428 lines
59 KiB
XML
428 lines
59 KiB
XML
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<mods:title id="72E1C20DC570519BDE5EC4E1A6A76954">Revision of Epironastes Carne, 1957 (Coleoptera: Scarabaeidae: Dynastinae Pentodontini: Pseudoryctina) with description of the female, behaviours, and habitats</mods:title>
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<mods:affiliation id="FC2891192C8BC2E53291D3C2C17D5BF3">Department of Agriculture and Food WA (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia, 6986, Australia. paul. hutchinson @ dpird. wa. gov. au; https: // orcid. org / 0000 - 0001 - 8747 - 1320</mods:affiliation>
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<taxonomicName id="4C364D44FFA1CF11F5FFFC78891B5E84" authority="(Endrodi, 1971)" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[151,602,942,969]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="demarzi" status="comb. nov.">
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<emphasis id="B942EAD5FFA1CF11F5FFFC788ACF5E85" bold="true" box="[151,398,942,969]" italics="true" pageId="7" pageNumber="435">Epironastes demarzi</emphasis>
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(
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<bibRefCitation id="EFA74B36FFA1CF11F4F5FC7989135E84" author="Endrodi, S." box="[413,594,942,969]" pageId="7" pageNumber="435" pagination="105 - 108" refId="ref10114" refString="Endrodi, S. (1971) Eine neue australische Pentodontine, Teinogenys demarzi n. sp. (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum Georg Frey, 22, 105 - 108. https: // doi. org / 10.1002 / mmnz. 19700460113" type="journal article" year="1971">Endrődi, 1971</bibRefCitation>
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)
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<taxonomicNameLabel id="A27157AEFFA1CF11F70AFC7888795E84" box="[610,824,942,969]" pageId="7" pageNumber="435" rank="species">new combination</taxonomicNameLabel>
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<paragraph id="8B8936C7FFA1CF11F5FFFC048A105EA1" blockId="7.[151,824,942,1005]" box="[151,337,978,1005]" pageId="7" pageNumber="435">
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(
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<figureCitation id="130D2A42FFA1CF11F5F7FC048A615EA1" box="[159,288,979,1005]" captionStart-0="FIGURES 19–20" captionStart-1="FIGURES 21–28" captionStart-2="FIGURE 29" captionStartId-0="8.[151,264,1332,1356]" captionStartId-1="9.[151,264,1943,1967]" captionStartId-2="10.[151,250,1086,1110]" captionTargetBox-0="[189,1398,309,1309]" captionTargetBox-1="[188,1385,187,1920]" captionTargetBox-2="[177,1411,181,1062]" captionTargetId-0="figure-86@8.[189,1398,309,1309]" captionTargetId-1="figure-25@9.[188,1399,181,1920]" captionTargetId-2="figure-19@10.[177,1411,181,1062]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 19–20. Epironastes demarzi (Endrődi 1971), holotype male. 19, dorsum; 20, labels." captionText-1="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." captionText-2="FIGURE 29. Epironastes demarzi (Endrődi 1971) habitat at 1 km south of Dedari Pumping Station, 80 km west of Kalgoorlie along the Perth-Kalgoorlie water pipeline." figureDoi-0="http://doi.org/10.5281/zenodo.4410051" figureDoi-1="http://doi.org/10.5281/zenodo.4410053" figureDoi-2="http://doi.org/10.5281/zenodo.4410055" httpUri-0="https://zenodo.org/record/4410051/files/figure.png" httpUri-1="https://zenodo.org/record/4410053/files/figure.png" httpUri-2="https://zenodo.org/record/4410055/files/figure.png" pageId="7" pageNumber="435">Figs.19–29</figureCitation>
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,
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<figureCitation id="130D2A42FFA1CF11F444FC058A085EA1" box="[300,329,978,1005]" captionStart="FIGURE 56" captionStartId="18.[151,250,2003,2027]" captionTargetBox="[177,1411,1209,1980]" captionTargetId="figure-471@18.[177,1411,1209,1980]" captionTargetPageId="18" captionText="FIGURE 56. Known distribution of Epironastes species in southern Western Australia." figureDoi="http://doi.org/10.5281/zenodo.4410071" httpUri="https://zenodo.org/record/4410071/files/figure.png" pageId="7" pageNumber="435">56</figureCitation>
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)
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<emphasis id="B942EAD5FFA1CF11F5FFFBCF8A255963" box="[151,356,1047,1071]" italics="true" pageId="7" pageNumber="435">Teinogenys demarzi</emphasis>
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<bibRefCitation id="EFA74B36FFA1CF11F405FBC089765963" author="Endrodi, S." box="[365,567,1047,1071]" pageId="7" pageNumber="435" pagination="105 - 108" refId="ref10114" refString="Endrodi, S. (1971) Eine neue australische Pentodontine, Teinogenys demarzi n. sp. (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum Georg Frey, 22, 105 - 108. https: // doi. org / 10.1002 / mmnz. 19700460113" type="journal article" year="1971">Endrődi, 1971: 105</bibRefCitation>
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(
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<figureCitation id="130D2A42FFA1CF11F720FBC089C95963" box="[584,648,1047,1071]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1486,1510]" captionTargetBox="[151,1435,558,1460]" captionTargetId="figure-198@3.[151,1435,558,1460]" captionTargetPageId="3" captionText="FIGURES 1–2. Epironastes abruptus Carne, 1957, holotype male. 1, dorsum; 2, labels." figureDoi="http://doi.org/10.5281/zenodo.4410043" httpUri="https://zenodo.org/record/4410043/files/figure.png" pageId="7" pageNumber="435">Fig. 1</figureCitation>
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:
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<specimenCount id="9D30FD4EFFA1CF11F7FFFBCF89EB5963" box="[663,682,1048,1071]" pageId="7" pageNumber="435" type="male">♂</specimenCount>
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head;
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Figs. 2–
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<specimenCount id="9D30FD4EFFA1CF11F622FBC088145963" box="[842,853,1047,1071]" pageId="7" pageNumber="435" type="male">3</specimenCount>
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♂ genitalia);
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(
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<figureCitation id="130D2A42FFA1CF11F1DBFBC08FBE5963" box="[1203,1279,1047,1071]" captionStart="FIGURE 29" captionStartId="10.[151,250,1086,1110]" captionTargetBox="[177,1411,181,1062]" captionTargetId="figure-19@10.[177,1411,181,1062]" captionTargetPageId="10" captionText="FIGURE 29. Epironastes demarzi (Endrődi 1971) habitat at 1 km south of Dedari Pumping Station, 80 km west of Kalgoorlie along the Perth-Kalgoorlie water pipeline." figureDoi="http://doi.org/10.5281/zenodo.4410055" httpUri="https://zenodo.org/record/4410055/files/figure.png" pageId="7" pageNumber="435">
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Fig.
