treatments-xml/data/03/A5/87/03A587BAFFAAFF95A2B2FD50C39CFDF3.xml

<document id="C8E3C6183EBCD0E7FA6E0DCC2D95FCB7" ID-CLB-Dataset="46723" ID-DOI="10.11646/zootaxa.3753.4.5" ID-GBIF-Dataset="bde64ad3-2af9-4db0-943a-707831f15913" ID-ISSN="1175-5326" ID-Zenodo-Dep="228608" ID-ZooBank="4556045E-C319-4549-995A-814011D9B684" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460365496199" checkinUser="plazi" docAuthor="Maghsoudlou, Abdolvahab &amp; Momtazi, Farzaneh" docDate="2014" docId="03A587BAFFAAFF95A2B2FD50C39CFDF3" docLanguage="en" docName="zt03753p374.pdf" docOrigin="Zootaxa 3753 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Imogine qeshmensis Maghsoudlou &amp; Momtazi, 2014, sp. nov." docType="treatment" docUuid="8E8539C5-CC88-4929-89BA-9B746A49B3F1" docUuidSource="ZooBank" docVersion="10" lastPageNumber="373" masterDocId="FF9CFFC2FFA8FF9CA225FFACC028FFEB" masterDocTitle="A new Imogine species (Turbellaria: Polycladida: Stylochidae) associated with rock oysters (Saccostrea cucullata) from the Persian Gulf, with a review of the genus" masterLastPageNumber="374" masterPageNumber="364" pageNumber="366" updateTime="1698295177128" updateUser="ExternalLinkService">
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<mods:title id="4B22A9BB696AEE42A67AF6846BE724C6">A new Imogine species (Turbellaria: Polycladida: Stylochidae) associated with rock oysters (Saccostrea cucullata) from the Persian Gulf, with a review of the genus</mods:title>
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<mods:namePart id="C04273ED705389F519BD59688F067AA5">Maghsoudlou, Abdolvahab</mods:namePart>
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<mods:date id="2C10DE7AD56EC858AD3CCAF099152E0D">2014</mods:date>
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<treatment id="03A587BAFFAAFF95A2B2FD50C39CFDF3" ID-DOI="http://doi.org/10.5281/zenodo.6129855" ID-GBIF-Taxon="119410730" ID-Zenodo-Dep="6129855" LSID="urn:lsid:zoobank.org:act:8E8539C5-CC88-4929-89BA-9B746A49B3F1" httpUri="http://treatment.plazi.org/id/03A587BAFFAAFF95A2B2FD50C39CFDF3" lastPageId="9" lastPageNumber="373" pageId="2" pageNumber="366">
<subSubSection id="C3166527FFAAFF9EA2B2FD50C106FCD3" pageId="2" pageNumber="366" type="nomenclature">
<paragraph id="8BB336ACFFAAFF9EA2B2FD50C1CBFCFE" blockId="2.[151,483,763,824]" box="[151,483,763,789]" pageId="2" pageNumber="366">
<heading id="D0FB81C0FFAAFF9EA2B2FD50C1CBFCFE" bold="true" box="[151,483,763,789]" fontSize="10" level="2" pageId="2" pageNumber="366" reason="2">
<emphasis id="B978EABEFFAAFF9EA2B2FD50C1CBFCFE" bold="true" box="[151,483,763,789]" pageId="2" pageNumber="366">
<taxonomicName id="4C0C4D2FFFAAFF9EA2B2FD50C1ABFCFF" ID-CoL="6N6F9" box="[151,387,763,788]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="2" pageNumber="366" phylum="Platyhelminthes" rank="species" species="qeshmensis" status="sp. nov.">
<emphasis id="B978EABEFFAAFF9EA2B2FD50C1ABFCFF" bold="true" box="[151,387,763,788]" italics="true" pageId="2" pageNumber="366">Imogine qeshmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C5FFAAFF9EA3ACFD50C1CBFCFE" box="[393,483,764,789]" pageId="2" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BB336ACFFAAFF9EA2B2FCB3C106FCD3" blockId="2.[151,483,763,824]" box="[151,302,799,824]" pageId="2" pageNumber="366">
(
<figureCitation id="13372A29FFAAFF9EA2BBFCB3C10FFCD3" box="[158,295,799,824]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="3.[151,248,1833,1855]" captionStartId-1="4.[151,248,1389,1411]" captionTargetBox-0="[159,1426,276,1806]" captionTargetBox-1="[173,1436,193,1363]" captionTargetId-0="figure@3.[151,1436,265,1811]" captionTargetId-1="figure@4.[161,1446,193,1367]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." captionText-1="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri-0="https://zenodo.org/record/228611/files/figure.png" httpUri-1="https://zenodo.org/record/228612/files/figure.png" pageId="2" pageNumber="366">Figures 3–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFAAFF9EA2B2FCCBC288FB2F" pageId="2" pageNumber="366" type="materials_examined">
<paragraph id="8BB336ACFFAAFF9EA2B2FCCBC103FC07" blockId="2.[151,1437,871,2013]" pageId="2" pageNumber="366">
<emphasis id="B978EABEFFAAFF9EA2B2FCCBC190FC6B" bold="true" box="[151,440,871,896]" pageId="2" pageNumber="366">Material and examined.</emphasis>
Six specimens were collected in 
<date id="FFB2106CFFAAFF9EA112FCCBC3C8FC6B" box="[823,992,871,896]" pageId="2" pageNumber="366" value="2011-02">February 2011</date>
: two mature specimens provided serial sagittal sections, to elucidate the histological structure of the reproductive systems; one specimen was used for a whole mount as an entire animal stained with Carmine Alum. The remaining three specimens were preserved in ethanol 70%.
</paragraph>
<paragraph id="8BB336ACFFAAFF9EA2E2FC5BC533FBDE" blockId="2.[151,1437,871,2013]" pageId="2" pageNumber="366">
<emphasis id="B978EABEFFAAFF9EA2E2FC5BC115FBFB" bold="true" box="[199,317,1015,1040]" pageId="2" pageNumber="366">
<typeStatus id="54B7880EFFAAFF9EA2E2FC5BC110FBFB" box="[199,312,1015,1040]" pageId="2" pageNumber="366" type="holotype">Holotype</typeStatus>
:
</emphasis>
one mature specimen, 
<collectionCode id="ED1DAE69FFAAFF9EA062FC54C2B8FBFB" box="[583,656,1016,1040]" pageId="2" pageNumber="366">ZUTC</collectionCode>
Platy 1265 HS, with the anterior half preserved in 
<collectionCode id="ED1DAE69FFAAFF9EA6F8FC54C50FFBFB" box="[1245,1319,1016,1040]" pageId="2" pageNumber="366">ETOH</collectionCode>
70%, and part of the posterior half containing the copulatory apparatus as series of sagittal sections (eight slides).
</paragraph>
<paragraph id="8BB336ACFFAAFF9EA2E2FB93C288FB2F" blockId="2.[151,1437,871,2013]" pageId="2" pageNumber="366">
<emphasis id="B978EABEFFAAFF9EA2E2FB93C160FBB3" bold="true" box="[199,328,1087,1112]" pageId="2" pageNumber="366">
<typeStatus id="54B7880EFFAAFF9EA2E2FB93C16BFBB3" box="[199,323,1087,1112]" pageId="2" pageNumber="366" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Five specimens: one sectioned specimen, 
<collectionCode id="ED1DAE69FFAAFF9EA104FBECC342FBB3" box="[801,874,1088,1112]" pageId="2" pageNumber="366">ZUTC</collectionCode>
Platy 1266 HS, with the anterior half preserved in 
<collectionCode id="ED1DAE69FFAAFF9EA2B2FBC8C0C9FB97" box="[151,225,1124,1148]" pageId="2" pageNumber="366">ETOH</collectionCode>
70 %, and part of the posterior half containing the copulatory apparatus as series of sagittal sections (seven slides); one entire specimen, 
<collectionCode id="ED1DAE69FFAAFF9EA3FAFB24C200FB4B" box="[479,552,1160,1184]" pageId="2" pageNumber="366">ZUTC</collectionCode>
Platy 1267 WM, cleared and stained with Carmine Alum; three entire worms, 
<collectionCode id="ED1DAE69FFAAFF9EA2B2FB00C0C8FB2F" box="[151,224,1196,1220]" pageId="2" pageNumber="366">ZUTC</collectionCode>
Platy 1268 S, preserved in 
<collectionCode id="ED1DAE69FFAAFF9EA031FB00C276FB2F" box="[532,606,1196,1220]" pageId="2" pageNumber="366">ETOH</collectionCode>
70%.
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFAAFF9EA2E2FB63C4F6FB03" box="[199,1246,1231,1256]" pageId="2" pageNumber="366" type="etymology">
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<emphasis id="B978EABEFFAAFF9EA2E2FB63C166FB03" bold="true" box="[199,334,1231,1256]" pageId="2" pageNumber="366">Etymology.</emphasis>
The specific epithet refers to 
<collectingCountry id="F31B763CFFAAFF9EA0BEFB63C2C2FB03" box="[667,746,1231,1256]" name="Iran" pageId="2" pageNumber="366">Qeshm</collectingCountry>
Island, where the specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFAAFF9EA2E2FB58C196FA93" pageId="2" pageNumber="366" type="materials_examined">
<paragraph id="8BB336ACFFAAFF9EA2E2FB58C196FA93" blockId="2.[151,1437,871,2013]" pageId="2" pageNumber="366">
<emphasis id="B978EABEFFAAFF9EA2E2FB58C141FAE6" bold="true" box="[199,361,1268,1293]" pageId="2" pageNumber="366">
<typeStatus id="54B7880EFFAAFF9EA2E2FB58C12AFAE6" box="[199,258,1268,1293]" pageId="2" pageNumber="366">Type</typeStatus>
locality.
