94175CFCB69766" box="[697,855,1304,1331]" country="Venezuela" name="Guarico" pageId="16" pageNumber="17">Guarico State</collectingRegion>
:
<geoCoordinate id="EE385036A873FFF40F4E175DFC0B9766" box="[867,1002,1305,1331]" degrees="8" direction="north" minutes="8.713" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="8.145217">8° 8.713’ N</geoCoordinate>
,
<geoCoordinate id="EE385036A873FFF40FDB175CFB7C9766" box="[1014,1181,1304,1331]" degrees="66" direction="west" minutes="20.598" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="-66.3433">66° 20.598’ W</geoCoordinate>
,
<quantity id="4CF49B14A873FFF40884175DFB3C9766" box="[1193,1245,1305,1331]" metricMagnitude="1" metricUnit="m" metricValue="7.4" pageId="16" pageNumber="17" unit="m" value="74.0">
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,
<collectingMunicipality id="6BD7AC8BA873FFF408C7175CFAD39767" box="[1258,1330,1304,1330]" pageId="16" pageNumber="17">Marsh</collectingMunicipality>
<locationDeviation id="9A012FD3A873FFF40914175DFEEC9703" location="San Nicolasito" pageId="16" pageNumber="17">E of San Nicolasito</locationDeviation>
,
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<collectingDate id="EFF6E9D9A873FFF40D311778FE669703" box="[284,391,1340,1366]" pageId="16" pageNumber="17" value="2009-01-11">11.i.2009</collectingDate>
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,
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; VZ09-0111-01G2” (
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</materialsCitation>
.
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<typeStatus id="54B78853A873FFF40F951779FBD19702" box="[952,1072,1341,1367]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
(102):
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:
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:
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AS-08-081A (1
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;
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:
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AS-06-019 (15
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:
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VZ09-0112-01A (4
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C (1
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<materialsCitation id="3B643CACA873FFF40CF417EDFE689796" ID-GBIF-Occurrence="2521263792" box="[217,393,1449,1475]" collectionCode="SEMC" country="Venezuela" pageId="16" pageNumber="17" specimenCode="02" specimenCount="1" stateProvince="Bolivar" typeStatus="paratype">
<specimenCode id="DBAA9E8AA873FFF40CF417EDFF149796" box="[217,245,1449,1475]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">02</specimenCode>
B (1
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)
</materialsCitation>
;
<materialsCitation id="3B643CACA873FFF40DBB17EDFD349796" ID-GBIF-Occurrence="2521263779" box="[406,725,1449,1475]" collectingDate="2001-10" collectionCode="SEMC" country="Venezuela" pageId="16" pageNumber="17" specimenCode="VZ09-0113" specimenCount="1" stateProvince="Bolivar" typeStatus="paratype">
<specimenCode id="DBAA9E8AA873FFF40DBB17EDFDFA9796" box="[406,539,1449,1475]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">VZ09-0113</specimenCode>
-
<date id="FFB21031A873FFF40E0F17EDFDB39796" box="[546,594,1449,1475]" pageId="16" pageNumber="17" value="2001-10">
<collectingDate id="EFF6E9D9A873FFF40E0F17EDFDB39796" box="[546,594,1449,1475]" pageId="16" pageNumber="17" value="2001-10">01X</collectingDate>
</date>
(3
<collectionCode id="ED1DAE34A873FFF40E5117EDFD2C9796" box="[636,717,1449,1475]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA873FFF40ECF17EDFC0097B2" ID-GBIF-Occurrence="2521263885" collectingDate="1989-01-25" collectionCode="NMNH" country="Venezuela" location="Rio Horero" municipality="Puerto Ayacucho" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Bolivar" typeStatus="paratype">
<collectingMunicipality id="6BD7AC8BA873FFF40ECF17EDFC439797" box="[738,930,1448,1475]" pageId="16" pageNumber="17">Puerto Ayacucho</collectingMunicipality>
