treatments-xml/data/03/9F/87/039F87A5676DFFB34D29FD94FE6AFCD7.xml

<document id="CAC7B0A5BBC1D2F66589598C82768E24" ID-CLB-Dataset="22144" ID-DOI="10.5852/ejt.2017.309" ID-GBIF-Dataset="4e7bcd4d-1cdb-45cf-b2c3-54da2466f171" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827189" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589468946001" checkinUser="carolina" docAuthor="Herbert, David G." docDate="2017" docId="039F87A5676DFFB34D29FD94FE6AFCD7" docLanguage="en" docName="ejt-2017-309.pdf.imf" docOrigin="European Journal of Taxonomy 309" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sheldonia fingolandensis Herbert 2017, sp. nov." docType="treatment" docVersion="6" lastPageNumber="48" masterDocId="FFA6FFDD6747FF9C4F79FFC5FFA2FFF9" masterDocTitle="A new genus and eight new species of tail-wagger snails from eastern South Africa, with a key to genera within Sheldonia s. l. (Gastropoda: Urocyclidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="43" updateTime="1698839690494" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="C6EC5EC1E4BB391B9D6E3CFC9F5AEFA9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A103CF4D5C6FC5180CC61D7DCBC30655">
<mods:title id="F73EBC3F9D459D3DD31F6BDCC826CFED">A new genus and eight new species of tail-wagger snails from eastern South Africa, with a key to genera within Sheldonia s. l. (Gastropoda: Urocyclidae)</mods:title>
</mods:titleInfo>
<mods:name id="D6E429F243B252E7E89B62EDA8DD477C" type="personal">
<mods:role id="E34EB63CBDE471C24087D77825B31E44">
<mods:roleTerm id="F9416479DC12C81F99BC17B0AB424889">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A3BFC49D10FE900E6E94A1ADD4E15ED">Herbert, David G.</mods:namePart>
</mods:name>
<mods:typeOfResource id="12714707CBF9289F9427160526AB1148">text</mods:typeOfResource>
<mods:relatedItem id="DB903635D51CDC8D729DC1EB485CB042" type="host">
<mods:titleInfo id="B58B43680162F8B4BDF75E7296D9EF04">
<mods:title id="EDFA286960F53ACF82FF06D9B9DA50C7">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="95D8649E46D9F62FA9A9BA014FF36232">
<mods:date id="94B2E2E7A8AB39D5084BD16CCCE2C6AE">2017</mods:date>
<mods:detail id="1F79798D5ED57B3C30DDA5DD46780E26" type="pubDate">
<mods:number id="28A2EBE9FED228661C7A8373A94B0CF1">2017-04-10</mods:number>
</mods:detail>
<mods:detail id="C6408BD5E26083F67480A95DFBCD33C1" type="issue">
<mods:number id="9A36972AAFB6F66AFC03B39FEB5871E3">309</mods:number>
</mods:detail>
<mods:extent id="FFC0F3C360188FD087717769DC52802B" unit="page">
<mods:start id="20E5FAD05368CD7F039F0D001B800F4C">1</mods:start>
<mods:end id="1D919A747882D7AC489A7C489C18388C">50</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1D6E32F7F39552E5D7F7435CE348BC03">journal article</mods:classification>
<mods:identifier id="7CC03B0DF4AF91CC3179B8D05F200BD2" type="CLB-Dataset">22144</mods:identifier>
<mods:identifier id="43C7E4E7A23D6BA41C4A1D7D6C2FE014" type="DOI">10.5852/ejt.2017.309</mods:identifier>
<mods:identifier id="BF81B42814383C771E210C00BA34136C" type="GBIF-Dataset">4e7bcd4d-1cdb-45cf-b2c3-54da2466f171</mods:identifier>
<mods:identifier id="61C432E75E329BF85D07D64E95B1D463" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="0609BA568F8B286653AA4BF2D953CC18" type="Zenodo-Dep">3827189</mods:identifier>
<mods:identifier id="576B91B962745B03135855ECEF5FB6D7" type="ZooBank">1E8FE779-D6E7-428E-9538-5E5F8ECFB271</mods:identifier>
</mods:mods>
<treatment id="039F87A5676DFFB34D29FD94FE6AFCD7" ID-DOI="http://doi.org/10.5281/zenodo.3846864" ID-GBIF-Taxon="164151033" ID-Zenodo-Dep="3846864" LSID="urn:lsid:plazi:treatment:039F87A5676DFFB34D29FD94FE6AFCD7" httpUri="http://treatment.plazi.org/id/039F87A5676DFFB34D29FD94FE6AFCD7" lastPageId="47" lastPageNumber="48" pageId="42" pageNumber="43">
<subSubSection id="C32C6538676DFFB64D29FD94FC46FD95" box="[592,996,593,620]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8B8936B3676DFFB64D29FD94FC46FD95" blockId="42.[367,1221,593,691]" box="[592,996,593,620]" pageId="42" pageNumber="43">
<heading id="D0C181DF676DFFB64D29FD94FC46FD95" box="[592,996,593,620]" centered="true" fontSize="11" level="2" pageId="42" pageNumber="43" reason="2">
<taxonomicName id="4C364D30676DFFB64D29FD94FC20FD95" ID-CoL="6YM8L" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[592,898,593,620]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA1676DFFB64D29FD94FC20FD95" bold="true" box="[592,898,593,620]" italics="true" pageId="42" pageNumber="43">Sheldonia fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA676DFFB64CF0FD97FC46FD95" box="[905,996,594,620]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32C6538676DFFB64E16FDB0FCD3FD4A" pageId="42" pageNumber="43" type="description">
<paragraph id="8B8936B3676DFFB64E16FDB0FB67FD69" blockId="42.[367,1221,593,691]" box="[367,1221,629,656]" pageId="42" pageNumber="43">
<uri id="FFA73AB1676DFFB64E16FDB0FB67FD69" box="[367,1221,629,656]" pageId="42" pageNumber="43">
urn:lsid:zoobank.org:act: 
<uuid id="FF900C66676DFFB64DF2FDB0FB67FD69" box="[651,1221,629,656]" pageId="42" pageNumber="43">8620960E-0BBC-49D2-9CF6-69D028EDB052</uuid>
</uri>
</paragraph>
<paragraph id="8B8936B3676DFFB64DBBFD5DFCD3FD4A" blockId="42.[367,1221,593,691]" box="[706,881,664,691]" pageId="42" pageNumber="43">
<figureCitation id="130D2A36676DFFB64DBBFD5DFCB9FD4A" box="[706,795,664,691]" captionStart="Fig" captionStartId="21.[189,232,1927,1953]" captionTargetBox="[265,1323,1094,1882]" captionTargetId="figure@21.[265,1323,1094,1882]" captionTargetPageId="21" captionText="Fig. 15. Distribution of Kerkophorus scrobicolus sp. nov. (asterisks), Kerkophorus terrestris sp. nov. (circumpuncta), Selatodryas luteosoma gen. et sp. nov. (pentagons), S. roseosoma gen. et sp. nov. (triangles) and Sheldonia fingolandensis sp. nov. (squares). Contours at 1000 and 1500 m." figureDoi="http://doi.org/10.5281/zenodo.3827223" httpUri="https://zenodo.org/record/3827223/files/figure.png" pageId="42" pageNumber="43">Figs 15</figureCitation>
, 
<figureCitation id="130D2A36676DFFB64C51FD5CFCD3FD4A" box="[808,881,665,691]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="43.[188,231,1928,1954]" captionStartId-1="44.[189,232,1928,1954]" captionStartId-2="45.[189,232,1929,1955]" captionStartId-3="46.[189,232,1895,1921]" captionTargetBox-0="[197,1397,511,1882]" captionTargetBox-1="[192,1396,1467,1883]" captionTargetBox-2="[193,1395,1113,1883]" captionTargetBox-3="[211,1378,763,1835]" captionTargetId-0="figure@43.[190,1400,503,1886]" captionTargetId-1="figure@44.[189,1399,1463,1886]" captionTargetId-2="figure@45.[189,1399,1107,1887]" captionTargetId-3="figure@46.[189,1399,750,1854]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionTargetPageId-2="45" captionTargetPageId-3="46" captionText-0="Fig. 31. Sheldonia fingolandensis sp. nov. A–D. Holotype, diameter 18.7 mm (NMSA W9688/T3383). E. Paratype, protoconch, punctate microsculpture with both spiral and radial alignment, scale bar = 200 μm (NMSA W3976/T3387)." captionText-1="Fig. 32. Sheldonia fingolandensis sp. nov., living specimens, Collywobbles, Mbhashe River valley, E Cape. A. Living specimens resting in situ on underside of Aloe ferox leaf. B. Crawling animal, shell diameter approx. 