</emphasis>
All specimens were collected from the upper intertidal rocky shore of Hamun jetty beach, 
<collectingCountry id="F31B763CFFAAFF9EA2B2FABBC0CEFADB" box="[151,230,1303,1328]" name="Iran" pageId="2" pageNumber="366">Qeshm</collectingCountry>
Island, Persian Gulf, 
<collectingCountry id="F31B763CFFAAFF9EA3F9FAB4C224FAC4" box="[476,524,1304,1327]" name="Iran" pageId="2" pageNumber="366">Iran</collectingCountry>
, 
<geoCoordinate id="EE38506BFFAAFF9EA03DFABBC2EEFADB" box="[536,710,1303,1328]" direction="north" orientation="latitude" pageId="2" pageNumber="366" precision="1" value="26.976473">26° 58´35.3&quot;N</geoCoordinate>
, 
<geoCoordinate id="EE38506BFFAAFF9EA0F7FABBC346FAC4" box="[722,878,1303,1328]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="15" value="56.248055">56° 14´53&quot; E</geoCoordinate>
(
<figureCitation id="13372A29FFAAFF9EA158FABBC3E8FADB" box="[893,960,1303,1328]" captionStart="FIGURE 1" captionStartId="1.[151,250,1260,1282]" captionTargetBox="[163,1436,376,1218]" captionTargetId="figure@1.[157,1442,373,1221]" captionTargetPageId="1" captionText="FIGURE 1. Map of the West Indo-Pacific and Persian Gulf. Type locality is Hamun jetty beach, Qeshm Island, the northern coasts of the Persian Gulf." httpUri="https://zenodo.org/record/228609/files/figure.png" pageId="2" pageNumber="366">Fig. 1</figureCitation>
). Habitat included rocky-muddy substrate (
<figureCitation id="13372A29FFAAFF9EA2BAFA90C0CAFABF" box="[159,226,1340,1365]" captionStart="FIGURE 2" captionStartId="1.[151,250,1858,1880]" captionTargetBox="[169,1433,1368,1825]" captionTargetId="figure@1.[169,1454,1362,1827]" captionTargetPageId="1" captionText="FIGURE 2. (A) Type locality of Imogine qeshmensis sp. nov., rocky substrate with rock oysters (B) Living specimen of I. qeshmensis." httpUri="https://zenodo.org/record/228610/files/figure.png" pageId="2" pageNumber="366">Fig. 2</figureCitation>
A) covered with rock oysters (
<taxonomicName id="4C0C4D2FFFAAFF9EA018FA91C30FFABE" box="[573,807,1340,1365]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreoida" pageId="2" pageNumber="366" phylum="Mollusca" rank="species" species="cucullata">
<emphasis id="B978EABEFFAAFF9EA018FA91C30FFABE" box="[573,807,1340,1365]" italics="true" pageId="2" pageNumber="366">Saccostrea cucullata</emphasis>
</taxonomicName>
), other associated animals with the specimens included sponges and red tunicates.
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFAAFF9EA2E2FA28C1E9F9C6" pageId="2" pageNumber="366" type="diagnosis">
<paragraph id="8BB336ACFFAAFF9EA2E2FA28C1E9F9C6" blockId="2.[151,1437,871,2013]" pageId="2" pageNumber="366">
<emphasis id="B978EABEFFAAFF9EA2E2FA28C169FA76" bold="true" box="[199,321,1412,1437]" pageId="2" pageNumber="366">Diagnosis.</emphasis>
Body thick, fleshy and broadly oval; rosy brown with darker brown spots, become smaller toward margin, body margin free of spots with white opaque band. With small white knobbed tentacles. Tentacular, cerebral and frontal eyes present. Marginal eyes encircle the whole body margin. Without spermiducal bulb, ejaculatory duct joins short prostatic duct in middle of penis. Penis papilla small; male and female antra ciliated. Gonopores well separated.
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFAAFF9EA2E2F99BC324F9BB" box="[199,780,1591,1616]" pageId="2" pageNumber="366" type="materials_examined">
<paragraph id="8BB336ACFFAAFF9EA2E2F99BC324F9BB" blockId="2.[151,1437,871,2013]" box="[199,780,1591,1616]" pageId="2" pageNumber="366">
<emphasis id="B978EABEFFAAFF9EA2E2F99BC1BFF9BB" bold="true" box="[199,407,1591,1616]" pageId="2" pageNumber="366">
<typeStatus id="54B7880EFFAAFF9EA2E2F99BC12AF9BB" box="[199,258,1591,1616]" pageId="2" pageNumber="366">Type</typeStatus>
of the genus
</emphasis>
: 
<taxonomicName id="4C0C4D2FFFAAFF9EA383F995C32FF9BB" authority="Girard, 1853" authorityName="Girard" authorityYear="1853" box="[422,775,1591,1616]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="2" pageNumber="366" phylum="Platyhelminthes" rank="species" species="oculifera">
<emphasis id="B978EABEFFAAFF9EA383F995C246F9BB" box="[422,622,1591,1616]" italics="true" pageId="2" pageNumber="366">Imogine oculifera</emphasis>
Girard, 1853
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFAAFF99A2E2F9F0C15FFCB7" lastPageId="5" lastPageNumber="369" pageId="2" pageNumber="366" type="description">
<paragraph id="8BB336ACFFAAFF9FA2E2F9F0C541FF3F" blockId="2.[151,1437,871,2013]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="367" pageId="2" pageNumber="366">
<emphasis id="B978EABEFFAAFF9EA2E2F9F0C275F99E" bold="true" box="[199,605,1628,1653]" pageId="2" pageNumber="366">Description. External morphology</emphasis>
. Body broadly oval, thick and fleshy with deep marginal ruffles (
<figureCitation id="13372A29FFAAFF9EA710F9F0C55FF99F" box="[1333,1399,1628,1653]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="2" pageNumber="366">Fig. 3</figureCitation>
A). 
<typeStatus id="54B7880EFFAAFF9EA2B2F9D3C0D7F973" box="[151,255,1663,1688]" pageId="2" pageNumber="366" type="holotype">Holotype</typeStatus>
25 × 
<quantity id="4CF49B49FFAAFF9EA366F92CC156F973" box="[323,382,1663,1688]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="2" pageNumber="366" unit="in" value="18.0">18 in</quantity>
size, the average size of 
<typeStatus id="54B7880EFFAAFF9EA0B8F92CC320F973" box="[669,776,1664,1688]" pageId="2" pageNumber="366" type="paratype">paratypes</typeStatus>
17 × 12. Dorsal body color of living animals rosy brown, lighter in the middle of the body around the pharynx (
<figureCitation id="13372A29FFAAFF9EA129F908C37AF957" box="[780,850,1700,1725]" captionStart="FIGURE 2" captionStartId="1.[151,250,1858,1880]" captionTargetBox="[169,1433,1368,1825]" captionTargetId="figure@1.[169,1454,1362,1827]" captionTargetPageId="1" captionText="FIGURE 2. (A) Type locality of Imogine qeshmensis sp. nov., rocky substrate with rock oysters (B) Living specimen of I. qeshmensis." httpUri="https://zenodo.org/record/228610/files/figure.png" pageId="2" pageNumber="366">Fig. 2</figureCitation>
B), with regular dark brown spots, become smaller toward margin; body margin with wide white opaque band, 
<quantity id="4CF49B49FFAAFF9EA164F96BC3B0F90B" box="[833,920,1735,1760]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="366" unit="mm" value="0.5">0.5 mm</quantity>
in thickness, free of spots (
<figureCitation id="13372A29FFAAFF9EA6F6F96BC53EF90B" box="[1235,1302,1735,1760]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="2" pageNumber="366">Fig. 3</figureCitation>
A), covered with eye spots (
<figureCitation id="13372A29FFAAFF9EA36CF940C1A3F8EF" box="[329,395,1772,1797]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="2" pageNumber="366">Fig. 3</figureCitation>
A, B). More or less translucent cream, in ventral view, vas deferens visible on eider sides of pharynx, as two dark, cream, coiled strips, extends anteriorly from the male gonopore to middle half of body (
<figureCitation id="13372A29FFAAFF9EA74BF8A3C08DF8A7" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="2" pageNumber="366">Fig. 3</figureCitation>
B). Nuchal tentacles small, rounded, retractile and white, 2.2 apart; tentacular eyes with 18–21 large eye spots at tip of each tentacle (
<figureCitation id="13372A29FFAAFF9EA3A3F8FBC1E3F89B" box="[390,459,1879,1904]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="2" pageNumber="366">Fig. 3</figureCitation>
C); small cerebral eyes embedded in epidermis between nuchal tentacles, difficult to observe, but readily observable on cleared anterior part of 
<typeStatus id="54B7880EFFAAFF9EA10DF8D0C3A2F87E" box="[808,906,1916,1941]" pageId="2" pageNumber="366" type="holotype">holotype</typeStatus>
and stained whole 
<typeStatus id="54B7880EFFAAFF9EA641F8D1C4ECF87E" box="[1124,1220,1917,1941]" pageId="2" pageNumber="366" type="paratype">paratype</typeStatus>
(ZUTC Platy.1267 WM), cerebral eye clusters more or less paired, more crowded posterior to nuchal tentacles with 55 eye spots in each posterior cluster, cerebral eyes extend anterior to nuchal tentacles; frontal eyes, with about 21 eyes in each cluster, scattered and subdivided (
<figureCitation id="13372A29FFABFF9FA032FF3BC272FF5B" box="[535,602,151,176]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="3" pageNumber="367">Fig. 3</figureCitation>
C), frontal eyes not reaching anterior margin. Marginal eyes extend along the entire margin, denser anteriorly, in four or five rows, reducing posteriorly into two or three rows (
<figureCitation id="13372A29FFABFF9FA721FF10C56EFF3F" box="[1284,1350,188,213]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="3" pageNumber="367">Fig. 3</figureCitation>
B).
</paragraph>
<caption id="DF736624FFABFF9FA2B2F885C1CBF833" httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="3" pageNumber="367" targetBox="[159,1426,276,1806]" targetPageId="3">
<paragraph id="8BB336ACFFABFF9FA2B2F885C1CBF833" blockId="3.[151,1436,1833,2008]" pageId="3" pageNumber="367">
<emphasis id="B978EABEFFABFF9FA2B2F885C138F8D5" bold="true" box="[151,272,1833,1855]" pageId="3" pageNumber="367">FIGURE 3.</emphasis>
<taxonomicName id="4C0C4D2FFFABFF9FA333F886C1F6F8D4" box="[278,478,1833,1855]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="3" pageNumber="367" phylum="Platyhelminthes" rank="species" species="qeshmensis" status="sp. nov.">
<emphasis id="B978EABEFFABFF9FA333F886C1F6F8D4" box="[278,478,1833,1855]" italics="true" pageId="3" pageNumber="367">Imogine qeshmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C5FFABFF9FA3C1F885C21CF8D5" box="[484,564,1833,1854]" pageId="3" pageNumber="367" rank="species">
<emphasis id="B978EABEFFABFF9FA3C1F885C21CF8D5" bold="true" box="[484,564,1833,1854]" pageId="3" pageNumber="367">sp. nov.</emphasis>
</taxonomicNameLabel>
(A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens.