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<specimenCount id="9D30FD4EFFA1CF11F18EFBC08FBE5963" box="[1254,1279,1047,1071]" pageId="7" pageNumber="435" type="male">29</specimenCount>
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♂ parameres);
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(Fig. 1376
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<specimenCount id="9D30FD4EFFA1CF11F493FBE1894F5901" box="[507,526,1078,1101]" pageId="7" pageNumber="435" type="male">♂</specimenCount>
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parameres; Plate XV
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<figureCitation id="130D2A42FFA1CF11F786FBE188685902" box="[750,809,1078,1102]" captionStart="FIGURES 3–10" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1404,193,1919]" captionTargetId="figure-19@4.[181,1407,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–10. Epironastes abruptus Carne, 1957, male specimen #0465. 3, dorsum; 4, venter; 5, lateral; 6, head, lateral; 7, head, dorsal; 8, pygidium; 9, parameres; 10, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410045" httpUri="https://zenodo.org/record/4410045/files/figure.png" pageId="7" pageNumber="435">
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Fig.
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<specimenCount id="9D30FD4EFFA1CF11F674FBE188685902" box="[796,809,1078,1102]" pageId="7" pageNumber="435" type="male">4</specimenCount>
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♂ adult);
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<treatmentCitation id="0A9710D6FFA1CF11F6F9FBE18FD35902" author="Cassis, G. & Weir, T. A." box="[913,1170,1078,1102]" page="416" pageId="7" pageNumber="435" year="1992">
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<bibRefCitation id="EFA74B36FFA1CF11F6F9FBE18FD35902" author="Cassis, G. & Weir, T. A." box="[913,1170,1078,1102]" pageId="7" pageNumber="435" refId="ref10030" refString="Cassis, G. & Weir, T. A. (1992) Dynastinae. In: Houston, W. W. K. (ed), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Volume 9. Australian Government Publishing Service, Canberra, pp. 383 - 425." type="book" year="1992">Cassis & Weir 1992: 416</bibRefCitation>
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;
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<treatmentCitation id="0A9710D6FFA1CF11F1F4FBE18E325902" author="Dechambre, R-P." box="[1180,1395,1078,1102]" page="61" pageId="7" pageNumber="435" year="2005">
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.
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<emphasis id="B942EAD5FFA1CF11F5FFFBAB8A6259DA" bold="true" box="[151,291,1148,1174]" pageId="7" pageNumber="435">Type series.</emphasis>
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|
||
<specimenCount id="9D30FD4EFFA1CF11F4F2FBAA8AEE59DB" box="[410,431,1149,1175]" pageId="7" pageNumber="435" type="male">♂</specimenCount>
|
||
(by original designation):
|
||
<location id="8EE9601CFFA1CF11F7B2FBAB883C59DA" LSID="urn:lsid:plazi:treatment:039F87D1FFA1CF1CF5FFFC7889B05B03:8EE9601CFFA1CF11F7B2FBAB883C59DA" box="[730,893,1148,1174]" latitude="-32.5746" longLatPrecision="6" longitude="120.3321" name="Lake Johnston" pageId="7" pageNumber="435">Lake Johnston</location>
|
||
[
|
||
<geoCoordinate id="EE025000FFA1CF11F6E3FBAB8F4759DB" box="[907,1030,1148,1175]" degrees="32.5746" direction="south" orientation="latitude" pageId="7" pageNumber="435" precision="5" value="-32.5746">32.5746°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE025000FFA1CF11F17AFBAC8FDB59DA" box="[1042,1178,1147,1174]" degrees="120.3321" direction="east" orientation="longitude" pageId="7" pageNumber="435" precision="5" value="120.3321">120.3321°E</geoCoordinate>
|
||
]
|
||
<collectorName id="26C35311FFA1CF11F1CFFBAB8E0559DB" box="[1191,1348,1148,1175]" pageId="7" pageNumber="435">W.A. Demarz</collectorName>
|
||
, 3.1965 //
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5E3C9AFFA1CF11F5C7FB778BAB5992" ID-GBIF-Occurrence="3031275315" collectionCode="NHMB" pageId="7" pageNumber="435" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="548D8865FFA1CF11F5C7FB778A6559F6" box="[175,292,1184,1210]" pageId="7" pageNumber="435" type="holotype">Holotypus</typeStatus>
|
||
<taxonomicName id="4C364D44FFA1CF11F443FB77891259F6" authority="Endr." authorityName="Endrodi" authorityYear="1971" box="[299,595,1184,1210]" class="Insecta" family="Dynastidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="demarzi">Teinogenys demarzi Endr.</taxonomicName>
|
||
// Museum Frey Tutzing // genitalia on card [in
|
||
<collectionCode id="ED27AE02FFA1CF11F112FB768F9059F7" box="[1146,1233,1185,1211]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="7" pageNumber="435">NHMB</collectionCode>
|
||
, examined] (
|
||
<figureCitation id="130D2A42FFA1CF11F00CFB778BA35992" captionStart="FIGURES 19–20" captionStartId="8.[151,264,1332,1356]" captionTargetBox="[189,1398,309,1309]" captionTargetId="figure-86@8.[189,1398,309,1309]" captionTargetPageId="8" captionText="FIGURES 19–20. Epironastes demarzi (Endrődi 1971), holotype male. 19, dorsum; 20, labels." figureDoi="http://doi.org/10.5281/zenodo.4410051" httpUri="https://zenodo.org/record/4410051/files/figure.png" pageId="7" pageNumber="435">Figs. 19–20</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFA1CF11F5AFFB3E88CC586B" blockId="7.[151,1437,1147,2039]" pageId="7" pageNumber="435">
|
||
<materialsCitation id="3B5E3C9AFFA1CF11F5AFFB3E88C8586B" ID-GBIF-Occurrence="3031275308" collectionCode="NHMB" location="Endrodi's" municipality="Hungarian Museum of Natural History" pageId="7" pageNumber="435" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
|
||
<typeStatus id="548D8865FFA1CF11F5AFFB3E8A79584F" box="[199,312,1257,1283]" pageId="7" pageNumber="435" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D30FD4EFFA1CF11F42BFB3F8A28584E" box="[323,361,1256,1282]" pageId="7" pageNumber="435" type="male">2♂</specimenCount>
