490 lines
64 KiB
XML
490 lines
64 KiB
XML
<document id="47B767DB3519DF000B46A5841C3EF628" ID-CLB-Dataset="8419" ID-DOI="10.11646/zootaxa.4852.4.2" ID-GBIF-Dataset="1430f8df-baac-425a-990d-e9db0b875e5d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4410041" ID-ZooBank="61DA8090-7DC0-4968-9503-C7FB3A113ED2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609596614450" checkinUser="plazi" docAuthor="Hutchinson, Paul M. & Allsopp, Peter G." docDate="2020" docId="039F87D1FFACCF18F5FFF9768E035CB8" docLanguage="en" docName="zootaxa.4852.4.2.pdf" docOrigin="Zootaxa 4852 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epironastes limbatus Carne 1957" docType="treatment" docVersion="7" lastPageNumber="442" masterDocId="FFA6FFA9FFA6CF16F568FFD78B415D4C" masterDocTitle="Revision of Epironastes Carne, 1957 (Coleoptera: Scarabaeidae: Dynastinae Pentodontini: Pseudoryctina) with description of the female, behaviours, and habitats" masterLastPageNumber="448" masterPageNumber="428" pageNumber="438" updateTime="1698851051965" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="05A54F154B40BC0CA3ED253CE7E95BEA">Revision of Epironastes Carne, 1957 (Coleoptera: Scarabaeidae: Dynastinae Pentodontini: Pseudoryctina) with description of the female, behaviours, and habitats</mods:title>
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<mods:namePart id="89EE0D1305F791C514B9357D250A87B6">Hutchinson, Paul M.</mods:namePart>
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<mods:affiliation id="2C50005AB6B83AEECBBC952E1EBB4335">Department of Agriculture and Food WA (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia, 6986, Australia. paul. hutchinson @ dpird. wa. gov. au; https: // orcid. org / 0000 - 0001 - 8747 - 1320</mods:affiliation>
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<taxonomicName id="4C364D44FFACCF1CF5FFF97689745BF6" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[151,565,1696,1723]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="limbatus">
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<emphasis id="B942EAD5FFACCF1CF5FFF97689745BF6" bold="true" box="[151,565,1696,1723]" pageId="10" pageNumber="438">
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<emphasis id="B942EAD5FFACCF1CF5FFF9768AD75BF7" bold="true" box="[151,406,1696,1723]" italics="true" pageId="10" pageNumber="438">Epironastes limbatus</emphasis>
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<bibRefCitation id="EFA74B36FFACCF1CF4F6F97789745BF6" author="Carne, P. B." box="[414,565,1696,1722]" pageId="10" pageNumber="438" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
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<paragraph id="8B8936C7FFACCF1CF5FFF9128A195B93" blockId="10.[151,565,1696,1759]" box="[151,344,1732,1759]" pageId="10" pageNumber="438">
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(
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<figureCitation id="130D2A42FFACCF1CF5F7F9128A665B93" box="[159,295,1733,1759]" captionStart-0="FIGURES 30–31" captionStart-1="FIGURES 32–39" captionStart-2="FIGURE 40" captionStartId-0="11.[151,264,1170,1194]" captionStartId-1="12.[151,264,1943,1967]" captionStartId-2="13.[151,250,1039,1063]" captionTargetBox-0="[172,1416,226,1132]" captionTargetBox-1="[161,1405,189,1920]" captionTargetBox-2="[164,1423,181,1015]" captionTargetId-0="figure-25@11.[151,1436,181,1146]" captionTargetId-1="figure-19@12.[161,1427,181,1920]" captionTargetId-2="figure-25@13.[164,1423,181,1015]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURES 30–31. Epironastes limbatus Carne, 1957, holotype male. 30, dorsum; 31, labels." captionText-1="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." captionText-2="FIGURE 40. Epironastes limbatus Carne, 1957 habitat at 10 km west of Yellowdine." figureDoi-0="http://doi.org/10.5281/zenodo.4410057" figureDoi-1="http://doi.org/10.5281/zenodo.4410059" figureDoi-2="http://doi.org/10.5281/zenodo.4410061" httpUri-0="https://zenodo.org/record/4410057/files/figure.png" httpUri-1="https://zenodo.org/record/4410059/files/figure.png" httpUri-2="https://zenodo.org/record/4410061/files/figure.png" pageId="10" pageNumber="438">Figs. 30–40</figureCitation>
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,
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<figureCitation id="130D2A42FFACCF1CF45BF9138A115B93" box="[307,336,1732,1759]" captionStart="FIGURE 56" captionStartId="18.[151,250,2003,2027]" captionTargetBox="[177,1411,1209,1980]" captionTargetId="figure-471@18.[177,1411,1209,1980]" captionTargetPageId="18" captionText="FIGURE 56. Known distribution of Epironastes species in southern Western Australia." figureDoi="http://doi.org/10.5281/zenodo.4410071" httpUri="https://zenodo.org/record/4410071/files/figure.png" pageId="10" pageNumber="438">56</figureCitation>
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)
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</paragraph>
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<emphasis id="B942EAD5FFACCF1CF5FFF8DD8A2C5A6D" box="[151,365,1801,1825]" italics="true" pageId="10" pageNumber="438">Epironastes limbatus</emphasis>
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<bibRefCitation id="EFA74B36FFACCF1CF41DF8DE89675A6D" author="Carne, P. B." box="[373,550,1801,1825]" pageId="10" pageNumber="438" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957: 126</bibRefCitation>
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</taxonomicName>
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(Fig.
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maxilla and mentum; Fig.
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<specimenCount id="9D30FD4EFFACCF1CF6D3F8DE88BA5A6D" box="[955,1019,1801,1825]" pageId="10" pageNumber="438" type="male">325 ♂</specimenCount>
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head; Fig.
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<specimenCount id="9D30FD4EFFACCF1CF107F8DE8FEE5A6D" box="[1135,1199,1801,1825]" pageId="10" pageNumber="438" type="male">330 ♂</specimenCount>
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parameres; Fig.
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<specimenCount id="9D30FD4EFFACCF1CF034F8DE8EDD5A6D" box="[1372,1436,1801,1825]" pageId="10" pageNumber="438" type="male">332 ♂</specimenCount>
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pygidium; Map 17);
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<bibRefCitation id="EFA74B36FFACCF1CF4F4F8FF89105A0C" author="Endrodi, S." box="[412,593,1832,1856]" pageId="10" pageNumber="438" pagination="17 - 70" refId="ref10164" refString="Endrodi, S. (1974) Monographie der Dynastinae 4. Tribus: Pentodontini von Australien und Neu-Zeeland (Coleoptera). Folia Entomologica Hungarica, Series Nova, 27, 17 - 70." type="journal article" year="1974">Endrődi 1974: 38</bibRefCitation>
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</treatmentCitation>
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, 65 (
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<figureCitation id="130D2A42FFACCF1CF7EDF8FF898F5A0C" box="[645,718,1832,1856]" captionStart="FIGURES 43–50" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[172,1415,217,1919]" captionTargetId="figure-25@15.[172,1415,181,1919]" captionTargetPageId="15" captionText="FIGURES 43–50. Epironastes nigrisetosus Carne, 1957, male, Dempster Inlet specimen. 43, dorsum; 44, venter; 45, lateral; 46, head, lateral; 47, head, dorsal; 48, pygidium; 49, parameres; 50, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410065" httpUri="https://zenodo.org/record/4410065/files/figure.png" pageId="10" pageNumber="438">
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Fig.