(66kmN),
<location id="8ED3602AA873FFF4080E17ECFB469796" LSID="urn:lsid:plazi:treatment:03A587E7A873FFF60CBA16E8FC4A93A2:8ED3602AA873FFF4080E17ECFB469796" box="[1059,1191,1448,1475]" country="Venezuela" municipality="Puerto Ayacucho" name="Rio Horero" pageId="16" pageNumber="17" stateProvince="Bolivar">Rio Horero</location>
,
<date id="FFB21031A873FFF4089917ECFAA29796" box="[1204,1347,1448,1475]" pageId="16" pageNumber="17" value="1989-01-25">
<collectingDate id="EFF6E9D9A873FFF4089917ECFAA29796" box="[1204,1347,1448,1475]" pageId="16" pageNumber="17" value="1989-01-25">25 Jan 1989</collectingDate>
</date>
, netted, stream edge, Collected by PJSpangler, RAFaitoute &amp; CBBarr (12
<collectionCode id="ED1DAE34A873FFF40F531789FC3897B2" box="[894,985,1485,1511]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B643CACA873FFF40FC61788FA7997B2" ID-GBIF-Occurrence="2521263794" box="[1003,1432,1484,1511]" collectingDate="2001-10" collectionCode="SEMC" country="Venezuela" location="Guarico" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
<emphasis id="B978EAE3A873FFF40FC61788FBB797B2" bold="true" box="[1003,1110,1484,1511]" pageId="16" pageNumber="17">
<collectingRegion id="49C8F813A873FFF40FC61788FBB097B2" box="[1003,1105,1484,1511]" country="Venezuela" name="Guarico" pageId="16" pageNumber="17">Guarico</collectingRegion>
:
</emphasis>
VZ09-0109-
<date id="FFB21031A873FFF408C71789FAF897B2" box="[1258,1305,1485,1511]" pageId="16" pageNumber="17" value="2001-10">
<collectingDate id="EFF6E9D9A873FFF408C71789FAF897B2" box="[1258,1305,1485,1511]" pageId="16" pageNumber="17" value="2001-10">01X</collectingDate>
</date>
(1
<collectionCode id="ED1DAE34A873FFF409121789FA7197B2" box="[1343,1424,1485,1511]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA873FFF40CBA17B5FEFA945E" ID-GBIF-Occurrence="2521263784" box="[151,283,1521,1547]" collectionCode="VZ" country="Venezuela" pageId="16" pageNumber="17" specimenCode="VZ09-0110" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
<specimenCode id="DBAA9E8AA873FFF40CBA17B5FEFA945E" box="[151,283,1521,1547]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">VZ09-0110</specimenCode>
</materialsCitation>
-
<materialsCitation id="3B643CACA873FFF40D0F17B5FD4B945E" ID-GBIF-Occurrence="2521263837" box="[290,682,1521,1547]" collectionCode="SEMC, MIZA, MALUZ" country="Venezuela" pageId="16" pageNumber="17" specimenCode="01" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
<specimenCode id="DBAA9E8AA873FFF40D0F17B5FEDD945E" box="[290,316,1521,1547]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">01</specimenCode>
A (21
<collectionCode id="ED1DAE34A873FFF40DAA17B5FE37945E" box="[391,470,1521,1547]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">SEMC</collectionCode>
,
<collectionCode id="ED1DAE34A873FFF40DCE17B5FDCC945E" box="[483,557,1521,1547]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="16" pageNumber="17" type="Museum">MIZA</collectionCode>
,
<collectionCode id="ED1DAE34A873FFF40E1717B5FD43945E" box="[570,674,1521,1547]" pageId="16" pageNumber="17">MALUZ</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B643CACA873FFF40E9817B5FBE1945E" ID-GBIF-Occurrence="2521263790" box="[693,1024,1521,1547]" collectingDate="2002-10" collectionCode="SEMC" country="Venezuela" pageId="16" pageNumber="17" specimenCode="VZ09-0110" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
<specimenCode id="DBAA9E8AA873FFF40E9817B5FCD8945E" box="[693,825,1521,1547]" collectionCode="MALUZ" pageId="16" pageNumber="17">VZ09-0110</specimenCode>
-
<date id="FFB21031A873FFF40F6D17B5FC90945E" box="[832,881,1521,1547]" pageId="16" pageNumber="17" value="2002-10">
<collectingDate id="EFF6E9D9A873FFF40F6D17B5FC90945E" box="[832,881,1521,1547]" pageId="16" pageNumber="17" value="2002-10">02X</collectingDate>
</date>
(17
<collectionCode id="ED1DAE34A873FFF40F8A17B5FC19945E" box="[935,1016,1521,1547]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA873FFF4082117B5FAA6945E" ID-GBIF-Occurrence="2521263778" box="[1036,1351,1521,1547]" collectingDate="2002-10" collectionCode="SEMC" country="Venezuela" pageId="16" pageNumber="17" specimenCode="VZ09-0111" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