18 mm." captionText-2="Fig. 33. Sheldonia fingolandensis sp. nov., paratype, radula (NMSA W4075/T3384). A. Rachidian and left lateral teeth. B. Rachidian and innermost lateral teeth. C. Inner left marginal teeth. D. Outer left marginal teeth. Scale bars: A = 100 μm; B–D = 50 μm." captionText-3="Fig. 34. Sheldonia fingolandensis sp. nov., genitalia and spermatophore. A. Entire genital tract. B. Genital atrium dissected to show irregularly folded stimulator. C–D. Apical and basal portions of penis lumen showing longitudinal and transverse folds respectively. E. Spermatophore. A–D. Paratype (NMSA W4169/T3385). E. Paratype (NMSA W6386/T3389)." figureDoi-0="http://doi.org/10.5281/zenodo.3827259" figureDoi-1="http://doi.org/10.5281/zenodo.3827261" figureDoi-2="http://doi.org/10.5281/zenodo.3827263" figureDoi-3="http://doi.org/10.5281/zenodo.3827265" httpUri-0="https://zenodo.org/record/3827259/files/figure.png" httpUri-1="https://zenodo.org/record/3827261/files/figure.png" httpUri-2="https://zenodo.org/record/3827263/files/figure.png" httpUri-3="https://zenodo.org/record/3827265/files/figure.png" pageId="42" pageNumber="43">31–34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32C6538676DFFB64FC4FD1FFDA9FCBE" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8B8936B3676DFFB64FC4FD1FFE94FD0D" blockId="42.[189,310,730,756]" box="[189,310,730,756]" pageId="42" pageNumber="43">
<heading id="D0C181DF676DFFB64FC4FD1FFE94FD0D" bold="true" box="[189,310,730,756]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B942EAA1676DFFB64FC4FD1FFE94FD0D" bold="true" box="[189,310,730,756]" pageId="42" pageNumber="43">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936B3676DFFB64FC4FCCCFDA9FCBE" blockId="42.[189,1399,777,839]" pageId="42" pageNumber="43">Characterised by the axially striped colour pattern, large tumescent shell, finely punctate protoconch sculpture and arboreal habit.</paragraph>
</subSubSection>
<subSubSection id="C32C6538676DFFB64FC4FCAAFAE3FC22" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8B8936B3676DFFB64FC4FCAAFEE6FC70" blockId="42.[189,324,879,905]" box="[189,324,879,905]" pageId="42" pageNumber="43">
<heading id="D0C181DF676DFFB64FC4FCAAFEE6FC70" bold="true" box="[189,324,879,905]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B942EAA1676DFFB64FC4FCAAFEE6FC70" bold="true" box="[189,324,879,905]" pageId="42" pageNumber="43">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936B3676DFFB64FC4FC5BFAE3FC22" blockId="42.[189,1398,926,988]" pageId="42" pageNumber="43">
From Fingoland, the territory between the Kei and Mbhashe rivers, E 
<collectingRegion id="49F2F851676DFFB64B6FFC5BFBF7FC41" box="[1046,1109,926,952]" country="South Africa" name="Western Cape" pageId="42" pageNumber="43">Cape</collectingRegion>
, inhabited by the Fingo people (amaFengu) after their flight south to escape the brutality of King Shaka in the early 1800s.
</paragraph>
</subSubSection>
<subSubSection id="C32C6538676DFFB74FC4FBC6FCC4FEB0" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8B8936B3676DFFB64FC4FBC6FE08FBE4" blockId="42.[189,426,1027,1053]" box="[189,426,1027,1053]" pageId="42" pageNumber="43">
<heading id="D0C181DF676DFFB64FC4FBC6FE08FBE4" bold="true" box="[189,426,1027,1053]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="B942EAA1676DFFB64FC4FBC6FE08FBE4" bold="true" box="[189,426,1027,1053]" pageId="42" pageNumber="43">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B5E3CEE676DFFB64F93FBF7FC69FB41" ID-GBIF-Occurrence="2618815307" collectingDate="2006-02-20" collectionCode="NMSA" collectorName="D. Herbert &amp; L. Davis" country="South Africa" elevation="585" latitude="-32.00396" location="Collywobbles" longLatPrecision="1" longitude="28.5953" municipality="Mbhashe River valley" pageId="42" pageNumber="43" specimenCode="NMSA W9688/T3383" specimenCount="1" stateProvince="Cape" typeStatus="holotype">
<paragraph id="8B8936B3676DFFB64F93FBF7FEFEFBB5" blockId="42.[189,1399,1074,1208]" box="[234,348,1074,1100]" pageId="42" pageNumber="43">
<emphasis id="B942EAA1676DFFB64F93FBF7FEFEFBB5" bold="true" box="[234,348,1074,1100]" pageId="42" pageNumber="43">
<typeStatus id="548D8811676DFFB64F93FBF7FEFEFBB5" box="[234,348,1074,1100]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8936B3676DFFB64FC4FB93FC69FB41" blockId="42.[189,1399,1074,1208]" pageId="42" pageNumber="43">
<collectingCountry id="F3217623676DFFB64FC4FB93FE3DFB89" box="[189,415,1110,1136]" name="South Africa" pageId="42" pageNumber="43">SOUTH AFRICA</collectingCountry>
: E 
<collectingRegion id="49F2F851676DFFB64EA0FB93FDBAFB89" box="[473,536,1110,1136]" country="South Africa" name="Western Cape" pageId="42" pageNumber="43">Cape</collectingRegion>
, 
<collectingMunicipality id="6BEDACC9676DFFB64D56FB93FCEBFB89" box="[559,841,1110,1136]" pageId="42" pageNumber="43">Mbhashe River valley</collectingMunicipality>
, 
<location id="8EE96068676DFFB64C26FB93FBA7FB89" LSID="urn:lsid:plazi:treatment:039F87A5676DFFB34D29FD94FE6AFCD7:8EE96068676DFFB64C26FB93FBA7FB89" box="[863,1029,1110,1136]" country="South Africa" latitude="-32.00396" longLatPrecision="1" longitude="28.5953" municipality="Mbhashe River valley" name="Collywobbles" pageId="42" pageNumber="43" stateProvince="Cape">Collywobbles</location>
, overlooking vulture colony, 
<geoCoordinate id="EE025074676DFFB64FC4FBBCFEECFB6D" box="[189,334,1145,1172]" degrees="32.00396" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-32.00396">32.00396° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64E20FBBCFE4EFB6D" box="[345,492,1145,1172]" degrees="28.59530" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="28.5953">28.59530° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64E8FFBBCFDE3FB6D" box="[502,577,1145,1172]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="42" pageNumber="43" unit="m" value="585.0">
<elevation id="001BD180676DFFB64E8FFBBCFDE3FB6D" box="[502,577,1145,1172]" metricMagnitude="2" metricUnit="m" metricValue="5.85" pageId="42" pageNumber="43" unit="m" value="585.0">585 m</elevation>
</quantity>
, sparse valley thicket invaded with 
<taxonomicName id="4C364D30676DFFB64CAEFBBFFB9BFB6A" box="[983,1081,1146,1171]" class="Magnoliopsida" family="Verbenaceae" genus="Lantana" higherTaxonomySource="CoL" kingdom="Plantae" order="Lamiales" pageId="42" pageNumber="43" phylum="Magnoliopsida" rank="genus">
<emphasis id="B942EAA1676DFFB64CAEFBBFFB9BFB6A" box="[983,1081,1146,1171]" italics="true" pageId="42" pageNumber="43">Lantana</emphasis>
</taxonomicName>
, living on aloes, 
<collectorName id="26C35365676DFFB64B82FBBFFAD5FB6A" box="[1275,1399,1145,1172]" pageId="42" pageNumber="43">D. Herbert</collectorName>
and 
<collectorName id="26C35365676DFFB64F96FB5BFEF1FB4E" box="[239,339,1181,1208]" pageId="42" pageNumber="43">L. Davis</collectorName>
leg., 
<date id="FF881073676DFFB64EECFB58FD8DFB4E" box="[405,559,1181,1207]" pageId="42" pageNumber="43" value="2006-02-20">
<collectingDate id="EFCCE99B676DFFB64EECFB58FD8DFB4E" box="[405,559,1181,1207]" pageId="42" pageNumber="43" value="2006-02-20">20 Feb. 2006</collectingDate>
</date>
(
<specimenCode id="DB909EC8676DFFB64D47FB58FCEAFB4E" box="[574,840,1181,1207]" pageId="42" pageNumber="43">NMSA W9688/T3383</specimenCode>
, dry shell).