|
||
, same data as holotype [in
|
||
<collectionCode id="ED27AE02FFA1CF11F7F4FB3089B7584E" box="[668,758,1255,1282]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="7" pageNumber="435">NHMB</collectionCode>
|
||
]. A further
|
||
<typeStatus id="548D8865FFA1CF11F61DFB3E8894584F" box="[885,981,1257,1283]" pageId="7" pageNumber="435" type="paratype">paratype</typeStatus>
|
||
was evidently retained in
|
||
<location id="8EE9601CFFA1CF11F197FB3F8E2D584E" LSID="urn:lsid:plazi:treatment:039F87D1FFA1CF1CF5FFFC7889B05B03:8EE9601CFFA1CF11F197FB3F8E2D584E" box="[1279,1388,1256,1282]" municipality="Hungarian Museum of Natural History" name="Endrodi's" pageId="7" pageNumber="435">Endrődi’s</location>
|
||
private collection (now in the
|
||
<collectingMunicipality id="6BEDACBDFFA1CF11F4A1FADB88C0586B" box="[457,897,1292,1319]" pageId="7" pageNumber="435">Hungarian Museum of Natural History</collectingMunicipality>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFA1CF11F5AFFAE78A8D58DE" blockId="7.[151,1437,1147,2039]" pageId="7" pageNumber="435">
|
||
<emphasis id="B942EAD5FFA1CF11F5AFFAE7894D5806" bold="true" box="[199,524,1328,1354]" pageId="7" pageNumber="435">Other specimen examined.</emphasis>
|
||
<materialsCitation id="3B5E3C9AFFA1CF11F77CFAE788C95822" ID-GBIF-Occurrence="3031275303" collectingDate="2016-01-27" collectorName="West. Aust. & P. Hutchinson" country="Australia" latitude="-31.094" location="Dedari" longLatPrecision="6" longitude="120.7001" pageId="7" pageNumber="435" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<collectingCountry id="F3217657FFA1CF11F77CFAE789F75806" box="[532,694,1328,1354]" name="Australia" pageId="7" pageNumber="435">AUSTRALIA</collectingCountry>
|
||
.
|
||
<collectingRegion id="49F2F825FFA1CF11F7ACFAE788B55806" box="[708,1012,1328,1354]" country="Australia" name="Western Australia" pageId="7" pageNumber="435">WESTERN AUSTRALIA</collectingRegion>
|
||
:
|
||
<specimenCount id="9D30FD4EFFA1CF11F16AFAE78F675806" box="[1026,1062,1328,1354]" pageId="7" pageNumber="435" type="male">1♂</specimenCount>
|
||
,
|
||
<location id="8EE9601CFFA1CF11F15EFAE78FC35806" LSID="urn:lsid:plazi:treatment:039F87D1FFA1CF1CF5FFFC7889B05B03:8EE9601CFFA1CF11F15EFAE78FC35806" box="[1078,1154,1328,1354]" country="Australia" latitude="-31.094" longLatPrecision="6" longitude="120.7001" name="Dedari" pageId="7" pageNumber="435" stateProvince="Western Australia">Dedari</location>
|
||
<quantity id="4CCE9B22FFA1CF11F1E5FAE78F885806" box="[1165,1225,1328,1354]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="435" unit="km" value="1.0">1 km</quantity>
|
||
S. of [
|
||
<geoCoordinate id="EE025000FFA1CF11F075FAE78ED95806" box="[1309,1432,1328,1354]" degrees="31.0940" direction="south" orientation="latitude" pageId="7" pageNumber="435" precision="5" value="-31.094">31.0940°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE025000FFA1CF11F5FFFA848A5E5822" box="[151,287,1363,1390]" degrees="120.7001" direction="east" orientation="longitude" pageId="7" pageNumber="435" precision="5" value="120.7001">120.7001°E</geoCoordinate>
|
||
]
|
||
<collectorName id="26C35311FFA1CF11F446FA828AEE5822" box="[302,431,1364,1391]" pageId="7" pageNumber="435">West. Aust.</collectorName>
|
||
<date id="FF881007FFA1CF11F4DFFA83897E5822" box="[439,575,1364,1390]" pageId="7" pageNumber="435" value="2016-01-27">
|
||
<collectingDate id="EFCCE9EFFFA1CF11F4DFFA83897E5822" box="[439,575,1364,1390]" pageId="7" pageNumber="435" value="2016-01-27">27.Jan.2016</collectingDate>
|
||
</date>
|
||
<collectorName id="26C35311FFA1CF11F72FFA8289A55822" box="[583,740,1364,1391]" pageId="7" pageNumber="435">P. Hutchinson</collectorName>
|
||
//
|
||
<taxonomicName id="4C364D44FFA1CF11F66DFA8388C95822" authorityName="Carne" authorityYear="1957" box="[773,904,1364,1390]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="genus">Epironastes</taxonomicName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5E3C9AFFA1CF11F6F8FA828A8858DE" ID-GBIF-Occurrence="3031275310" collectionCode="PMH" collectorName="Coll & Dyn" country="Australia" pageId="7" pageNumber="435" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D30FD4EFFA1CF11F6F8FA8288E65823" box="[912,935,1365,1391]" pageId="7" pageNumber="435" type="male">♂</specimenCount>
|
||
, in storm puddle // PMH
|
||
<collectorName id="26C35311FFA1CF11F1A5FA838FBC5822" box="[1229,1277,1364,1390]" pageId="7" pageNumber="435">Coll</collectorName>
|
||
#
|
||
<collectorName id="26C35311FFA1CF11F07CFA828E055823" box="[1300,1348,1365,1391]" pageId="7" pageNumber="435">Dyn</collectorName>
|
||
1713 // genitalia on card [in
|
||
<collectionCode id="ED27AE02FFA1CF11F417FAA08A8358DE" box="[383,450,1399,1426]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14877" name="City Museum and Records Office" pageId="7" pageNumber="435" type="Herbarium">PMH</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C654CFFA1CF1EF5AFFA4B8ADB5B05" lastPageId="8" lastPageNumber="436" pageId="7" pageNumber="435" type="description">
|
||
<paragraph id="8B8936C7FFA1CF1EF5AFFA4B8E475C51" blockId="7.[151,1437,1147,2039]" lastBlockId="8.[151,1437,151,285]" lastPageId="8" lastPageNumber="436" pageId="7" pageNumber="435">
|
||
<emphasis id="B942EAD5FFA1CF11F5AFFA4B896C58FA" bold="true" box="[199,557,1436,1462]" pageId="7" pageNumber="435">
|
||
Redescription. Male
|
||
<typeStatus id="548D8865FFA1CF11F4A8FA4B896858FA" box="[448,553,1436,1462]" pageId="7" pageNumber="435" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Body (
|
||
<figureCitation id="130D2A42FFA1CF11F7E9FA4B884658FA" box="[641,775,1436,1462]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Figs. 21–22</figureCitation>
|
||
) black with antennal club ventrally light-brown over black and dorsally black,
|
||
<quantity id="4CCE9B22FFA1CF11F41FFA178A9C5897" box="[375,477,1472,1499]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="7" pageNumber="435" unit="mm" value="14.8">14.8 mm</quantity>
|
||
long,
|
||
<quantity id="4CCE9B22FFA1CF11F74FFA17893F5897" box="[551,638,1472,1499]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="7" pageNumber="435" unit="mm" value="8.8">8.8 mm</quantity>
|
||
wide across elytra post median. Mentum weakly reflexed at apex, bilobed with indistinct medial groove with ligula reflexed, apically attenuate with apex weakly bilobed.Antennal club (
|
||
<figureCitation id="130D2A42FFA1CF11F5F7F9DF8A645B6F" box="[159,293,1544,1571]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Figs. 21–25</figureCitation>
|
||
) only slightly longer than shaft, inner face setose, shallowly foveate on apical third; outer face of club bitexturous with ventral and dorsal portions subequal, club straight, in outline with ventral margin strongly bowed. Clypeus (
|
||
<figureCitation id="130D2A42FFA1CF11F469F9878AF15B27" box="[257,432,1616,1643]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Figs. 21, 24–25</figureCitation>
|
||
) attenuate with arcuate reflexed apex, abruptly declivous from broadly rounded ridge joining antennal angles, surface rugulose; clypeofrontal ridge distinct. Frons (
|
||
<figureCitation id="130D2A42FFA1CF11F6B0F9A38FCA5BC3" box="[984,1163,1652,1679]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Figs. 21, 24–25</figureCitation>
|
||
) rugulose; ocular canthi with anterior margin weakly arcuate, laterally truncate, anterior margin bearing long orange setae. Pronotum (
|
||
<figureCitation id="130D2A42FFA1CF11F00CF94F8A495B9B" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Figs. 21, 23–25</figureCitation>
|
||
) with anterior lateral angles slightly obtuse, widest in anterior half, about one-third distance from anterior margin; anteromedian horn stout, erect, preceding a transverse impression confined to anterior third with abrupt posterior margin, basal convexity uninterrupted; surface sparsely and finely punctate, punctures becoming coarser, denser towards margins often transverse along anterolateral margins; simple punctures tending transverse in impression;, basal ridge complete. Scutellum (
|
||
<figureCitation id="130D2A42FFA1CF11F726F89B89DF5A2A" box="[590,670,1868,1894]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Fig. 21</figureCitation>
|
||
) with sparse moderate punctures medially in basal half. Each elytron (
|
||
<figureCitation id="130D2A42FFA1CF11F5F7F8A78A685AC6" box="[159,297,1904,1930]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Figs. 21, 23</figureCitation>
|
||
) with sutural striae punctate to apex, elytral disc with small, shallow, umbilicate punctures forming well-defined striae, intervals with random punctures, punctures continuous to apical calli, lateral declivity sparsely rugulose. Pygidium (
|
||
<figureCitation id="130D2A42FFA1CF11F4EAF86F8A955A9F" box="[386,468,1976,2003]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="7" pageNumber="435">Fig. 26</figureCitation>
|
||
) moderately convex, weakly microreticulate, concentrically punctate, with dense long setae across base and sparsely punctate apical zone, apical ridge not thickened medially, bearing dense fringe of long setae across median third. Abdominal sternites 2–4 (
|
||
<figureCitation id="130D2A42FFAECF1EF7B3FF40886A5DFD" box="[731,811,151,177]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="8" pageNumber="436">Fig. 22</figureCitation>
|
||
) laterally setose over entire surface, last sternite bearing continuous fringe setae on posterior margin. Coxae (
|
||
<figureCitation id="130D2A42FFAECF1EF7B6FF6C886E5D99" box="[734,815,187,213]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="8" pageNumber="436">Fig. 22</figureCitation>
|
||
) bearing orange vestiture; metatibiae with basal carinae bearing long, acute setae, apical carinae bearing long, fine closely set ciliae on outer surface; metatibial spurs (
|
||
<figureCitation id="130D2A42FFAECF1EF007FF088BF55C51" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="8" pageNumber="436">Fig. 22</figureCitation>
|
||
) of unequal length, long, subparallel, with longer spur slightly curved. Genitalia as in
|
||
<figureCitation id="130D2A42FFAECF1EF113FED48E425C51" box="[1147,1283,259,285]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="8" pageNumber="436">Figs. 27–28</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF49664FFFAECF1EF5FFFAE38F1C5801" ID-DOI="http://doi.org/10.5281/zenodo.4410051" ID-Zenodo-Dep="4410051" box="[151,1117,1332,1357]" httpUri="https://zenodo.org/record/4410051/files/figure.png" pageId="8" pageNumber="436" startId="8.[151,264,1332,1356]" targetBox="[189,1398,309,1309]" targetPageId="8">
|
||
<paragraph id="8B8936C7FFAECF1EF5FFFAE38F1C5801" blockId="8.[151,1117,1332,1357]" box="[151,1117,1332,1357]" pageId="8" pageNumber="436">
|
||
<emphasis id="B942EAD5FFAECF1EF5FFFAE38A155800" bold="true" box="[151,340,1332,1356]" pageId="8" pageNumber="436">FIGURES 19–20.</emphasis>
|
||
<taxonomicName id="4C364D44FFAECF1EF432FAE2898F5801" authority="(Endrodi 1971)" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[346,718,1333,1357]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="demarzi">
|
||
<emphasis id="B942EAD5FFAECF1EF432FAE2896A5801" box="[346,555,1333,1357]" italics="true" pageId="8" pageNumber="436">Epironastes demarzi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA74B36FFAECF1EF75FFAE289865801" author="Endrodi, S." box="[567,711,1333,1357]" pageId="8" pageNumber="436" pagination="105 - 108" refId="ref10114" refString="Endrodi, S. (1971) Eine neue australische Pentodontine, Teinogenys demarzi n. sp. (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum Georg Frey, 22, 105 - 108. https: // doi. org / 10.1002 / mmnz. 19700460113" type="journal article" year="1971">Endrődi 1971</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, holotype male.
|
||
<emphasis id="B942EAD5FFAECF1EF61EFAE388D45800" bold="true" box="[886,917,1332,1356]" pageId="8" pageNumber="436">19,</emphasis>
|
||
dorsum;
|
||
<emphasis id="B942EAD5FFAECF1EF69DFAE38F4F5800" bold="true" box="[1013,1038,1332,1356]" pageId="8" pageNumber="436">20</emphasis>
|
||
, labels.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8936C7FFAECF1EF5AFFAAC8EC35B69" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="436">
|
||
<emphasis id="B942EAD5FFAECF1EF5AFFAAC8AC258D9" bold="true" box="[199,387,1403,1429]" pageId="8" pageNumber="436">Male variation.</emphasis>
|
||
Body
|
||
<quantity id="4CCE9B22FFAECF1EF4BDFAAC893A58D9" box="[469,635,1403,1430]" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="1.75" metricValueMin="1.48" pageId="8" pageNumber="436" unit="mm" value="16.15" valueMax="17.5" valueMin="14.8">14.8–17.5 mm</quantity>
|
||
long,
|
||
<quantity id="4CCE9B22FFAECF1EF7A1FAAC882358D9" box="[713,866,1403,1429]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="10.1" metricValueMin="8.8" pageId="8" pageNumber="436" unit="mm" value="9.45" valueMax="10.1" valueMin="8.8">8.8–10.1 mm</quantity>
|
||
long (elytra post median). Mentum with distinct groove; ligula apically broadened and weakly bilobed. Antennal club with ventral portion of outer face black. Clypeus abruptly declivous. Impression on pronotum with abrupt and steep posterior margin, basal convexity with trace of median impression, basal ridge obsolete in median half. The specimen from near Dedari is larger than the
|
||
<typeStatus id="548D8865FFAECF1EF5FFF9DC8BB85B69" box="[151,249,1547,1573]" pageId="8" pageNumber="436" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="548D8865FFAECF1EF458F9DC8ADA5B69" box="[304,411,1547,1573]" pageId="8" pageNumber="436" type="paratype">paratypes</typeStatus>
|
||
we examined and with above structures more defined yet having elytral punctation finer.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFAECF1EF5AFF9F88ADB5B05" blockId="8.[151,1437,1403,2041]" box="[199,410,1583,1609]" pageId="8" pageNumber="436">
|
||
<emphasis id="B942EAD5FFAECF1EF5AFF9F88A645B05" bold="true" box="[199,293,1583,1609]" pageId="8" pageNumber="436">Female.</emphasis>
|
||
Unknown
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C654CFFAECF1CF5AFF98489B05B03" lastPageId="10" lastPageNumber="438" pageId="8" pageNumber="436" type="diagnosis">
|
||
<paragraph id="8B8936C7FFAECF1EF5AFF9848AFD5A6D" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="436">
|
||
<emphasis id="B942EAD5FFAECF1EF5AFF9848A035B21" bold="true" box="[199,322,1619,1645]" pageId="8" pageNumber="436">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C364D44FFAECF1EF42FF983896D5B21" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[327,556,1619,1645]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="demarzi">
|
||
<emphasis id="B942EAD5FFAECF1EF42FF983896D5B21" box="[327,556,1619,1645]" italics="true" pageId="8" pageNumber="436">Epironastes demarzi</emphasis>
|
||
</taxonomicName>
|
||
can be recognised by having: orange vestiture; clypeal apex attenuate to rounded reflexed apex; antennal club with ventral margin strongly bowed; pronotal impression transverse and confined to anterior third with well-defined posterior margin; pygidium bearing dense long setae across base with non-broadened apical ridge that bears a dense fringe across medial third. It differs from
|
||
<taxonomicName id="4C364D44FFAECF1EF14AF9178F8A5B95" authorityName="Carne" authorityYear="1957" box="[1058,1227,1728,1753]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="nigrisetosus">
|
||
<emphasis id="B942EAD5FFAECF1EF14AF9178F8A5B95" box="[1058,1227,1728,1753]" italics="true" pageId="8" pageNumber="436">E. nigrisetosus</emphasis>
|
||
</taxonomicName>
|
||
by having orange vestiture, from
|
||
<taxonomicName id="4C364D44FFAECF1EF42FF9338A8D5BB1" authorityName="Carne" authorityYear="1957" box="[327,460,1763,1789]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="abruptus">
|
||
<emphasis id="B942EAD5FFAECF1EF42FF9338A8D5BB1" box="[327,460,1763,1789]" italics="true" pageId="8" pageNumber="436">E. abruptus</emphasis>
|
||
</taxonomicName>
|
||
by having the pygidium setose across the base, and from
|
||
<taxonomicName id="4C364D44FFAECF1EF10EF9338FA65BB1" authorityName="Carne" authorityYear="1957" box="[1126,1255,1763,1789]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B942EAD5FFAECF1EF10EF9338FA65BB1" box="[1126,1255,1763,1789]" italics="true" pageId="8" pageNumber="436">E. limbatus</emphasis>
|
||
</taxonomicName>
|
||
by the pygidial apical ridge bearing setae.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFAECF1EF5AFF8FC8AEF5AB5" blockId="8.[151,1437,1403,2041]" pageId="8" pageNumber="436">
|
||
<bibRefCitation id="EFA74B36FFAECF1EF5AFF8FC8A325A09" author="Endrodi, S." box="[199,371,1835,1861]" pageId="8" pageNumber="436" pagination="105 - 108" refId="ref10114" refString="Endrodi, S. (1971) Eine neue australische Pentodontine, Teinogenys demarzi n. sp. (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum Georg Frey, 22, 105 - 108. https: // doi. org / 10.1002 / mmnz. 19700460113" type="journal article" year="1971">Endrődi (1971)</bibRefCitation>
|
||
hesitantly placed this species in
|
||
<taxonomicName id="4C364D44FFAECF1EF789F8FB88AE5A09" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[737,1007,1835,1861]" class="Insecta" family="Dynastidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFAECF1EF789F8FB881F5A09" box="[737,862,1836,1861]" italics="true" pageId="8" pageNumber="436">Teinogenys</emphasis>
|
||
Sharp, 1873
|
||
</taxonomicName>
|
||
, but did not mention in his description the generic defining character of
|
||
<taxonomicName id="4C364D44FFAECF1EF779F88789D75A25" authorityName="Carne" authorityYear="1957" box="[529,662,1872,1897]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFAECF1EF779F88789D75A25" box="[529,662,1872,1897]" italics="true" pageId="8" pageNumber="436">Epironastes</emphasis>
|
||
</taxonomicName>
|
||
of having a ridge joining the antennal insertions anterior to the clypeofrontal suture, nor is it evident in his illustration of the head. Later, he (
|
||
<bibRefCitation id="EFA74B36FFAECF1EF6BCF8A48F2E5AC1" author="Endrodi, S." box="[980,1135,1907,1933]" pageId="8" pageNumber="436" pagination="17 - 70" refId="ref10164" refString="Endrodi, S. (1974) Monographie der Dynastinae 4. Tribus: Pentodontini von Australien und Neu-Zeeland (Coleoptera). Folia Entomologica Hungarica, Series Nova, 27, 17 - 70." type="journal article" year="1974">Endrődi 1974</bibRefCitation>
|
||
) excluded
|
||
<taxonomicName id="4C364D44FFAECF1EF182F8A38E1E5AC1" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[1258,1375,1907,1933]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="demarzi">
|
||
<emphasis id="B942EAD5FFAECF1EF182F8A38E1E5AC1" box="[1258,1375,1907,1933]" italics="true" pageId="8" pageNumber="436">T. demarzi</emphasis>
|
||
</taxonomicName>
|
||
from his key to male
|
||
<taxonomicName id="4C364D44FFAECF1EF421F84F8A875AFD" authorityName="Sharp" authorityYear="1873" box="[329,454,1944,1969]" class="Insecta" family="Dynastidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFAECF1EF421F84F8A875AFD" box="[329,454,1944,1969]" italics="true" pageId="8" pageNumber="436">Teinogenys</emphasis>
|
||
</taxonomicName>
|
||
and noted it differed from all described species of
|
||
<taxonomicName id="4C364D44FFAECF1EF16AF84F8F3E5AFD" authorityName="Sharp" authorityYear="1873" box="[1026,1151,1944,1969]" class="Insecta" family="Dynastidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFAECF1EF16AF84F8F3E5AFD" box="[1026,1151,1944,1969]" italics="true" pageId="8" pageNumber="436">Teinogenys</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C364D44FFAECF1EF1CDF8408ED95AFE" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[1189,1432,1943,1970]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFAECF1EF1CDF8408E465AFD" box="[1189,1287,1943,1969]" italics="true" pageId="8" pageNumber="436">Cavonus</emphasis>
|
||
Sharp, 1875
|
||
</taxonomicName>
|
||
, indicating he was uncertain of its generic placement. However, in
|
||
<bibRefCitation id="EFA74B36FFAECF1EF61EF86C8F605A9A" author="Endrodi, S." box="[886,1057,1979,2006]" pageId="8" pageNumber="436" refId="ref10200" refString="Endrodi, S. (1985) The Dynastinae of the World. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi (1985)</bibRefCitation>
|
||
he did place
|
||
<taxonomicName id="4C364D44FFAECF1EF1DDF86B8E6B5A99" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[1205,1322,1979,2005]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="demarzi">
|
||
<emphasis id="B942EAD5FFAECF1EF1DDF86B8E6B5A99" box="[1205,1322,1979,2005]" italics="true" pageId="8" pageNumber="436">T. demarzi</emphasis>
|
||
</taxonomicName>
|
||
in his key to species of
|
||
<taxonomicName id="4C364D44FFAECF1EF442F8378AE65AB5" authorityName="Sharp" authorityYear="1873" box="[298,423,2016,2041]" class="Insecta" family="Dynastidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFAECF1EF442F8378AE65AB5" box="[298,423,2016,2041]" italics="true" pageId="8" pageNumber="436">Teinogenys</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF49664FFFAFCF1FF5FFF84088D95A98" ID-DOI="http://doi.org/10.5281/zenodo.4410053" ID-Zenodo-Dep="4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="9" pageNumber="437" startId="9.[151,264,1943,1967]" targetBox="[188,1385,187,1920]" targetPageId="9">
|
||
<paragraph id="8B8936C7FFAFCF1FF5FFF84088D95A98" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="437">
|
||
<emphasis id="B942EAD5FFAFCF1FF5FFF8408A175AE3" bold="true" box="[151,342,1943,1967]" pageId="9" pageNumber="437">FIGURES 21–28.</emphasis>
|
||
<taxonomicName id="4C364D44FFAFCF1FF435F84F89975AFC" authority="(Endrodi 1971)" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[349,726,1944,1968]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="demarzi">
|
||
<emphasis id="B942EAD5FFAFCF1FF435F84F896E5AFC" box="[349,559,1944,1968]" italics="true" pageId="9" pageNumber="437">Epironastes demarzi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA74B36FFAFCF1FF756F84F898E5AFC" author="Endrodi, S." box="[574,719,1944,1968]" pageId="9" pageNumber="437" pagination="105 - 108" refId="ref10114" refString="Endrodi, S. (1971) Eine neue australische Pentodontine, Teinogenys demarzi n. sp. (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum Georg Frey, 22, 105 - 108. https: // doi. org / 10.1002 / mmnz. 19700460113" type="journal article" year="1971">Endrődi 1971</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male specimen #1713.