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<specimenCount id="9D30FD4EFFACCF1CF7DDF8FF898F5A0C" box="[693,718,1832,1856]" pageId="10" pageNumber="438" type="male">49</specimenCount>
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♂ parameres);
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</treatmentCitation>
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(Fig. 1396
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<specimenCount id="9D30FD4EFFACCF1CF1CFF8FF8FFB5A73" box="[1191,1210,1832,1855]" pageId="10" pageNumber="438" type="male">♂</specimenCount>
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parameres);
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<treatmentCitation id="0A9710D6FFACCF1CF028F8FF8A265A13" author="Cassis, G. & Weir, T. A." page="401" pageId="10" pageNumber="438" year="1992">
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<bibRefCitation id="EFA74B36FFACCF1CF028F8FF8A265A13" author="Cassis, G. & Weir, T. A." pageId="10" pageNumber="438" refId="ref10030" refString="Cassis, G. & Weir, T. A. (1992) Dynastinae. In: Houston, W. W. K. (ed), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Volume 9. Australian Government Publishing Service, Canberra, pp. 383 - 425." type="book" year="1992">Cassis & Weir 1992: 401</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A9710D6FFACCF1CF41AF89189075A12" author="Dechambre, R-P." box="[370,582,1862,1886]" page="68" pageId="10" pageNumber="438" year="2005">
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<bibRefCitation id="EFA74B36FFACCF1CF41AF89189075A12" author="Dechambre, R-P." box="[370,582,1862,1886]" pageId="10" pageNumber="438" refId="ref10085" refString="Dechambre, R-P. (2005) Les Coleopteres du Monde. Vol. 30. Dynastidae Australiens et Oceaniens. Hillside Books, Canterbury, 132 pp." type="book" year="2005">Dechambre 2005: 68</bibRefCitation>
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</treatmentCitation>
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(Fig.
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<specimenCount id="9D30FD4EFFACCF1CF7EBF89089D75A12" box="[643,662,1863,1886]" pageId="10" pageNumber="438" type="male">♂</specimenCount>
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adult; map; Plate 2 Fig.
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<specimenCount id="9D30FD4EFFACCF1CF6F8F89088825A12" box="[912,963,1863,1887]" pageId="10" pageNumber="438" type="male">83 ♂</specimenCount>
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adult);
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<bibRefCitation id="EFA74B36FFACCF1CF179F8918FF55A13" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1041,1204,1862,1887]" pageId="10" pageNumber="438" pagination="1919" refId="ref10501" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae Macleay 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Volume 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton, pp. 516 - 530." type="book chapter" year="2019">
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Weir
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<emphasis id="B942EAD5FFACCF1CF120F8908F3A5A12" box="[1096,1147,1862,1886]" italics="true" pageId="10" pageNumber="438">et al.</emphasis>
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2019
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</bibRefCitation>
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(Plate 54O
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<specimenCount id="9D30FD4EFFACCF1CF045F8908E015A12" box="[1325,1344,1863,1886]" pageId="10" pageNumber="438" type="male">♂</specimenCount>
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adult).
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</paragraph>
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<paragraph id="8B8936C7FFACCF1CF5FFF85A89AA5AA3" blockId="10.[151,1437,1932,2031]" pageId="10" pageNumber="438">
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<emphasis id="B942EAD5FFACCF1CF5FFF85A8A645AEA" bold="true" box="[151,293,1932,1959]" pageId="10" pageNumber="438">Type series.</emphasis>
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<typeStatus id="548D8865FFACCF1CF446F85A8AD75AEB" box="[302,406,1933,1959]" pageId="10" pageNumber="438" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D30FD4EFFACCF1CF4C8F85A8AF45AEB" box="[416,437,1933,1959]" pageId="10" pageNumber="438" type="male">♂</specimenCount>
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(by original designation): Type // Marloo Stn. [
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,
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] Wurarga W.A. 1931-1941 A. Goerling //
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<typeStatus id="548D8865FFACCF1CF4D1F86689605A87" box="[441,545,1969,1995]" pageId="10" pageNumber="438" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D30FD4EFFACCF1CF740F866897C5A87" box="[552,573,1969,1995]" pageId="10" pageNumber="438" type="male">♂</specimenCount>
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<taxonomicName id="4C364D44FFACCF1CF72CF866886C5A87" authorityName="Carne" authorityYear="1957" box="[580,813,1969,1995]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="limbatus">Epironastes limbatus</taxonomicName>
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mihi P.B. Carne det. 1952 //
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||
<materialsCitation id="3B5E3C9AFFACCF1CF11AF86689A65AA3" ID-GBIF-Occurrence="3031275328" collectionCode="ANIC" pageId="10" pageNumber="438" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="548D8865FFACCF1CF11AF8668F9B5A87" box="[1138,1242,1969,1995]" pageId="10" pageNumber="438" type="holotype">Holotype</typeStatus>
|
||
// ANIC image // genitalia on card [in
|
||
<collectionCode id="ED27AE02FFACCF1CF415F8028A855AA3" box="[381,452,2005,2031]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="438">ANIC</collectionCode>
|
||
, examined] (
|
||
<figureCitation id="130D2A42FFACCF1CF73DF802899E5AA3" box="[597,735,2005,2031]" captionStart="FIGURES 30–31" captionStartId="11.[151,264,1170,1194]" captionTargetBox="[172,1416,226,1132]" captionTargetId="figure-25@11.[151,1436,181,1146]" captionTargetPageId="11" captionText="FIGURES 30–31. Epironastes limbatus Carne, 1957, holotype male. 30, dorsum; 31, labels." figureDoi="http://doi.org/10.5281/zenodo.4410057" httpUri="https://zenodo.org/record/4410057/files/figure.png" pageId="10" pageNumber="438">Figs. 30–31</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF49664FFFADCF1DF5FFFB458F0259E6" ID-DOI="http://doi.org/10.5281/zenodo.4410057" ID-Zenodo-Dep="4410057" box="[151,1091,1170,1194]" httpUri="https://zenodo.org/record/4410057/files/figure.png" pageId="11" pageNumber="439" startId="11.[151,264,1170,1194]" targetBox="[172,1416,226,1132]" targetPageId="11">
|
||
<paragraph id="8B8936C7FFADCF1DF5FFFB458F0259E6" blockId="11.[151,1091,1170,1194]" box="[151,1091,1170,1194]" pageId="11" pageNumber="439">
|
||
<emphasis id="B942EAD5FFADCF1DF5FFFB458A1559E6" bold="true" box="[151,340,1170,1194]" pageId="11" pageNumber="439">FIGURES 30–31.</emphasis>
|
||
<taxonomicName id="4C364D44FFADCF1DF432FB4489F559E6" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[346,692,1170,1194]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="439" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B942EAD5FFADCF1DF432FB44897159E6" box="[346,560,1170,1194]" italics="true" pageId="11" pageNumber="439">Epironastes limbatus</emphasis>
|
||
<bibRefCitation id="EFA74B36FFADCF1DF75EFB4589F559E6" author="Carne, P. B." box="[566,692,1170,1194]" pageId="11" pageNumber="439" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype male.