</paragraph>
</materialsCitation>
<paragraph id="8B8936B3676DFFB64F93FB1AFD57FB00" blockId="42.[189,1399,1247,1879]" box="[234,757,1247,1273]" pageId="42" pageNumber="43">
<emphasis id="B942EAA1676DFFB64F93FB1AFEC5FB00" bold="true" box="[234,359,1247,1273]" pageId="42" pageNumber="43">
<typeStatus id="548D8811676DFFB64F93FB1AFEC5FB00" box="[234,359,1247,1273]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
</emphasis>
(listed north to south, all E 
<collectingRegion id="49F2F851676DFFB64DD6FB1AFD4FFB00" box="[687,749,1247,1273]" country="South Africa" name="Western Cape" pageId="42" pageNumber="43">Cape</collectingRegion>
)
</paragraph>
<paragraph id="8B8936B3676DFFB64FC4FAC6FDF2F8AE" blockId="42.[189,1399,1247,1879]" pageId="42" pageNumber="43">
<collectingCountry id="F3217623676DFFB64FC4FAC6FE34FAE4" box="[189,406,1283,1309]" name="South Africa" pageId="42" pageNumber="43">SOUTH AFRICA</collectingCountry>
: Mbhashe River valley, near N2 bridge, 
<geoCoordinate id="EE025074676DFFB64C00FAC7FBACFAE4" box="[889,1038,1282,1309]" degrees="31.92461" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-31.92461">31.92461° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64B65FAC7FB10FAE4" box="[1052,1202,1282,1309]" degrees="28.45113" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="28.45113">28.45113° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64BB9FAC6FAA9FAE4" box="[1216,1291,1283,1309]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="42" pageNumber="43" unit="m" value="470.0">470 m</quantity>
, riverine thicket, living on aloe leaves, A. Moussalli and D. Stuart-Fox leg., 
<date id="FF881073676DFFB64CC6FAE3FBFEFAB8" box="[959,1116,1318,1345]" pageId="42" pageNumber="43" value="2005-11-23">23 Nov. 2005</date>
(
<collectionCode id="ED27AE76676DFFB64B10FAE3FB1CFAB9" box="[1129,1214,1318,1344]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="42" pageNumber="43" type="Museum">NMSA</collectionCode>
W4169/T3385, six dry shells with six bodies and eight whole specimens in ethanol); Mbhashe River valley, near N2 bridge, 
<geoCoordinate id="EE025074676DFFB64E60FAA8FE0DFA7E" box="[281,431,1389,1415]" degrees="31.92461" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-31.92461">31.92461° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64EC6FAA8FDF4FA71" box="[447,598,1389,1416]" degrees="28.45113" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="28.45113">28.45113° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64D1CFAA8FD17FA71" box="[613,693,1389,1416]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="42" pageNumber="43" unit="m" value="470.0">470 m</quantity>
, riverine thicket, dead in leaf-litter and under fallen aloes, A. Moussalli and D. Stuart-Fox leg., 
<date id="FF881073676DFFB64D0EFA54FCBAFA55" box="[631,792,1425,1452]" pageId="42" pageNumber="43" value="2005-11-23">23 Nov. 2005</date>
(
<collectionCode id="ED27AE76676DFFB64C51FA54FCDFFA52" box="[808,893,1425,1451]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="42" pageNumber="43" type="Museum">NMSA</collectionCode>
W5373/T3386, two dry shells); same data as 
<typeStatus id="548D8811676DFFB64FA4FA71FEE6FA37" box="[221,324,1460,1486]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED27AE76676DFFB64E2CFA70FE66FA36" box="[341,452,1461,1487]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="42" pageNumber="43" type="Museum">NHMUK</collectionCode>
20160245, one dry shell; 
<collectionCode id="ED27AE76676DFFB64D81FA70FCE9FA36" box="[760,843,1461,1487]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="42" pageNumber="43" type="Museum">NMSA</collectionCode>
W4075/T3384, seven dry shells with bodies in ethanol; 
<collectionCode id="ED27AE76676DFFB64E5FFA1DFE20FA0A" box="[294,386,1496,1523]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="42" pageNumber="43" type="Museum">RMNH</collectionCode>
.5004187, one dry shell); Mbhashe River valley, Collywobbles, slope below carpark on neck, north-facing, valley thicket, under leaves of 
<taxonomicName id="4C364D30676DFFB64C3EFA39FC60F9EF" box="[839,962,1532,1558]" class="Magnoliopsida" family="Xanthorrhoeaceae" genus="Aloe" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ferox">
<emphasis id="B942EAA1676DFFB64C3EFA39FC60F9EF" box="[839,962,1532,1558]" italics="true" pageId="42" pageNumber="43">Aloe ferox</emphasis>
</taxonomicName>
, 
<geoCoordinate id="EE025074676DFFB64CADFA39FBC8F9EF" box="[980,1130,1532,1558]" degrees="23.00436" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-23.00436">23.00436° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64B03FA39FAB0F9EF" box="[1146,1298,1532,1559]" degrees="28.58169" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="28.58169">28.58169° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64A5BFA39FAD3F9EF" box="[1314,1393,1532,1559]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="42" pageNumber="43" unit="m" value="510.0">510 m</quantity>
, M. Bursey leg., 
<date id="FF881073676DFFB64EFAF9DAFD81F9C0" box="[387,547,1567,1593]" pageId="42" pageNumber="43" value="2006-02-20">20 Feb. 2006</date>
(
<collectionCode id="ED27AE76676DFFB64D4CF9DAFDD7F9C0" box="[565,629,1567,1593]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="42" pageNumber="43">ELM</collectionCode>
W2927/T164, five dry shells with bodies in ethanol); Mbhashe River valley, Collywobbles, beside thatched picnic site, 
<geoCoordinate id="EE025074676DFFB64C15F986FBB0F9A4" box="[876,1042,1603,1629]" degrees="32.004417" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-32.004417">32.004417° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64B5AF986FB1FF9A7" box="[1059,1213,1603,1630]" degrees="28.58172" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="28.58172">28.58172° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64BB7F986FABCF9A7" box="[1230,1310,1603,1630]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="42" pageNumber="43" unit="m" value="510.0">510 m</quantity>
, valley thicket, living under green leaves of aloes, D. Herbert and L. Davis leg., 
<date id="FF881073676DFFB64B6AF9A2FB0FF979" box="[1043,1197,1638,1665]" pageId="42" pageNumber="43" value="2006-02-20">20 Feb. 2006</date>
(
<collectionCode id="ED27AE76676DFFB64BC5F9A2FAB3F978" box="[1212,1297,1639,1665]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="42" pageNumber="43" type="Museum">NMSA</collectionCode>
W3976/ T3387, 20 dry shells with 14 bodies in ethanol); Kei River valley, NNE of Stutterheim, Wedgeley Farm, Shed Camp, 
<geoCoordinate id="EE025074676DFFB64E2BF96BFE46F931" box="[338,484,1710,1736]" degrees="32.28986" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-32.28986">32.28986° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64E89F96BFD26F931" box="[496,644,1710,1737]" degrees="27.54674" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="27.54674">27.54674° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64DE9F96BFD79F931" box="[656,731,1710,1737]" metricMagnitude="2" metricUnit="m" metricValue="8.35" pageId="42" pageNumber="43" unit="m" value="835.0">835 m</quantity>
, valley thicket, D. Herbert and L. Davis leg., 
<date id="FF881073676DFFB64B93F96BFAD5F931" box="[1258,1399,1710,1736]" pageId="42" pageNumber="43" value="2007-03-04">4 Mar. 2007</date>
(
<collectionCode id="ED27AE76676DFFB64FBCF917FEB8F915" box="[197,282,1746,1772]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="42" pageNumber="43" type="Museum">NMSA</collectionCode>
W5345/T3388, one dry shell); Sihota, west bank of Kei River, approx. 
<quantity id="4CCE9B56676DFFB64B18F914FB10F912" box="[1121,1202,1745,1772]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="42" pageNumber="43" unit="km" value="2.5">2.5 km</quantity>
upstream of Kei Bridge, below Moordenaars Kop, 
<geoCoordinate id="EE025074676DFFB64D28F930FD47F8F6" box="[593,741,1781,1808]" degrees="32.50445" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-32.50445">32.50445° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64D8AF930FC2BF8E9" box="[755,905,1781,1808]" degrees="27.94945" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="27.94945">27.94945° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64CEFF930FC41F8E9" box="[918,995,1781,1808]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="m" value="200.0">200 m</quantity>
, under 
<taxonomicName id="4C364D30676DFFB64B42F930FB16F8F6" box="[1083,1204,1781,1807]" class="Magnoliopsida" family="Xanthorrhoeaceae" genus="Aloe" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="species" species="ferox">
<emphasis id="B942EAA1676DFFB64B42F930FB16F8F6" box="[1083,1204,1781,1807]" italics="true" pageId="42" pageNumber="43">Aloe ferox</emphasis>
</taxonomicName>
leaves, M. Cole leg., 
<date id="FF881073676DFFB64F8FF8DCFED9F8CA" box="[246,379,1817,1843]" pageId="42" pageNumber="43" value="2008-01-09">9 Jan. 2008</date>
(
<collectionCode id="ED27AE76676DFFB64EF2F8DCFE6BF8CA" box="[395,457,1817,1843]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="42" pageNumber="43">ELM</collectionCode>
W3263/T165, one dry shell with body in ethanol; 
<collectionCode id="ED27AE76676DFFB64B6DF8DCFBC5F8CA" box="[1044,1127,1817,1843]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="42" pageNumber="43" type="Museum">NMSA</collectionCode>
W6386/T3389, six dry shells with two bodies in ethanol).
</paragraph>
<paragraph id="8B8936B3676DFFB64FC4F8BBFCA4F861" blockId="42.[189,774,1918,1945]" box="[189,774,1918,1945]" pageId="42" pageNumber="43">
<emphasis id="B942EAA1676DFFB64FC4F8BBFEDBF861" bold="true" box="[189,377,1918,1944]" pageId="42" pageNumber="43">Other material</emphasis>
(listed north to south, all E 
<collectingRegion id="49F2F851676DFFB64DB9F8BBFD5CF861" box="[704,766,1918,1944]" country="South Africa" name="Western Cape" pageId="42" pageNumber="43">Cape</collectingRegion>
)
</paragraph>
<paragraph id="8B8936B3676DFFB64FC4F868FAD4F812" blockId="42.[189,1398,1965,2027]" pageId="42" pageNumber="43">
<collectingCountry id="F3217623676DFFB64FC4F868FE38F83E" box="[189,410,1965,1991]" name="South Africa" pageId="42" pageNumber="43">SOUTH AFRICA</collectingCountry>
: Kei Pass, 
<geoCoordinate id="EE025074676DFFB64D50F868FD60F83E" box="[553,706,1965,1991]" degrees="32.49364" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="-32.49364">32.49364° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676DFFB64DADF868FCCDF831" box="[724,879,1965,1992]" degrees="27.99175" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="27.99175">27.99175° E</geoCoordinate>
, 
<quantity id="4CCE9B56676DFFB64CF9F868FC70F831" box="[896,978,1965,1992]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="42" pageNumber="43" unit="m" value="282.0">282 m</quantity>
, valley thicket, dead under rocks, A. Moussalli and D. Stuart-Fox leg., 
<date id="FF881073676DFFB64D01F814FCB8F812" box="[632,794,2000,2027]" pageId="42" pageNumber="43" value="2005-11-23">23 Nov. 2005</date>
(
<collectionCode id="ED27AE76676DFFB64C53F814FCDDF812" box="[810,895,2001,2027]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="42" pageNumber="43" type="Museum">NMSA</collectionCode>
W5486); Mncotsho, Buffalo River valley,
</paragraph>
<paragraph id="8B8936B3676CFFB74FC4FECEFCC4FEB0" blockId="43.[189,1398,267,329]" pageId="43" pageNumber="44">
<geoCoordinate id="EE025074676CFFB74FC4FECEFE94FEDC" box="[189,310,267,293]" degrees="32.917" direction="south" orientation="latitude" pageId="43" pageNumber="44" precision="55" value="-32.917">32.917° S</geoCoordinate>
, 
<geoCoordinate id="EE025074676CFFB74E3EFECEFE60FEDF" box="[327,450,267,294]" degrees="27.583" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="55" value="27.583">27.583° E</geoCoordinate>
, 
<quantity id="4CCE9B56676CFFB74EABFECEFD80FEDF" box="[466,546,267,294]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="43" pageNumber="44" unit="m" value="360.0">360 m</quantity>
, under scattered bushes (esp. 
<emphasis id="B942EAA1676CFFB74CE8FECEFB8CFEDC" box="[913,1070,267,293]" italics="true" pageId="43" pageNumber="44">Coddia rudis</emphasis>
) in grassland near beacon, M. Cole and C. Vernon leg., 
<date id="FF881073676CFFB74D77FEEBFD3CFEB0" box="[526,670,302,329]" pageId="43" pageNumber="44" value="2007-11-06">6 Nov. 2007</date>
(
<collectionCode id="ED27AE76676CFFB74DD4FEEAFCA0FEB0" box="[685,770,303,329]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="43" pageNumber="44" type="Museum">NMSA</collectionCode>
P0630).
</paragraph>
</subSubSection>
<subSubSection id="C32C6538676CFFB24FC4FEB5FECDFEB1" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44" type="description">
<paragraph id="8B8936B3676CFFB74FC4FEB5FEECFE73" blockId="43.[189,334,368,394]" box="[189,334,368,394]" pageId="43" pageNumber="44">
<heading id="D0C181DF676CFFB74FC4FEB5FEECFE73" bold="true" box="[189,334,368,394]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="B942EAA1676CFFB74FC4FEB5FEECFE73" bold="true" box="[189,334,368,394]" pageId="43" pageNumber="44">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936B3676CFFB04FC4FE5AFE8FFCA0" blockId="43.[189,1398,414,476]" lastBlockId="44.[189,1399,267,857]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
SHELL (
<figureCitation id="130D2A36676CFFB74E6FFE5BFECCFE40" box="[278,366,414,441]" captionStart="Fig" captionStartId="43.[188,231,1928,1954]" captionTargetBox="[197,1397,511,1882]" captionTargetId="figure@43.[190,1400,503,1886]" captionTargetPageId="43" captionText="Fig. 31. Sheldonia fingolandensis sp. nov. A–D. Holotype, diameter 18.7 mm (NMSA W9688/T3383). E. Paratype, protoconch, punctate microsculpture with both spiral and radial alignment, scale bar = 200 μm (NMSA W3976/T3387)." figureDoi="http://doi.org/10.5281/zenodo.3827259" httpUri="https://zenodo.org/record/3827259/files/figure.png" pageId="43" pageNumber="44">Fig. 31</figureCitation>
). Subglobose to globose, spire prominent; periphery at or close to mid-whorl, evenly rounded; H:D 0.72–0.83 (N=9); suture rather distinctly indented, inserting just above periphery; thin, translucent straw-brown when fresh, sometimes with a greenish tint and with conspicuous pattern of unevenly spaced, opaque, pale buff-white axial lines and stripes; apical surface satin-like, base more glossy. Protoconch diameter 
<quantity id="4CCE9B56676BFFB04D6BFE97FD02FE95" box="[530,672,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="44" pageNumber="45" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
(N=6); junction with teleoconch sometimes clear, sometimes weakly marked; sculpture very fine, comprising numerous close-set, microscopic spiral lirae, these initially composed of microscopic punctae (
<figureCitation id="130D2A36676BFFB04DB8FE5DFC8EFE4B" box="[705,812,408,434]" captionStart="Fig" captionStartId="43.[188,231,1928,1954]" captionTargetBox="[197,1397,511,1882]" captionTargetId="figure@43.[190,1400,503,1886]" captionTargetPageId="43" captionText="Fig. 31. Sheldonia fingolandensis sp. nov. A–D. Holotype, diameter 18.7 mm (NMSA W9688/T3383). E. Paratype, protoconch, punctate microsculpture with both spiral and radial alignment, scale bar = 200 μm (NMSA W3976/T3387)." figureDoi="http://doi.org/10.5281/zenodo.3827259" httpUri="https://zenodo.org/record/3827259/files/figure.png" pageId="44" pageNumber="45">Fig. 31E</figureCitation>
); punctae also with an axial alignment, producing a secondary collabral pattern; latter half whorl of protoconch with spiral lirae only (no longer punctate). Teleoconch of up to almost 3.0 whorls; early part of first whorl with fine spiral sculpture similar to that on protoconch, but finer; remainder of shell with only finest traces of close-set, microscopic lirae, producing satin-like shell surface; an axial component not evident in microsculpture, but uneven growth-lines frequent and appearing to coincide with axial variation in colour pattern; occasional sparse malleation also evident, but insignificant; base more glossy, microsculpture becoming obsolete below periphery. Umbilicus completely absent in adult specimens, edge of columella lip reflected and slightly thickened with whitish callus, running into axis of coiling rather than fusing with parietal region; juvenile and subadult specimens often minutely rimate, columella lip lacking callus thickening and umbilicus remaining as a narrow open channel inside reflected lip. Aperture subcircular, but indented by penultimate whorl in parietal region. Diameter up to 
<quantity id="4CCE9B56676BFFB04C2FFCDEFC60FCCF" box="[854,962,795,822]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="44" pageNumber="45" unit="mm" value="20.9">20.9 mm</quantity>
; 
<typeStatus id="548D8811676BFFB04CB6FCDEFB9BFCCC" box="[975,1081,795,821]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
, diameter 
<quantity id="4CCE9B56676BFFB04BCDFCDEFA83FCCF" box="[1204,1313,795,822]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="44" pageNumber="45" unit="mm" value="18.7">18.7 mm</quantity>
, height 15.0 mm.
</paragraph>
<caption id="DF49663B676CFFB74FC5F84DFD9CF810" ID-DOI="http://doi.org/10.5281/zenodo.3827259" ID-Zenodo-Dep="3827259" httpUri="https://zenodo.org/record/3827259/files/figure.png" pageId="43" pageNumber="44" startId="43.[188,231,1928,1954]" targetBox="[197,1397,511,1882]" targetPageId="43">
<paragraph id="8B8936B3676CFFB74FC5F84DFD9CF810" blockId="43.[188,1400,1928,2025]" pageId="43" pageNumber="44">
<emphasis id="B942EAA1676CFFB74FC5F84DFEBBF85B" bold="true" box="[188,281,1928,1954]" pageId="43" pageNumber="44">Fig. 31.</emphasis>
<taxonomicName id="4C364D30676CFFB74E58F84DFDE5F85B" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[289,583,1928,1954]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA1676CFFB74E58F84DFDE5F85B" box="[289,583,1928,1954]" italics="true" pageId="43" pageNumber="44">Sheldonia fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA676CFFB74D29F84DFD09F85B" box="[592,683,1928,1954]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B942EAA1676CFFB74DCBF84DFD49F85B" bold="true" box="[690,747,1928,1954]" pageId="43" pageNumber="44">A–D</emphasis>
. Holotype, diameter 18.7 mm (NMSA W9688/T3383). 
<emphasis id="B942EAA1676CFFB74FC5F86EFF72F83C" bold="true" box="[188,208,1963,1989]" pageId="43" pageNumber="44">E</emphasis>
. Paratype, protoconch, punctate microsculpture with both spiral and radial alignment, scale bar = 200 μm (NMSA W3976/T3387).
</paragraph>
</caption>
<paragraph id="8B8936B3676BFFB04FC4FC45FEA4FBFA" blockId="44.[189,1399,896,1027]" pageId="44" pageNumber="45">
LIVING ANIMAL (
<figureCitation id="130D2A36676BFFB04E0DFC45FE68FC63" box="[372,458,896,922]" captionStart="Fig" captionStartId="44.[189,232,1928,1954]" captionTargetBox="[192,1396,1467,1883]" captionTargetId="figure@44.[189,1399,1463,1886]" captionTargetPageId="44" captionText="Fig. 32. Sheldonia fingolandensis sp. nov., living specimens, Collywobbles, Mbhashe River valley, E Cape. A. Living specimens resting in situ on underside of Aloe ferox leaf. B. Crawling animal, shell diameter approx. 18 mm." figureDoi="http://doi.org/10.5281/zenodo.3827261" httpUri="https://zenodo.org/record/3827261/files/figure.png" pageId="44" pageNumber="45">Fig. 32</figureCitation>
). Head-foot mostly pale yellowish-buff, neck and mantle edge somewhat darker orange-buff, tentacles similarly pale, but caudal appendage slightly darker; coloration of spire viscera generally not visible through shell, but cream blotches sometimes evident overlying darker digestive gland.
</paragraph>
<paragraph id="8B8936B3676BFFB04FC4FBE9FE7CFAE1" blockId="44.[189,1398,1067,1304]" pageId="44" pageNumber="45">
RADULA (
<figureCitation id="130D2A36676BFFB04E55FBEEFE27FBBC" box="[300,389,1067,1094]" captionStart="Fig" captionStartId="45.[189,232,1929,1955]" captionTargetBox="[193,1395,1113,1883]" captionTargetId="figure@45.[189,1399,1107,1887]" captionTargetPageId="45" captionText="Fig. 33. Sheldonia fingolandensis sp. nov., paratype, radula (NMSA W4075/T3384). A. Rachidian and left lateral teeth. B. Rachidian and innermost lateral teeth. C. Inner left marginal teeth. D. Outer left marginal teeth. Scale bars: A = 100 μm; B–D = 50 μm." figureDoi="http://doi.org/10.5281/zenodo.3827263" httpUri="https://zenodo.org/record/3827263/files/figure.png" pageId="44" pageNumber="45">Fig. 33</figureCitation>
). Formula R+17+1+(60–70); rachidian tricuspid; laterals essentially bicuspid with a mesocone and strong basal ectocone, but also with a minute endocone on side of mesocone (scarcely more than a low bump); laterals followed by one intermediary tooth and then a long series of marginals; marginals curved, bearing a large terminal cusp with a smaller subterminal cusp on outer (concave) margin, additional irregular cusps may be present on shaft, but these normally obscured by overlapping outer neighbouring tooth; marginals progressively decreasing in size toward edge of radula, but otherwise morphologically similar.
</paragraph>
<paragraph id="8B8936B3676BFFB04FC4FA85FAD5FA84" blockId="44.[189,1399,1344,1405]" pageId="44" pageNumber="45">
DISTAL GENITALIA (
<figureCitation id="130D2A36676BFFB04EF7FA85FE42FAA3" box="[398,480,1344,1370]" captionStart="Fig" captionStartId="46.[189,232,1895,1921]" captionTargetBox="[211,1378,763,1835]" captionTargetId="figure@46.[189,1399,750,1854]" captionTargetPageId="46" captionText="Fig. 34. Sheldonia fingolandensis sp. nov., genitalia and spermatophore. A. Entire genital tract. B. Genital atrium dissected to show irregularly folded stimulator. C–D. Apical and basal portions of penis lumen showing longitudinal and transverse folds respectively. E. Spermatophore. A–D. Paratype (NMSA W4169/T3385). E. Paratype (NMSA W6386/T3389)." figureDoi="http://doi.org/10.5281/zenodo.3827265" httpUri="https://zenodo.org/record/3827265/files/figure.png" pageId="44" pageNumber="45">Fig. 34</figureCitation>
A–D). Penis elongate and cylindrical, but conspicuously looped upon itself and surrounded by a thin sheath, 2–3 times longer when removed from sheath; retractor muscle attached to
</paragraph>
<caption id="DF49663B676BFFB04FC4F84DFE47F810" ID-DOI="http://doi.org/10.5281/zenodo.3827261" ID-Zenodo-Dep="3827261" httpUri="https://zenodo.org/record/3827261/files/figure.png" pageId="44" pageNumber="45" startId="44.[189,232,1928,1954]" targetBox="[192,1396,1467,1883]" targetPageId="44">
<paragraph id="8B8936B3676BFFB04FC4F84DFE47F810" blockId="44.[189,1398,1928,2025]" pageId="44" pageNumber="45">
<emphasis id="B942EAA1676BFFB04FC4F84DFEBFF85B" bold="true" box="[189,285,1928,1954]" pageId="44" pageNumber="45">Fig. 32.</emphasis>
<taxonomicName id="4C364D30676BFFB04E50F84DFDF1F85B" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[297,595,1928,1954]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA1676BFFB04E50F84DFDF1F85B" box="[297,595,1928,1954]" italics="true" pageId="44" pageNumber="45">Sheldonia fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA676BFFB04D19F84DFD63F85B" box="[608,705,1928,1954]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
, living specimens, Collywobbles, Mbhashe River valley, E Cape. 
<emphasis id="B942EAA1676BFFB04E5AF86EFE9AF83C" bold="true" box="[291,312,1963,1989]" pageId="44" pageNumber="45">A</emphasis>
. Living specimens resting 
<emphasis id="B942EAA1676BFFB04D01F869FD63F83C" box="[632,705,1964,1989]" italics="true" pageId="44" pageNumber="45">in situ</emphasis>
on underside of 
<taxonomicName id="4C364D30676BFFB04CF3F86EFBA1F83C" box="[906,1027,1963,1989]" class="Magnoliopsida" family="Xanthorrhoeaceae" genus="Aloe" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="ferox">
<emphasis id="B942EAA1676BFFB04CF3F86EFBA1F83C" box="[906,1027,1963,1989]" italics="true" pageId="44" pageNumber="45">Aloe ferox</emphasis>
</taxonomicName>
leaf. 
<emphasis id="B942EAA1676BFFB04B3EF86EFBF9F83C" bold="true" box="[1095,1115,1963,1989]" pageId="44" pageNumber="45">B</emphasis>
. Crawling animal, shell diameter approx. 18 mm.
</paragraph>
</caption>
<paragraph id="8B8936B3676AFFB14FC4FECEFD37FCEB" blockId="45.[189,1399,267,786]" pageId="45" pageNumber="46">
penis apex; basal half of penis relatively thin-walled, its lumen lined with close-set, irregularly sinuous transverse folds (
<figureCitation id="130D2A36676AFFB14EF0FEEBFE5AFEB0" box="[393,504,302,329]" captionStart="Fig" captionStartId="46.[189,232,1895,1921]" captionTargetBox="[211,1378,763,1835]" captionTargetId="figure@46.[189,1399,750,1854]" captionTargetPageId="46" captionText="Fig. 34. Sheldonia fingolandensis sp. nov., genitalia and spermatophore. A. Entire genital tract. B. Genital atrium dissected to show irregularly folded stimulator. C–D. Apical and basal portions of penis lumen showing longitudinal and transverse folds respectively. E. Spermatophore. A–D. Paratype (NMSA W4169/T3385). E. Paratype (NMSA W6386/T3389)." figureDoi="http://doi.org/10.5281/zenodo.3827265" httpUri="https://zenodo.org/record/3827265/files/figure.png" pageId="45" pageNumber="46">Fig. 34D</figureCitation>
); apical half of penis with thicker wall, lumen lined by longitudinal ridges, these largely smooth near junction with epiphallus, but progressively stronger and more zig-zag toward mid-region of penis (
<figureCitation id="130D2A36676AFFB14EC1FEB0FD87FE76" box="[440,549,373,399]" captionStart="Fig" captionStartId="46.[189,232,1895,1921]" captionTargetBox="[211,1378,763,1835]" captionTargetId="figure@46.[189,1399,750,1854]" captionTargetPageId="46" captionText="Fig. 34. Sheldonia fingolandensis sp. nov., genitalia and spermatophore. A. Entire genital tract. B. Genital atrium dissected to show irregularly folded stimulator. C–D. Apical and basal portions of penis lumen showing longitudinal and transverse folds respectively. E. Spermatophore. A–D. Paratype (NMSA W4169/T3385). E. Paratype (NMSA W6386/T3389)." figureDoi="http://doi.org/10.5281/zenodo.3827265" httpUri="https://zenodo.org/record/3827265/files/figure.png" pageId="45" pageNumber="46">Fig. 34C</figureCitation>
). Epiphallus short, with a stout, recurved caecum; caecum arising close to middle of epiphallus; lumen of caecum coarsely and irregularly papillate. Flagellum well developed, basal portion (f1) broader and with distinct transverse internal structure; distal portion (f2) longer and more slender, twisted into a spiral of several revolutions with little internal structure evident; junction of epiphallus and flagellum at insertion of vas deferens, with white contents; vas deferens simple and slender, in life running beside vagina and free oviduct to join spermoviduct. Genital atrium globose, lumen almost filled with a well-developed stimulator appearing as an irregularly folded and ridged mass, flaccid rather than muscular, its contracted shape variable between individuals, but occupying most of lumen wall (
<figureCitation id="130D2A36676AFFB14E0CFD4BFE40FD50" box="[373,482,654,681]" captionStart="Fig" captionStartId="46.[189,232,1895,1921]" captionTargetBox="[211,1378,763,1835]" captionTargetId="figure@46.[189,1399,750,1854]" captionTargetPageId="46" captionText="Fig. 34. Sheldonia fingolandensis sp. nov., genitalia and spermatophore. A. Entire genital tract. B. Genital atrium dissected to show irregularly folded stimulator. C–D. Apical and basal portions of penis lumen showing longitudinal and transverse folds respectively. E. Spermatophore. A–D. Paratype (NMSA W4169/T3385). E. Paratype (NMSA W6386/T3389)." figureDoi="http://doi.org/10.5281/zenodo.3827265" httpUri="https://zenodo.org/record/3827265/files/figure.png" pageId="45" pageNumber="46">Fig. 34B</figureCitation>
). Vagina short; gametolytic sac ovate, thin-walled, its duct short and thickerwalled; free oviduct greatly hypertrophied and convoluted, pale apricot in fresh material, occupying much of body cavity posterior to buccal mass, both dorsal and ventral to gut; spermoviduct divided into distinct prostatic and oviductal portions.
</paragraph>
<paragraph id="8B8936B3676AFFB24FC4FCFFFECDFEB1" blockId="45.[189,1399,825,1028]" lastBlockId="46.[189,1398,267,328]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
SPERMATOPHORE (
<figureCitation id="130D2A36676AFFB14EFAFCFCFE4DFCAD" box="[387,495,825,852]" captionStart="Fig" captionStartId="46.[189,232,1895,1921]" captionTargetBox="[211,1378,763,1835]" captionTargetId="figure@46.[189,1399,750,1854]" captionTargetPageId="46" captionText="Fig. 34. Sheldonia fingolandensis sp. nov., genitalia and spermatophore. A. Entire genital tract. B. Genital atrium dissected to show irregularly folded stimulator. C–D. Apical and basal portions of penis lumen showing longitudinal and transverse folds respectively. E. Spermatophore. A–D. Paratype (NMSA W4169/T3385). E. Paratype (NMSA W6386/T3389)." figureDoi="http://doi.org/10.5281/zenodo.3827265" httpUri="https://zenodo.org/record/3827265/files/figure.png" pageId="45" pageNumber="46">Fig. 34E</figureCitation>
). Elbowed, with an elongate capsule and a long, spirally coiled tail of ca 6 revolutions (coiled length of tail approx. 
<quantity id="4CCE9B56676AFFB14DCBFC98FCB7FC8E" box="[690,789,861,887]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="45" pageNumber="46" unit="mm" value="4.1">4.1 mm</quantity>
); proximal portion of tail (first whorl) with paired branching spines; spines asymmetric, larger on one side; spines in middle part of this whorl largest, progressively smaller distally; spines arching outward and with branches on one side only; branch tips bluntly bifid, ending in short recurved hooks; mid-region of tail (± 1.5 whorls) lacking spines; subsequent whorls (± 2.5) of tail somewhat flattened and with short stout projections on both sides, these curve over flattened area delimiting a U-shaped channel that spirals along tail; tip of tail (± one whorl) without spines.
</paragraph>
</subSubSection>
<caption id="DF49663B676AFFB14FC4F84CFC9DF813" ID-DOI="http://doi.org/10.5281/zenodo.3827263" ID-Zenodo-Dep="3827263" httpUri="https://zenodo.org/record/3827263/files/figure.png" pageId="45" pageNumber="46" startId="45.[189,232,1929,1955]" targetBox="[193,1395,1113,1883]" targetPageId="45">
<paragraph id="8B8936B3676AFFB14FC4F84CFC9DF813" blockId="45.[189,1398,1929,2026]" pageId="45" pageNumber="46">
<emphasis id="B942EAA1676AFFB14FC4F84CFEB8F85A" bold="true" box="[189,282,1929,1955]" pageId="45" pageNumber="46">Fig. 33.</emphasis>
<taxonomicName id="4C364D30676AFFB14E5BF84CFDEAF85A" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[290,584,1929,1955]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA1676AFFB14E5BF84CFDEAF85A" box="[290,584,1929,1955]" italics="true" pageId="45" pageNumber="46">Sheldonia fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA676AFFB14D29F84FFD0CF85D" box="[592,686,1930,1956]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, radula (NMSA W4075/T3384). 
<emphasis id="B942EAA1676AFFB14BD0F84CFB1CF85A" bold="true" box="[1193,1214,1929,1955]" pageId="45" pageNumber="46">A</emphasis>
. Rachidian and left lateral teeth. 
<emphasis id="B942EAA1676AFFB14EF2F869FE3DF83F" bold="true" box="[395,415,1964,1990]" pageId="45" pageNumber="46">B</emphasis>
. Rachidian and innermost lateral teeth. 
<emphasis id="B942EAA1676AFFB14CF8F869FC34F83F" bold="true" box="[897,918,1964,1990]" pageId="45" pageNumber="46">C</emphasis>
. Inner left marginal teeth. 
<emphasis id="B942EAA1676AFFB14BA5F869FB53F83F" bold="true" box="[1244,1265,1964,1990]" pageId="45" pageNumber="46">D</emphasis>
. Outer left marginal teeth. Scale bars: A = 100 μm; B–D = 50 μm.
</paragraph>
</caption>
<subSubSection id="C32C65386769FFB24FC4FEB5FDDBFE06" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8B8936B36769FFB24FC4FEB5FE66FE72" blockId="46.[189,452,368,395]" box="[189,452,368,395]" pageId="46" pageNumber="47">
<heading id="D0C181DF6769FFB24FC4FEB5FE66FE72" box="[189,452,368,395]" pageId="46" pageNumber="47">
<emphasis id="B942EAA16769FFB24FC4FEB5FEF4FE73" bold="true" box="[189,342,368,394]" pageId="46" pageNumber="47">Distribution</emphasis>
(
<figureCitation id="130D2A366769FFB24E1FFEB5FE19FE72" box="[358,443,368,395]" captionStart="Fig" captionStartId="21.[189,232,1927,1953]" captionTargetBox="[265,1323,1094,1882]" captionTargetId="figure@21.[265,1323,1094,1882]" captionTargetPageId="21" captionText="Fig. 15. Distribution of Kerkophorus scrobicolus sp. nov. (asterisks), Kerkophorus terrestris sp. nov. (circumpuncta), Selatodryas luteosoma gen. et sp. nov. (pentagons), S. roseosoma gen. et sp. nov. (triangles) and Sheldonia fingolandensis sp. nov. (squares). Contours at 1000 and 1500 m." figureDoi="http://doi.org/10.5281/zenodo.3827223" httpUri="https://zenodo.org/record/3827223/files/figure.png" pageId="46" pageNumber="47">Fig. 15</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8B8936B36769FFB24FC4FE5AFDDBFE06" blockId="46.[189,1398,414,512]" pageId="46" pageNumber="47">
A relatively narrow-range endemic, known only from the lower catchments of the Mbhashe and Kei Rivers (formerly known as Fingoland) and southward to the Buffalo River valley near East London; from 
<quantity id="4CCE9B566769FFB24F84FE20FEEAFE06" box="[253,328,485,511]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="46" pageNumber="47" unit="m" value="200.0">200 m</quantity>
to 
<quantity id="4CCE9B566769FFB24E14FE20FE15FE06" box="[365,439,485,512]" metricMagnitude="2" metricUnit="m" metricValue="8.35" pageId="46" pageNumber="47" unit="m" value="835.0">835 m</quantity>
above sea level.
</paragraph>
</subSubSection>
<subSubSection id="C32C65386769FFB24FC4FDE3FD86FD6B" pageId="46" pageNumber="47" type="biology_ecology">
<paragraph id="8B8936B36769FFB24FC4FDE3FEBFFDB9" blockId="46.[189,285,550,576]" box="[189,285,550,576]" pageId="46" pageNumber="47">
<heading id="D0C181DF6769FFB24FC4FDE3FEBFFDB9" bold="true" box="[189,285,550,576]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B942EAA16769FFB24FC4FDE3FEBFFDB9" bold="true" box="[189,285,550,576]" pageId="46" pageNumber="47">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936B36769FFB24FC4FD90FD86FD6B" blockId="46.[189,1398,597,658]" pageId="46" pageNumber="47">
Mostly known from Eastern Valley Bushveld (
<bibRefCitation id="EFA74B426769FFB24D90FD90FB9AFD89" author="Mucina L. &amp; Rutherford M. C." box="[745,1080,597,624]" pageId="46" pageNumber="47" refId="ref24274" refString="Mucina L. &amp; Rutherford M. C. (eds) 2006. The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19, South African National Biodiversity Institute, Pretoria." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
); living amongst the green leaf-bases of candelabra aloes.
</paragraph>
</subSubSection>
<caption id="DF49663B6769FFB24FC4F8A2FC31F815" ID-DOI="http://doi.org/10.5281/zenodo.3827265" ID-Zenodo-Dep="3827265" httpUri="https://zenodo.org/record/3827265/files/figure.png" pageId="46" pageNumber="47" startId="46.[189,232,1895,1921]" targetBox="[211,1378,763,1835]" targetPageId="46">
<paragraph id="8B8936B36769FFB24FC4F8A2FC31F815" blockId="46.[189,1399,1895,2028]" pageId="46" pageNumber="47">
<emphasis id="B942EAA16769FFB24FC4F8A2FE82F87B" bold="true" box="[189,288,1895,1922]" pageId="46" pageNumber="47">Fig. 34.</emphasis>
<taxonomicName id="4C364D306769FFB24E49F8ADFDFCF87B" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[304,606,1896,1922]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA16769FFB24E49F8ADFDFCF87B" box="[304,606,1896,1922]" italics="true" pageId="46" pageNumber="47">Sheldonia fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA6769FFB24D17F8ADFD71F87B" box="[622,723,1896,1922]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, genitalia and spermatophore. 
<emphasis id="B942EAA16769FFB24B20F8A2FBCCF878" bold="true" box="[1113,1134,1895,1921]" pageId="46" pageNumber="47">A</emphasis>
. Entire genital tract. 
<emphasis id="B942EAA16769FFB24FC4F84EFF73F85C" bold="true" box="[189,209,1931,1957]" pageId="46" pageNumber="47">B</emphasis>
. Genital atrium dissected to show irregularly folded stimulator. 
<emphasis id="B942EAA16769FFB24CA1F84EFBB3F85C" bold="true" box="[984,1041,1931,1957]" pageId="46" pageNumber="47">C–D</emphasis>
. Apical and basal portions of penis lumen showing longitudinal and transverse folds respectively. 
<emphasis id="B942EAA16769FFB24C9DF86BFC5AF831" bold="true" box="[996,1016,1966,1992]" pageId="46" pageNumber="47">E</emphasis>
. Spermatophore. A–D. Paratype (NMSA W4169/T3385). E. Paratype (NMSA W6386/T3389).
</paragraph>
</caption>
<subSubSection id="C32C65386768FFB34FC4FECEFB07FD60" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8B8936B36768FFB34FC4FECEFE8DFEDC" blockId="47.[189,303,267,293]" box="[189,303,267,293]" pageId="47" pageNumber="48">
<heading id="D0C181DF6768FFB34FC4FECEFE8DFEDC" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B942EAA16768FFB34FC4FECEFE8DFEDC" bold="true" box="[189,303,267,293]" pageId="47" pageNumber="48">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936B36768FFB34FC4FEFFFB07FD60" blockId="47.[189,1399,314,665]" pageId="47" pageNumber="48">
With its rather globose profile, the shell of 
<taxonomicName id="4C364D306768FFB34D88FEFFFBBDFEAD" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[753,1055,314,340]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA16768FFB34D88FEFFFBBDFEAD" box="[753,1055,314,340]" italics="true" pageId="47" pageNumber="48">Sheldonia fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA6768FFB34B57FEFEFB32FEAC" box="[1070,1168,315,341]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
resembles that of 
<taxonomicName id="4C364D306768FFB34FC4FE9AFD86FE81" authority="(Benson, 1848)" baseAuthorityName="Benson" baseAuthorityYear="1848" box="[189,548,350,377]" class="Magnoliopsida" family="Salicaceae" genus="Helix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="trotteriana">
<emphasis id="B942EAA16768FFB34FC4FE9AFEFEFE81" box="[189,348,351,377]" italics="true" pageId="47" pageNumber="48">S. trotteriana</emphasis>
(Benson, 1848)
</taxonomicName>
, but it is less elevated, has a more strongly patterned shell and more rapidly expanding whorls. Furthermore, whereas 
<taxonomicName id="4C364D306768FFB34C77FE46FC75FE65" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[782,983,386,413]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA16768FFB34C77FE46FC75FE65" box="[782,983,386,413]" italics="true" pageId="47" pageNumber="48">S. fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA6768FFB34C9BFE46FB9DFE64" box="[994,1087,387,413]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
is arboreal, 
<taxonomicName id="4C364D306768FFB34BA1FE46FAD4FE65" baseAuthorityName="Benson" baseAuthorityYear="1848" box="[1240,1398,387,413]" class="Magnoliopsida" family="Salicaceae" genus="Helix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="trotteriana">
<emphasis id="B942EAA16768FFB34BA1FE46FAD4FE65" box="[1240,1398,387,413]" italics="true" pageId="47" pageNumber="48">S. trotteriana</emphasis>
</taxonomicName>
is evidently exclusively ground-dwelling. The aloe-dwelling habit of the new species is shared with 
<taxonomicName id="4C364D306768FFB34FC4FE0EFD34FE1D" authority="(Melvill &amp; Ponsonby, 1890)" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1890" box="[189,662,458,485]" class="Gastropoda" family="Vitrinidae" genus="Vitrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="aloicola">
<emphasis id="B942EAA16768FFB34FC4FE0EFE9EFE1D" box="[189,316,458,485]" italics="true" pageId="47" pageNumber="48">S. aloicola</emphasis>
(Melvill &amp; Ponsonby, 1890)
</taxonomicName>
, but that species has a more depressed, more translucent shell of a more uniform colour. It seems likely, however, that the two are closely related, as the morphology of their distal genitalia is similar. 
<taxonomicName id="4C364D306768FFB34D3EFDD7FCAFFDD5" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[583,781,530,556]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA16768FFB34D3EFDD7FCAFFDD5" box="[583,781,530,556]" italics="true" pageId="47" pageNumber="48">S. fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA6768FFB34C6CFDD6FCD2FDD4" box="[789,880,531,557]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
may well have originated as a north-eastern isolate of the more southerly and more widespread 
<taxonomicName id="4C364D306768FFB34C76FDF3FC29FDA9" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1890" box="[783,907,566,592]" class="Gastropoda" family="Vitrinidae" genus="Vitrina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="aloicola">
<emphasis id="B942EAA16768FFB34C76FDF3FC29FDA9" box="[783,907,566,592]" italics="true" pageId="47" pageNumber="48">S. aloicola</emphasis>
</taxonomicName>
. The shell of 
<taxonomicName id="4C364D306768FFB34B50FDF3FAD4FDA8" authority="(Pfeiffer, 1846)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1846" box="[1065,1398,566,593]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="natalensis">
<emphasis id="B942EAA16768FFB34B50FDF3FB1EFDA9" box="[1065,1212,566,592]" italics="true" pageId="47" pageNumber="48">S. natalensis</emphasis>
(Pfeiffer, 1846)
</taxonomicName>
may also have a colour pattern of narrow axial stripes, but it is more globose, has a smaller protoconch (diameter 
<quantity id="4CCE9B566768FFB34E4CFDBBFE66FD60" box="[309,452,638,665]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="47" pageNumber="48" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
) and never attains such a large adult size (diameter &lt;
<quantity id="4CCE9B566768FFB34B4AFDBBFB3CFD60" box="[1075,1182,638,665]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="47" pageNumber="48" unit="mm" value="16.5">16.5 mm</quantity>
)
</paragraph>
</subSubSection>
<subSubSection id="C32C65386768FFB34FC4FD05FE6AFCD7" pageId="47" pageNumber="48" type="conservation">
<paragraph id="8B8936B36768FFB34FC4FD05FEC7FD23" blockId="47.[189,357,704,730]" box="[189,357,704,730]" pageId="47" pageNumber="48">
<heading id="D0C181DF6768FFB34FC4FD05FEC7FD23" bold="true" box="[189,357,704,730]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B942EAA16768FFB34FC4FD05FEC7FD23" bold="true" box="[189,357,704,730]" pageId="47" pageNumber="48">Conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936B36768FFB34FC4FD2AFE6AFCD7" blockId="47.[189,1398,751,814]" pageId="47" pageNumber="48">
The observations made above for 
<taxonomicName id="4C364D306768FFB34D1CFD35FCD6FCF0" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[613,884,752,778]" class="Gastropoda" family="Urocyclidae" genus="Kerkophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="terrestris" status="sp. nov.">
<emphasis id="B942EAA16768FFB34D1CFD35FCD6FCF0" box="[613,884,752,778]" italics="true" pageId="47" pageNumber="48">Kerkophorus terrestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA6768FFB34CF8FD35FC43FCF3" box="[897,993,752,778]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
are also applicable to 
<taxonomicName id="4C364D306768FFB34A78FD35FEC4FCD7" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="fingolandensis" status="sp. nov.">
<emphasis id="B942EAA16768FFB34A78FD35FEC4FCD7" italics="true" pageId="47" pageNumber="48">Sheldonia fingolandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DA6768FFB34E17FCD1FE6AFCD7" box="[366,456,788,814]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>