</paragraph>
</caption>
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<paragraph id="8BB336ACFFACFF98A2B2FAC1C246F9F6" blockId="4.[151,1436,1389,1565]" pageId="4" pageNumber="368">
<emphasis id="B978EABEFFACFF98A2B2FAC1C13BFA68" bold="true" box="[151,275,1389,1411]" pageId="4" pageNumber="368">FIGURE 4.</emphasis>
Haematoxylin-eosin stained sections of 
<taxonomicName id="4C0C4D2FFFACFF98A09FFAC3C3ACFA68" box="[698,900,1389,1412]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="4" pageNumber="368" phylum="Platyhelminthes" rank="species" species="qeshmensis" status="sp. nov.">
<emphasis id="B978EABEFFACFF98A09FFAC3C3ACFA68" box="[698,900,1389,1412]" italics="true" pageId="4" pageNumber="368">Imogine qeshmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C5FFACFF98A1A9FAC2C3C8FA68" box="[908,992,1390,1411]" pageId="4" pageNumber="368" rank="species">
<emphasis id="B978EABEFFACFF98A1A9FAC2C3C8FA68" bold="true" box="[908,992,1390,1411]" pageId="4" pageNumber="368">sp. nov.</emphasis>
</taxonomicNameLabel>
(Holotype, ZUTC Platy.1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe.
</paragraph>
</caption>
<paragraph id="8BB336ACFFACFF98A2E2F9E5C319F941" blockId="4.[151,1436,1609,2031]" pageId="4" pageNumber="368">
Ruffled pharynx located centrally, 4.7 from the anterior end, 7.3 from posterior end of the body, 
<quantity id="4CF49B49FFACFF98A73CF9E5C556F98A" box="[1305,1406,1609,1634]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="368" unit="mm" value="12.5">12.5 mm</quantity>
in length, with about 12 pairs of lateral folds, mouth located in anterior part of middle third of pharynx (
<figureCitation id="13372A29FFACFF98A711F9C1C551F96D" box="[1332,1401,1645,1670]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="4" pageNumber="368">Fig. 3</figureCitation>
B). Male pore located posterior to pharynx, 3.8 apart (
<figureCitation id="13372A29FFACFF98A0EFF93DC325F941" box="[714,781,1681,1706]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="4" pageNumber="368">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="8BB336ACFFACFF98A2E2F919C1D5F869" blockId="4.[151,1436,1609,2031]" pageId="4" pageNumber="368">
<emphasis id="B978EABEFFACFF98A2E2F919C22DF925" bold="true" box="[199,517,1717,1742]" pageId="4" pageNumber="368">Epidermis and body wall.</emphasis>
Ventral epidermis three times thicker than dorsal epidermis, 95–100 µm thick (
<figureCitation id="13372A29FFACFF98A2BAF975C0C6F919" box="[159,238,1753,1778]" captionStart="FIGURE 4" captionStartId="4.[151,248,1389,1411]" captionTargetBox="[173,1436,193,1363]" captionTargetId="figure@4.[161,1446,193,1367]" captionTargetPageId="4" captionText="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri="https://zenodo.org/record/228612/files/figure.png" pageId="4" pageNumber="368">Figs. 4</figureCitation>
B, D). Cellular epidermis consists of thick-walled (18.5 µm) columnar cells and rhabdites, basal membrane 6.7 µm in thickness; arrangement of muscle fibers beneath basement membrane: outer, very thin circular muscle layer (3.6 µm), followed internally by thick longitudinal fibers (11.5 µm), then well-developed diagonal muscles (31 µm) and innermost longitudinal muscles merging with diagonal muscle fibers (26 µm). Transverse muscle fibers well developed (
<figureCitation id="13372A29FFACFF98A3B2F8C5C1F3F869" box="[407,475,1897,1922]" captionStart="FIGURE 4" captionStartId="4.[151,248,1389,1411]" captionTargetBox="[173,1436,193,1363]" captionTargetId="figure@4.[161,1446,193,1367]" captionTargetPageId="4" captionText="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri="https://zenodo.org/record/228612/files/figure.png" pageId="4" pageNumber="368">Fig. 4</figureCitation>
B).
</paragraph>
<paragraph id="8BB336ACFFACFF99A2E2F821C23BFF3F" blockId="4.[151,1436,1609,2031]" lastBlockId="5.[151,1436,151,2013]" lastPageId="5" lastPageNumber="369" pageId="4" pageNumber="368">
Dorsal epidermis about 36 µm thick (
<figureCitation id="13372A29FFACFF98A054F821C29EF84D" box="[625,694,1933,1958]" captionStart="FIGURE 4" captionStartId="4.[151,248,1389,1411]" captionTargetBox="[173,1436,193,1363]" captionTargetId="figure@4.[161,1446,193,1367]" captionTargetPageId="4" captionText="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri="https://zenodo.org/record/228612/files/figure.png" pageId="4" pageNumber="368">Fig. 4</figureCitation>
A); cellular epidermis consists of tall, thin-walled columnar cells, ciliated, with intra-epithelial nuclei and rhabdites; basal membrane about 7 µm in thickness; body wall musculature consists of outer, very thin circular layer, followed by inner longitudinal layer (12 µm). Clusters of granular pigment scattered beneath dorsal and ventral wall throughout the parenchyma; few sparse rhabdites observed below dorsal epidermis (
<figureCitation id="13372A29FFADFF99A38EFF10C1C7FF3F" box="[427,495,188,213]" captionStart="FIGURE 4" captionStartId="4.[151,248,1389,1411]" captionTargetBox="[173,1436,193,1363]" captionTargetId="figure@4.[161,1446,193,1367]" captionTargetPageId="4" captionText="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri="https://zenodo.org/record/228612/files/figure.png" pageId="5" pageNumber="369">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="8BB336ACFFADFF99A2E2FF73C451FD27" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="369">
<emphasis id="B978EABEFFADFF99A2E2FF73C1C6FF13" bold="true" box="[199,494,223,248]" pageId="5" pageNumber="369">Reproductive structures</emphasis>
. Male reproductive system with tripartite seminal vesicle, free, elongated oval prostatic vesicle, without spermiducal bulb, and penis papilla housed in a ciliated antrum (
<figureCitation id="13372A29FFADFF99A6B3FEA8C4CBFEF7" box="[1174,1251,260,285]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Figs. 3</figureCitation>
D &amp; 4C). Testes scattered ventrally throughout body length. In ventral view, vasa deferentia arising in mid-body, converging posteriorly on either side of pharynx (
<figureCitation id="13372A29FFADFF99A074FEE0C2BFFE8F" box="[593,663,332,357]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Fig. 3</figureCitation>
B). In sagittal view, vas deferens recurrent below prostatic vesicle, opens into lateral lobes of seminal vesicle (
<figureCitation id="13372A29FFADFF99A0ADFEC3C2E4FE63" box="[648,716,367,392]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Fig. 3</figureCitation>
D). Seminal vesicle 620 µm wide, enclosed in strong muscular layer (100 µm in thickness), located anterior to prostatic vesicle, oriented horizontally in 
<typeStatus id="54B7880EFFADFF99A6B0FE38C4DFFE46" box="[1173,1271,404,429]" pageId="5" pageNumber="369" type="holotype">holotype</typeStatus>
(
<figureCitation id="13372A29FFADFF99A72DFE38C565FE47" box="[1288,1357,404,429]" captionStart="FIGURE 4" captionStartId="4.[151,248,1389,1411]" captionTargetBox="[173,1436,193,1363]" captionTargetId="figure@4.[161,1446,193,1367]" captionTargetPageId="4" captionText="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri="https://zenodo.org/record/228612/files/figure.png" pageId="5" pageNumber="369">Fig. 4</figureCitation>
C) but, oriented slightly dorsally in sectioned 
<typeStatus id="54B7880EFFADFF99A062FE14C283FE3B" box="[583,683,440,464]" pageId="5" pageNumber="369" type="paratype">paratype</typeStatus>
. Lateral lobes of seminal vesicle equal in size, 860 µm long × 200 µm wide, central lobe larger than lateral lobes, 
<quantity id="4CF49B49FFADFF99A08AFE70C32EFE1F" box="[687,774,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="369" unit="mm" value="1.2">1.2 mm</quantity>
long × 200 µm wide (
<figureCitation id="13372A29FFADFF99A620FE70C47BFE1F" box="[1029,1107,476,501]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Figs. 3</figureCitation>