|
||
<emphasis id="B942EAD5FFADCF1DF634FB45883A59E6" bold="true" box="[860,891,1170,1194]" pageId="11" pageNumber="439">30,</emphasis>
|
||
dorsum;
|
||
<emphasis id="B942EAD5FFADCF1DF6B3FB4588B559E6" bold="true" box="[987,1012,1170,1194]" pageId="11" pageNumber="439">31</emphasis>
|
||
, labels.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8936C7FFADCF1DF5AFFB0E89B958EA" blockId="11.[151,1437,1240,2023]" pageId="11" pageNumber="439">
|
||
<typeStatus id="548D8865FFADCF1DF5AFFB0E8A7959BF" box="[199,312,1241,1267]" pageId="11" pageNumber="439" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D30FD4EFFADCF1DF42AFB0E8A2759BF" box="[322,358,1241,1267]" pageId="11" pageNumber="439" type="male">1♂</specimenCount>
|
||
, Cunderdin [
|
||
<geoCoordinate id="EE025000FFADCF1DF49CFB0F892E59BF" box="[500,623,1240,1267]" degrees="31.6510" direction="south" orientation="latitude" pageId="11" pageNumber="439" precision="5" value="-31.651">31.6510°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE025000FFADCF1DF712FB0F884059BF" box="[634,769,1240,1267]" degrees="117.2376" direction="east" orientation="longitude" pageId="11" pageNumber="439" precision="5" value="117.2376">117.2376°E</geoCoordinate>
|
||
] W.A.
|
||
<date id="FF881007FFADCF1DF625FB0F88F759BE" box="[845,950,1240,1266]" pageId="11" pageNumber="439" value="1914-02">Feb.1914</date>
|
||
425 // Isodon Curtus Burm. [
|
||
<taxonomicName id="4C364D44FFADCF1DF192FB0F8AD4585B" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" class="Insecta" family="Dynastidae" genus="Trissodon" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="439" phylum="Arthropoda" rank="species" species="curtus">
|
||
<emphasis id="B942EAD5FFADCF1DF192FB0F8BF9585A" italics="true" pageId="11" pageNumber="439">Trissodon curtus</emphasis>
|
||
(Burmeister, 1847)
|
||
</taxonomicName>
|
||
] Id by A.M. Lea //
|
||
<typeStatus id="548D8865FFADCF1DF704FB2A898F585B" box="[620,718,1277,1303]" pageId="11" pageNumber="439" type="paratype">Paratype</typeStatus>
|
||
<specimenCount id="9D30FD4EFFADCF1DF7BDFB2A89AB585B" box="[725,746,1277,1303]" pageId="11" pageNumber="439" type="male">♂</specimenCount>
|
||
<taxonomicName id="4C364D44FFADCF1DF799FB2B889A585A" authorityName="Carne" authorityYear="1957" box="[753,987,1276,1302]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="439" phylum="Arthropoda" rank="species" species="limbatus">Epironastes limbatus</taxonomicName>
|
||
mihi P.B. Carne det. 1952 //
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF048FB2A89415877" ID-GBIF-Occurrence="3031275331" collectionCode="WAM" pageId="11" pageNumber="439" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D30FD4EFFADCF1DF048FB2A8E74585B" box="[1312,1333,1277,1303]" pageId="11" pageNumber="439" type="male">♂</specimenCount>
|
||
<typeStatus id="548D8865FFADCF1DF054FB2A8EDD585B" box="[1340,1436,1277,1303]" pageId="11" pageNumber="439" type="paratype">paratype</typeStatus>
|
||
// mouthparts on card [in
|
||
<collectionCode id="ED27AE02FFADCF1DF4C7FAF78ABB5877" box="[431,506,1312,1339]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="439" type="Museum">WAM</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D30FD4EFFADCF1DF763FAF6896E5877" box="[523,559,1313,1339]" pageId="11" pageNumber="439" type="male">1♂</specimenCount>
|
||
, 37-814 Kukerin [
|
||
<geoCoordinate id="EE025000FFADCF1DF791FAF688355877" box="[761,884,1313,1339]" degrees="33.1919" direction="south" orientation="latitude" pageId="11" pageNumber="439" precision="5" value="-33.1919">33.1919°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE025000FFADCF1DF617FAF78F465877" box="[895,1031,1312,1339]" degrees="118.0818" direction="east" orientation="longitude" pageId="11" pageNumber="439" precision="5" value="118.0818">118.0818°E</geoCoordinate>
|
||
] // W.A. Mus A. Douglas //
|
||
<typeStatus id="548D8865FFADCF1DF052FAF68EDD5877" box="[1338,1436,1313,1339]" pageId="11" pageNumber="439" type="paratype">Paratype</typeStatus>
|
||
//
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF5C6FA9289B558EA" ID-GBIF-Occurrence="3031275312" collectionCode="WAM" country="Australia" determinerName="P. B. Carne" location="Western Australia" pageId="11" pageNumber="439" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<typeStatus id="548D8865FFADCF1DF5C6FA928A515813" box="[174,272,1349,1375]" pageId="11" pageNumber="439" type="paratype">Paratype</typeStatus>
|
||
<specimenCount id="9D30FD4EFFADCF1DF47FFA928A6D5813" box="[279,300,1349,1375]" pageId="11" pageNumber="439" type="male">♂</specimenCount>
|
||
<taxonomicName id="4C364D44FFADCF1DF45BFA93895C5812" authorityName="Carne" authorityYear="1957" box="[307,541,1348,1374]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="439" phylum="Arthropoda" rank="species" species="limbatus">Epironastes limbatus</taxonomicName>
|
||
mihi
|
||
<determinerName id="7E55DA03FFADCF1DF737FA9289975813" box="[607,726,1349,1375]" pageId="11" pageNumber="439">P.B. Carne</determinerName>
|
||
det. 1952 // mouthparts on card [in
|
||
<collectionCode id="ED27AE02FFADCF1DF100FA938FF25813" box="[1128,1203,1348,1375]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="439" type="Museum">WAM</collectionCode>
|
||
].
|
||
<bibRefCitation id="EFA74B36FFADCF1DF1ACFA928E195813" author="Carne, P. B." box="[1220,1368,1348,1375]" pageId="11" pageNumber="439" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne (1957)</bibRefCitation>
|
||
listed two additional male
|
||
<typeStatus id="548D8865FFADCF1DF414FABE8AA658CF" box="[380,487,1385,1411]" pageId="11" pageNumber="439" type="paratype">paratypes</typeStatus>
|
||
in SAM from “
|
||
<collectingCountry id="F3217657FFADCF1DF7FDFABF884058CE" box="[661,769,1384,1410]" name="Australia" pageId="11" pageNumber="439">Australia</collectingCountry>
|
||
” and “Swan R.”, respectively; the latter was often used as a general descriptor for southwestern
|
||
<collectingRegion id="49F2F825FFADCF1DF741FA5A89B558EA" box="[553,756,1420,1447]" country="Australia" name="Western Australia" pageId="11" pageNumber="439">Western Australia</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFADCF1DF5AFFA6789055B8B" blockId="11.[151,1437,1240,2023]" pageId="11" pageNumber="439">
|
||
<emphasis id="B942EAD5FFADCF1DF5AFFA67894A5886" bold="true" box="[199,523,1456,1482]" pageId="11" pageNumber="439">Other specimens examined.</emphasis>
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF767FA668ED958A3" ID-GBIF-Occurrence="3031275333" collectingDate="1972-03-21" collectionCode="DAFWA" collectorName="K. T. Richards" country="Australia" latitude="-29.4399" location="Western Australia" longLatPrecision="6" longitude="116.2916" municipality="Perenjori" pageId="11" pageNumber="439" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<collectingCountry id="F3217657FFADCF1DF767FA6689F05887" box="[527,689,1457,1483]" name="Australia" pageId="11" pageNumber="439">AUSTRALIA</collectingCountry>
|
||
.
