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<mods:title id="69B492B5A03C1AE5A90F1843E7440FE3">A Revision Of The Indochinese Genera Cheirochela And Gestroiella Heteroptera Naucoridae And A Review Of The Tribe Cheirochelini</mods:title>
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KEY TO SPECIES OF
<taxonomicName id="4C364D52E57BFFCDFE0BFDA8319AFDCF" ID-CoL="3MJZ" authorityName="Hope" authorityYear="1841" box="[497,691,545,569]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="B942EAC3E57BFFCDFE0BFDA8319AFDCF" bold="true" box="[497,691,545,569]" italics="true" pageId="4" pageNumber="259">CHEIROCHELA</emphasis>
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<subSubSection id="C32C655AE57BFFCDFF70FDE9302EF829" pageId="4" pageNumber="259" type="key">
<keyStep id="30C22E74E57BFFCDFF70FDE9302DFBFE" pageId="4" pageNumber="259">
<paragraph id="8B8936D1E57BFFCDFF70FDE9302CFCC9" blockId="4.[138,777,608,2015]" pageId="4" pageNumber="259">
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1. Large, robust species, body length usually exceeding
<quantity id="4CCE9B34E57BFFCDFD47FDE9302DFD83" box="[701,772,608,629]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="259" unit="mm" value="26.0">26 mm</quantity>
; overall body form broad (
<figureCitation id="130D2A54E57BFFCDFE49FDF432D8FD64" box="[435,497,637,658]" captionStart="Fig" captionStartId="4.[836,871,2065,2086]" captionTargetBox="[844,1475,1140,2016]" captionTargetId="figure-14@4.[819,1504,1121,2034]" captionTargetPageId="4" captionText="Fig. 2. Cheirochela tonkina Polhemus &amp; Polhemus. Male, dorsal habitus. Drawn by Young Sohn." figureDoi="http://doi.org/10.5281/zenodo.5340374" httpUri="https://zenodo.org/record/5340374/files/figure.png" pageId="4" pageNumber="259">Fig. 2</figureCitation>
), ratio of total body length/ body width across embolia falling in range of 1.501.65/1.00; posterolateral angles of pronotum highly produced and acuminate, ratio of maximum pronotal width/pronotal length along midline in submacropterous individuals falling in range of 3.784.21; male paramere shape as in
<figureCitation id="130D2A54E57BFFCDFDB8FC843152FCD4" box="[578,635,781,802]" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 711. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="4" pageNumber="259">Fig. 8</figureCitation>
.................... ................................................................
<taxonomicName id="4C364D52E57BFFCDFDC6FCA3302CFCC9" ID-CoL="TTYX" authority="Montandon" authorityName="Montandon" authorityYear="1897" box="[572,773,810,831]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="feana">
<emphasis id="B942EAC3E57BFFCDFDC6FCA331A7FCC9" box="[572,654,810,831]" italics="true" pageId="4" pageNumber="259">C. feana</emphasis>
Montandon
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<keyLead id="30C795E4E57BFFCDFF70FCCE302DFBFE" pageId="4" pageNumber="259">
Slightly smaller and more elongate species, body length usually less than
<quantity id="4CCE9B34E57BFFCDFEF0FCEA3278FC8E" box="[266,337,867,888]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="259" unit="mm" value="26.0">26 mm</quantity>
; overall form more elongate (
<figureCitation id="130D2A54E57BFFCDFD89FCEA3186FC8E" box="[627,687,867,888]" captionStart="Fig" captionStartId="4.[836,871,2065,2086]" captionTargetBox="[844,1475,1140,2016]" captionTargetId="figure-14@4.[819,1504,1121,2034]" captionTargetPageId="4" captionText="Fig. 2. Cheirochela tonkina Polhemus &amp; Polhemus. Male, dorsal habitus. Drawn by Young Sohn." figureDoi="http://doi.org/10.5281/zenodo.5340374" httpUri="https://zenodo.org/record/5340374/files/figure.png" pageId="4" pageNumber="259">Fig. 2</figureCitation>
), ratio of total body length/body width across embolia falling in range of 1.741.95/1.00; posterolateral angles of pronotum only moderately produced, not highly acuminate, ratio of maximum pronotal width/pronotal length along midline in submacropterous individuals falling in range of 3.153.60 ............................ 2
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<paragraph id="8B8936D1E57BFFCDFF70FB99302DFB43" blockId="4.[138,777,608,2015]" pageId="4" pageNumber="259">
<keyLead id="30C795E4E57BFFCDFF70FB99302DFB43" pageId="4" pageNumber="259">
2. Abdominal sternite V lacking a posteriorly directed medial tubercle; medial tubercle on sternite IV small and flattened, usually not projecting posteriorly over the base of sternite V when viewed laterally; preocular head 0.8 × the inner length of an eye or greater (
<figureCitation id="130D2A54E57BFFCDFE78FB0A3296FB6E" box="[386,447,1155,1176]" captionStart="Fig" captionStartId="4.[836,871,2065,2086]" captionTargetBox="[844,1475,1140,2016]" captionTargetId="figure-14@4.[819,1504,1121,2034]" captionTargetPageId="4" captionText="Fig. 2. Cheirochela tonkina Polhemus &amp; Polhemus. Male, dorsal habitus. Drawn by Young Sohn." figureDoi="http://doi.org/10.5281/zenodo.5340374" httpUri="https://zenodo.org/record/5340374/files/figure.png" pageId="4" pageNumber="259">Fig. 2</figureCitation>
); male paramere shape as in
<figureCitation id="130D2A54E57BFFCDFD18FB0A33E0FB43" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 711. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="4" pageNumber="259">Fig. 10</figureCitation>
; northern
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........................
<taxonomicName id="4C364D52E57BFFCDFDD9FB2931A0FB43" ID-CoL="TTZ2" authority="Dan &amp; Polhemus &amp; John &amp; Polhemus &amp; Sites, 2008" authorityName="Dan &amp; Polhemus &amp; John &amp; Polhemus &amp; Sites" authorityYear="2008" box="[547,649,1184,1205]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="tonkina" status="sp. nov.">
<emphasis id="B942EAC3E57BFFCDFDD9FB2931A0FB43" box="[547,649,1184,1205]" italics="true" pageId="4" pageNumber="259">C. tonkina</emphasis>
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<taxonomicNameLabel id="A27157B8E57BFFCDFD6AFB29302DFB43" box="[656,772,1184,1205]" pageId="4" pageNumber="259" rank="species">new species</taxonomicNameLabel>
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<keyLead id="30C795E4E57BFFCDFF70FB34302DFA94" pageId="4" pageNumber="259">
Abdominal sternite V with a well developed, posteriorly developed medial tubercle, tip of this tubercle sharply angulate (
<figureCitation id="130D2A54E57BFFCDFF4CFB7E3239FAFA" box="[182,272,1271,1292]" captionStart="Figs" captionStartId="5.[115,159,2065,2086]" captionTargetBox="[118,1452,1214,2007]" captionTargetId="figure-16@5.[85,1470,1181,2031]" captionTargetPageId="5" captionText="Figs. 36. Cheirochela assamensis Hope, holotype male, structural details: 3, Dorsal habitus; 4, Left antenna, dorsal view; 5, Terminal abdomen, ventral view; 6, Medial processes on abdominal ventrites IV and V, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5340376" httpUri="https://zenodo.org/record/5340376/files/figure.png" pageId="4" pageNumber="259">Figs. 5, 6</figureCitation>
); medial tubercle on segment IV also well developed, raised and angulate (
<figureCitation id="130D2A54E57BFFCDFE94FA9A328CFADE" box="[366,421,1299,1320]" captionStart="Figs" captionStartId="5.[115,159,2065,2086]" captionTargetBox="[118,1452,1214,2007]" captionTargetId="figure-16@5.[85,1470,1181,2031]" captionTargetPageId="5" captionText="Figs. 36. Cheirochela assamensis Hope, holotype male, structural details: 3, Dorsal habitus; 4, Left antenna, dorsal view; 5, Terminal abdomen, ventral view; 6, Medial processes on abdominal ventrites IV and V, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5340376" httpUri="https://zenodo.org/record/5340376/files/figure.png" pageId="4" pageNumber="259">Fig. 6</figureCitation>
), often projecting over base of sternite V when viewed laterally; preocular head 0.7 X the inner length of an eye or less;
<collectingCountry id="F3217641E57BFFCDFEA4FAC4328BFA94" box="[350,418,1357,1378]" name="Myanmar" pageId="4" pageNumber="259">Burma</collectingCountry>
,
<collectingCountry id="F3217641E57BFFCDFE57FAC4312CFA94" box="[429,517,1357,1378]" name="Thailand" pageId="4" pageNumber="259">Thailand</collectingCountry>
, or northeastern
<collectingCountry id="F3217641E57BFFCDFD52FAC431F0FA94" box="[680,729,1357,1378]" name="India" pageId="4" pageNumber="259">India</collectingCountry>
.. 3
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<paragraph id="8B8936D1E57BFFCDFF70FAE3302EF9DA" blockId="4.[138,777,608,2015]" pageId="4" pageNumber="259">
<keyLead id="30C795E4E57BFFCDFF70FAE3302EF9DA" pageId="4" pageNumber="259">
3. Smaller species, body length less than
<quantity id="4CCE9B34E57BFFCDFDE7FAE3314BFA89" box="[541,610,1386,1407]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="259" unit="mm" value="22.0">22 mm</quantity>
; medial tubercles on abdominal sternites IV and V not strongly produced posteriorly or projecting over basomedial sections of subsequent sternites (
<figureCitation id="130D2A54E57BFFCDFEF1FA49324CFA23" box="[267,357,1472,1493]" captionStart="Figs" captionStartId="5.[115,159,2065,2086]" captionTargetBox="[118,1452,1214,2007]" captionTargetId="figure-16@5.[85,1470,1181,2031]" captionTargetPageId="5" captionText="Figs. 36. Cheirochela assamensis Hope, holotype male, structural details: 3, Dorsal habitus; 4, Left antenna, dorsal view; 5, Terminal abdomen, ventral view; 6, Medial processes on abdominal ventrites IV and V, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5340376" httpUri="https://zenodo.org/record/5340376/files/figure.png" pageId="4" pageNumber="259">Figs. 5, 6</figureCitation>
); male paramere bearing a short, thumb-like process on basal half separated from main body of paramere by a deep incision (
<figureCitation id="130D2A54E57BFFCDFEB2FA7332ABF9F9" box="[328,386,1530,1551]" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 711. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="4" pageNumber="259">Fig. 7</figureCitation>
); Bramaputra River basin of northeastern
<collectingCountry id="F3217641E57BFFCDFF55F99E33C9F9DA" box="[175,224,1559,1580]" name="India" pageId="4" pageNumber="259">India</collectingCountry>
........................................................
<taxonomicName id="4C364D52E57BFFCDFDB9F99E302EF9DA" ID-CoL="TTYV" authority="Hope" authorityName="Hope" authorityYear="1841" box="[579,775,1559,1580]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B942EAC3E57BFFCDFDB9F99E31E5F9DA" box="[579,716,1559,1580]" italics="true" pageId="4" pageNumber="259">C. assamensis</emphasis>
Hope
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<paragraph id="8B8936D1E57BFFCDFF70F9BA302EF92E" blockId="4.[138,777,608,2015]" pageId="4" pageNumber="259">
<keyLead id="30C795E4E57BFFCDFF70F9BA302EF92E" pageId="4" pageNumber="259">
Larger species, body length exceeding
<quantity id="4CCE9B34E57BFFCDFDAEF9BA319CF9BE" box="[596,693,1587,1608]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="4" pageNumber="259" unit="mm" value="23.5">23.5 mm</quantity>
; medial tubercles on abdominal sternites IV and V produced posteriorly over bases of following segments when viewed laterally; male paramere lacking a short, thumb-like process on basal half separated from main body of paramere by a deep incision (
<figureCitation id="130D2A54E57BFFCDFD22F92E33CAF92E" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 711. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="4" pageNumber="259">Figs. 9, 11</figureCitation>
);
<collectingCountry id="F3217641E57BFFCDFF0CF94A3213F92E" box="[246,314,1731,1752]" name="Myanmar" pageId="4" pageNumber="259">Burma</collectingCountry>
,
<collectingCountry id="F3217641E57BFFCDFEBFF94A325DF92E" box="[325,372,1731,1752]" name="Laos" pageId="4" pageNumber="259">Laos</collectingCountry>
and
<collectingCountry id="F3217641E57BFFCDFE5FF94A32D3F92E" box="[421,506,1731,1752]" name="Thailand" pageId="4" pageNumber="259">Thailand</collectingCountry>
........................................ 4
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<keyLead id="30C795E4E57BFFCDFF70F969302EF8BA" pageId="4" pageNumber="259">
4. Anterolateral angles of pronotum strongly produced and spinose; male paramere lacking a deep notch at apex (
<figureCitation id="130D2A54E57BFFCDFD8FF974319BF8E4" box="[629,690,1789,1810]" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 711. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="4" pageNumber="259">Fig. 9</figureCitation>
); medial process of genital capsule notched at apex (Fig. 13);
<collectingCountry id="F3217641E57BFFCDFD3DF8933021F8D9" box="[711,776,1818,1839]" name="Myanmar" pageId="4" pageNumber="259">Burma</collectingCountry>
and
<collectingCountry id="F3217641E57BFFCDFF23F8BE3221F8BA" box="[217,264,1847,1868]" name="Laos" pageId="4" pageNumber="259">Laos</collectingCountry>
....................................
<taxonomicName id="4C364D52E57BFFCDFE02F8BE302EF8BA" ID-CoL="TTYW" authority="Montandon" authorityName="Montandon" authorityYear="1897" box="[504,775,1847,1868]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="birmaniensis">
<emphasis id="B942EAC3E57BFFCDFE02F8BE31B8F8BA" box="[504,657,1847,1868]" italics="true" pageId="4" pageNumber="259">C. birmaniensis</emphasis>
Montandon
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8936D1E57BFFCDFF70F8DA302EF829" blockId="4.[138,777,608,2015]" pageId="4" pageNumber="259">
<keyLead id="30C795E4E57BFFCDFF70F8DA302EF829" pageId="4" pageNumber="259">
Anterolateral angles of pronotum sharply angulate, but not strongly produced and spinose; male paramere bearing a deep notch at apex (
<figureCitation id="130D2A54E57BFFCDFEC7F80432ACF854" box="[317,389,1933,1954]" captionStart="Figs" captionStartId="6.[138,182,2065,2086]" captionTargetBox="[145,1490,188,2025]" captionTargetId="figure-14@6.[112,1498,155,2045]" captionTargetPageId="6" captionText="Figs. 711. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype." figureDoi="http://doi.org/10.5281/zenodo.5340378" httpUri="https://zenodo.org/record/5340378/files/figure.png" pageId="4" pageNumber="259">Fig. 11</figureCitation>
); medial process of genital capsule with apex truncate, not notched (Fig. 15);
<collectingCountry id="F3217641E57BFFCDFDE1F8233159F849" box="[539,624,1962,1983]" name="Thailand" pageId="4" pageNumber="259">Thailand</collectingCountry>
and
<collectingCountry id="F3217641E57BFFCDFD5BF82331F9F849" box="[673,720,1962,1983]" name="Laos" pageId="4" pageNumber="259">Laos</collectingCountry>
....... .....................................................
<taxonomicName id="4C364D52E57BFFCDFE00F84331A2F829" ID-CoL="TTYZ" authority="Dan &amp; Polhemus &amp; John &amp; Polhemus &amp; Sites, 2008" authorityName="Dan &amp; Polhemus &amp; John &amp; Polhemus &amp; Sites" authorityYear="2008" box="[506,651,1994,2015]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="thailandana" status="sp. nov.">
<emphasis id="B942EAC3E57BFFCDFE00F84331A2F829" box="[506,651,1994,2015]" italics="true" pageId="4" pageNumber="259">C. thailandana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157B8E57BFFCDFD68F843302EF829" box="[658,775,1994,2015]" pageId="4" pageNumber="259" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>