D &amp; 4C). Central lobe tapers posteriorly below the prostatic vesicle to form straight ejaculatory duct, near distal end of prostatic vesicle; ejaculatory duct joins short prostatic duct in middle of penis. Prostatic vesicle surrounded by muscular layer, 80 µm in thickness, 
<quantity id="4CF49B49FFADFF99A30EFDE4C144FDB4" box="[299,364,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="369" unit="mm" value="1.0">1 mm</quantity>
long × 400 µm width (
<figureCitation id="13372A29FFADFF99A054FDEBC296FD8B" box="[625,702,583,608]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Figs. 3</figureCitation>
D &amp; 4C). Wall of prostatic vesicle with numerous narrow regular transverse ducts (extra-vesicular gland canals) leading into lumen, with polyglandular lining, glandular cell-bodies of extra-vesicular glands localized more or less distant in parenchyma surrounding male apparatus (
<figureCitation id="13372A29FFADFF99A6D7FD23C51EFD43" box="[1266,1334,655,680]" captionStart="FIGURE 4" captionStartId="4.[151,248,1389,1411]" captionTargetBox="[173,1436,193,1363]" captionTargetId="figure@4.[161,1446,193,1367]" captionTargetPageId="4" captionText="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri="https://zenodo.org/record/228612/files/figure.png" pageId="5" pageNumber="369">Fig. 4</figureCitation>
D). Penis papilla small, 170 × 250 µm, housed in ciliated, deep and narrow male antrum (
<figureCitation id="13372A29FFADFF99A637FD18C47DFD27" box="[1042,1109,692,717]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="8BB336ACFFADFF99A2E2FD7BC15FFCB7" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="369">
Female gonopore 230 µm posterior to male gonopore; vagina narrow, somewhat muscular and curved, with cuboidal epithelium, vagina proceeds dorsally, and then curves backward to receive uteri; vagina surrounded by numerous cement glands, with ciliated female antrum (
<figureCitation id="13372A29FFADFF99A0D8FCB3C368FCD3" box="[765,832,799,824]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Fig. 3</figureCitation>
D). Lang's vesicle absent. Male and female gonopores separated (
<figureCitation id="13372A29FFADFF99A335FCE8C17AFCB7" box="[272,338,836,861]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="5" pageNumber="369">Fig. 3</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="C3166527FFADFF95A2E2FCCBC39CFDF3" lastPageId="9" lastPageNumber="373" pageId="5" pageNumber="369" type="discussion">
<paragraph id="8BB336ACFFADFF99A2E2FCCBC3C1FB97" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="369">
<emphasis id="B978EABEFFADFF99A2E2FCCBC113FC6B" bold="true" box="[199,315,871,896]" pageId="5" pageNumber="369">Remarks.</emphasis>
<bibRefCitation id="EF9D4B5DFFADFF99A362FCCBC1C4FC6B" author="Faubel" box="[327,492,871,896]" pageId="5" pageNumber="369" refString="Faubel, A. (1983) The Polycladida, Turbellaria. Proposal and establishment of a new system. Part I. The Acotylea. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 80, 17 - 121." type="journal article" year="1983">Faubel (1983)</bibRefCitation>
, in his classification, emended the family 
<taxonomicName id="4C0C4D2FFFADFF99A1CEFCCBC447FC6B" box="[1003,1135,871,896]" class="Rhabditophora" family="Stylochidae" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="family">Stylochidae</taxonomicName>
and introduced the main diagnostic character of the family as: the ridged glandular lining of the prostatic vesicle with extra-vesicular glands. Recently, 
<bibRefCitation id="EF9D4B5DFFADFF99A323FC03C1CCFC23" box="[262,484,943,968]" pageId="5" pageNumber="369" refString="Bulnes, V. N., Faubel, A. &amp; Park, J. K. (2005) Two new marine species from South Korea with remarks on the family Stylochidae (Acotylea, Polycladida, Plathelminthes). Journal of Natural History, 39 (23), 2089 - 2107. http: // dx. doi. org / 10.1080 / 00222930500081997" type="journal article">
Bulnes 
<emphasis id="B978EABEFFADFF99A37EFC1DC1A3FC23" box="[347,395,943,968]" italics="true" pageId="5" pageNumber="369">et al</emphasis>
. (2005)
</bibRefCitation>
defined the family 
<taxonomicName id="4C0C4D2FFFADFF99A098FC03C369FC23" box="[701,833,943,968]" class="Rhabditophora" family="Stylochidae" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="family">Stylochidae</taxonomicName>
based on two new 
<typeStatus id="54B7880EFFADFF99A633FC1CC479FC23" box="[1046,1105,944,968]" pageId="5" pageNumber="369">types</typeStatus>
of prostatic linings: the polyglandular and the mono-glandular 
<typeStatus id="54B7880EFFADFF99A01CFC79C250FC06" box="[569,632,981,1005]" pageId="5" pageNumber="369">types</typeStatus>
. In the case of mono-glandular 
<typeStatus id="54B7880EFFADFF99A623FC79C41EFC06" box="[1030,1078,981,1005]" pageId="5" pageNumber="369">type</typeStatus>
each extra-vesicular gland is connected via a glandular duct with a single inner tube, so the number of extra-vesicular glands and inner tubes is equal. In poly-glandular 
<typeStatus id="54B7880EFFADFF99A39FFBB1C1C5FBDE" box="[442,493,1053,1077]" pageId="5" pageNumber="369">type</typeStatus>
, each long-fingered extension is connected via several glandular ducts to extravesicular glands (
<bibRefCitation id="EF9D4B5DFFADFF99A37EFB93C205FBB3" author="Bulnes" box="[347,557,1087,1112]" pageId="5" pageNumber="369" refString="Bulnes, V. N., Faubel, A. &amp; Park, J. K. (2005) Two new marine species from South Korea with remarks on the family Stylochidae (Acotylea, Polycladida, Plathelminthes). Journal of Natural History, 39 (23), 2089 - 2107. http: // dx. doi. org / 10.1080 / 00222930500081997" type="journal article" year="2005">Bulnes et al. 2005</bibRefCitation>
). However, extra vesicular glands were not observed by Newman and Cannon (1996a, b) in some members of the family including 
<taxonomicName id="4C0C4D2FFFADFF99A0C3FBC9C36AFB97" box="[742,834,1125,1148]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A0C3FBC9C36AFB97" box="[742,834,1125,1148]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0C4D2FFFADFF99A15DFBC8C3CAFB96" box="[888,994,1124,1149]" class="Rhabditophora" family="Stylochidae" genus="Stylochus" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A15DFBC8C3CAFB96" box="[888,994,1124,1149]" italics="true" pageId="5" pageNumber="369">Stylochus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB336ACFFADFF99A2E2FB2BC52BFADB" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="369">
Based on the presence / absence of a tripartite seminal vesicle, Marcus and Marcus (1968) divided the genus 
<taxonomicName id="4C0C4D2FFFADFF99A2B2FB00C12AFB2E" box="[151,258,1196,1221]" class="Rhabditophora" family="Stylochidae" genus="Stylochus" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A2B2FB00C12AFB2E" box="[151,258,1196,1221]" italics="true" pageId="5" pageNumber="369">Stylochus</emphasis>
</taxonomicName>
within the 
<taxonomicName id="4C0C4D2FFFADFF99A3A2FB00C225FB2E" box="[391,525,1196,1221]" class="Rhabditophora" family="Stylochidae" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="family">Stylochidae</taxonomicName>
, into the two subgenera: 
<taxonomicName id="4C0C4D2FFFADFF99A117FB01C401FB2E" authority="Girard, 1853" authorityName="Girard" authorityYear="1853" box="[818,1065,1196,1221]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A117FB01C3A6FB2F" box="[818,910,1197,1220]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
Girard, 1853
</taxonomicName>
and 
<taxonomicName id="4C0C4D2FFFADFF99A641FB00C5B0FB2F" authority="Ehrenberg, 1831" authorityName="Ehrenberg" authorityYear="1831" box="[1124,1432,1196,1221]" class="Rhabditophora" family="Stylochidae" genus="Stylochus" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A641FB00C4E6FB2E" box="[1124,1230,1196,1221]" italics="true" pageId="5" pageNumber="369">Stylochus</emphasis>
Ehrenberg, 1831
</taxonomicName>
. 