|
||
<collectingRegion id="49F2F825FFADCF1DF7D1FA6688A25887" box="[697,995,1457,1483]" country="Australia" name="Western Australia" pageId="11" pageNumber="439">WESTERN AUSTRALIA</collectingRegion>
|
||
:
|
||
<specimenCount id="9D30FD4EFFADCF1DF683FA668F4E5887" box="[1003,1039,1457,1483]" pageId="11" pageNumber="439" type="male">1♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BEDACBDFFADCF1DF171FA678FC15886" box="[1049,1152,1456,1482]" pageId="11" pageNumber="439">Perenjori</collectingMunicipality>
|
||
[
|
||
<geoCoordinate id="EE025000FFADCF1DF1E3FA668E475887" box="[1163,1286,1457,1483]" degrees="29.4399" direction="south" orientation="latitude" pageId="11" pageNumber="439" precision="5" value="-29.4399">29.4399°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE025000FFADCF1DF067FA678ED75887" box="[1295,1430,1456,1483]" degrees="116.2916" direction="east" orientation="longitude" pageId="11" pageNumber="439" precision="5" value="116.2916">116.2916°E</geoCoordinate>
|
||
]
|
||
<collectorName id="26C35311FFADCF1DF5FFFA028BB858A3" box="[151,249,1493,1519]" pageId="11" pageNumber="439">W. Aust.</collectorName>
|
||
<date id="FF881007FFADCF1DF468FA038A3858A2" box="[256,377,1492,1518]" pageId="11" pageNumber="439" value="1972-03-21">
|
||
<collectingDate id="EFCCE9EFFFADCF1DF468FA038A3858A2" box="[256,377,1492,1518]" pageId="11" pageNumber="439" value="1972-03-21">21.iii.1972</collectingDate>
|
||
</date>
|
||
<collectorName id="26C35311FFADCF1DF4E9FA02895F58A2" box="[385,542,1492,1519]" pageId="11" pageNumber="439">K.T. Richards</collectorName>
|
||
//
|
||
<collectorName id="26C35311FFADCF1DF754FA0389FC58A2" box="[572,701,1492,1518]" pageId="11" pageNumber="439">Agriculture</collectorName>
|
||
(Dept)
|
||
<location id="8EE9601CFFADCF1DF67DFA02889F58A2" LSID="urn:lsid:plazi:treatment:039F87D1FFACCF18F5FFF9768E035CB8:8EE9601CFFADCF1DF67DFA02889F58A2" box="[789,990,1492,1519]" country="Australia" latitude="-29.4399" longLatPrecision="6" longitude="116.2916" municipality="Perenjori" name="Western Australia" pageId="11" pageNumber="439" stateProvince="Western Australia">Western Australia</location>
|
||
9071 // genitalia on card [in
|
||
<collectionCode id="ED27AE02FFADCF1DF040FA038ED358A3" box="[1320,1426,1492,1519]" pageId="11" pageNumber="439">DAFWA</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF5FFFA2E8F805B5E" ID-GBIF-Occurrence="3031275319" box="[151,1217,1528,1555]" collectingDate="2016-01-26" collectorName="W. Aust. & P. Hutchinson" country="Australia" latitude="-31.3325" location="Southern Cross" longLatPrecision="6" longitude="118.9689" pageId="11" pageNumber="439" specimenCount="11" specimenCount-male="11" stateProvince="Western Australia">
|
||
<specimenCount id="9D30FD4EFFADCF1DF5FFFA2E8B885B5F" box="[151,201,1529,1555]" pageId="11" pageNumber="439" type="male">11♂</specimenCount>
|
||
,
|
||
<quantity id="4CCE9B22FFADCF1DF5BCFA2F8A5B5B5E" box="[212,282,1528,1554]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="11" pageNumber="439" unit="km" value="36.0">36 km</quantity>
|
||
W. of
|
||
<location id="8EE9601CFFADCF1DF40AFA2F894F5B5F" LSID="urn:lsid:plazi:treatment:039F87D1FFACCF18F5FFF9768E035CB8:8EE9601CFFADCF1DF40AFA2F894F5B5F" box="[354,526,1528,1555]" country="Australia" latitude="-31.3325" longLatPrecision="6" longitude="118.9689" name="Southern Cross" pageId="11" pageNumber="439" stateProvince="Western Australia">Southern Cross</location>
|
||
[
|
||
<geoCoordinate id="EE025000FFADCF1DF772FA2F89D45B5F" box="[538,661,1528,1555]" degrees="31.3325" direction="south" orientation="latitude" pageId="11" pageNumber="439" precision="5" value="-31.3325">31.3325°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE025000FFADCF1DF7C8FA2F88665B5F" box="[672,807,1528,1555]" degrees="118.9689" direction="east" orientation="longitude" pageId="11" pageNumber="439" precision="5" value="118.9689">118.9689°E</geoCoordinate>
|
||
]
|
||
<collectorName id="26C35311FFADCF1DF65BFA2E88D25B5F" box="[819,915,1529,1555]" pageId="11" pageNumber="439">W. Aust.</collectorName>
|
||
<date id="FF881007FFADCF1DF6F1FA2F8F615B5E" box="[921,1056,1528,1554]" pageId="11" pageNumber="439" value="2016-01-26">
|
||
<collectingDate id="EFCCE9EFFFADCF1DF6F1FA2F8F615B5E" box="[921,1056,1528,1554]" pageId="11" pageNumber="439" value="2016-01-26">26.Jan.2016</collectingDate>
|
||
</date>
|
||
<collectorName id="26C35311FFADCF1DF14EFA2E8F805B5E" box="[1062,1217,1528,1555]" pageId="11" pageNumber="439">P. Hutchinson</collectorName>
|
||
</materialsCitation>
|
||
|
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF1BBFA2E8AD25B17" ID-GBIF-Occurrence="3031275306" collectionCode="PMH, QM, WAM" country="Australia" location="Dyn" municipality="Coll" pageId="11" pageNumber="439" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D30FD4EFFADCF1DF1BBFA2E8FA95B5F" box="[1235,1256,1529,1555]" pageId="11" pageNumber="439" type="male">♂</specimenCount>
|
||
in storm puddle // PMH
|
||
<collectingMunicipality id="6BEDACBDFFADCF1DF59AF9CB8A635B7A" box="[242,290,1564,1590]" pageId="11" pageNumber="439">Coll</collectingMunicipality>
|
||
#
|
||
<location id="8EE9601CFFADCF1DF450F9CA8A295B7B" LSID="urn:lsid:plazi:treatment:039F87D1FFACCF18F5FFF9768E035CB8:8EE9601CFFADCF1DF450F9CA8A295B7B" box="[312,360,1565,1591]" country="Australia" municipality="Coll" name="Dyn" pageId="11" pageNumber="439" stateProvince="Western Australia">Dyn</location>
|
||
1693, 1694, 1695, 1703 [genitalia on cards], 1701, 1702, 1697, 1698 [in
|
||
<collectionCode id="ED27AE02FFADCF1DF1CCF9CB8FA75B7B" box="[1188,1254,1564,1591]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14877" name="City Museum and Records Office" pageId="11" pageNumber="439" type="Herbarium">PMH</collectionCode>
|
||
], 1696 [in
|
||
<collectionCode id="ED27AE02FFADCF1DF037F9CB8ED05B7B" box="[1375,1425,1564,1591]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="439" type="Museum">QM</collectionCode>
|
||
], 1699, 1700 [in
|
||
<collectionCode id="ED27AE02FFADCF1DF42AF9978ACC5B17" box="[322,397,1600,1627]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="439" type="Museum">WAM</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF4F7F9968BFF5B33" ID-GBIF-Occurrence="3031275324" collectingDate="2016-01-26" collectorName="West. Aust." country="Australia" latitude="-31.2773" location="Yellowdine" longLatPrecision="6" longitude="119.5551" pageId="11" pageNumber="439" specimenCount="7" specimenCount-male="7" stateProvince="Western Australia">
