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<document id="F2B20FE07FA0EBE767B50ECEBC141374" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA77FFBCCBAFBC32FB97F94A" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Agladrillia rhodochroa Dautzenberg 1900" docType="treatment" docVersion="13" lastPageNumber="21" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="19" updateTime="1698184378282" updateUser="plazi">
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<mods:title id="96DBEC40D1AC5852C39E0ECE32527E29">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:namePart id="9755F0B9FC893FB8026A843E467ACE01">Fallon, Phillip J.</mods:namePart>
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<mods:date id="077FB479A27D8E3178E82F16085EFD6A">2016</mods:date>
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<treatment id="039F87C4FA77FFBCCBAFBC32FB97F94A" ID-DOI="http://doi.org/10.5281/zenodo.6076243" ID-GBIF-Taxon="117911508" ID-Zenodo-Dep="6076243" LSID="urn:lsid:plazi:treatment:039F87C4FA77FFBCCBAFBC32FB97F94A" httpUri="http://treatment.plazi.org/id/039F87C4FA77FFBCCBAFBC32FB97F94A" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="C32C6559FA77FFBACBAFBC32FD3CFA0E" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B8936D2FA77FFBACBAFBC32FD5BFAE9" blockId="18.[151,691,1321,1381]" box="[151,691,1321,1347]" pageId="18" pageNumber="19">
<heading id="D0C181BEFA77FFBACBAFBC32FD5BFAE9" bold="true" box="[151,691,1321,1347]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C364D51FA77FFBACBAFBC32FD5BFAE9" ID-CoL="5TRN6" authority="Dautzenberg, 1900" authorityName="Dautzenberg" authorityYear="1900" box="[151,691,1321,1347]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA77FFBACBAFBC32FD5BFAE9" bold="true" box="[151,691,1321,1347]" pageId="18" pageNumber="19">
<emphasis id="B942EAC0FA77FFBACBAFBC32FE46FAE9" bold="true" box="[151,430,1321,1347]" italics="true" pageId="18" pageNumber="19">Agladrillia rhodochroa</emphasis>
(Dautzenberg, 1900)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8936D2FA77FFBACBAFBC57FF1EFACF" blockId="18.[151,691,1321,1381]" box="[151,246,1356,1381]" pageId="18" pageNumber="19">
(
<figureCitation id="130D2A57FA77FFBACBA7BC57FF05FACF" box="[159,237,1356,1381]" captionStart="PLATE 2" captionStartId="19.[151,234,1812,1834]" captionTargetBox="[242,1315,195,1773]" captionTargetId="figure@19.[242,1315,195,1773]" captionTargetPageId="19" captionText="PLATE 2. Agladrillia rhodochroa (Dautzenberg, 1900). Upper panel. Fig. 1: lectotype, Rio Hacha, Guajira Dept., Colombia (RBINS / MT / 524919), ventral, lateral &amp; dorsal views. Images by Yves Barett, © RBINS, and courtesy of Thierry Backeljau. Figs. 2 3: same specimen in Fig. 5. Fig. 2: enlarged view of protoconch (not to scale); Fig. 3: apical view, V = varix, L = edge of outer lip. Lower panel. Fig. 4: reproduction of Dautzenberg (1900: pl. 9, fig. 3); Figs. 5 6: Tayrona National Park, Colombia (authors coll.); Fig. 7: Cartagena, Bolivar Dept., Colombia (USNM 900073); Figs. 8 10: Bahia Concha, Magdalena Dept., Colombia (P. Stahlschmidt coll.); Figs. 11 12: Punta de Betin, Santa Marta, Magdalena Dept., Colombia (P. Stahlschmidt coll.)." httpUri="https://zenodo.org/record/263301/files/figure.png" pageId="18" pageNumber="19">Plate 2</figureCitation>
)
</paragraph>
<paragraph id="8B8936D2FA77FFBACBAFBC95FD3CFA0E" blockId="18.[151,1437,1422,1659]" box="[151,724,1422,1444]" pageId="18" pageNumber="19">
<taxonomicName id="4C364D51FA77FFBACBAFBC95FDBCFA0E" authority="Dautzenberg, 1900: 165" authorityName="Dautzenberg" authorityPageNumber="165" authorityYear="1900" box="[151,596,1422,1444]" class="Gastropoda" family="Drilliidae" genus="Drillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA77FFBACBAFBC95FEB0FA0E" box="[151,344,1422,1444]" italics="true" pageId="18" pageNumber="19">Drillia rhodochroa</emphasis>
Dautzenberg, 1900: 165
</taxonomicName>
, pl. 9, fig. 3.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA77FFBACBAFBCB6FAADF951" pageId="18" pageNumber="19" type="description">
<paragraph id="8B8936D2FA77FFBACBAFBCB6FD02FA48" blockId="18.[151,1437,1422,1659]" pageId="18" pageNumber="19">
<taxonomicName id="4C364D51FA77FFBACBAFBCB6FDB4FA69" authority="Dautzenberg, 1900" authorityName="Dautzenberg" authorityYear="1900" box="[151,604,1453,1475]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA77FFBACBAFBCB6FE6AFA69" box="[151,386,1453,1475]" italics="true" pageId="18" pageNumber="19">Agladrillia rhodochroa</emphasis>
(Dautzenberg, 1900)
</taxonomicName>
: Díaz (1994: 40); Díaz &amp; Puyana (1994: 224, fig. 885); Williams (2009: 1503 [left photo only]); Massemin
<emphasis id="B942EAC0FA77FFBACAF8BCD6FE1CFA48" box="[448,500,1484,1506]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2009: 203, text photo).