<bibRefCitation id="EF9D4B5DFFADFF99A2B2FB63C16EFB03" author="Prudhoe" box="[151,326,1231,1256]" pageId="5" pageNumber="369" refString="Prudhoe, S. (1989) Polyclad turbellarians recorded from African waters. Bulletin of the British Museum (Natural History), 55, 47 - 96." type="journal article" year="1989">Prudhoe (1989)</bibRefCitation>
and 
<bibRefCitation id="EF9D4B5DFFADFF99A358FB7CC25CFB03" box="[381,628,1231,1256]" pageId="5" pageNumber="369" refString="Newman, L. J., Cannon, L. R. G. &amp; Govan, H. (1993) Stylochus (Imogene) matatasi n. sp. (Platyhelminthes, Polycladida): pest of cultured giant clams and pearl oysters from Solomon Islands. Hydrobiologia, 257 (3), 185 - 189. http: // dx. doi. org / 10.1007 / bf 00765011" type="journal article">
Newman 
<emphasis id="B978EABEFFADFF99A3CCFB7DC232FB03" box="[489,538,1231,1256]" italics="true" pageId="5" pageNumber="369">et al</emphasis>
. (1993)
</bibRefCitation>
recognized 28 species of 
<taxonomicName id="4C0C4D2FFFADFF99A1B2FB63C429FB03" box="[919,1025,1231,1256]" class="Rhabditophora" family="Stylochidae" genus="Stylochus" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A1B2FB63C429FB03" box="[919,1025,1231,1256]" italics="true" pageId="5" pageNumber="369">Stylochus</emphasis>
</taxonomicName>
within the subgenus 
<taxonomicName id="4C0C4D2FFFADFF99A6D4FB7DC565FB03" box="[1265,1357,1233,1256]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A6D4FB7DC565FB03" box="[1265,1357,1233,1256]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
</taxonomicName>
. Later, 
<bibRefCitation id="EF9D4B5DFFADFF99A2B2FB58C1D0FAE6" author="Jennings" box="[151,504,1268,1293]" pageId="5" pageNumber="369" refString="Jennings, K. A. &amp; Newman, L. J. (1996 b) Two new Stylochid flatworms (Platyhelminthes: Polycladida) from the southern Great Barrier Reef, Australia. The Raffles Bulletin of Zoology, 44, 135 - 142." type="journal article" year="1996" yearSuffix="b">Jennings and Newman (1996b)</bibRefCitation>
elevated the 
<taxonomicName id="4C0C4D2FFFADFF99A0B5FB59C2C4FAE7" box="[656,748,1269,1292]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A0B5FB59C2C4FAE7" box="[656,748,1269,1292]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
</taxonomicName>
to generic rank. Then, several new species were assigned to this genus around the world (
<bibRefCitation id="EF9D4B5DFFADFF99A3F8FABBC246FADB" author="Bulnes" box="[477,622,1303,1328]" pageId="5" pageNumber="369" refString="Bulnes, V. N. (2010) Five new Polycladida (Platyhelminthes: Acotylea) species from the Aegean Sea with remarks on the prostatic structures. Journal of Natural History, 44, 515 - 544. http: // dx. doi. org / 10.1080 / 00222930903497535" type="journal article" year="2010">Bulnes 2010</bibRefCitation>
; 
<bibRefCitation id="EF9D4B5DFFADFF99A05FFABBC39CFADB" author="Jennings" box="[634,948,1303,1328]" pageId="5" pageNumber="369" refString="Jennings, K. A. &amp; Newman, L. J. (1996 a) Four new Stylochid flatworms (Platyhelminthes: Polycladida) associated with commercial oysters from Moreton Bay, southeast Queensland, Australia. The Raffles Bulletin of Zoology, 44, 493 - 508." type="journal article" year="1996" yearSuffix="a">Jennings &amp; Newman 1996a</bibRefCitation>
; Lee 2006; 
<bibRefCitation id="EF9D4B5DFFADFF99A612FABBC4DFFADB" author="Sluys" box="[1079,1271,1303,1328]" pageId="5" pageNumber="369" refString="Sluys, R., Faubel, A., Rajagopal, S. &amp; van der Velde, G. (2005) A new and alien species of &quot; oyster leech &quot; (Platyhelminthes, Polycladida, Stylochidae) from the brackish North Sea Canal, The Netherlands. Helgoland Marine Research, 59 (4), 310 - 314." type="journal article" year="2005">
Sluys 
<emphasis id="B978EABEFFADFF99A659FAB5C49CFADB" box="[1148,1204,1303,1328]" italics="true" pageId="5" pageNumber="369">et al.</emphasis>
2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8BB336ACFFADFF99A2E2FA90C2AEF9C7" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="369">
The genus 
<taxonomicName id="4C0C4D2FFFADFF99A367FA91C1B6FABF" box="[322,414,1341,1364]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A367FA91C1B6FABF" box="[322,414,1341,1364]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
</taxonomicName>
is defined by a tripartite and highly muscular seminal vesicle and a prostatic vesicle of the polyglandular 
<typeStatus id="54B7880EFFADFF99A31CFACDC141FA92" box="[313,361,1377,1401]" pageId="5" pageNumber="369">type</typeStatus>
(
<bibRefCitation id="EF9D4B5DFFADFF99A352FAF3C26FFA93" author="Bulnes" box="[375,583,1375,1400]" pageId="5" pageNumber="369" refString="Bulnes, V. N., Faubel, A. &amp; Park, J. K. (2005) Two new marine species from South Korea with remarks on the family Stylochidae (Acotylea, Polycladida, Plathelminthes). Journal of Natural History, 39 (23), 2089 - 2107. http: // dx. doi. org / 10.1080 / 00222930500081997" type="journal article" year="2005">Bulnes et al. 2005</bibRefCitation>
). Within the genus, the species are discriminated by their color pattern, body shape and consistency, presence or absence of tentacles, the number and pattern of tentacular and cerebral eyes, and the presence of frontal and marginal eyes. Due to the phenotypic variability of these characters, additional features such as the position of pharynx and mouth, the presence of spermiducal bulbs, the position of the junction between the ejaculatory duct and the prostatic duct, the form and size of the penis papilla and the histology of the male atrium have been considered (
<bibRefCitation id="EF9D4B5DFFADFF99A3C3F9B8C252F9C7" author="Bulnes" box="[486,634,1556,1581]" pageId="5" pageNumber="369" refString="Bulnes, V. N. (2010) Five new Polycladida (Platyhelminthes: Acotylea) species from the Aegean Sea with remarks on the prostatic structures. Journal of Natural History, 44, 515 - 544. http: // dx. doi. org / 10.1080 / 00222930903497535" type="journal article" year="2010">Bulnes 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BB336ACFFADFF99A2E2F99BC39DF8EF" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="369">
The presence of a tripartite seminal vesicle clearly places the specimen described here as a member of the genus 
<taxonomicName id="4C0C4D2FFFADFF99A2C7F9F1C116F99F" box="[226,318,1629,1652]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A2C7F9F1C116F99F" box="[226,318,1629,1652]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
</taxonomicName>
. Currently, this genus contains 36 species (
<bibRefCitation id="EF9D4B5DFFADFF99A116F9F0C3FBF99F" author="Bolanos" box="[819,979,1628,1653]" pageId="5" pageNumber="369" refString="Bolanos, D. M. (2010) Polycladida lifedesks. Available from: http: // polycladida. lifedesks. org / pages / 3179 (accessed 5 June 2012)" type="journal article" year="2010">Bolaños 2010</bibRefCitation>
) with two taxonomic groups within the species: those with eyes around the entire body margin and those with only anterior eyes (
<bibRefCitation id="EF9D4B5DFFADFF99A684F92CC5B8F973" author="Newman" box="[1185,1424,1663,1688]" pageId="5" pageNumber="369" refString="Newman, L. J., Cannon, L. R. G. &amp; Govan, H. (1993) Stylochus (Imogene) matatasi n. sp. (Platyhelminthes, Polycladida): pest of cultured giant clams and pearl oysters from Solomon Islands. Hydrobiologia, 257 (3), 185 - 189. http: // dx. doi. org / 10.1007 / bf 00765011" type="journal article" year="1993">
Newman 
<emphasis id="B978EABEFFADFF99A734F92DC563F973" box="[1297,1355,1663,1688]" italics="true" pageId="5" pageNumber="369">et al.</emphasis>
1993
</bibRefCitation>
). Sixteen of them, like the species describe here, have eyes around the entire body margin (Table 1). 
<bibRefCitation id="EF9D4B5DFFADFF99A72CF908C173F90B" author="Jennings" pageId="5" pageNumber="369" refString="Jennings, K. A. &amp; Newman, L. J. (1996 a) Four new Stylochid flatworms (Platyhelminthes: Polycladida) associated with commercial oysters from Moreton Bay, southeast Queensland, Australia. The Raffles Bulletin of Zoology, 44, 493 - 508." type="journal article" year="1996" yearSuffix="a">Jennings and Newman (1996a)</bibRefCitation>
have summarized the main diagnostic characters of 
<taxonomicName id="4C0C4D2FFFADFF99A197F965C426F90B" box="[946,1038,1737,1760]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A197F965C426F90B" box="[946,1038,1737,1760]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
</taxonomicName>
species with eyes around the body margin. These characters with some modifications are listed in Table 1.
</paragraph>
<paragraph id="8BB336ACFFADFF9AA2E2F8BDC0C7FF3E" blockId="5.[151,1436,151,2013]" lastBlockId="6.[151,1436,151,2013]" lastPageId="6" lastPageNumber="370" pageId="5" pageNumber="369">
In terms of geographical distribution (Table 1), the aforementioned sixteen species occur in the following regions: Atlantic Ocean (eight species), Mediterranean Sea (one species), north Pacific (one species) and 
<collectingCountry id="F31B763CFFADFF99A70EF898C576F8A6" box="[1323,1374,1844,1869]" name="India" pageId="5" pageNumber="369">Indo</collectingCountry>
West Pacific (six species). 
<bibRefCitation id="EF9D4B5DFFADFF99A3ADF8FBC2C1F89B" author="Jennings" box="[392,745,1879,1904]" pageId="5" pageNumber="369" refString="Jennings, K. A. &amp; Newman, L. J. (1996 a) Four new Stylochid flatworms (Platyhelminthes: Polycladida) associated with commercial oysters from Moreton Bay, southeast Queensland, Australia. The Raffles Bulletin of Zoology, 44, 493 - 508." type="journal article" year="1996" yearSuffix="a">Jennings and Newman (1996 a</bibRefCitation>
, 
<bibRefCitation id="EF9D4B5DFFADFF99A0D0F8FBC32CF89B" author="Jennings" box="[757,772,1879,1904]" pageId="5" pageNumber="369" refString="Jennings, K. A. &amp; Newman, L. J. (1996 b) Two new Stylochid flatworms (Platyhelminthes: Polycladida) from the southern Great Barrier Reef, Australia. The Raffles Bulletin of Zoology, 44, 135 - 142." type="journal article" year="1996" yearSuffix="b">b</bibRefCitation>
) considered the geographical distribution as a character to discriminate their five new species of 
<taxonomicName id="4C0C4D2FFFADFF99A067F8D1C2B6F87F" box="[578,670,1917,1940]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="5" pageNumber="369" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFADFF99A067F8D1C2B6F87F" box="[578,670,1917,1940]" italics="true" pageId="5" pageNumber="369">Imogine</emphasis>
</taxonomicName>
inhabiting the Indo-West Pacific from all other congeners with eyes around the entire body margin. We believe that history and intensity of the maritime activities in the Persian Gulf provides the possibility (through discharge of ballast water) to introduce species from other parts of world’s ocean into the region. Therefore, we compared our species from the Persian Gulf with all of the above mentioned sixteen species.
</paragraph>
<paragraph id="8BB336ACFFAEFF9AA2E2FF73C54DFA93" blockId="6.[151,1436,151,2013]" pageId="6" pageNumber="370">
Based on the information given in Table 1, and compared with the 
<taxonomicName id="4C0C4D2FFFAEFF9AA1FCFF4DC41DFF13" box="[985,1077,225,248]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFAEFF9AA1FCFF4DC41DFF13" box="[985,1077,225,248]" italics="true" pageId="6" pageNumber="370">Imogine</emphasis>
</taxonomicName>
species inhabiting the Atlantic Ocean, the species described here is distinguished from: (1) 
<taxonomicName id="4C0C4D2FFFAEFF9AA16CFEA9C43AFEF7" box="[841,1042,261,284]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="arenosus">
<emphasis id="B978EABEFFAEFF9AA16CFEA9C43AFEF7" box="[841,1042,261,284]" italics="true" pageId="6" pageNumber="370">Imogine arenosus</emphasis>
</taxonomicName>
by its color pattern, tentacle color and number of tentacular eyes. 
<taxonomicName id="4C0C4D2FFFAEFF9AA022FE85C2ADFEAB" box="[519,645,297,320]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="arenosus">
<emphasis id="B978EABEFFAEFF9AA022FE85C2ADFEAB" box="[519,645,297,320]" italics="true" pageId="6" pageNumber="370">I. arenosus</emphasis>
</taxonomicName>
is dorsally reddish brown with white and black dots, has numerous tentacular eyes at the base of tentacles, while the species described here is dorsally rosy brown with regular dark brown spots, these spots become smaller towards the margin; the nuchal tentacles are small, retractile and white in the fixed specimens; with 18–21 tentacular eyes at the tip of each tentacle (2) 
<taxonomicName id="4C0C4D2FFFAEFF9AA63AFE39C45CFE47" box="[1055,1140,405,428]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="catus">
<emphasis id="B978EABEFFAEFF9AA63AFE39C45CFE47" box="[1055,1140,405,428]" italics="true" pageId="6" pageNumber="370">I. catus</emphasis>
</taxonomicName>
in terms of color pattern, shape of tentacles, shape of the prostatic vesicle and body size. The body in 
<taxonomicName id="4C0C4D2FFFAEFF9AA1C1FE15C41DFE3B" box="[996,1077,441,464]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="catus">
<emphasis id="B978EABEFFAEFF9AA1C1FE15C41DFE3B" box="[996,1077,441,464]" italics="true" pageId="6" pageNumber="370">I. catus</emphasis>
</taxonomicName>
dorsally has brown spots in two lateral bands, unlike the species described here. The species presented here is 25 × 
<quantity id="4CF49B49FFAEFF9AA677FE70C4A5FE1E" box="[1106,1165,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="6" pageNumber="370" unit="in" value="18.0">18 in</quantity>
size, with small nuchal tentacles, and elongated oval prostatic, while 
<taxonomicName id="4C0C4D2FFFAEFF9AA0B7FDADC2CCFDF3" box="[658,740,513,536]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="catus">
<emphasis id="B978EABEFFAEFF9AA0B7FDADC2CCFDF3" box="[658,740,513,536]" italics="true" pageId="6" pageNumber="370">I. catus</emphasis>
</taxonomicName>
is 65 × 
<quantity id="4CF49B49FFAEFF9AA118FDACC3A4FDFC" box="[829,908,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="370" unit="mm" value="32.0">32 mm</quantity>
in size with elongated tentacles and a spherical prostatic vesicle (3) The presence of cilia in the male and female antra of the species described here distinguishes this species form 
<taxonomicName id="4C0C4D2FFFAEFF9AA37AFDE5C1DDFD8B" box="[351,501,583,608]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="meridianus">
<emphasis id="B978EABEFFAEFF9AA37AFDE5C1DDFD8B" box="[351,501,583,608]" italics="true" pageId="6" pageNumber="370">I. meridianus</emphasis>
</taxonomicName>
that lacks ciliated male and female antra (
<bibRefCitation id="EF9D4B5DFFAEFF9AA1F2FDEBC443FD8B" author="Bulnes" box="[983,1131,583,608]" pageId="6" pageNumber="370" refString="Bulnes, V. N. (2010) Five new Polycladida (Platyhelminthes: Acotylea) species from the Aegean Sea with remarks on the prostatic structures. Journal of Natural History, 44, 515 - 544. http: // dx. doi. org / 10.1080 / 00222930903497535" type="journal article" year="2010">Bulnes 2010</bibRefCitation>
); in the present species the ejaculatory duct joins the prostatic duct in the middle of the penis, while in 
<taxonomicName id="4C0C4D2FFFAEFF9AA1C0FDC1C452FD6E" box="[997,1146,620,645]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="meridianus">
<emphasis id="B978EABEFFAEFF9AA1C0FDC1C452FD6E" box="[997,1146,620,645]" italics="true" pageId="6" pageNumber="370">I. meridianus</emphasis>
</taxonomicName>
the ejaculatory duct joins the prostatic duct at the proximal end of the penis, (4) 
<taxonomicName id="4C0C4D2FFFAEFF9AA0D7FD3DC340FD43" box="[754,872,657,680]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="minimus">
<emphasis id="B978EABEFFAEFF9AA0D7FD3DC340FD43" box="[754,872,657,680]" italics="true" pageId="6" pageNumber="370">I. minimus</emphasis>
</taxonomicName>
differs from the species described here in terms of body size, color pattern, and size of the prostatic vesicle with respect to the seminal vesicle. The body of 
<taxonomicName id="4C0C4D2FFFAEFF9AA703FD19C5B4FD27" box="[1318,1436,693,716]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="minimus">
<emphasis id="B978EABEFFAEFF9AA703FD19C5B4FD27" box="[1318,1436,693,716]" italics="true" pageId="6" pageNumber="370">I. minimus</emphasis>
</taxonomicName>
is relatively small, 3 × 
<quantity id="4CF49B49FFAEFF9AA383FD74C1C4FD04" box="[422,492,728,752]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="370" unit="mm" value="3.0">3 mm</quantity>
, the dorsal surface is light brown with dark grey flecks and brown spots, and the prostatic vesicle is twice the size of the seminal vesicle, while in the species described here seminal vesicle is equal in size with the prostatic vesicle; (5) 
<taxonomicName id="4C0C4D2FFFAEFF9AA067FC8DC299FCD3" box="[578,689,799,824]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="pulcher">
<emphasis id="B978EABEFFAEFF9AA067FC8DC299FCD3" box="[578,689,799,824]" italics="true" pageId="6" pageNumber="370">I. pulcher</emphasis>
</taxonomicName>
differs from the species presented here in terms of color pattern, numerous tentacular and frontal eyes, and marginal eyes few and scattered in the posterior half of the body, while in the present species tentacular eyes numbered 18–21 per tentacle, with about 40 frontal eye spots, and marginal eyes extending well posteriorly as two visible rows. 
<taxonomicName id="4C0C4D2FFFAEFF9AA0C1FC21C379FC4E" box="[740,849,908,933]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="pulcher">
<emphasis id="B978EABEFFAEFF9AA0C1FC21C379FC4E" box="[740,849,908,933]" italics="true" pageId="6" pageNumber="370">I. pulcher</emphasis>
</taxonomicName>
has a pyriform male antrum and an extremely small penis papilla, while in our species the male antrum is deep and narrow, and the penis papilla is not extremely small (170 µm long × 250 µm wide); (6) 
<taxonomicName id="4C0C4D2FFFAEFF9AA00DFC79C2BEFC07" box="[552,662,980,1004]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="refertus">
<emphasis id="B978EABEFFAEFF9AA00DFC79C2BEFC07" box="[552,662,980,1004]" italics="true" pageId="6" pageNumber="370">I. refertus</emphasis>
</taxonomicName>
has a dorsal surface which is brown with a bright orange mesh, few cerebral eyes that differ in number and distribution from our species; (7) 
<taxonomicName id="4C0C4D2FFFAEFF9AA1D0FC55C466FBFB" box="[1013,1102,1015,1040]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="zebra">
<emphasis id="B978EABEFFAEFF9AA1D0FC55C466FBFB" box="[1013,1102,1015,1040]" italics="true" pageId="6" pageNumber="370">I. zebra</emphasis>
</taxonomicName>
is clearly different from the species presented here in terms of color pattern, number of tentacular eyes, frontal eyes extending to the anterior margin, having a spherical and erect prostatic vesicle, while the prostatic vesicle in the species described here is elongated, oval and horizontal; (8) 
<taxonomicName id="4C0C4D2FFFAEFF9AA001FBC9C2F6FB96" box="[548,734,1124,1149]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="mediterraneus">
<emphasis id="B978EABEFFAEFF9AA001FBC9C2F6FB96" box="[548,734,1124,1149]" italics="true" pageId="6" pageNumber="370">I. mediterraneus</emphasis>
</taxonomicName>
is elongated and oval unlike the species presented here that is broadly oval in form, the cerebral eyes number 50 eyes in each cluster, the prostatic vesicle is nearly twice the size of the seminal vesicle, the ejaculatory duct joins distally to the prostatic duct, while in our species the cerebral eyes numbered 91 eyes; the prostatic vesicle is equal in size to the seminal vesicle and the ejaculatory duct joins the prostatic duct in the middle of the penis papilla; (8) 
<taxonomicName id="4C0C4D2FFFAEFF9AA0C0FB59C346FAE6" box="[741,878,1268,1293]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="oculiferus">
<emphasis id="B978EABEFFAEFF9AA0C0FB59C346FAE6" box="[741,878,1268,1293]" italics="true" pageId="6" pageNumber="370">I. oculiferus</emphasis>
</taxonomicName>
, as 
<typeStatus id="54B7880EFFAEFF9AA1B9FB59C3E4FAE6" box="[924,972,1269,1293]" pageId="6" pageNumber="370">type</typeStatus>
of the genus, is dorsally cream to brown with red or pink spots, with a moderately thick muscular coat and a pyriform prostatic vesicle, an ejaculatory duct which joins the prostatic duct at the base of penis, and with a spermiducal bulb (
<bibRefCitation id="EF9D4B5DFFAEFF9AA608FA90C4EDFABF" author="Hyman" box="[1069,1221,1340,1364]" pageId="6" pageNumber="370" refString="Hyman, L. H. (1940) The polyclad flatworms of the Atlantic coast of the United States and Canada. Proceedings of the U. S. National Museum, 89, 449 - 495. http: // dx. doi. org / 10.5479 / si. 00963801.89 - 3101.449" type="journal article" year="1940">Hyman 1940</bibRefCitation>
), while the species described here has an elongated oval and highly musculature prostatic vesicle,, and lacks a spermiducal bulb.