|
||
<specimenCount id="9D30FD4EFFADCF1DF4F7F9968A855B17" box="[415,452,1601,1627]" pageId="11" pageNumber="439" type="male">7♂</specimenCount>
|
||
,
|
||
<quantity id="4CCE9B22FFADCF1DF4B8F99689565B16" box="[464,535,1600,1627]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="439" unit="km" value="10.0">10 km</quantity>
|
||
W. of
|
||
<location id="8EE9601CFFADCF1DF709F99789A05B16" LSID="urn:lsid:plazi:treatment:039F87D1FFACCF18F5FFF9768E035CB8:8EE9601CFFADCF1DF709F99789A05B16" box="[609,737,1600,1626]" country="Australia" latitude="-31.2773" longLatPrecision="6" longitude="119.5551" name="Yellowdine" pageId="11" pageNumber="439" stateProvince="Western Australia">Yellowdine</location>
|
||
[
|
||
<geoCoordinate id="EE025000FFADCF1DF786F99688285B17" box="[750,873,1601,1627]" degrees="31.2773" direction="south" orientation="latitude" pageId="11" pageNumber="439" precision="5" value="-31.2773">31.2773°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE025000FFADCF1DF61DF99788BD5B17" box="[885,1020,1600,1627]" degrees="119.5551" direction="east" orientation="longitude" pageId="11" pageNumber="439" precision="5" value="119.5551">119.5551°E</geoCoordinate>
|
||
]
|
||
<collectorName id="26C35311FFADCF1DF161F9968FC85B17" box="[1033,1161,1601,1627]" pageId="11" pageNumber="439">West. Aust.</collectorName>
|
||
<date id="FF881007FFADCF1DF1F8F9978E565B16" box="[1168,1303,1600,1626]" pageId="11" pageNumber="439" value="2016-01-26">
|
||
<collectingDate id="EFCCE9EFFFADCF1DF1F8F9978E565B16" box="[1168,1303,1600,1626]" pageId="11" pageNumber="439" value="2016-01-26">26.Jan.2016</collectingDate>
|
||
</date>
|
||
P. Hutchinson
|
||
</materialsCitation>
|
||
|
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF5B0F9B28BA05BEF" ID-GBIF-Occurrence="3031275332" collectionCode="PMH" country="Australia" location="Dyn" municipality="Coll" pageId="11" pageNumber="439" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D30FD4EFFADCF1DF5B0F9B28BAC5B33" box="[216,237,1637,1663]" pageId="11" pageNumber="439" type="male">♂</specimenCount>
|
||
in storm puddle // PMH
|
||
<collectingMunicipality id="6BEDACBDFFADCF1DF771F9B389085B32" box="[537,585,1636,1662]" pageId="11" pageNumber="439">Coll</collectingMunicipality>
|
||
#
|
||
<location id="8EE9601CFFADCF1DF709F9B289D05B33" LSID="urn:lsid:plazi:treatment:039F87D1FFACCF18F5FFF9768E035CB8:8EE9601CFFADCF1DF709F9B289D05B33" box="[609,657,1637,1663]" country="Australia" municipality="Coll" name="Dyn" pageId="11" pageNumber="439" stateProvince="Western Australia">Dyn</location>
|
||
1704, 1708 [genitalia on cards], 1705, 1706, 1707, 1711, 1712 [in
|
||
<collectionCode id="ED27AE02FFADCF1DF5FFF95F8B9B5BEF" box="[151,218,1672,1699]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14877" name="City Museum and Records Office" pageId="11" pageNumber="439" type="Herbarium">PMH</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF584F95E88A35BEE" ID-GBIF-Occurrence="3031275342" box="[236,994,1672,1699]" collectingDate="2016-01-26" collectorName="P. Hutchinson" country="Australia" location="Yellowdine West. Aust." pageId="11" pageNumber="439" specimenCount="2" specimenCount-male="2" stateProvince="Western Australia">
|
||
<specimenCount id="9D30FD4EFFADCF1DF584F95E8A4E5BEF" box="[236,271,1673,1699]" pageId="11" pageNumber="439" type="male">2♂</specimenCount>
|
||
<quantity id="4CCE9B22FFADCF1DF47FF95E8A1F5BEE" box="[279,350,1672,1699]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="439" unit="km" value="10.0">10 km</quantity>
|
||
W. of
|
||
<location id="8EE9601CFFADCF1DF4C1F95F89F05BEF" LSID="urn:lsid:plazi:treatment:039F87D1FFACCF18F5FFF9768E035CB8:8EE9601CFFADCF1DF4C1F95F89F05BEF" box="[425,689,1672,1699]" country="Australia" name="Yellowdine West. Aust." pageId="11" pageNumber="439" stateProvince="Western Australia">Yellowdine West. Aust.</location>
|
||
<date id="FF881007FFADCF1DF7DFF95F887E5BEE" box="[695,831,1672,1698]" pageId="11" pageNumber="439" value="2016-01-26">
|
||
<collectingDate id="EFCCE9EFFFADCF1DF7DFF95F887E5BEE" box="[695,831,1672,1698]" pageId="11" pageNumber="439" value="2016-01-26">26.Jan.2016</collectingDate>
|
||
</date>
|
||
<collectorName id="26C35311FFADCF1DF62EF95E88A35BEE" box="[838,994,1672,1699]" pageId="11" pageNumber="439">P. Hutchinson</collectorName>
|
||
</materialsCitation>
|
||
|
|
||
<materialsCitation id="3B5E3C9AFFADCF1DF69FF95E89005B8B" ID-GBIF-Occurrence="3031275330" collectionCode="MHC" country="Australia" location="Dyn" municipality="Coll" pageId="11" pageNumber="439" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D30FD4EFFADCF1DF69FF95E8F4D5BEF" box="[1015,1036,1673,1699]" pageId="11" pageNumber="439" type="male">♂</specimenCount>
|
||
in storm puddle // PMH
|
||
<collectingMunicipality id="6BEDACBDFFADCF1DF04EF95F8E175BEE" box="[1318,1366,1672,1698]" pageId="11" pageNumber="439">Coll</collectingMunicipality>
|
||
#
|
||
<location id="8EE9601CFFADCF1DF004F95E8EDD5BEF" LSID="urn:lsid:plazi:treatment:039F87D1FFACCF18F5FFF9768E035CB8:8EE9601CFFADCF1DF004F95E8EDD5BEF" box="[1388,1436,1673,1699]" country="Australia" municipality="Coll" name="Dyn" pageId="11" pageNumber="439" stateProvince="Western Australia">Dyn</location>
|
||
1709[10347], 1710[10348] [in
|
||
<collectionCode id="ED27AE02FFADCF1DF49CF97B897B5B8B" box="[500,570,1708,1735]" pageId="11" pageNumber="439">MHC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C654CFFADCF1BF5AFF9078AE35B9F" lastPageId="13" lastPageNumber="441" pageId="11" pageNumber="439" type="description">
|
||
<paragraph id="8B8936C7FFADCF1BF5AFF9078E3258B0" blockId="11.[151,1437,1240,2023]" lastBlockId="13.[151,1437,1109,2035]" lastPageId="13" lastPageNumber="441" pageId="11" pageNumber="439">
|
||
<emphasis id="B942EAD5FFADCF1DF5AFF90789705BA7" bold="true" box="[199,561,1744,1771]" pageId="11" pageNumber="439">
|
||
Redescription. Male
|
||
<typeStatus id="548D8865FFADCF1DF4ABF906896D5BA7" box="[451,556,1745,1771]" pageId="11" pageNumber="439" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Body (