</paragraph>
<paragraph id="8B8936D2FA77FFBACBAFBCF0FC63F994" blockId="18.[151,1437,1422,1659]" pageId="18" pageNumber="19">
<taxonomicName id="4C364D51FA77FFBACBAFBCF0FE9CF9AB" box="[151,372,1515,1537]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="scissurata">
<emphasis id="B942EAC0FA77FFBACBAFBCF0FE9CF9AB" box="[151,372,1515,1537]" italics="true" pageId="18" pageNumber="19">Agladrillia scissurata</emphasis>
</taxonomicName>
auct. non (Dall, 1890) are misidentifications by Williams (2009: 1502 [right photo only]) and Daccarett &amp; Bossio (2011: 129, fig. 778) that may be this species.
<taxonomicName id="4C364D51FA77FFBAC814BF12FB54F9B5" authority="Dall, 1890" authorityName="Dall" authorityYear="1890" box="[812,1212,1545,1567]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="scissurata">
<emphasis id="B942EAC0FA77FFBAC814BF12FBDBF9B5" box="[812,1075,1545,1567]" italics="true" pageId="18" pageNumber="19">Syntomodrillia scissurata</emphasis>
(Dall, 1890)
</taxonomicName>
is a fossil of the St. Petersburg, Caloosahatchie Formation [Florida Pliocene-Pleistocene].
</paragraph>
<paragraph id="8B8936D2FA77FFBACBAFBF5DFE8CF9D1" blockId="18.[151,1437,1422,1659]" pageId="18" pageNumber="19">
<taxonomicName id="4C364D51FA77FFBACBAFBF5DFEBEF9F6" box="[151,342,1606,1628]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="lassula">
<emphasis id="B942EAC0FA77FFBACBAFBF5DFEBEF9F6" box="[151,342,1606,1628]" italics="true" pageId="18" pageNumber="19">Agladrillia lassula</emphasis>
</taxonomicName>
auct non Jung, 1969: Daccarett &amp; Bossio (2011: 128129) erroneously place
<taxonomicName id="4C364D51FA77FFBACF44BF5DFAE7F9F6" box="[1148,1295,1606,1628]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA77FFBACF44BF5DFAE7F9F6" box="[1148,1295,1606,1628]" italics="true" pageId="18" pageNumber="19">A. rhodochroa</emphasis>
</taxonomicName>
in synonymy with
<taxonomicName id="4C364D51FA77FFBACBC2BF7EFEB6F9D1" box="[250,350,1637,1659]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="lassula">
<emphasis id="B942EAC0FA77FFBACBC2BF7EFEB6F9D1" box="[250,350,1637,1659]" italics="true" pageId="18" pageNumber="19">A. lassula</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8936D2FA77FFBACBAFBFB2FCD0F914" blockId="18.[151,1349,1704,1787]" box="[151,824,1704,1726]" pageId="18" pageNumber="19">
<emphasis id="B942EAC0FA77FFBACBAFBFB2FCD0F914" bold="true" box="[151,824,1704,1726]" pageId="18" pageNumber="19">Not this species or unconfirmed fossil and extralimital reports.</emphasis>
</paragraph>
<paragraph id="8B8936D2FA77FFBACBAFBFDDFAADF951" blockId="18.[151,1349,1704,1787]" pageId="18" pageNumber="19">
<taxonomicName id="4C364D51FA77FFBACBAFBFDDFEEEF976" box="[151,262,1734,1756]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA77FFBACBAFBFDDFEEEF976" box="[151,262,1734,1756]" italics="true" pageId="18" pageNumber="19">Agladrillia</emphasis>
</taxonomicName>
(?)