</paragraph>
<paragraph id="8BB336ACFFAEFF9AA2E2FA28C52BF99E" blockId="6.[151,1436,151,2013]" pageId="6" pageNumber="370">
Compared with the Indo-West Pacific species of 
<taxonomicName id="4C0C4D2FFFAEFF9AA0D4FA29C365FA77" box="[753,845,1413,1436]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFAEFF9AA0D4FA29C365FA77" box="[753,845,1413,1436]" italics="true" pageId="6" pageNumber="370">Imogine</emphasis>
</taxonomicName>
, the species described here is distinguished from all other congeners except 
<taxonomicName id="4C0C4D2FFFAEFF9AA387FA05C219FA2B" box="[418,561,1447,1472]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="pardalotus">
<emphasis id="B978EABEFFAEFF9AA387FA05C219FA2B" box="[418,561,1447,1472]" italics="true" pageId="6" pageNumber="370">I. pardalotus</emphasis>
</taxonomicName>
by: having cerebral eyes as two parallel clusters, while the Indo-West Pacific species described by 
<bibRefCitation id="EF9D4B5DFFAEFF9AA3AAFA60C2D2FA0F" author="Jennings" box="[399,762,1484,1509]" pageId="6" pageNumber="370" refString="Jennings, K. A. &amp; Newman, L. J. (1996 a) Four new Stylochid flatworms (Platyhelminthes: Polycladida) associated with commercial oysters from Moreton Bay, southeast Queensland, Australia. The Raffles Bulletin of Zoology, 44, 493 - 508." type="journal article" year="1996" yearSuffix="a">Jennings and Newman (1996 a</bibRefCitation>
, 
<bibRefCitation id="EF9D4B5DFFAEFF9AA12CFA60C330FA0E" author="Jennings" box="[777,792,1484,1509]" pageId="6" pageNumber="370" refString="Jennings, K. A. &amp; Newman, L. J. (1996 b) Two new Stylochid flatworms (Platyhelminthes: Polycladida) from the southern Great Barrier Reef, Australia. The Raffles Bulletin of Zoology, 44, 135 - 142." type="journal article" year="1996" yearSuffix="b">b</bibRefCitation>
) have a cerebral eyes as single cluster. The number of tentacular eyes, 18–21 per tentacle, of the species described here is considerably lower than all of the species of 
<taxonomicName id="4C0C4D2FFFAEFF9AA2B2F9B9C0DBF9C7" box="[151,243,1557,1580]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFAEFF9AA2B2F9B9C0DBF9C7" box="[151,243,1557,1580]" italics="true" pageId="6" pageNumber="370">Imogine</emphasis>
</taxonomicName>
from the Indo-West Pacific. As indicated in Table 1, the number of tentacular eyes per tentacle is: 100 eyes for 
<taxonomicName id="4C0C4D2FFFAEFF9AA325F995C177F9BB" box="[256,351,1591,1616]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="kimae">
<emphasis id="B978EABEFFAEFF9AA325F995C177F9BB" box="[256,351,1591,1616]" italics="true" pageId="6" pageNumber="370">I. kimae</emphasis>
</taxonomicName>
, 50 eyes for 
<taxonomicName id="4C0C4D2FFFAEFF9AA027F995C24CF9BB" box="[514,612,1591,1616]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="lesteri">
<emphasis id="B978EABEFFAEFF9AA027F995C24CF9BB" box="[514,612,1591,1616]" italics="true" pageId="6" pageNumber="370">I. lesteri</emphasis>
</taxonomicName>
, 30 eyes for 
<taxonomicName id="4C0C4D2FFFAEFF9AA12DF995C3A4F9BB" box="[776,908,1591,1616]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="mcgrathi">
<emphasis id="B978EABEFFAEFF9AA12DF995C339F9BB" box="[776,785,1593,1616]" italics="true" pageId="6" pageNumber="370">I</emphasis>
. 
<emphasis id="B978EABEFFAEFF9AA101F99BC3A4F9BB" box="[804,908,1591,1616]" italics="true" pageId="6" pageNumber="370">mcgrathi</emphasis>
</taxonomicName>
, 50 eyes for 
<taxonomicName id="4C0C4D2FFFAEFF9AA609F995C485F9BB" box="[1068,1197,1593,1616]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="meganae">
<emphasis id="B978EABEFFAEFF9AA609F995C485F9BB" box="[1068,1197,1593,1616]" italics="true" pageId="6" pageNumber="370">I. meganae</emphasis>
</taxonomicName>
and 100 eyes for 
<emphasis id="B978EABEFFAEFF9AA7A9F995C10EF99F" italics="true" pageId="6" pageNumber="370">
<taxonomicName id="4C0C4D2FFFAEFF9AA7A9F995C13BF99E" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="pardalotus">I. pardalotus</taxonomicName>
. 
<taxonomicName id="4C0C4D2FFFAEFF9AA338F9F1C10EF99F" box="[285,294,1629,1652]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="mcgrathi">I</taxonomicName>
</emphasis>
. 
<emphasis id="B978EABEFFAEFF9AA311F9F0C1B1F99E" box="[308,409,1628,1653]" italics="true" pageId="6" pageNumber="370">mcgrathi</emphasis>
also differs from the species described here by having numerous frontal eyes.
</paragraph>
<paragraph id="8BB336ACFFAEFF9AA2E2F92DC10DF89B" blockId="6.[151,1436,151,2013]" pageId="6" pageNumber="370">
<emphasis id="B978EABEFFAEFF9AA2E2F92DC148F973" box="[199,352,1663,1688]" italics="true" pageId="6" pageNumber="370">
<taxonomicName id="4C0C4D2FFFAEFF9AA2E2F92DC173F973" box="[199,347,1663,1688]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="pardalotus">I. pardalotus</taxonomicName>
,
</emphasis>
unlike the species described here, has an elongated oval body, 15 × 
<quantity id="4CF49B49FFAEFF9AA65FF9D3C497F973" box="[1146,1215,1663,1688]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="370" unit="mm" value="6.0">6 mm</quantity>
, the dorsal color is beige with a leopard spotted pattern of greenish brown spots, irregular in shape and color, with elongate tentacles and the ejaculatory duct joins the prostatic duct at the proximal end of penis papilla (Table 1). 
<taxonomicName id="4C0C4D2FFFAEFF9AA6C8F965C544F90B" box="[1261,1388,1737,1760]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="meganae">
<emphasis id="B978EABEFFAEFF9AA6C8F965C544F90B" box="[1261,1388,1737,1760]" italics="true" pageId="6" pageNumber="370">I. meganae</emphasis>
</taxonomicName>
has yellow margins, with elongate yellow tentacles, and is clearly distinguished from the species described here that has no marginal band and small, retractile, white tentacles. 
<taxonomicName id="4C0C4D2FFFAEFF9AA160F8BDC38AF8C3" box="[837,930,1807,1832]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="kimae">
<emphasis id="B978EABEFFAEFF9AA160F8BDC38AF8C3" box="[837,930,1807,1832]" italics="true" pageId="6" pageNumber="370">I. kimae</emphasis>
</taxonomicName>
has a prostatic vesicle twice the size of the seminal vesicle and the ejaculatory duct joins the prostatic duct at the proximal end of the penis papilla, unlike our new species.
</paragraph>
<paragraph id="8BB336ACFFAEFF9AA2E2F8D0C39DF837" blockId="6.[151,1436,151,2013]" pageId="6" pageNumber="370">
Based on the 
<figureCitation id="13372A29FFAEFF9AA348F8D0C1DDF87F" box="[365,501,1916,1941]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1260,1282]" captionStartId-1="1.[151,250,1858,1880]" captionStartId-2="3.[151,248,1833,1855]" captionStartId-3="4.[151,248,1389,1411]" captionTargetBox-0="[163,1436,376,1218]" captionTargetBox-1="[169,1433,1368,1825]" captionTargetBox-2="[159,1426,276,1806]" captionTargetBox-3="[173,1436,193,1363]" captionTargetId-0="figure@1.[157,1442,373,1221]" captionTargetId-1="figure@1.[169,1454,1362,1827]" captionTargetId-2="figure@3.[151,1436,265,1811]" captionTargetId-3="figure@4.[161,1446,193,1367]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Map of the West Indo-Pacific and Persian Gulf. Type locality is Hamun jetty beach, Qeshm Island, the northern coasts of the Persian Gulf." captionText-1="FIGURE 2. (A) Type locality of Imogine qeshmensis sp. nov., rocky substrate with rock oysters (B) Living specimen of I. qeshmensis." captionText-2="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." captionText-3="FIGURE 4. Haematoxylin-eosin stained sections of Imogine qeshmensis sp. nov. (Holotype, ZUTC Platy. 1265): (A) Dorsal epidermis; (B) ventral epidermis (C) male reproductive structures; (D) close up of prostatic vesicle and ejaculatory duct. Abbreviations: bm, basement membrane; cm, circular muscles; cma, ciliated male antrum; de, dorsal epidermis; dm, diagonal muscle; ed, ejaculatory duct; exc, extra-vesicular gland canal; exg, extra-vesicular gland, gp, granular pigment; gl, glandular lining; lm, longitudinal muscle; ml, muscular layer; pv, prostatic vesicle; rb, rhabdite; sv, seminal vesicle; tm, transverse muscle; ve, ventral epidermis; vl, ventral lobe." httpUri-0="https://zenodo.org/record/228609/files/figure.png" httpUri-1="https://zenodo.org/record/228610/files/figure.png" httpUri-2="https://zenodo.org/record/228611/files/figure.png" httpUri-3="https://zenodo.org/record/228612/files/figure.png" pageId="6" pageNumber="370">Figures 1–4</figureCitation>
of 
<bibRefCitation id="EF9D4B5DFFAEFF9AA003F8D0C3A5F87E" author="Jennings" box="[550,909,1916,1941]" pageId="6" pageNumber="370" refString="Jennings, K. A. &amp; Newman, L. J. (1996 a) Four new Stylochid flatworms (Platyhelminthes: Polycladida) associated with commercial oysters from Moreton Bay, southeast Queensland, Australia. The Raffles Bulletin of Zoology, 44, 493 - 508." type="journal article" year="1996" yearSuffix="a">Jennings and Newman (1996a)</bibRefCitation>
for 
<taxonomicName id="4C0C4D2FFFAEFF9AA1E3F8D1C40CF87E" box="[966,1060,1916,1941]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="kimae">
<emphasis id="B978EABEFFAEFF9AA1E3F8D1C40CF87E" box="[966,1060,1916,1941]" italics="true" pageId="6" pageNumber="370">I. kimae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0C4D2FFFAEFF9AA61DF8D1C491F87E" box="[1080,1209,1916,1941]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="mcgrathi">
<emphasis id="B978EABEFFAEFF9AA61DF8D1C469F87F" box="[1080,1089,1917,1940]" italics="true" pageId="6" pageNumber="370">I</emphasis>
. 