|
||
<figureCitation id="130D2A42FFADCF1DF7EFF907884E5BA7" box="[647,783,1744,1771]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="11" pageNumber="439">Figs. 32–33</figureCitation>
|
||
) black, sometimes with red tinge, dorsal half of antennal club reddish brown and ventral half dull fawn,
|
||
<quantity id="4CCE9B22FFADCF1DF7C8F92388425A43" box="[672,771,1780,1807]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="11" pageNumber="439" unit="mm" value="13.5">13.5 mm</quantity>
|
||
long,
|
||
<quantity id="4CCE9B22FFADCF1DF620F92388DC5A43" box="[840,925,1780,1807]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="439" unit="mm" value="7.5">7.5 mm</quantity>
|
||
wide. Mentum strongly and abruptly reflexed through 90°, surface flat, narrowed to ligula; ligula abruptly constricted before divergent and truncate apex. Antennal club (
|
||
<figureCitation id="130D2A42FFADCF1DF468F8EB8AC95A1B" box="[256,392,1852,1879]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="11" pageNumber="439">Figs. 33–36</figureCitation>
|
||
) only slightly longer than shaft, inner face setose, shallowly foveate in apical half, outer face bitexturous. Clypeus (
|
||
<figureCitation id="130D2A42FFADCF1DF4FCF8B789085A37" box="[404,585,1888,1915]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="11" pageNumber="439">Figs. 32, 35–36</figureCitation>
|
||
) gently declivous from low rounded ridge joining antennal angles, lateral ridges distinct, apex arcuate and reflexed, surface rugulose; clypeofrontal suture distinct. Frons (
|
||
<figureCitation id="130D2A42FFADCF1DF1A8F8538E2C5AD3" box="[1216,1389,1924,1951]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="11" pageNumber="439">Figs. 32, 35–36</figureCitation>
|
||
) rugulose; ocular canthi broad, anterior margin obtusely angulate, laterally subequal to margin of eye, surface coarsely punctate with anterior margin bearing setae. Pronotum (
|
||
<figureCitation id="130D2A42FFADCF1DF672F81B88915AAB" box="[794,976,1996,2023]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="11" pageNumber="439">Figs. 32, 34–36</figureCitation>
|
||
) with anterolateral angles forming right angles, widest slightly anterior to midlength, anteromedial tubercle small, preceding a shallow subcircular impression restricted to anterior half and with a shallower depression extending to basal margin, disc weakly convex and without distinct basal convexity, surface with distinct sparse fine punctures becoming larger and denser towards margins, tending transverse in impression. Scutellum (
|
||
<figureCitation id="130D2A42FFABCF1BF799FB1688005990" box="[753,833,1217,1244]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="13" pageNumber="441">Fig. 32</figureCitation>
|
||
) bearing large punctures over entire surface. Each elytron (
|
||
<figureCitation id="130D2A42FFABCF1BF5BCFB328A1A584C" box="[212,347,1253,1280]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="13" pageNumber="441">Figs. 32, 34</figureCitation>
|
||
) with sutural striae linear-punctate, sutural costa distinctly raised, elytral disc with broad, deeply impressed somewhat annulate punctures forming conspicuous striae, intervals with numerous random large punctures, micropunctures and surface microreticulate. Pygidium (
|
||
<figureCitation id="130D2A42FFABCF1BF624FAFA88DC5804" box="[844,925,1325,1352]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="13" pageNumber="441">Fig. 37</figureCitation>
|
||
) moderately convex, concentrically punctate, weakly microreticulate, setose across base, apical ridge glabrous. Abdominal sternites 2–4 (
|
||
<figureCitation id="130D2A42FFABCF1BF1FDFA868FA75820" box="[1173,1254,1361,1388]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="13" pageNumber="441">Fig. 33</figureCitation>
|
||
) laterally setose over entire surface and glabrous medially, sternite 5 laterally setose over entire surface and glabrous across middle, last sternite across entire posterior border with dense long setae. Coxae (
|
||
<figureCitation id="130D2A42FFABCF1BF6D2FA4E8F4B58F8" box="[954,1034,1433,1460]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="13" pageNumber="441">Fig. 33</figureCitation>
|
||
) bearing yellow vestiture; metatibia with basal carina bearing long, acute setae, distal carina with short, stout, closely set ciliae on outer surface; metatibial spurs (
|
||
<figureCitation id="130D2A42FFABCF1BF47BFA368A2558B0" box="[275,356,1505,1532]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="13" pageNumber="441">Fig. 37</figureCitation>
|
||
) of unequal length, long, subparallel, both spurs slightly curved. Genitalia as in
|
||
<figureCitation id="130D2A42FFABCF1BF18FFA368E2F58B0" box="[1255,1390,1505,1532]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="13" pageNumber="441">Figs. 38–39</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF49664FFFAACF1AF5FFF840883F5A98" ID-DOI="http://doi.org/10.5281/zenodo.4410059" ID-Zenodo-Dep="4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="12" pageNumber="440" startId="12.[151,264,1943,1967]" targetBox="[161,1405,189,1920]" targetPageId="12">
|
||
<paragraph id="8B8936C7FFAACF1AF5FFF840883F5A98" blockId="12.[151,1436,1943,2004]" pageId="12" pageNumber="440">
|
||
<emphasis id="B942EAD5FFAACF1AF5FFF8408A145AE3" bold="true" box="[151,341,1943,1967]" pageId="12" pageNumber="440">FIGURES 32–39.</emphasis>
|
||
<taxonomicName id="4C364D44FFAACF1AF433F84F89F65AE3" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[347,695,1943,1968]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="440" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B942EAD5FFAACF1AF433F84F89705AFC" box="[347,561,1944,1968]" italics="true" pageId="12" pageNumber="440">Epironastes limbatus</emphasis>
|
||
<bibRefCitation id="EFA74B36FFAACF1AF751F84F89F65AE3" author="Carne, P. B." box="[569,695,1943,1968]" pageId="12" pageNumber="440" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
, male specimen #1703;
|
||
<emphasis id="B942EAD5FFAACF1AF6C6F84088865AE3" bold="true" box="[942,967,1943,1967]" pageId="12" pageNumber="440">32</emphasis>
|
||
, dorsum;
|
||
<emphasis id="B942EAD5FFAACF1AF147F8408F095AE3" bold="true" box="[1071,1096,1943,1967]" pageId="12" pageNumber="440">33</emphasis>