<specimenCode id="DBAA9E8AA873FFF4082117B5FB6F945E" box="[1036,1166,1521,1547]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">VZ09-0111</specimenCode>
-
<date id="FFB21031A873FFF408B817B5FB27945E" box="[1173,1222,1521,1547]" pageId="16" pageNumber="17" value="2002-10">
<collectingDate id="EFF6E9D9A873FFF408B817B5FB27945E" box="[1173,1222,1521,1547]" pageId="16" pageNumber="17" value="2002-10">02X</collectingDate>
</date>
(1
<collectionCode id="ED1DAE34A873FFF408C317B5FADE945E" box="[1262,1343,1521,1547]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="17" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B643CACA873FFF4097917B5FCEE947A" ID-GBIF-Occurrence="2521263775" collectingDate="1969-02-11" collectionCode="NMNH" collectorName="P. &amp; P. Spangler" country="Venezuela" location="Calabozo" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
<quantity id="4CF49B14A873FFF4097917B5FA7C945F" box="[1364,1437,1520,1547]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="17" unit="km" value="32.0">32 km</quantity>
SW.
<location id="8ED3602AA873FFF40CE01450FED6947B" LSID="urn:lsid:plazi:treatment:03A587E7A873FFF60CBA16E8FC4A93A2:8ED3602AA873FFF40CE01450FED6947B" box="[205,311,1556,1582]" country="Venezuela" name="Calabozo" pageId="16" pageNumber="17" stateProvince="Guarico">Calabozo</location>
<date id="FFB21031A873FFF40D101450FE59947B" box="[317,440,1556,1582]" pageId="16" pageNumber="17" value="1969-02-11">
<collectingDate id="EFF6E9D9A873FFF40D101450FE59947B" box="[317,440,1556,1582]" pageId="16" pageNumber="17" value="1969-02-11">II-11-1969</collectingDate>
</date>
,
<collectorName id="26F95327A873FFF40DE91451FD98947B" box="[452,633,1556,1583]" pageId="16" pageNumber="17">P. &amp; P. Spangler</collectorName>
(18
<collectionCode id="ED1DAE34A873FFF40E811451FCE6947A" box="[684,775,1557,1583]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB336F1A873FFF40CEA147CFEF39552" blockId="16.[151,1437,1304,2014]" pageId="16" pageNumber="17">
<emphasis id="B978EAE3A873FFF40CEA147CFE359406" bold="true" box="[199,468,1592,1619]" pageId="16" pageNumber="17">Differential Diagnosis.</emphasis>
Among Venezuelan species, recognized by the combination of moderately large size (ca.
<quantity id="4CF49B14A873FFF40CEA1418FE8F9422" box="[199,366,1628,1655]" metricMagnitude="-3" metricUnit="m" metricValue="3.615" metricValueMax="4.08" metricValueMin="3.15" pageId="16" pageNumber="17" unit="mm" value="3.615" valueMax="4.08" valueMin="3.15">3.15–4.08 mm</quantity>
), the shallow pronotal depressions, the comparatively long elytra, the lack of an elevation of the 5
<superScript id="7C799BB9A873FFF40C88143BFF5394D8" attach="left" box="[165,178,1663,1677]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
elytral interstria, and the male genitalia (
<figureCitation id="13372A74A873FFF40EA014C4FD0394CE" box="[653,738,1664,1691]" captionStart="FIGURE 12" captionStartId="28.[151,250,1701,1726]" captionTargetBox="[171,1431,215,1649]" captionTargetId="figure@28.[151,1435,181,1676]" captionTargetPageId="28" captionText="FIGURE 12. Hydrochus formosus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587205" httpUri="https://zenodo.org/record/3587205/files/figure.png" pageId="16" pageNumber="17">Fig. 12</figureCitation>
; described below). Additionally, males are recognized by the sharply pointed tubercles, one each on the midline of abdominal ventrites three and four; this appears to be a unique sexual dimorphism in Neotropical
<taxonomicName id="4C0C4D72A873FFF40E32148CFD7894B7" authorityName="Leach" authorityYear="1817" box="[543,665,1736,1762]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAE3A873FFF40E32148CFD7894B7" box="[543,665,1736,1762]" italics="true" pageId="16" pageNumber="17">Hydrochus</emphasis>
</taxonomicName>
. The male genitalia are unique in the pattern of sclerotization of the parameres.
</paragraph>
</subSubSection>
<subSubSection id="C316657AA873FFF60CEA1554FF159386" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="8BB336F1A873FFF40CEA1554FF5E958B" blockId="16.[151,1437,1304,2014]" pageId="16" pageNumber="17">
<emphasis id="B978EAE3A873FFF40CEA1554FEB0957E" bold="true" box="[199,337,1808,1835]" pageId="16" pageNumber="17">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="13372A74A873FFF40E761554FD4C957E" box="[603,685,1808,1835]" captionStart="FIGURE 12" captionStartId="28.[151,250,1701,1726]" captionTargetBox="[171,1431,215,1649]" captionTargetId="figure@28.[151,1435,181,1676]" captionTargetPageId="28" captionText="FIGURE 12. Hydrochus formosus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587205" httpUri="https://zenodo.org/record/3587205/files/figure.png" pageId="16" pageNumber="17">Fig. 12</figureCitation>
) Size:
<typeStatus id="54B78853A873FFF40ED61554FCBC957F" box="[763,861,1808,1834]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 4.04 /1.38; head width 0.96; pronotum l/w 0.85/0.92; PA 0.88; PB 0.85; elytra 2.65/1.38. Body size moderately large (ca.
<quantity id="4CF49B14A873FFF40D2B151CFE4C9526" box="[262,429,1880,1907]" metricMagnitude="-3" metricUnit="m" metricValue="3.615" metricValueMax="4.08" metricValueMin="3.15" pageId="16" pageNumber="17" unit="mm" value="3.615" valueMax="4.08" valueMin="3.15">3.15–4.08 mm</quantity>
). Dorsum grey, with very slight iridescence, elytra with black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 1.5–
<date id="FFB21031A873FFF40F2F1538FCFF95C2" box="[770,798,1916,1943]" pageId="16" pageNumber="17">2x</date>
that of pronotum. Elytra interstriae ca.
<date id="FFB21031A873FFF408F51539FB1595C2" box="[1240,1268,1917,1943]" pageId="16" pageNumber="17">2x</date>
strial puncture diameter. Interstria 9
<superScript id="7C799BB9A873FFF40DAF15DBFE6E95F8" attach="left" box="[386,399,1951,1965]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
more convex than others, overhanging 10
<superScript id="7C799BB9A873FFF40F4315DBFC9A95F8" attach="left" box="[878,891,1951,1965]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
interstria. Interstria 5
<superScript id="7C799BB9A873FFF4085C15DBFB9F95F8" attach="left" box="[1137,1150,1951,1965]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
without elevation or callus.
</paragraph>
<caption id="DF736679A872FFF50CBA1491FDC89547" ID-DOI="http://doi.org/10.5281/zenodo.3587189" ID-Zenodo-Dep="3587189" httpUri="https://zenodo.org/record/3587189/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,1749,1774]" targetBox="[227,1390,212,1676]" targetPageId="17">
<paragraph id="8BB336F1A872FFF50CBA1491FDC89547" blockId="17.[151,1436,1749,1810]" pageId="17" pageNumber="18">
<emphasis id="B978EAE3A872FFF50CBA1491FEB194BB" bold="true" box="[151,336,1749,1774]" pageId="17" pageNumber="18">FIGURES 4A–B.</emphasis>
Male genitalia characters of (A)
<taxonomicName id="4C0C4D72A872FFF50EB51493FC6C94BB" authority="Oliva" authorityName="Oliva" authorityYear="1996" box="[664,909,1749,1774]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pseudosecretus">
<emphasis id="B978EAE3A872FFF50EB51493FCB194BB" box="[664,848,1750,1774]" italics="true" pageId="17" pageNumber="18">H. pseudosecretus</emphasis>
Oliva
</taxonomicName>
(ventral); (B)
<taxonomicName id="4C0C4D72A872FFF508301493FB4794BB" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[1053,1190,1751,1774]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sagittarius" status="sp. nov.">
<emphasis id="B978EAE3A872FFF508301493FB4794BB" box="[1053,1190,1751,1774]" italics="true" pageId="17" pageNumber="18">H. sagittarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5798A872FFF508811493FB0094BB" box="[1196,1249,1751,1774]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
(ventral; diagrammatical, showing locations of muscles).
</paragraph>
</caption>
<paragraph id="8BB336F1A872FFF50CEA1505FE3295F7" blockId="17.[151,1436,1856,2027]" pageId="17" pageNumber="18">Pronotum slightly wider than long, widest at or slightly behind anterior margin, narrowed at base, sides sinuate, smooth; anterior margin slightly bisinuate; depressions shallow, only very slightly more densely punctate than low reliefs between depressions.</paragraph>
<paragraph id="8BB336F1A872FFF50CEA15E8FC5595BF" blockId="17.[151,1436,1856,2027]" pageId="17" pageNumber="18">
Elytra with spaces between strial punctures ca. 0.5–
<date id="FFB21031A872FFF50F2215E8FCC69592" box="[783,807,1964,1991]" pageId="17" pageNumber="18">1x</date>
their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with very slight angulation.
</paragraph>
<paragraph id="8BB336F1A871FFF60CEA12DDFACD92E6" blockId="18.[151,1436,153,503]" box="[199,1324,153,179]" pageId="18" pageNumber="19">Males with sharply pointed tubercles, one each on the midline of abdominal ventrites three and four.</paragraph>
<paragraph id="8BB336F1A871FFF60CBA12F9FB2E9282" blockId="18.[151,1436,153,503]" box="[151,1231,189,215]" pageId="18" pageNumber="19">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8BB336F1A871FFF60CEA12A5FB04934A" blockId="18.[151,1436,153,503]" pageId="18" pageNumber="19">
Male genitalia (
<figureCitation id="13372A74A871FFF60D5412A5FE2A92AE" box="[377,459,225,251]" captionStart="FIGURE 12" captionStartId="28.[151,250,1701,1726]" captionTargetBox="[171,1431,215,1649]" captionTargetId="figure@28.[151,1435,181,1676]" captionTargetPageId="28" captionText="FIGURE 12. Hydrochus formosus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587205" httpUri="https://zenodo.org/record/3587205/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
) general characters: paramere tips in dorsal and ventral views arcuate toward midline; in lateral view narrow and tapering, very lightly sclerotized; aedeagus basically lanceolate shaped.
</paragraph>
<paragraph id="8BB336F1A871FFF60CBA136DFB7D9316" blockId="18.[151,1436,153,503]" box="[151,1180,297,323]" pageId="18" pageNumber="19">Dorsal surface: pdmm slightly sinuate and at points slightly separated from alm; adbl small.</paragraph>
<paragraph id="8BB336F1A871FFF60CEA1309FF159386" blockId="18.[151,1436,153,503]" pageId="18" pageNumber="19">Ventral surface: each paramere with distinctive sclerotization pattern in distal ½; basal membranous area large; pseudoparameres extending almost to midlength of parameres, rather large in lateral view; avtl distinctive, subapical; basal piece distal membranous area large, with distinctively shaped sclerotization line, produced to point in midline.</paragraph>
</subSubSection>
<subSubSection id="C316657AA871FFF60CEA1399FC4A93A2" box="[199,939,477,503]" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BB336F1A871FFF60CEA1399FC4A93A2" blockId="18.[151,1436,153,503]" box="[199,939,477,503]" pageId="18" pageNumber="19">
<emphasis id="B978EAE3A871FFF60CEA1399FEAF93A2" bold="true" box="[199,334,477,503]" pageId="18" pageNumber="19">Etymology.</emphasis>
Named in reference to the pleasing body proportions.
</paragraph>
</subSubSection>
</treatment>
</document>
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