<emphasis id="B978EABEFFAEFF9AA671F8D0C491F87E" box="[1108,1209,1916,1941]" italics="true" pageId="6" pageNumber="370">mcgrathi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0C4D2FFFAEFF9AA6EEF8D1C563F87F" box="[1227,1355,1917,1940]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="meganae">I. meganae</taxonomicName>
and 
<taxonomicName id="4C0C4D2FFFAEFF9AA7A9F8D1C125F852" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="370" phylum="Platyhelminthes" rank="species" species="pardalotus">
<emphasis id="B978EABEFFAEFF9AA7A9F8D1C125F852" italics="true" pageId="6" pageNumber="370">I. pardalotus</emphasis>
</taxonomicName>
, the seminal vesicle is located ventrally to the prostatic vesicle, while in the species described here the seminal vesicle is entirely anterior to the prostatic vesicle (
<figureCitation id="13372A29FFAEFF9AA13DF868C34BF837" box="[792,867,1988,2013]" captionStart="FIGURE 3" captionStartId="3.[151,248,1833,1855]" captionTargetBox="[159,1426,276,1806]" captionTargetId="figure@3.[151,1436,265,1811]" captionTargetPageId="3" captionText="FIGURE 3. Imogine qeshmensis sp. nov. (A) Dorsal view of paratype (sectioned specimen, ZUTC platy 1266); (B) Drawing of a cleared whole specimen; (C) pattern of cerebral and tentacular eyes; (D) Sagittal reconstruction of the male and female reproductive systems. Abbreviations: ce, cerebral eyes; cg, cement glands; cma, ciliated male antrum; ed, ejaculatory duct; exc, extra-vesicular gland canal; fg, female gonpore; gl, glandular lining; i, intestine; m, mouth; me, marginal eyes; mg, male gonopore; o, ovaries; ov, oviduct; pd, prostatic duct; ph, pharynx; pv, prostatic vesicle; sv, seminal vesicle; te, tentacular eyes;; ts, testis; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/228611/files/figure.png" pageId="6" pageNumber="370">Figs. 3</figureCitation>
D, 4C).
</paragraph>
<paragraph id="8BB336ACFFA1FF95A2E2FF3BC3A7FEF6" blockId="9.[151,1436,151,536]" pageId="9" pageNumber="373">
The ejaculatory duct joins the prostatic duct at the proximal end of the penis papilla in 
<taxonomicName id="4C0C4D2FFFA1FF95A681FF35C52AFF5B" box="[1188,1282,151,176]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="9" pageNumber="373" phylum="Platyhelminthes" rank="species" species="lesteri">
<emphasis id="B978EABEFFA1FF95A681FF35C52AFF5B" box="[1188,1282,151,176]" italics="true" pageId="9" pageNumber="373">I. lesteri</emphasis>
</taxonomicName>
, the situation which is not similar to our species. 
<taxonomicName id="4C0C4D2FFFA1FF95A003FF11C2ADFF3F" box="[550,645,189,212]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="9" pageNumber="373" phylum="Platyhelminthes" rank="species" species="exigus">
<emphasis id="B978EABEFFA1FF95A003FF11C2ADFF3F" box="[550,645,189,212]" italics="true" pageId="9" pageNumber="373">I. exigus</emphasis>
</taxonomicName>
is relatively smaller (7 × 
<quantity id="4CF49B49FFA1FF95A182FF10C3C2FF3F" box="[935,1002,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="373" unit="mm" value="4.0">4 mm</quantity>
) than present species (25 × 
<quantity id="4CF49B49FFA1FF95A700FF10C550FF3F" box="[1317,1400,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="373" unit="mm" value="18.0">18 mm</quantity>
). 
<taxonomicName id="4C0C4D2FFFA1FF95A7A9FF11C0F7FF13" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="9" pageNumber="373" phylum="Platyhelminthes" rank="species" species="exigus">
<emphasis id="B978EABEFFA1FF95A7A9FF11C0F7FF13" italics="true" pageId="9" pageNumber="373">I. exigus</emphasis>
</taxonomicName>
also has fewer cerebral eyes (7 eyes) compared with the species described here (about 91 eyes). Based on the above diagnostic features, 
<taxonomicName id="4C0C4D2FFFA1FF95A3CEFEA9C2E4FEF6" box="[491,716,260,285]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="9" pageNumber="373" phylum="Platyhelminthes" rank="species" species="qeshmensis">
<emphasis id="B978EABEFFA1FF95A3CEFEA9C2E4FEF6" box="[491,716,260,285]" italics="true" pageId="9" pageNumber="373">Imogine qeshmensis</emphasis>
</taxonomicName>
is a new species.
</paragraph>
<paragraph id="8BB336ACFFA1FF95A2E2FE84C39CFDF3" blockId="9.[151,1436,151,536]" pageId="9" pageNumber="373">
In terms of habitat, members of the genus 
<taxonomicName id="4C0C4D2FFFA1FF95A087FE85C2D6FEAB" box="[674,766,297,320]" class="Turbellaria" family="Stylochidae" genus="Imogine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polycladida" pageId="9" pageNumber="373" phylum="Platyhelminthes" rank="genus">
<emphasis id="B978EABEFFA1FF95A087FE85C2D6FEAB" box="[674,766,297,320]" italics="true" pageId="9" pageNumber="373">Imogine</emphasis>
</taxonomicName>
are known to be associated with or feed on giant clams and rock oysters (
<bibRefCitation id="EF9D4B5DFFA1FF95A315FEE0C258FE8E" author="Jennings" box="[304,624,332,357]" pageId="9" pageNumber="373" refString="Jennings, K. A. &amp; Newman, L. J. (1996 a) Four new Stylochid flatworms (Platyhelminthes: Polycladida) associated with commercial oysters from Moreton Bay, southeast Queensland, Australia. The Raffles Bulletin of Zoology, 44, 493 - 508." type="journal article" year="1996" yearSuffix="a">Jennings &amp; Newman 1996a</bibRefCitation>
; 
<bibRefCitation id="EF9D4B5DFFA1FF95A058FEE1C34FFE8F" author="Newman" box="[637,871,332,357]" pageId="9" pageNumber="373" refString="Newman, L. J., Cannon, L. R. G. &amp; Govan, H. (1993) Stylochus (Imogene) matatasi n. sp. (Platyhelminthes, Polycladida): pest of cultured giant clams and pearl oysters from Solomon Islands. Hydrobiologia, 257 (3), 185 - 189. http: // dx. doi. org / 10.1007 / bf 00765011" type="journal article" year="1993">
Newman 
<emphasis id="B978EABEFFA1FF95A0CEFEE1C30CFE8E" box="[747,804,332,357]" italics="true" pageId="9" pageNumber="373">et al.</emphasis>
1993
</bibRefCitation>
; 
<bibRefCitation id="EF9D4B5DFFA1FF95A150FEE0C411FE8E" author="Sluys" box="[885,1081,332,357]" pageId="9" pageNumber="373" refString="Sluys, R., Faubel, A., Rajagopal, S. &amp; van der Velde, G. (2005) A new and alien species of &quot; oyster leech &quot; (Platyhelminthes, Polycladida, Stylochidae) from the brackish North Sea Canal, The Netherlands. Helgoland Marine Research, 59 (4), 310 - 314." type="journal article" year="2005">
Sluys 
<emphasis id="B978EABEFFA1FF95A19EFEE1C3DDFE8E" box="[955,1013,332,357]" italics="true" pageId="9" pageNumber="373">et al.</emphasis>
2005
</bibRefCitation>
), or other invertebrates such as barnacles (Lee 
<emphasis id="B978EABEFFA1FF95A364FEDDC151FE63" box="[321,377,367,392]" italics="true" pageId="9" pageNumber="373">et al.</emphasis>
2006). All specimens of the species described here associated with rock oysters. Our sampling area located at 
<collectingCountry id="F31B763CFFA1FF95A36DFE38C1BFFE46" box="[328,407,404,429]" name="Iran" pageId="9" pageNumber="373">Qeshm</collectingCountry>
Island is a free economic zone. The island is used for mariculture such as cultivation of shellfish and pearl oysters. The presence of the new species described here should be considered by managers of oyster farms because these flatworms kill and consume cultured bivalves, significantly contributing to mortalities (
<bibRefCitation id="EF9D4B5DFFA1FF95A2BBFE53C15CFDF3" author="Galleni" box="[158,372,511,536]" pageId="9" pageNumber="373" refString="Galleni, L., Tongiorgi, P., Ferrero, E. &amp; Salghetti, U. (1980) Stylochus mediterraneus (Turbellaria: Polycladida), predator on the mussel Mytilus galloprovincialis. Marine Biology, 55 (4), 317 - 326." type="journal article" year="1980">
Galleni 
<emphasis id="B978EABEFFA1FF95A2DFFDADC11AFDF3" box="[250,306,511,536]" italics="true" pageId="9" pageNumber="373">et al.</emphasis>
1980
</bibRefCitation>
; 
<bibRefCitation id="EF9D4B5DFFA1FF95A35AFE53C29CFDF3" author="Littlewood" box="[383,692,511,536]" pageId="9" pageNumber="373" refString="Littlewood, D. &amp; Marsbe, L. (1990) Predation on cultivated oysters (Crassostrea rhizophorae) by the polyclad turbellarian flatworm (Stylochus (Stylochus) frontalis) Aquaculture, 88 (2), 145 - 150. http: // dx. doi. org / 10.1016 / 0044 - 8486 (90) 90289 - y" type="journal article" year="1990">Littlewood &amp; Marsbe 1990</bibRefCitation>
; 
<bibRefCitation id="EF9D4B5DFFA1FF95A0E5FDACC38FFDF3" author="Newman" box="[704,935,511,536]" pageId="9" pageNumber="373" refString="Newman, L. J., Cannon, L. R. G. &amp; Govan, H. (1993) Stylochus (Imogene) matatasi n. sp. (Platyhelminthes, Polycladida): pest of cultured giant clams and pearl oysters from Solomon Islands. Hydrobiologia, 257 (3), 185 - 189. http: // dx. doi. org / 10.1007 / bf 00765011" type="journal article" year="1993">
Newman 
<emphasis id="B978EABEFFA1FF95A109FDADC34CFDF3" box="[812,868,511,536]" italics="true" pageId="9" pageNumber="373">et al.</emphasis>
1993
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>