|
||
, venter;
|
||
<emphasis id="B942EAD5FFAACF1AF1CBF8408FFD5AE3" bold="true" box="[1187,1212,1943,1967]" pageId="12" pageNumber="440">34</emphasis>
|
||
, lateral;
|
||
<emphasis id="B942EAD5FFAACF1AF07FF8408E715AE3" bold="true" box="[1303,1328,1943,1967]" pageId="12" pageNumber="440">35</emphasis>
|
||
, head, lateral;
|
||
<emphasis id="B942EAD5FFAACF1AF5A3F86C8BA55A9F" bold="true" box="[203,228,1979,2003]" pageId="12" pageNumber="440">36</emphasis>
|
||
, head, dorsal;
|
||
<emphasis id="B942EAD5FFAACF1AF410F86C8AD05A9F" bold="true" box="[376,401,1979,2003]" pageId="12" pageNumber="440">37</emphasis>
|
||
, pygidium;
|
||
<emphasis id="B942EAD5FFAACF1AF765F86C89675A9F" bold="true" box="[525,550,1979,2003]" pageId="12" pageNumber="440">38</emphasis>
|
||
, parameres;
|
||
<emphasis id="B942EAD5FFAACF1AF7C0F86C89805A9F" bold="true" box="[680,705,1979,2003]" pageId="12" pageNumber="440">39</emphasis>
|
||
, aedeagus, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF49664FFFABCF1BF5FFFBD888B8596B" ID-DOI="http://doi.org/10.5281/zenodo.4410061" ID-Zenodo-Dep="4410061" box="[151,1017,1039,1063]" httpUri="https://zenodo.org/record/4410061/files/figure.png" pageId="13" pageNumber="441" startId="13.[151,250,1039,1063]" targetBox="[164,1423,181,1015]" targetPageId="13">
|
||
<paragraph id="8B8936C7FFABCF1BF5FFFBD888B8596B" blockId="13.[151,1017,1039,1063]" box="[151,1017,1039,1063]" pageId="13" pageNumber="441">
|
||
<emphasis id="B942EAD5FFABCF1BF5FFFBD88A61596B" bold="true" box="[151,288,1039,1063]" pageId="13" pageNumber="441">FIGURE 40.</emphasis>
|
||
<taxonomicName id="4C364D44FFABCF1BF44EFBC7893F596B" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[294,638,1039,1063]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="441" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B942EAD5FFABCF1BF44EFBC78ABA596B" box="[294,507,1039,1063]" italics="true" pageId="13" pageNumber="441">Epironastes limbatus</emphasis>
|
||
<bibRefCitation id="EFA74B36FFABCF1BF76AFBD8893F596B" author="Carne, P. B." box="[514,638,1039,1063]" pageId="13" pageNumber="441" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
habitat at 10 km west of Yellowdine.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8936C7FFABCF1BF5AFF9D188F25BE3" blockId="13.[151,1437,1109,2035]" pageId="13" pageNumber="441">
|
||
<emphasis id="B942EAD5FFABCF1BF5AFF9D18A3E5B53" bold="true" box="[199,383,1541,1568]" pageId="13" pageNumber="441">Male variation.</emphasis>
|
||
Body dorsally dark brown to black, ventral surfaces and legs light brown, tending dark brown at margins,
|
||
<quantity id="4CCE9B22FFABCF1BF475F9FD8AF25B08" box="[285,435,1578,1604]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.31" metricValueMin="0.97" pageId="13" pageNumber="441" unit="mm" value="11.399999999999999" valueMax="13.1" valueMin="9.7">9.7–13.1 mm</quantity>
|
||
long, 6.0–
|
||
<quantity id="4CCE9B22FFABCF1BF747F9FE89C55B08" box="[559,644,1577,1604]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="441" unit="mm" value="8.1">8.1 mm</quantity>
|
||
wide. Antennal club with inner face foveate in apical third and bitexurous. Clypeal apex not reflexed; clypeofrontal suture faint. Ocular canthi anterior margin strongly arcuate. Pronotal armature as low swelling, medial depression posterior to impression obsolete. Abdominal sternite 5 sparsely setose at midlength across middle. Metatibia with longer spur slightly curved.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFABCF1BF5AFF96D8AE35B9F" blockId="13.[151,1437,1109,2035]" box="[199,418,1721,1748]" pageId="13" pageNumber="441">
|
||
<emphasis id="B942EAD5FFABCF1BF5AFF96D8A645B98" bold="true" box="[199,293,1722,1748]" pageId="13" pageNumber="441">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C654CFFABCF18F5AFF90A8E035CB8" lastPageId="14" lastPageNumber="442" pageId="13" pageNumber="441" type="diagnosis">
|
||
<paragraph id="8B8936C7FFABCF1BF5AFF90A8AC15ABF" blockId="13.[151,1437,1109,2035]" pageId="13" pageNumber="441">
|
||
<emphasis id="B942EAD5FFABCF1BF5AFF90A8A035BBB" bold="true" box="[199,322,1757,1783]" pageId="13" pageNumber="441">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C364D44FFABCF1BF422F90989765BBB" authorityName="Carne" authorityYear="1957" box="[330,567,1757,1783]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="441" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B942EAD5FFABCF1BF422F90989765BBB" box="[330,567,1757,1783]" italics="true" pageId="13" pageNumber="441">Epironastes limbatus</emphasis>
|
||
</taxonomicName>
|
||
can be recognised by: small size, body
|
||
<quantity id="4CCE9B22FFABCF1BF168F9098FD75BB4" box="[1024,1174,1758,1784]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" metricValueMax="1.31" metricValueMin="0.97" pageId="13" pageNumber="441" unit="mm" value="11.399999999999999" valueMax="13.1" valueMin="9.7">9.7–13.1 mm</quantity>
|
||
long, vestiture yellow; mentum abruptly reflexed to ligula with ligula constricted prior to divergent and truncate apex; ocular canthi with anterior margin angulate or strongly arcuate; pronotum with a small tubercle and shallow subcircular impression; sutural costa on elytra distinctly raised; pygidium setose across base and last sternite with continuous dense, long fringe setae along posterior margin. It differs from
|
||
<taxonomicName id="4C364D44FFABCF1BF788F8B988C65ACB" authorityName="Carne" authorityYear="1957" box="[736,903,1902,1927]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="441" phylum="Arthropoda" rank="species" species="nigrisetosus">
|
||
<emphasis id="B942EAD5FFABCF1BF788F8B988C65ACB" box="[736,903,1902,1927]" italics="true" pageId="13" pageNumber="441">E. nigrisetosus</emphasis>
|
||
</taxonomicName>
|
||
by its much smaller size and yellow vestiture, from
|
||
<taxonomicName id="4C364D44FFABCF1BF5BDF8458A195AE7" authorityName="Carne" authorityYear="1957" box="[213,344,1937,1963]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="441" phylum="Arthropoda" rank="species" species="abruptus">
|
||
<emphasis id="B942EAD5FFABCF1BF5BDF8458A195AE7" box="[213,344,1937,1963]" italics="true" pageId="13" pageNumber="441">E. abruptus</emphasis>
|
||
</taxonomicName>
|
||