<taxonomicName id="4C364D51FA77FFBACA17BFDDFE04F976" authority="Jung, 1969" authorityName="Jung" authorityYear="1969" box="[303,492,1734,1756]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="lassula">
<emphasis id="B942EAC0FA77FFBACA17BFDDFE9FF976" box="[303,375,1734,1756]" italics="true" pageId="18" pageNumber="19">lassula</emphasis>
Jung, 1969
</taxonomicName>
[fossil]: is placed in synonymy with
<taxonomicName id="4C364D51FA77FFBAC85CBFDDFC1EF976" box="[868,1014,1734,1756]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA77FFBAC85CBFDDFC1EF976" box="[868,1014,1734,1756]" italics="true" pageId="18" pageNumber="19">A. rhodochroa</emphasis>
</taxonomicName>
by Diaz &amp; Puyana (1994: 224).
<taxonomicName id="4C364D51FA77FFBACBAFBFFEFE6AF951" box="[151,386,1765,1787]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA77FFBACBAFBFFEFE6AF951" box="[151,386,1765,1787]" italics="true" pageId="18" pageNumber="19">Agladrillia rhodochroa</emphasis>
</taxonomicName>
(auct. non Dautzenberg, 1900): Williams (2009: 1503, right photo only [=
<taxonomicName id="4C364D51FA77FFBACF4CBFFEFAE2F951" box="[1140,1290,1765,1787]" class="Gastropoda" family="Drilliidae" genus="Syntomodrillia" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA77FFBACF4CBFFEFAE2F951" box="[1140,1290,1765,1787]" italics="true" pageId="18" pageNumber="19">Syntomodrillia</emphasis>
</taxonomicName>
sp.]).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA77FFBCCBAFBE3FFA99FE5F" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8B8936D2FA77FFBACBAFBE3FFDDDF845" blockId="18.[151,1437,1827,2031]" pageId="18" pageNumber="19">
<emphasis id="B942EAC0FA77FFBACBAFBE3FFEADF896" bold="true" box="[151,325,1827,1852]" pageId="18" pageNumber="19">
<typeStatus id="548D8870FA77FFBACBAFBE3FFF3AF891" box="[151,210,1828,1851]" pageId="18" pageNumber="19">Type</typeStatus>
material.
</emphasis>
The illustrated
<typeStatus id="548D8870FA77FFBACACDBE3FFDCDF891" box="[501,549,1828,1851]" pageId="18" pageNumber="19">type</typeStatus>
(note on label accompanying specimen),
<date id="FF881012FA77FFBAC8C1BE38FBD6F891" box="[1017,1086,1827,1851]" pageId="18" pageNumber="19" value="2011-10-04">
11 x
<quantity id="4CCE9B37FA77FFBACF08BE38FBD6F891" box="[1072,1086,1827,1851]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="19" unit="mm" value="4.0">4</quantity>
</date>
mm, is in RBINS/MT/524919. It was not examined; instead, high quality photographs were provided by the museum that are reproduced in Pl. 2,
<figureCitation id="130D2A57FA77FFBACBAFBE70FF37F829" box="[151,223,1899,1924]" captionStart="Fig. 1" captionStartId="66.[151,188,1159,1181]" captionTargetBox="[151,1436,375,1146]" captionTargetId="figure@66.[151,1436,375,1146]" captionTargetPageId="66" captionText="Fig. 1: ventral, lateral &amp; dorsal views; Fig. 2: enlarged view of protoconch (not to scale); Fig. 3: apical view, V = varix, L = edge of outer lip." httpUri="https://zenodo.org/record/263325/files/figure.png" pageId="18" pageNumber="19">Fig. 1</figureCitation>
. Inasmuch as this specimen is the illustrated
<typeStatus id="548D8870FA77FFBAC83EBE77FCD1F829" box="[774,825,1900,1923]" pageId="18" pageNumber="19">type</typeStatus>
, it is here declared the
<typeStatus id="548D8870FA77FFBACF65BE70FB2FF82E" box="[1117,1223,1899,1924]" pageId="18" pageNumber="19" type="lectotype">lectotype</typeStatus>
. The existence of
<typeStatus id="548D8870FA77FFBACBAFBE95FED1F80D" box="[151,313,1934,1959]" pageId="18" pageNumber="19" type="paralectotype">paralectotypes</typeStatus>
is surmised from Dautzenbergs list of stations at which the species was reportedly taken. Dautzenbergs original illustration is reproduced in Pl. 2, Fig. 4 and differs principally in the number of ribs, probably a liberty taken by the artist.
</paragraph>
<caption id="DF49665AFA76FFBBCBAFBE0FFEB5F858" httpUri="https://zenodo.org/record/263301/files/figure.png" pageId="19" pageNumber="20" targetBox="[242,1315,195,1773]" targetPageId="19">
<paragraph id="8B8936D2FA76FFBBCBAFBE0FFEB5F858" blockId="19.[151,1437,1812,2034]" pageId="19" pageNumber="20">
<emphasis id="B942EAC0FA76FFBBCBAFBE0FFEEBF880" bold="true" box="[151,259,1812,1834]" pageId="19" pageNumber="20">PLATE 2.</emphasis>
<taxonomicName id="4C364D51FA76FFBBCA33BE0FFD3BF880" authority="Dautzenberg, 1900" authorityName="Dautzenberg" authorityYear="1900" box="[267,723,1812,1834]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA76FFBBCA33BE0FFE1FF880" box="[267,503,1812,1834]" italics="true" pageId="19" pageNumber="20">Agladrillia rhodochroa</emphasis>
(Dautzenberg, 1900)
</taxonomicName>
.
<emphasis id="B942EAC0FA76FFBBC9E5BE0EFC8CF880" bold="true" box="[733,868,1813,1834]" pageId="19" pageNumber="20">Upper panel</emphasis>
. Fig. 1: lectotype, Rio Hacha, Guajira Dept., Colombia (RBINS/MT/524919), ventral, lateral &amp; dorsal views. Images by Yves Barett, © RBINS, and courtesy of Thierry Backeljau. Figs. 23: same specimen in Fig. 5. Fig. 2: enlarged view of protoconch (not to scale); Fig. 3: apical view, V = varix, L = edge of outer lip.
<emphasis id="B942EAC0FA76FFBBCA1BBE62FE5DF824" bold="true" box="[291,437,1913,1934]" pageId="19" pageNumber="20">Lower panel.</emphasis>
Fig. 4: reproduction of Dautzenberg (1900: pl. 9, fig. 3); Figs. 56: Tayrona National Park, Colombia (authors coll.); Fig. 7: Cartagena, Bolivar Dept., Colombia (USNM 900073); Figs. 810: Bahia Concha, Magdalena Dept., Colombia (P. Stahlschmidt coll.); Figs. 1112: Punta de Betin, Santa Marta, Magdalena Dept., Colombia (P. Stahlschmidt coll.).
</paragraph>
</caption>
<paragraph id="8B8936D2FA71FFBCCBFFB983FC50FF52" blockId="20.[151,1437,151,1760]" pageId="20" pageNumber="21">
<emphasis id="B942EAC0FA71FFBCCBFFB983FE84FF1A" bold="true" box="[199,364,151,176]" pageId="20" pageNumber="21">
<typeStatus id="548D8870FA71FFBCCBFFB983FEEAFF05" box="[199,258,152,175]" pageId="20" pageNumber="21">Type</typeStatus>
locality.
</emphasis>
Rio Hacha [Guajira Dept.]
<emphasis id="B942EAC0FA71FFBCC9F9B98CFCCDFF1A" box="[705,805,151,176]" italics="true" pageId="20" pageNumber="21">Chazalie</emphasis>
Station 52,
<collectingCountry id="F3217642FA71FFBCC885B98CFBC6FF1A" box="[957,1070,151,176]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
, at
<quantity id="4CCE9B37FA71FFBCCF59B983FB7BFF05" box="[1121,1171,152,176]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="m" value="6.0">6 m</quantity>
depth (locality of the
<typeStatus id="548D8870FA71FFBCCBAFB9A7FEEBFF7F" box="[151,259,188,213]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
). Other specimens from the
<typeStatus id="548D8870FA71FFBCC96DB9A6FD6DFF7E" box="[597,645,189,212]" pageId="20" pageNumber="21">type</typeStatus>
series were collected at
<emphasis id="B942EAC0FA71FFBCC897B9A7FBFBFF7F" box="[943,1043,188,213]" italics="true" pageId="20" pageNumber="21">Chazalie</emphasis>
Stations 41, 42, 44, Santa
<collectingCountry id="F3217642FA71FFBCCE61B9A6FA74FF7E" box="[1369,1436,189,212]" name="Malta" pageId="20" pageNumber="21">Marta</collectingCountry>
[Magdalena Dept.],
<collectingCountry id="F3217642FA71FFBCCA40B9C4FE01FF52" box="[376,489,223,248]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
, at 15, 40, and
<quantity id="4CCE9B37FA71FFBCC9ABB9FBFD25FF5D" box="[659,717,224,248]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="21" unit="m" value="23.0">23 m</quantity>
depths, respectively.
</paragraph>
<paragraph id="8B8936D2FA71FFBCCBFFB81FFB8FFE07" blockId="20.[151,1437,151,1760]" pageId="20" pageNumber="21">
<emphasis id="B942EAC0FA71FFBCCBFFB81FFE58FEB7" bold="true" box="[199,432,260,285]" pageId="20" pageNumber="21">Material examined.</emphasis>
An total of
<specimenCount id="9D30FD5BFA71FFBCC90DB81FFD57FEB7" box="[565,703,260,285]" pageId="20" pageNumber="21" type="generic">8 specimens</specimenCount>
were examined: 2 spec., 10.2 x 4.1 &amp; 10.3 x 4.0 mm, from in soft sand at
<quantity id="4CCE9B37FA71FFBCCBD5B833FECFFE95" box="[237,295,296,320]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="20" pageNumber="21" unit="m" value="18.0">18 m</quantity>
, Tayrona Park, Santa
<collectingCountry id="F3217642FA71FFBCC924B833FD8AFE95" box="[540,610,296,319]" name="Malta" pageId="20" pageNumber="21">Marta</collectingCountry>
, Magdalena Dept.,
<collectingCountry id="F3217642FA71FFBCC804B83CFC42FEEA" box="[828,938,295,320]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
(authors coll.); 2 spec., 13.6 x 4.5 &amp; 12.7 x
<quantity id="4CCE9B37FA71FFBCCBAFB857FF07FECE" box="[151,239,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="20" pageNumber="21" unit="mm" value="4.8">4.8 mm</quantity>
, in
<quantity id="4CCE9B37FA71FFBCCA2EB857FE9DFECE" box="[278,373,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="20" pageNumber="21" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">26 mm</quantity>
, Punta de Betin, Santa
<collectingCountry id="F3217642FA71FFBCC94FB856FD55FECE" box="[631,701,333,356]" name="Malta" pageId="20" pageNumber="21">Marta</collectingCountry>
, Magdalena Dept.,
<collectingCountry id="F3217642FA71FFBCC8AEB857FBECFECF" box="[918,1028,332,357]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
(P. Stahlschmidt coll.); 3 spec., 12.6 x 4.6, 10.9 x 3.9 &amp; 10.3 x
<quantity id="4CCE9B37FA71FFBCCA85B86BFDFDFE2D" box="[445,533,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="20" pageNumber="21" unit="mm" value="3.9">3.9 mm</quantity>
, in
<quantity id="4CCE9B37FA71FFBCC905B86BFD4AFE2D" box="[573,674,368,392]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="20" pageNumber="21" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">2030 m</quantity>
, Bahia Concha, Magdalena Dept.,
<collectingCountry id="F3217642FA71FFBCCF10B874FB7EFE22" box="[1064,1174,367,392]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
(P. Stahlschmidt coll.); 1 spec., 11.8 x
<quantity id="4CCE9B37FA71FFBCCA06B88FFE7EFE06" box="[318,406,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="20" pageNumber="21" unit="mm" value="4.6">4.6 mm</quantity>
, in
<quantity id="4CCE9B37FA71FFBCCA87B88FFE12FE06" box="[447,506,404,428]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="20" pageNumber="21" unit="m" value="37.0">37 m</quantity>
, Cartagena, Bolivar Dept.,
<collectingCountry id="F3217642FA71FFBCC814B88FFC72FE07" box="[812,922,404,429]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
(USNM 900073).
</paragraph>
<paragraph id="8B8936D2FA71FFBCCBFFB8A3FA99FE5F" blockId="20.[151,1437,151,1760]" pageId="20" pageNumber="21">
<emphasis id="B942EAC0FA71FFBCCBFFB8A3FE46FE7A" bold="true" box="[199,430,439,464]" pageId="20" pageNumber="21">Range and habitat.</emphasis>
<collectingCountry id="F3217642FA71FFBCCA8EB8ACFDCCFE7A" box="[438,548,439,464]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
(Bolivar, Magdalena &amp; Guajira Depts.), at depths between 2 and
<specimenCount id="9D30FD5BFA71FFBCCE2DB8A3FABEFE65" box="[1301,1366,440,464]" pageId="20" pageNumber="21" type="generic">
<quantity id="4CCE9B37FA71FFBCCE2DB8A3FAA6FE65" box="[1301,1358,440,464]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="21" unit="m" value="60.0">60 m</quantity>
.
</specimenCount>
If the specimen illustrated by Massemin
<emphasis id="B942EAC0FA71FFBCC924B8C6FDBCFE5F" box="[540,596,476,501]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
is this species, then its southern range would extend to
<collectingCountry id="F3217642FA71FFBCCFFBB8C7FA85FE5F" box="[1219,1389,476,501]" name="French Guiana" pageId="20" pageNumber="21">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA71FFBCCBFFB8E4FEC7FB42" pageId="20" pageNumber="21" type="description">
<paragraph id="8B8936D2FA71FFBCCBFFB8E4FEC7FB42" blockId="20.[151,1437,151,1760]" pageId="20" pageNumber="21">
<emphasis id="B942EAC0FA71FFBCCBFFB8E4FE77FDB2" bold="true" box="[199,415,511,536]" pageId="20" pageNumber="21">
Description.
<emphasis id="B942EAC0FA71FFBCCA5CB8E4FE77FDB2" bold="true" box="[356,415,511,536]" italics="true" pageId="20" pageNumber="21">Shell</emphasis>
</emphasis>
small (to
<quantity id="4CCE9B37FA71FFBCC925BB1BFD6EFDBD" box="[541,646,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="20" pageNumber="21" unit="mm" value="13.6">13.6 mm</quantity>
in total length), glossy, fusiform, up to about 10¼ slightly convex whorls, the last approximately 53% of total length, flattened dorso-ventrally and rapidly tapering, “pinched” when viewed ventrally, to a narrow anterior canal. Spire whorls appear compressed—proportionally shorter than wide. Shell sculpture of ribs with deep intercostal spiral grooves.
<emphasis id="B942EAC0FA71FFBCC842BB77FC17FD2F" bold="true" box="[890,1023,620,645]" italics="true" pageId="20" pageNumber="21">Protoconch</emphasis>
of approximately 1½1¾ smooth translucent whorls, the first not immersed in the second.
<emphasis id="B942EAC0FA71FFBCC82FBB94FC20FD02" bold="true" box="[791,968,655,680]" italics="true" pageId="20" pageNumber="21">Axial sculpture</emphasis>
of narrow opisthocline ribs that run from suture-to-suture on spire whorls, evanesce at anterior fasciole of last whorl and are absent on dorsum of last; intercostal spaces greater than rib width. Ribs recurved in sulcus, sigmoid on ventrum of last whorl, number 1214 on penultimate, 79 on last whorl to varix.
<emphasis id="B942EAC0FA71FFBCC9B2BBE7FCACFCBF" bold="true" box="[650,836,764,789]" italics="true" pageId="20" pageNumber="21">Spiral sculpture</emphasis>
of evenly spaced deep grooves in intercostal spaces (rib crests are smooth) more closely spaced in sulcus. Grooves present across the dorsum of the last whorl. Grooves give the appearance of closely packed spiral cords, number about 8 on penultimate and 12 on shell base, and 7 on anterior fasciole.
<emphasis id="B942EAC0FA71FFBCCA62BA7CFE40FC2A" bold="true" box="[346,424,871,896]" italics="true" pageId="20" pageNumber="21">Sulcus</emphasis>
with recurved ribs that are lower than the ribs on the whorl face, ribs overridden by spiral grooves giving sulcus the appearance of having 34 spiral cords.
<emphasis id="B942EAC0FA71FFBCC84DBA97FC59FC0F" bold="true" box="[885,945,908,933]" italics="true" pageId="20" pageNumber="21">Varix</emphasis>
hump-like, positioned ½-turn from edge of outer lip; its position causing the body whorl to appear compressed dorso-laterally.
<emphasis id="B942EAC0FA71FFBCCF03BAB4FB4AFC62" bold="true" box="[1083,1186,943,968]" italics="true" pageId="20" pageNumber="21">Outer lip</emphasis>
thin, but strengthened by up to 4 axial folds clustered on about the last ¼-turn, the one closest to the edge the strongest; edge forms a low almost flat arch from anal sinus to stromboid notch where it is bent slightly inward. Stromboid notch deep. The dorsum of the last whorl thin, translucent, and except for their presence in the sulcus, ribs obsolete.
<emphasis id="B942EAC0FA71FFBCCE25BD07FA74FB9F" bold="true" box="[1309,1436,1052,1077]" italics="true" pageId="20" pageNumber="21">Anal sinus</emphasis>
deep, U-shaped, entrance constricted by parietal lobe and projection of outer lip, somewhat tubular and projecting laterally.
<emphasis id="B942EAC0FA71FFBCCA3EBD7EFE9BFBD7" bold="true" box="[262,371,1124,1149]" italics="true" pageId="20" pageNumber="21">Inner lip</emphasis>
edge free, erect anteriorly, recumbent on parietal wall, and formed into a parietal lobe posteriorly, arching over the rear of the aperture.
<emphasis id="B942EAC0FA71FFBCC985BD9CFC8EFB0A" bold="true" box="[701,870,1159,1184]" italics="true" pageId="20" pageNumber="21">Anterior canal</emphasis>
moderately long, open and narrow, turned slightly to the right and dorsally; deeply notched.
<emphasis id="B942EAC0FA71FFBCC952BDB7FD43FB6F" bold="true" box="[618,683,1196,1221]" italics="true" pageId="20" pageNumber="21">Color</emphasis>
pink, rib crests lighter, varix white, apex violet, and dorsum of last whorl brown.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA71FFBCCBFFBDEEFB97F94A" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B8936D2FA71FFBCCBFFBDEEFB97F94A" blockId="20.[151,1437,151,1760]" pageId="20" pageNumber="21">
<emphasis id="B942EAC0FA71FFBCCBFFBDEEFE52FAA6" bold="true" box="[199,442,1269,1292]" pageId="20" pageNumber="21">
Remarks.
<emphasis id="B942EAC0FA71FFBCCA7CBDEEFE52FAA6" bold="true" box="[324,442,1269,1292]" italics="true" pageId="20" pageNumber="21">Taxonomy</emphasis>
</emphasis>
.
<taxonomicName id="4C364D51FA71FFBCCAF3BDEFFD3AFAA7" box="[459,722,1268,1293]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA71FFBCCAF3BDEFFD3AFAA7" box="[459,722,1268,1293]" italics="true" pageId="20" pageNumber="21">Agladrillia rhodochroa</emphasis>
</taxonomicName>
has the distinguishing characteristics of the genus: a varix ½- turn from the edge of the outer lip, a strongly constricted anterior canal, and a smooth dorsum on last whorl. Dautzenberg described his specimens, which were from
<collectingCountry id="F3217642FA71FFBCC82ABC27FC6BFAFF" box="[786,899,1340,1365]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
, as having a pink base, ribs a purplish rose, and the back of the last whorl a wide brown spot. The specimens depicted in
<figureCitation id="130D2A57FA71FFBCC8F1BC44FBF0FAD2" box="[969,1048,1375,1400]" captionStart="PLATE 2" captionStartId="19.[151,234,1812,1834]" captionTargetBox="[242,1315,195,1773]" captionTargetId="figure@19.[242,1315,195,1773]" captionTargetPageId="19" captionText="PLATE 2. Agladrillia rhodochroa (Dautzenberg, 1900). Upper panel. Fig. 1: lectotype, Rio Hacha, Guajira Dept., Colombia (RBINS / MT / 524919), ventral, lateral &amp; dorsal views. Images by Yves Barett, © RBINS, and courtesy of Thierry Backeljau. Figs. 2 3: same specimen in Fig. 5. Fig. 2: enlarged view of protoconch (not to scale); Fig. 3: apical view, V = varix, L = edge of outer lip. Lower panel. Fig. 4: reproduction of Dautzenberg (1900: pl. 9, fig. 3); Figs. 5 6: Tayrona National Park, Colombia (authors coll.); Fig. 7: Cartagena, Bolivar Dept., Colombia (USNM 900073); Figs. 8 10: Bahia Concha, Magdalena Dept., Colombia (P. Stahlschmidt coll.); Figs. 11 12: Punta de Betin, Santa Marta, Magdalena Dept., Colombia (P. Stahlschmidt coll.)." httpUri="https://zenodo.org/record/263301/files/figure.png" pageId="20" pageNumber="21">Plate 2</figureCitation>
, all from
<collectingCountry id="F3217642FA71FFBCCFBDBC44FB1EFAD2" box="[1157,1270,1375,1400]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
, closely match his description and drawing.
<emphasis id="B942EAC0FA71FFBCCAE4BC9FFDB2FA37" bold="true" box="[476,602,1412,1437]" italics="true" pageId="20" pageNumber="21">Variability.</emphasis>
The average length of 9 measured specimens is
<quantity id="4CCE9B37FA71FFBCCF44BC9FFB06FA36" box="[1148,1262,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.137" pageId="20" pageNumber="21" unit="mm" value="11.37">11.37 mm</quantity>
(
<quantity id="4CCE9B37FA71FFBCCFC5BC9FFA7CFA37" box="[1277,1428,1412,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.3599999999999999" metricValueMin="0.99" pageId="20" pageNumber="21" unit="mm" value="11.75" valueMax="13.6" valueMin="9.9">9.913.6 mm</quantity>
) and average W/
<taxonomicName id="4C364D51FA71FFBCCA71BCBCFE7DFA6A" box="[329,405,1447,1472]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="ratio">L ratio</taxonomicName>
is 0.376. Some difference in color and form is seen between the Cartagena and Magdalena Departments specimens that is believed at this time to be geographic variation, not rising to the level of species.
<emphasis id="B942EAC0FA71FFBCCBAFBCF4FDA0F9A2" box="[151,584,1519,1544]" italics="true" pageId="20" pageNumber="21">
<emphasis id="B942EAC0FA71FFBCCBAFBCF4FED3F9A2" bold="true" box="[151,315,1519,1544]" italics="true" pageId="20" pageNumber="21">Identification.</emphasis>
<taxonomicName id="4C364D51FA71FFBCCA7BBCF4FDA0F9A2" box="[323,584,1519,1544]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="rhodochroa">Agladrillia rhodochroa</taxonomicName>
</emphasis>
is most similar to
<taxonomicName id="4C364D51FA71FFBCC823BCEBFC79F9A2" box="[795,913,1519,1544]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="aureola" status="sp. nov.">
<emphasis id="B942EAC0FA71FFBCC823BCEBFC79F9A2" box="[795,913,1519,1544]" italics="true" pageId="20" pageNumber="21">A. aureola</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFA71FFBCC898BCEBFBC3F9A2" box="[928,1067,1519,1544]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
. It differs in color, which is pink with a brown varical spot and not golden yellow with an orange brown varical spot, and in relative whorl height, which is higher in
<taxonomicName id="4C364D51FA71FFBCCA43BF23FE1DF9FA" box="[379,501,1591,1616]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="B942EAC0FA71FFBCCA43BF23FE1DF9FA" box="[379,501,1591,1616]" italics="true" pageId="20" pageNumber="21">A. aureola</emphasis>
</taxonomicName>
. Also, whorls are less convex and axial ribs more oblique than in
<taxonomicName id="4C364D51FA71FFBCCE23BF23FA7DF9FA" box="[1307,1429,1591,1616]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="B942EAC0FA71FFBCCE23BF23FA7DF9FA" box="[1307,1429,1591,1616]" italics="true" pageId="20" pageNumber="21">A. aureola</emphasis>
</taxonomicName>
.
<taxonomicName id="4C364D51FA71FFBCCBAFBF47FE77F9DF" box="[151,415,1628,1653]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="rhodochroa">
<emphasis id="B942EAC0FA71FFBCCBAFBF47FE77F9DF" box="[151,415,1628,1653]" italics="true" pageId="20" pageNumber="21">Agladrillia rhodochroa</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C364D51FA71FFBCC904BF47FCA8F9DF" box="[572,832,1628,1653]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="tobagoensis" status="sp. nov.">
<emphasis id="B942EAC0FA71FFBCC904BF47FCA8F9DF" box="[572,832,1628,1653]" italics="true" pageId="20" pageNumber="21">Calliclava tobagoensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFA71FFBCC869BF46FC35F9DF" box="[849,989,1628,1653]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
, in possessing a smooth protoconch, a greater constriction of the anterior canal and a varix located ½-turn from the edge of the outer lip. The latter species has a carinate protoconch, a less-constricted anterior canal, and a varix located about ⅓-turn from the edge of the outer lip. Both species of
<taxonomicName id="4C364D51FA71FFBCCA8EBFDCFDD9F94A" box="[438,561,1735,1760]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA71FFBCCA8EBFDCFDD9F94A" box="[438,561,1735,1760]" italics="true" pageId="20" pageNumber="21">Agladrillia</emphasis>
</taxonomicName>
and of
<taxonomicName id="4C364D51FA71FFBCC9BEBFDCFD13F94A" box="[646,763,1735,1760]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA71FFBCC9BEBFDCFD13F94A" box="[646,763,1735,1760]" italics="true" pageId="20" pageNumber="21">Calliclava</emphasis>
</taxonomicName>
are show in the lineup in
<figureCitation id="130D2A57FA71FFBCCF25BFDCFB93F94A" box="[1053,1147,1735,1760]" captionStart="PLATE 20" captionStartId="53.[151,234,1615,1637]" captionTargetBox="[151,1436,195,1588]" captionTargetId="figure@53.[151,1436,195,1610]" captionTargetPageId="53" captionText="PLATE 20. Agladrillia and Calliclava spp. map and lineup. Fig. 1. Approximate known localities. Fig. 2. Holotype, 21.6 x 6.3 mm, A. torquata, new species (MNHN IM- 2013 - 56291). Fig. 3. Holotype, 16.1 x 6.0 mm, C. fasciata, new species (UF 470272). Fig. 4. Holotype, 14.9 x 6.0 mm, C. vigorata, new species (ANSP 324013). Fig. 5. Holotype, 12.0 x 4.3 mm, A. aureola, new species (USNM 1291318). Fig. 6. Holotype, 11.3 x 4.3 mm, C. tobagoensis, new species (USNM 1291324). Fig. 7. Lectotype, 9.9 x 3.9 mm, A. rhodochroa (Dautzenberg, 1900) (RBINS / MT / 524919). Image by Yves Barett, © RBINS, and courtesy of Thierry Backeljau of the RBINS." httpUri="https://zenodo.org/record/263319/files/figure.png" pageId="20" pageNumber="21">Plate 20</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>