|
||
<emphasis id="B942EAD5FFAFCF1FF6B8F84088A85AE3" bold="true" box="[976,1001,1943,1967]" pageId="9" pageNumber="437">21</emphasis>
|
||
, dorsum;
|
||
<emphasis id="B942EAD5FFAFCF1FF13BF8408F2D5AE3" bold="true" box="[1107,1132,1943,1967]" pageId="9" pageNumber="437">22</emphasis>
|
||
, venter;
|
||
<emphasis id="B942EAD5FFAFCF1FF1A1F8408FA35AE3" bold="true" box="[1225,1250,1943,1967]" pageId="9" pageNumber="437">23</emphasis>
|
||
, lateral;
|
||
<emphasis id="B942EAD5FFAFCF1FF056F8408E165AE3" bold="true" box="[1342,1367,1943,1967]" pageId="9" pageNumber="437">24</emphasis>
|
||
, head, lateral;
|
||
<emphasis id="B942EAD5FFAFCF1FF58CF86C8BBC5A9F" bold="true" box="[228,253,1979,2003]" pageId="9" pageNumber="437">25</emphasis>
|
||
, head, dorsal;
|
||
<emphasis id="B942EAD5FFAFCF1FF4FAF86C8AEA5A9F" bold="true" box="[402,427,1979,2003]" pageId="9" pageNumber="437">26</emphasis>
|
||
, pygidium;
|
||
<emphasis id="B942EAD5FFAFCF1FF74EF86C897E5A9F" bold="true" box="[550,575,1979,2003]" pageId="9" pageNumber="437">27</emphasis>
|
||
, parameres;
|
||
<emphasis id="B942EAD5FFAFCF1FF7AAF86C899A5A9F" bold="true" box="[706,731,1979,2003]" pageId="9" pageNumber="437">28</emphasis>
|
||
, aedeagus, lateral.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF49664FFFACCF1CF5FFFBE989025936" ID-DOI="http://doi.org/10.5281/zenodo.4410055" ID-Zenodo-Dep="4410055" httpUri="https://zenodo.org/record/4410055/files/figure.png" pageId="10" pageNumber="438" startId="10.[151,250,1086,1110]" targetBox="[177,1411,181,1062]" targetPageId="10">
|
||
<paragraph id="8B8936C7FFACCF1CF5FFFBE989025936" blockId="10.[151,1436,1086,1146]" pageId="10" pageNumber="438">
|
||
<emphasis id="B942EAD5FFACCF1CF5FFFBE98A61591A" bold="true" box="[151,288,1086,1110]" pageId="10" pageNumber="438">FIGURE 29.</emphasis>
|
||
<taxonomicName id="4C364D44FFACCF1CF44EFBE889D9591B" authority="(Endrodi 1971)" authorityName="Hutchinson & Allsopp" authorityYear="2020" baseAuthorityName="Endrodi" baseAuthorityYear="1971" box="[294,664,1086,1111]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="438" phylum="Arthropoda" rank="species" species="demarzi">
|
||
<emphasis id="B942EAD5FFACCF1CF44EFBE88AB6591A" box="[294,503,1086,1110]" italics="true" pageId="10" pageNumber="438">Epironastes demarzi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA74B36FFACCF1CF76CFBE989D3591B" author="Endrodi, S." box="[516,658,1086,1111]" pageId="10" pageNumber="438" pagination="105 - 108" refId="ref10114" refString="Endrodi, S. (1971) Eine neue australische Pentodontine, Teinogenys demarzi n. sp. (Col. Melolonthidae). Entomologische Arbeiten aus dem Museum Georg Frey, 22, 105 - 108. https: // doi. org / 10.1002 / mmnz. 19700460113" type="journal article" year="1971">Endrődi 1971</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
habitat at 1 km south of Dedari Pumping Station, 80 km west of Kalgoorlie along the Perth-Kalgoorlie water pipeline.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8936C7FFACCF1CF5AFFB7E89C5583B" blockId="10.[151,1437,1193,1615]" pageId="10" pageNumber="438">
|
||
The absence of a description of the clypeal ridge, omission in the accompanying illustration and placement in the
|
||
<bibRefCitation id="EFA74B36FFACCF1CF5AAFB1A8A3159AB" author="Endrodi, S." box="[194,368,1228,1255]" pageId="10" pageNumber="438" refId="ref10200" refString="Endrodi, S. (1985) The Dynastinae of the World. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi (1985)</bibRefCitation>
|
||
key did not allow subsequent researchers (
|
||
<bibRefCitation id="EFA74B36FFACCF1CF637FB1A8F2F59AB" author="Allsopp, P. G. & Carne, P. B." box="[863,1134,1229,1255]" pageId="10" pageNumber="438" pagination="89 - 93" refId="ref9853" refString="Allsopp, P. G. & Carne, P. B. (1986) Teinogenys mooniensis sp. n. and new locality records for other species of Teinogenys Sharp (Coleoptera: Scarabaeidae: Dynastinae). Journal of the Australian Entomological Society, 25, 89 - 93. https: // doi. org / 10.1111 / j. 1440 - 6055.1986. tb 01081. x" type="journal article" year="1986">Allsopp & Carne, 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA74B36FFACCF1CF114FB1A8E2459AB" author="Cassis, G. & Weir, T. A." box="[1148,1381,1229,1255]" pageId="10" pageNumber="438" refId="ref10030" refString="Cassis, G. & Weir, T. A. (1992) Dynastinae. In: Houston, W. W. K. (ed), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Volume 9. Australian Government Publishing Service, Canberra, pp. 383 - 425." type="book" year="1992">Cassis & Weir 1992</bibRefCitation>
|
||
; Dechambre 2005;
|
||
<bibRefCitation id="EFA74B36FFACCF1CF42BFB268AB65847" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[323,503,1265,1291]" pageId="10" pageNumber="438" pagination="1919" refId="ref10501" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae Macleay 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Volume 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton, pp. 516 - 530." type="book chapter" year="2019">
|
||
Weir
|
||
<emphasis id="B942EAD5FFACCF1CF417FB268AF65847" box="[383,439,1265,1291]" italics="true" pageId="10" pageNumber="438">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
) to recognise the error, particularly given the absence of available types in
|
||
<collectingCountry id="F3217657FFACCF1CF05EFB268EDC5847" box="[1334,1437,1265,1291]" name="Australia" pageId="10" pageNumber="438">Australia</collectingCountry>
|
||
and restrictions on access to the Frey collection following his death in 1976 and its housing in NHMB in 1997. Our examination of the
|
||
<typeStatus id="548D8865FFACCF1CF41BFAEE8A94581F" box="[371,469,1337,1363]" pageId="10" pageNumber="438" type="holotype">holotype</typeStatus>
|
||
confirms the generic misplacement and the match of the distinct genitalia shape with the specimen collected near Dedari in 2016.
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFACCF1CF5AFFA578E5958AF" blockId="10.[151,1437,1193,1615]" pageId="10" pageNumber="438">
|
||
<emphasis id="B942EAD5FFACCF1CF5AFFA578A9C58D6" bold="true" box="[199,477,1408,1435]" pageId="10" pageNumber="438">Habitat and behaviour.</emphasis>
|
||
The two collection localities, Lake Johnston and Dedari, are both east of the Western Australian Wheatbelt (
|
||
<figureCitation id="130D2A42FFACCF1CF4FCFA728AA458F3" box="[404,485,1444,1471]" captionStart="FIGURE 56" captionStartId="18.[151,250,2003,2027]" captionTargetBox="[177,1411,1209,1980]" captionTargetId="figure-471@18.[177,1411,1209,1980]" captionTargetPageId="18" captionText="FIGURE 56. Known distribution of Epironastes species in southern Western Australia." figureDoi="http://doi.org/10.5281/zenodo.4410071" httpUri="https://zenodo.org/record/4410071/files/figure.png" pageId="10" pageNumber="438">Fig. 56</figureCitation>
|
||
) and are mallee-dominated yellow sandplains with annual rainfalls of about
|
||
<quantity id="4CCE9B22FFACCF1CF051FA728ED858F3" box="[1337,1433,1445,1471]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="438" unit="mm" value="300.0">300 mm</quantity>
|
||
. The collection site near Dedari has a
|
||
<taxonomicName id="4C364D44FFACCF1CF75FFA1E89EC58AF" box="[567,685,1481,1507]" class="Agaricomycetes" family="Tricholomataceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="438" phylum="Basidiomycota" rank="genus">
|
||
<emphasis id="B942EAD5FFACCF1CF75FFA1E89EC58AF" box="[567,685,1481,1507]" italics="true" pageId="10" pageNumber="438">Melaleuca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D44FFACCF1CF78CFA1E88C258AF" box="[740,899,1481,1507]" class="Magnoliopsida" family="Casuarinaceae" genus="Allocasuarina" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="438" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B942EAD5FFACCF1CF78CFA1E88C258AF" box="[740,899,1481,1507]" italics="true" pageId="10" pageNumber="438">Allocasuarina</emphasis>
|
||
</taxonomicName>
|
||
understorey with
|
||
<taxonomicName id="4C364D44FFACCF1CF125FA1E8FE658AF" authorityName="Linnaeus" authorityYear="1771" box="[1101,1191,1481,1507]" class="Liliopsida" family="Poaceae" genus="Spinifex" kingdom="Plantae" order="Poales" pageId="10" pageNumber="438" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B942EAD5FFACCF1CF125FA1E8FE658AF" box="[1101,1191,1481,1507]" italics="true" pageId="10" pageNumber="438">Spinifex</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130D2A42FFACCF1CF1DFFA1E8E4A58AF" box="[1207,1291,1481,1507]" captionStart="FIGURE 29" captionStartId="10.[151,250,1086,1110]" captionTargetBox="[177,1411,181,1062]" captionTargetId="figure-19@10.[177,1411,181,1062]" captionTargetPageId="10" captionText="FIGURE 29. Epironastes demarzi (Endrődi 1971) habitat at 1 km south of Dedari Pumping Station, 80 km west of Kalgoorlie along the Perth-Kalgoorlie water pipeline." figureDoi="http://doi.org/10.5281/zenodo.4410055" httpUri="https://zenodo.org/record/4410055/files/figure.png" pageId="10" pageNumber="438">Fig. 29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8936C7FFACCF1CF5AFFA3A89B05B03" blockId="10.[151,1437,1193,1615]" pageId="10" pageNumber="438">
|
||
The diurnal activity pattern of this species is not known, with collections of adults in late January and March. On
|
||
<date id="FF881007FFACCF1CF5A9F9C68A3C5B67" box="[193,381,1553,1579]" pageId="10" pageNumber="438" value="2016-01-27">27 January 2016</date>
|
||
near Dedari a large storm puddle contained a dead male, almost certainly having emerged in response to rainfall. No light trapping was conducted.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |