140 lines
16 KiB
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140 lines
16 KiB
XML
<document id="30C36DD3F30B0C80ECBF38D551C267DA" ID-DOI="10.5281/zenodo.5340370" ID-ISSN="2345-7600" ID-Zenodo-Dep="5340370" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630186249002" checkinUser="diego" docAuthor="Dan, Polhemus, John, Polhemus & Sites, Robert" docDate="2008" docId="039F87C7E57DFFCBFC3FFB183771F83F" docLanguage="en" docName="56rbz255-279.pdf" docOrigin="Raffles Bulletin of Zoology 56 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Coptocatus Montandon 1909" docType="treatment" docVersion="4" lastPageNumber="257" masterDocId="FFA6FFBFE57FFFC9FFFAFF893329FFF6" masterDocTitle="A Revision Of The Indochinese Genera Cheirochela And Gestroiella Heteroptera Naucoridae And A Review Of The Tribe Cheirochelini" masterLastPageNumber="279" masterPageNumber="255" pageNumber="257" updateTime="1699179848367" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5D18182BF469F6EBBECB6F4EE71D0C7B">A Revision Of The Indochinese Genera Cheirochela And Gestroiella Heteroptera Naucoridae And A Review Of The Tribe Cheirochelini</mods:title>
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<mods:namePart id="FA29729162CC16EF933892BF56511A7F">Polhemus</mods:namePart>
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<mods:namePart id="E4854AF49E42DAAD7BEBA658C9DDC9EB">John</mods:namePart>
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<mods:namePart id="E3174388FEA2F1FF11CE334BF60A9D7B">Polhemus</mods:namePart>
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<mods:namePart id="C3E0D735A38851CDC7C61E2BD9E3335E">Sites, Robert</mods:namePart>
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<mods:title id="6F94D302F7C351415687DA52BCC6FF8D">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="3678DD38596C9BDA060B752DFD5E2EBB">2008</mods:date>
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<treatment id="039F87C7E57DFFCBFC3FFB183771F83F" ID-DOI="http://doi.org/10.5281/zenodo.5416662" ID-Zenodo-Dep="5416662" LSID="urn:lsid:plazi:treatment:039F87C7E57DFFCBFC3FFB183771F83F" httpUri="http://treatment.plazi.org/id/039F87C7E57DFFCBFC3FFB183771F83F" lastPageNumber="257" pageId="2" pageNumber="257">
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<subSubSection id="C32C655AE57DFFCBFC3FFB18366AFB5F" box="[965,1347,1169,1193]" pageId="2" pageNumber="257" type="nomenclature">
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<paragraph id="8B8936D1E57DFFCBFC3FFB18366AFB5F" blockId="2.[965,1347,1169,1193]" box="[965,1347,1169,1193]" pageId="2" pageNumber="257">
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<heading id="D0C181BDE57DFFCBFC3FFB18366AFB5F" bold="true" box="[965,1347,1169,1193]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="257" reason="2">
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<taxonomicName id="4C364D52E57DFFCBFC3FFB18366AFB5F" ID-CoL="3TSG" authority="Montandon, 1909" authorityName="Montandon" authorityYear="1909" box="[965,1347,1169,1193]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EAC3E57DFFCBFC3FFB18366AFB5F" bold="true" box="[965,1347,1169,1193]" pageId="2" pageNumber="257">
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<emphasis id="B942EAC3E57DFFCBFC3FFB18375BFB5F" bold="true" box="[965,1138,1169,1193]" italics="true" pageId="2" pageNumber="257">COPTOCATUS</emphasis>
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<bibRefCitation id="EFA74B20E57DFFCBFB80FB18366AFB5F" author="Montandon, A" box="[1146,1347,1169,1193]" pageId="2" pageNumber="257" pagination="43 - 61" refId="ref21235" refString="Montandon, A. L., 1909. Naucoridae descriptions d'especes nouvelles. Bulletin de la Societe de Sciences de Bucarest- Roumanie, 18 (1): 43 - 61." type="journal article" year="1909">Montandon, 1909</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C655AE57DFFCBFCBFFB5837F7F9DF" pageId="2" pageNumber="257" type="diagnosis">
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<paragraph id="8B8936D1E57DFFCBFCBFFB5837F7F9DF" blockId="2.[837,1476,1233,1577]" pageId="2" pageNumber="257">
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<emphasis id="B942EAC3E57DFFCBFCBFFB5830F9FB1F" bold="true" box="[837,976,1233,1257]" italics="true" pageId="2" pageNumber="257">Diagnosis. –</emphasis>
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Form elongate-oval. (
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<figureCitation id="130D2A54E57DFFCBFB3CFB5B362EFB1F" box="[1222,1287,1234,1257]" captionStart="Fig" captionStartId="3.[813,848,1597,1618]" captionTargetBox="[814,1457,647,1554]" captionTargetId="figure-635@3.[797,1472,630,1575]" captionTargetPageId="3" captionText="Fig. 1. Coptocatus kinabalu Polhemus. Male, dorsal habitus. Drawn by Young Sohn." figureDoi="http://doi.org/10.5281/zenodo.5340372" httpUri="https://zenodo.org/record/5340372/files/figure.png" pageId="2" pageNumber="257">Fig. 1</figureCitation>
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) Abdominal and thoracic venter with fine pile of short, closely appressed hairs. Pronotum not markedly expanded posteriorly (except in
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<taxonomicName id="4C364D52E57DFFCBFC99FABB30E6FABF" authorityName="Nieser & Chen" authorityYear="1991" box="[867,975,1330,1353]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="stereos">
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<emphasis id="B942EAC3E57DFFCBFC99FABB305CFABF" box="[867,885,1330,1353]" italics="true" pageId="2" pageNumber="257">C</emphasis>
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.
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<emphasis id="B942EAC3E57DFFCBFC7EFABB30E6FABF" box="[900,975,1330,1353]" italics="true" pageId="2" pageNumber="257">stereos</emphasis>
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</taxonomicName>
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), posterior width less than 2× width of head; anterolateral angles of pronotum simple, not forming ventrallydirected cup-shaped depressions at apices. Hemelytra of brachypterous forms short and truncate, apices rounded, not reaching to abdominal tergite VI. Parameres variable in shape, either deeply cleft medially and asymmetrically bidentate, or uniramous and tapering on distal half, sometimes with a fringe of long setae on outer margins.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C655AE57DFFCBFCBFF9D83771F83F" pageId="2" pageNumber="257" type="discussion">
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<paragraph id="8B8936D1E57DFFCBFCBFF9D83771F83F" blockId="2.[837,1476,1617,1993]" pageId="2" pageNumber="257">
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<emphasis id="B942EAC3E57DFFCBFCBFF9D830FEF99F" bold="true" box="[837,983,1617,1641]" italics="true" pageId="2" pageNumber="257">Discussion. –</emphasis>
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The genus
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<taxonomicName id="4C364D52E57DFFCBFBAFF9DB37E7F99F" authorityName="Montandon" authorityYear="1909" box="[1109,1230,1618,1641]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EAC3E57DFFCBFBAFF9DB37E7F99F" box="[1109,1230,1618,1641]" italics="true" pageId="2" pageNumber="257">Coptocatus</emphasis>
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</taxonomicName>
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is confined to Borneo, and was proposed by
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<bibRefCitation id="EFA74B20E57DFFCBFBC1F9FB3623F97F" author="Montandon, A" box="[1083,1290,1650,1673]" pageId="2" pageNumber="257" pagination="43 - 61" refId="ref21235" refString="Montandon, A. L., 1909. Naucoridae descriptions d'especes nouvelles. Bulletin de la Societe de Sciences de Bucarest- Roumanie, 18 (1): 43 - 61." type="journal article" year="1909">Montandon (1909)</bibRefCitation>
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to hold a single species,
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<taxonomicName id="4C364D52E57DFFCBFC59F91B3715F95F" authorityName="Montandon" authorityYear="1909" box="[931,1084,1682,1705]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="oblongulus">
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<emphasis id="B942EAC3E57DFFCBFC59F91B309CF95F" box="[931,949,1682,1705]" italics="true" pageId="2" pageNumber="257">C</emphasis>
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.
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<emphasis id="B942EAC3E57DFFCBFC3FF91B3715F95F" box="[965,1084,1682,1705]" italics="true" pageId="2" pageNumber="257">oblongulus</emphasis>
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</taxonomicName>
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, collected near Mt. Kinabalu. Five additional species have been described subsequently by
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<bibRefCitation id="EFA74B20E57DFFCBFCBFF95B30D6F91F" author="Polhemus, D" box="[837,1023,1746,1769]" pageId="2" pageNumber="257" pagination="248 - 256" refId="ref21307" refString="Polhemus, D. A., 1986. A review of the genus Coptocatus Montandon (Hemiptera: Naucoridae). Pan-Pacific Entomologist, 62 (3): 248 - 256." type="journal article" year="1986">Polhemus (1986)</bibRefCitation>
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, Nieser & Chen (1991) and
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<bibRefCitation id="EFA74B20E57DFFCBFACAF95B36E9F91F" author="Zettel, H" box="[1328,1472,1746,1769]" pageId="2" pageNumber="257" pagination="171 - 178" refId="ref21521" refString="Zettel, H., 2002. Beschreibungen von zwei neuen Arten der Gattung Coptocatus Montandon (Heteroptera: Naucoridae: Cheirochelinae) aus Sarawak (Borneo), mit einem Bestimmungsschlusssel zu den Arten der Gattung. Entomological Problems, 32: 171 - 178." type="journal article" year="2002">Zettel (2002)</bibRefCitation>
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. Although clearly members of the tribe
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<taxonomicName id="4C364D52E57DFFCBFB17F97B3657F8FF" authorityName="Montandon" authorityYear="1897" box="[1261,1406,1778,1801]" pageId="2" pageNumber="257" rank="tribe" tribe="Cheirochelini">Cheirochelini</taxonomicName>
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based on synapomorphies of the head and ventral abdomen (see
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<bibRefCitation id="EFA74B20E57DFFCBFCBFF8BB30D8F8BF" author="Polhemus, D" box="[837,1009,1842,1865]" pageId="2" pageNumber="257" pagination="248 - 256" refId="ref21307" refString="Polhemus, D. A., 1986. A review of the genus Coptocatus Montandon (Hemiptera: Naucoridae). Pan-Pacific Entomologist, 62 (3): 248 - 256." type="journal article" year="1986">Polhemus, 1986</bibRefCitation>
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,
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<figureCitation id="130D2A54E57DFFCBFC00F8BB3773F8BF" box="[1018,1114,1842,1865]" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 7–11. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="2" pageNumber="257">Figs. 7, 8</figureCitation>
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),
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<taxonomicName id="4C364D52E57DFFCBFB94F8BB3610F8BF" box="[1134,1337,1842,1865]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B942EAC3E57DFFCBFB94F8BB37CEF8BF" box="[1134,1255,1842,1865]" italics="true" pageId="2" pageNumber="257">Coptocatus</emphasis>
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species
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</taxonomicName>
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are narrower and more elongate in comparison to those of
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<taxonomicName id="4C364D52E57DFFCBFAD4F8DB368CF89F" authorityName="Montandon" authorityYear="1897" box="[1326,1445,1874,1897]" class="Insecta" family="Naucoridae" genus="Gestroiella" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EAC3E57DFFCBFAD4F8DB368CF89F" box="[1326,1445,1874,1897]" italics="true" pageId="2" pageNumber="257">Gestroiella</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C364D52E57DFFCBFCBFF8FB30E1F87F" box="[837,968,1906,1929]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EAC3E57DFFCBFCBFF8FB30E1F87F" box="[837,968,1906,1929]" italics="true" pageId="2" pageNumber="257">Cheirochela</emphasis>
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</taxonomicName>
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(compare
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<figureCitation id="130D2A54E57DFFCBFBC4F8FB37A1F87F" box="[1086,1160,1906,1929]" captionStart="Fig" captionStartId="3.[813,848,1597,1618]" captionTargetBox="[814,1457,647,1554]" captionTargetId="figure-635@3.[797,1472,630,1575]" captionTargetPageId="3" captionText="Fig. 1. Coptocatus kinabalu Polhemus. Male, dorsal habitus. Drawn by Young Sohn." figureDoi="http://doi.org/10.5281/zenodo.5340372" httpUri="https://zenodo.org/record/5340372/files/figure.png" pageId="2" pageNumber="257">Figs. 1</figureCitation>
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,
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<figureCitation id="130D2A54E57DFFCBFB62F8FB378CF87F" box="[1176,1189,1906,1929]" captionStart="Fig" captionStartId="4.[836,871,2065,2086]" captionTargetBox="[844,1475,1140,2016]" captionTargetId="figure-14@4.[819,1504,1121,2034]" captionTargetPageId="4" captionText="Fig. 2. Cheirochela tonkina Polhemus & Polhemus. Male, dorsal habitus. Drawn by Young Sohn." figureDoi="http://doi.org/10.5281/zenodo.5340374" httpUri="https://zenodo.org/record/5340374/files/figure.png" pageId="2" pageNumber="257">2</figureCitation>
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and
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<figureCitation id="130D2A54E57DFFCBFB24F8FB37D2F87F" box="[1246,1275,1906,1929]" captionStart="Fig" captionStartId="12.[837,872,2065,2086]" captionTargetBox="[916,1399,1474,2029]" captionTargetId="figure-785@12.[903,1409,1461,2049]" captionTargetPageId="12" captionText="Fig. 18. Gestroiella siamensis new species, female, dorsal habitus. Drawn by Young Sohn." figureDoi="http://doi.org/10.5281/zenodo.5340386" httpUri="https://zenodo.org/record/5340386/files/figure.png" pageId="2" pageNumber="257">18</figureCitation>
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). The key below, modified from
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<bibRefCitation id="EFA74B20E57DFFCBFC19F81B3759F85F" author="Zettel, H" box="[995,1136,1938,1961]" pageId="2" pageNumber="257" pagination="171 - 178" refId="ref21521" refString="Zettel, H., 2002. Beschreibungen von zwei neuen Arten der Gattung Coptocatus Montandon (Heteroptera: Naucoridae: Cheirochelinae) aus Sarawak (Borneo), mit einem Bestimmungsschlusssel zu den Arten der Gattung. Entomological Problems, 32: 171 - 178." type="journal article" year="2002">Zettel (2002)</bibRefCitation>
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, will permit identification of the species described to date.
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</paragraph>
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</subSubSection>
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</treatment>
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