by having the clypeus gently declivous anterior to antennal angles and the pygidium being setose across base, and from
|
||
<taxonomicName id="4C364D44FFABCF1BF4FCF861894C5A83" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[404,525,1973,1999]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="441" phylum="Arthropoda" rank="species" species="demarzi">
|
||
<emphasis id="B942EAD5FFABCF1BF4FCF861894C5A83" box="[404,525,1973,1999]" italics="true" pageId="13" pageNumber="441">E. demarzi</emphasis>
|
||
</taxonomicName>
|
||
by having a shallow, subcircular, weakly-defined pronotal impression and a glabrous pygidial ridge.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFA8CF18F5AFFF418A725DB5" blockId="14.[151,1437,150,501]" pageId="14" pageNumber="442">
|
||
<emphasis id="B942EAD5FFA8CF18F5AFFF418A9E5DFC" bold="true" box="[199,479,150,176]" pageId="14" pageNumber="442">Habitat and behaviour.</emphasis>
|
||
This species occurs within the Western Australian Wheatbelt in mallee areas with a
|
||
<taxonomicName id="4C364D44FFA8CF18F5FFFF6D8A4C5D98" box="[151,269,186,212]" class="Agaricomycetes" family="Tricholomataceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="14" pageNumber="442" phylum="Basidiomycota" rank="genus">
|
||
<emphasis id="B942EAD5FFA8CF18F5FFFF6D8A4C5D98" box="[151,269,186,212]" italics="true" pageId="14" pageNumber="442">Melaleuca</emphasis>
|
||
</taxonomicName>
|
||
-dominant understorey on yellow sandplains (
|
||
<figureCitation id="130D2A42FFA8CF18F670FF6D882D5D98" box="[792,876,186,212]" captionStart="FIGURE 40" captionStartId="13.[151,250,1039,1063]" captionTargetBox="[164,1423,181,1015]" captionTargetId="figure-25@13.[164,1423,181,1015]" captionTargetPageId="13" captionText="FIGURE 40. Epironastes limbatus Carne, 1957 habitat at 10 km west of Yellowdine." figureDoi="http://doi.org/10.5281/zenodo.4410061" httpUri="https://zenodo.org/record/4410061/files/figure.png" pageId="14" pageNumber="442">Fig. 40</figureCitation>
|
||
) and with an average annual rainfall of
|
||
<quantity id="4CCE9B22FFA8CF18F052FF6D8B825DB5" metricMagnitude="-1" metricUnit="m" metricValue="3.2" metricValueMax="3.6" metricValueMin="2.8" pageId="14" pageNumber="442" unit="mm" value="320.0" valueMax="360.0" valueMin="280.0">280–360 mm</quantity>
|
||
(
|
||
<figureCitation id="130D2A42FFA8CF18F5BAFF098A675DB5" box="[210,294,222,249]" captionStart="FIGURE 56" captionStartId="18.[151,250,2003,2027]" captionTargetBox="[177,1411,1209,1980]" captionTargetId="figure-471@18.[177,1411,1209,1980]" captionTargetPageId="18" captionText="FIGURE 56. Known distribution of Epironastes species in southern Western Australia." figureDoi="http://doi.org/10.5281/zenodo.4410071" httpUri="https://zenodo.org/record/4410071/files/figure.png" pageId="14" pageNumber="442">Fig. 56</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFA8CF18F5AFFED48E035CB8" blockId="14.[151,1437,150,501]" pageId="14" pageNumber="442">
|
||
It is diurnally active with males flying around 14:00 hr and emergence is in response to summer rainfall (Janu-ary–March). On
|
||
<date id="FF881007FFA8CF18F431FEF1895A5C0C" box="[345,539,294,321]" pageId="14" pageNumber="442" value="2016-01-26">26 January 2016</date>
|
||
, thunderstorms over the previous 1–2 days resulted in storm puddles through much of the range of
|
||
<emphasis id="B942EAD5FFA8CF18F4E4FE9C89555C28" box="[396,532,330,356]" italics="true" pageId="14" pageNumber="442">
|
||
<taxonomicName id="4C364D44FFA8CF18F4E4FE9C89515C28" authorityName="Carne" authorityYear="1957" box="[396,528,330,356]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="442" phylum="Arthropoda" rank="species" species="limbatus">E. limbatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Live and dead males were collected from puddles near Southern Cross at about 14:00 hr, with further live specimens seen floating on later inspections that day, indicating diurnal flights. Later that day, culverts with storm water in them near Yellowdine yielded more males. Light-trapping there (
|
||
<figureCitation id="130D2A42FFA8CF18F069FE458E155CE0" box="[1281,1364,402,428]" captionStart="FIGURE 40" captionStartId="13.[151,250,1039,1063]" captionTargetBox="[164,1423,181,1015]" captionTargetId="figure-25@13.[164,1423,181,1015]" captionTargetPageId="13" captionText="FIGURE 40. Epironastes limbatus Carne, 1957 habitat at 10 km west of Yellowdine." figureDoi="http://doi.org/10.5281/zenodo.4410061" httpUri="https://zenodo.org/record/4410061/files/figure.png" pageId="14" pageNumber="442">Fig. 40</figureCitation>
|
||
) from 19:30 (dusk) to 21:30 hr (moon rise) yielded no
|
||
<taxonomicName id="4C364D44FFA8CF18F7DCFE6088725C9C" authorityName="Carne" authorityYear="1957" box="[692,819,438,464]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="442" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B942EAD5FFA8CF18F7DCFE6088725C9C" box="[692,819,438,464]" italics="true" pageId="14" pageNumber="442">E. limbatus</emphasis>
|
||
</taxonomicName>
|
||
attracted to light, although adults of the dynastines
|
||
<taxonomicName id="4C364D44FFA8CF18F010FE60891C5CB8" authority="Blackburn, 1888" authorityName="Blackburn" authorityYear="1888" class="Insecta" family="Dynastidae" genus="Semanopterus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="442" phylum="Arthropoda" rank="species" species="angustatus">
|
||
<emphasis id="B942EAD5FFA8CF18F010FE608ADA5CB8" italics="true" pageId="14" pageNumber="442">Semanopterus angustatus</emphasis>
|
||
Blackburn, 1888
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D44FFA8CF18F7FCFE0C8F575CB9" authority="(Carne, 1957)" baseAuthorityName="Carne" baseAuthorityYear="1957" box="[660,1046,474,501]" class="Insecta" family="Dynastidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="442" phylum="Arthropoda" rank="species" species="psilus">
|
||
<emphasis id="B942EAD5FFA8CF18F7FCFE0C88325CB8" box="[660,883,474,501]" italics="true" pageId="14" pageNumber="442">Carneoryctes psilus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA74B36FFA8CF18F6EAFE0D8F4C5CB9" author="Carne, P. B." box="[898,1037,474,501]" pageId="14" pageNumber="442" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
were attracted to the light.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |