treatments-xml/data/03/9F/87/039F87A3FFBEFFA3FF27FF46DC54FA30.xml

<document id="65280AC59BC0061885A470696B87652D" ID-DOI="10.11646/zootaxa.3841.1.4" ID-GBIF-Dataset="8a6fc357-377e-4e49-ab5d-3291a965710c" ID-ISSN="1175-5326" ID-Zenodo-Dep="228924" ID-ZooBank="F5B71746-8A42-4BE6-BCBC-39449735EA42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460460922801" checkinUser="plazi" docAuthor="Manuel, Michael" docDate="2014" docId="039F87A3FFBEFFA3FF27FF46DC54FA30" docLanguage="en" docName="zt03841p106.pdf" docOrigin="Zootaxa 3841 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydroporus rifensis Manuel, 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="99" masterDocId="FFA6FFDBFFBDFFAAFFB0FFD1DD59FFF0" masterDocTitle="A new species of the genus Hydroporus Clairville, 1806 from the Central Rif mountains of northern Morocco (Coleoptera: Dytiscidae)" masterLastPageNumber="106" masterPageNumber="90" pageNumber="93" updateTime="1698303867716" updateUser="plazi">
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<mods:title id="8A621240F67D5A2680FBD194675C2C68">A new species of the genus Hydroporus Clairville, 1806 from the Central Rif mountains of northern Morocco (Coleoptera: Dytiscidae)</mods:title>
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<mods:date id="DFC97796AA3478907F1138874CA63044">2014</mods:date>
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<taxonomicName id="4C364D36FFBEFFA9FF27FF46DCD1FF41" ID-CoL="8LVDR" box="[151,392,151,177]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFBEFFA9FF27FF46DCD1FF41" bold="true" box="[151,392,151,177]" italics="true" pageId="3" pageNumber="93">Hydroporus rifensis</emphasis>
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<taxonomicNameLabel id="A27157DCFFBEFFA9FE20FF49DC8BFF42" box="[400,466,152,178]" pageId="3" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
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<figureCitation id="130D2A30FFBEFFA9FF27FF6BDC59FF23" box="[151,256,186,211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1842,1864]" captionTargetBox="[184,1398,783,1798]" captionTargetId="figure@3.[170,1417,767,1820]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Hydroporus rifensis sp. n. (left) (holotype, male) and Hydroporus gyllenhalii (right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987)." httpUri="https://zenodo.org/record/228925/files/figure.png" pageId="3" pageNumber="93">Figures 1</figureCitation>
, 
<figureCitation id="130D2A30FFBEFFA9FEBEFF6ADC0FFF23" box="[270,342,186,211]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,552,575]" captionTargetBox="[267,1317,196,525]" captionTargetId="figure@5.[264,1322,193,532]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Microsculpture of pronotal disc (2, 3), anterior juxta-sutural region of elytral disc (4, 5) and antero-lateral region of metacoxal plate (6, 7) of (2, 4, 6) Hydroporus rifensis sp. n. (2, 4: paratype, male; 6: paratype, female) and (3, 5, 7) H. gyllenhalii (3, 5: male specimen from Rigny-Ussé, Touraine, France; 7: female specimen from same locality). Figures 2 – 5 were done using a stereomicroscope with a nearly horizontal light source; pictures 6 – 7 are SEM views. All pictures shown at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228926/files/figure.png" pageId="3" pageNumber="93">2, 4, 6</figureCitation>
, 
<figureCitation id="130D2A30FFBEFFA9FED2FF6ADCC5FF23" box="[354,412,187,211]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[267,1320,745,1895]" captionTargetId="figure@5.[264,1322,741,1898]" captionTargetPageId="5" captionText="FIGURES 8 – 13. SEM views of selected structures of the ventral side of (8 – 10) Hydroporus rifensis sp. n. (8, 10: paratype, female; 9: paratype, male) and (11 – 13) H. gyllenhalii (female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm." httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="3" pageNumber="93">8–10</figureCitation>
, 
<figureCitation id="130D2A30FFBEFFA9FE18FF6ADCA9FF23" box="[424,496,186,211]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="3" pageNumber="93">14, 15</figureCitation>
, 
<figureCitation id="130D2A30FFBEFFA9FE4CFF6ADF1DFF23" box="[508,580,187,211]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[274,1284,1260,1890]" captionTargetId="figure@6.[264,1323,1235,1890]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Drawings of fifth protarsomere and anterior claw (top) and posterior claw (bottom) in male (18, 20) and female (19, 21) of (18 – 19) Hydroporus rifensis sp. n. and (20 – 21) H. gyllenhalii. All drawings at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228929/files/figure.png" pageId="3" pageNumber="93">18, 19</figureCitation>
, 
<figureCitation id="130D2A30FFBEFFA9FDE0FF6BDFC0FF23" box="[592,665,186,211]" captionStart="FIGURES 22 – 31" captionStartId="7.[151,264,1037,1060]" captionTargetBox="[180,1414,199,1006]" captionTargetId="figure@7.[170,1417,193,1017]" captionTargetPageId="7" captionText="FIGURES 22 – 31. Male and female genitalia of (22 – 26) Hydroporus rifensis sp. n. (paratypes) and (27 – 31) H. gyllenhalii (specimens from Plévenon, Bretagne, France). (22, 27) Median lobe of aedeagus in profile view; (23, 28) median lobe of aedeagus in ventral view; (24, 29) left paramere (external view); (25, 30) left gonocoxosternum (external view); (26, 31) gonocoxa (ventral view). Homologous structures shown at same scale for both species. Scale bars: 22 – 25, 27 – 30: 0.1 mm; 26, 31: 0.05 mm." httpUri="https://zenodo.org/record/228930/files/figure.png" pageId="3" pageNumber="93">22–26</figureCitation>
.
</paragraph>
</subSubSection>
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<typeStatus id="548D8817FFBEFFA9FF27FED3DD8BFEEB" box="[151,210,258,283]" pageId="3" pageNumber="93">Type</typeStatus>
locality
</emphasis>
. Northern 
<collectingCountry id="F3217625FFBEFFA9FE17FED2DF56FEEA" box="[423,527,259,282]" name="Morocco" pageId="3" pageNumber="93">Morocco</collectingCountry>
, Central Rif, Anasar, ca. 
<quantity id="4CCE9B50FFBEFFA9FC98FED2DE36FEEB" box="[808,879,258,283]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="93" unit="km" value="10.0">10 km</quantity>
WNW Bab Berred and 
<quantity id="4CCE9B50FFBEFFA9FBCBFED2D99BFEEB" box="[1147,1218,258,283]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="93" unit="km" value="30.0">30 km</quantity>
ESE Chefchaouen, GPS coordinates 
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<geoCoordinate id="EE025072FFBEFFA9FE48FEF7DFD1FECF" box="[504,648,294,319]" direction="west" orientation="longitude" pageId="3" pageNumber="93" precision="1" value="-5.0003333">5°00'01.2''W</geoCoordinate>
, altitude 
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.
</paragraph>
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material
</emphasis>
. 
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<emphasis id="B942EAA7FFBEFFA9FECEFE9BDCB2FE93" bold="true" box="[382,491,330,355]" pageId="3" pageNumber="93">Holotype</emphasis>
</typeStatus>
(♂): “ 
<date id="FF881075FFBEFFA9FD86FE9BDFE0FE93" box="[566,697,330,355]" pageId="3" pageNumber="93" value="2013-05-18">18.05.2013</date>
, 
<collectingCountry id="F3217625FFBEFFA9FD75FE9ADE74FE92" box="[709,813,331,354]" name="Morocco" pageId="3" pageNumber="93">Morocco</collectingCountry>
, Central Rif, Anasar ca. 
<quantity id="4CCE9B50FFBEFFA9FBF8FE9AD9C8FE93" box="[1096,1169,330,355]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="93" unit="km" value="10.0">10 km</quantity>
WNW Bab Berred, ca. 
<quantity id="4CCE9B50FFBEFFA9FF27FEBEDD87FE77" box="[151,222,366,391]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="93" unit="km" value="30.0">30 km</quantity>
ESE Chefchaouen, 
<geoCoordinate id="EE025072FFBEFFA9FE0EFEBFDF0FFE77" box="[446,598,366,391]" direction="north" orientation="latitude" pageId="3" pageNumber="93" precision="1" value="35.019276">35°01'09.4''N</geoCoordinate>
<geoCoordinate id="EE025072FFBEFFA9FDEDFEBFDFB4FE77" box="[605,749,366,391]" direction="west" orientation="longitude" pageId="3" pageNumber="93" precision="1" value="-5.0003333">5°00'01.2''W</geoCoordinate>
, alt. 
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, among decaying vegetal material under trees on the margin of a large pond with 
<taxonomicName id="4C364D36FFBEFFA9FE4EFE42DFDBFE5B" box="[510,642,403,427]" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="B942EAA7FFBEFFA9FE4EFE42DFDBFE5B" box="[510,642,403,427]" italics="true" pageId="3" pageNumber="93">Ranunculus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C364D36FFBEFFA9FD09FE42DE6BFE5B" box="[697,818,403,427]" class="Magnoliopsida" family="Plantaginaceae" genus="Callitriche" kingdom="Animalia" order="Lamiales" pageId="3" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="B942EAA7FFBEFFA9FD09FE42DE6BFE5B" box="[697,818,403,427]" italics="true" pageId="3" pageNumber="93">Callitriche</emphasis>
</taxonomicName>
plants, Manuel leg.” [printed], “ 
<typeStatus id="548D8817FFBEFFA9FB2EFE43D852FE5B" box="[1182,1291,402,427]" pageId="3" pageNumber="93" type="holotype">Holotype</typeStatus>
, 
<taxonomicName id="4C364D36FFBEFFA9FAA6FE43DDB3FE3F" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">Hydroporus rifensis</taxonomicName>
<taxonomicNameLabel id="A27157DCFFBEFFA9FF42FE66DC68FE3E" box="[242,305,439,462]" pageId="3" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
, Manuel det. 2014” [red, printed] (
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). 
<typeStatus id="548D8817FFBEFFA9FC84FE67DEF5FE3F" box="[820,940,438,463]" pageId="3" pageNumber="93" type="paratype">
<emphasis id="B942EAA7FFBEFFA9FC84FE67DEF5FE3F" bold="true" box="[820,940,438,463]" pageId="3" pageNumber="93">Paratypes</emphasis>
</typeStatus>
: 38 exs (18 ♂♂, 20 ♀♀), same data as the 
<typeStatus id="548D8817FFBEFFA9FF27FE0BDDA0FE03" box="[151,249,474,499]" pageId="3" pageNumber="93" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED27AE70FFBEFFA9FEBAFE0ADC3DFE03" LSID="urn:lsid:biocol.org:col:34988" box="[266,356,475,499]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="93">MNHN</collectionCode>
, 
<collectionCode id="ED27AE70FFBEFFA9FEC2FE0ADC90FE03" LSID="urn:lsid:biocol.org:col:35275" box="[370,457,475,499]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="93">MNCN</collectionCode>
, 
<collectionCode id="ED27AE70FFBEFFA9FE66FE0ADF44FE03" box="[470,541,475,499]" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi  (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="3" pageNumber="93">CMM</collectionCode>
, 
<collectionCode id="ED27AE70FFBEFFA9FD9BFE0ADF3FFE03" box="[555,614,475,499]" pageId="3" pageNumber="93">CHF</collectionCode>
), including three DNA-extracted vouchers kept in 
<collectionCode id="ED27AE70FFBEFFA9FB1EFE0AD9ABFE03" box="[1198,1266,475,499]" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi  (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="3" pageNumber="93">CMM</collectionCode>
and two goldcoated 
<typeStatus id="548D8817FFBEFFA9FF5CFE2EDC01FDE7" box="[236,344,511,535]" pageId="3" pageNumber="93" type="paratype">paratypes</typeStatus>
mounted upside down (one male and one female) available at the 
<collectionCode id="ED27AE70FFBEFFA9FBCCFE2ED988FDE6" LSID="urn:lsid:biocol.org:col:34988" box="[1148,1233,511,534]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="93">MNHN</collectionCode>
for further SEM investigations. All 
<typeStatus id="548D8817FFBEFFA9FEDBFDF2DC8FFDCB" box="[363,470,547,571]" pageId="3" pageNumber="93" type="paratype">paratypes</typeStatus>
with the respective red label.
</paragraph>
</subSubSection>
<subSubSection id="C32C653EFFBEFFA3FF77FD97D9BDFD3C" lastPageId="9" lastPageNumber="99" pageId="3" pageNumber="93" type="description">
<paragraph id="8B8936B5FFBEFFA9FF77FD97DC42FD3B" blockId="3.[151,1437,258,715]" pageId="3" pageNumber="93">
<emphasis id="B942EAA7FFBEFFA9FF77FD97DC08FDAF" bold="true" box="[199,337,582,607]" pageId="3" pageNumber="93">Description</emphasis>
. The following description relates exclusively to the male 
<typeStatus id="548D8817FFBEFFA9FC45FD97D90EFDAF" box="[1013,1111,582,607]" pageId="3" pageNumber="93" type="holotype">holotype</typeStatus>
until paragraph “Aedeagus” inclusively (note however that some of the pictures cited to illustrate 
<typeStatus id="548D8817FFBEFFA9FC71FDBBD97AFD73" box="[961,1059,618,643]" pageId="3" pageNumber="93" type="holotype">holotype</typeStatus>
characters have been elaborated using 
<typeStatus id="548D8817FFBEFFA9FF6FFD5EDC09FD57" box="[223,336,655,679]" pageId="3" pageNumber="93" type="paratype">paratypes</typeStatus>
). Variations of characters in the 
<typeStatus id="548D8817FFBEFFA9FD65FD5EDE5CFD57" box="[725,773,655,679]" pageId="3" pageNumber="93">type</typeStatus>
series are given in a separate section at the end of the description.
</paragraph>
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<paragraph id="8B8936B5FFBEFFA9FF27F8E3DED0F896" blockId="3.[151,1436,1842,1894]" pageId="3" pageNumber="93">
<emphasis id="B942EAA7FFBEFFA9FF27F8E3DC57F8B7" bold="true" box="[151,270,1842,1864]" pageId="3" pageNumber="93">FIGURE 1</emphasis>
. Habitus of 
<taxonomicName id="4C364D36FFBEFFA9FE3FF8E3DF05F8B8" box="[399,604,1842,1864]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFBEFFA9FE3FF8E3DF05F8B8" box="[399,604,1842,1864]" italics="true" pageId="3" pageNumber="93">Hydroporus rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFBEFFA9FDD3F8E3DFC4F8B7" box="[611,669,1842,1863]" pageId="3" pageNumber="93" rank="species">
<emphasis id="B942EAA7FFBEFFA9FDD3F8E3DFC4F8B7" bold="true" box="[611,669,1842,1863]" pageId="3" pageNumber="93">sp. n.</emphasis>
</taxonomicNameLabel>
(left) (holotype, male) and 
<taxonomicName id="4C364D36FFBEFFA9FC0DF8E3D9F3F8B8" box="[957,1194,1842,1864]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="species" species="gyllenhalii">
<emphasis id="B942EAA7FFBEFFA9FC0DF8E3D9F3F8B8" box="[957,1194,1842,1864]" italics="true" pageId="3" pageNumber="93">Hydroporus gyllenhalii</emphasis>
</taxonomicName>
(right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987).
</paragraph>
</caption>
<paragraph id="8B8936B5FFBEFFA9FF77F85FD8C4F81F" blockId="3.[151,1437,1934,2031]" pageId="3" pageNumber="93">
<emphasis id="B942EAA7FFBEFFA9FF77F85FDC7FF857" bold="true" box="[199,294,1934,1959]" pageId="3" pageNumber="93">Habitus</emphasis>
(
<figureCitation id="130D2A30FFBEFFA9FE88F85FDC27F857" box="[312,382,1934,1959]" captionStart="FIGURE 1" captionStartId="3.[151,250,1842,1864]" captionTargetBox="[184,1398,783,1798]" captionTargetId="figure@3.[170,1417,767,1820]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Hydroporus rifensis sp. n. (left) (holotype, male) and Hydroporus gyllenhalii (right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987)." httpUri="https://zenodo.org/record/228925/files/figure.png" pageId="3" pageNumber="93">Fig. 1</figureCitation>
, left) oblong-elongate, with lateral outline slightly discontinuous at pronotal base. Head of moderate size; ratio between interocular distance and maximum pronotal width about 0.45. Body dorsally regularly and rather strongly convex. Elytron sides regularly curved; rather strongly attenuated towards posterior extremity.
</paragraph>
<paragraph id="8B8936B5FFB9FFAEFF27FF46DF96FEB0" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="94">Maximum width of body slightly before middle of total length, at about first third of elytral length. Ratio between maximum pronotal width and maximum body width about 0.85; at posterior angles pronotal width equalling width of elytra at shoulders. Pronotum dark ferrugineous with broadly testaceous margins; elytra brown with lateral borders and apex paler; head paler than pronotum and elytron. Entire dorsal surface microreticulated and rather strongly shining, with well-developed pubescence.</paragraph>
<paragraph id="8B8936B5FFB9FFAEFF77FE9DDE06FD90" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="94">
<emphasis id="B942EAA7FFB9FFAEFF77FE9DDC5EFE95" bold="true" box="[199,263,332,357]" pageId="4" pageNumber="94">Head</emphasis>
rufo-testaceous with two wide brownish markings between eyes and posterior part of vertex brown. Clypeus anteriorly rather strongly expanded; its anterior outline regularly rounded. Antero-lateral head outline discontinuous between clypeus and eye. Clypeus with two wide subcircular and densely punctated depressions. Entire head surface microreticulated with well-impressed polygonal meshes, these smaller and strongly transverse on vertex behind eyes. Punctures on head moderately coarse and dense, heterogeneous in size (their diameter equalling 1 to 2 meshes of microreticulation); behind eyes punctures very small and sparse; beside inner eye margin with a discrete irregular row of small punctures and meshes of microreticulation smaller. Small setae present only in clypeal depressions and along inner eye margin.
</paragraph>
<paragraph id="8B8936B5FFB9FFAEFF77FDBDDE39FB84" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="94">
<emphasis id="B942EAA7FFB9FFAEFF77FDBDDC19FD75" bold="true" box="[199,320,620,645]" pageId="4" pageNumber="94">Pronotum</emphasis>
dark ferrugineous with lateral margins broadly testaceous; transition to lateral paler colour rather sharp (
<figureCitation id="130D2A30FFB9FFAEFF55FD5EDC70FD58" box="[229,297,655,680]" captionStart="FIGURE 1" captionStartId="3.[151,250,1842,1864]" captionTargetBox="[184,1398,783,1798]" captionTargetId="figure@3.[170,1417,767,1820]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Hydroporus rifensis sp. n. (left) (holotype, male) and Hydroporus gyllenhalii (right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987)." httpUri="https://zenodo.org/record/228925/files/figure.png" pageId="4" pageNumber="94">Fig. 1</figureCitation>
, left). Posterior angles sharp and very slightly obtuse (almost right). Sides of pronotum converging from posterior angles; in posterior third straight then curving inwards. Posterior margin of pronotum medially strongly expanded backwards. Lateral margin with well-defined testaceous bead of moderate thickness (visible on 
<figureCitation id="130D2A30FFB9FFAEFF27FD28DD80FCE2" box="[151,217,761,786]" captionStart="FIGURE 1" captionStartId="3.[151,250,1842,1864]" captionTargetBox="[184,1398,783,1798]" captionTargetId="figure@3.[170,1417,767,1820]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Hydroporus rifensis sp. n. (left) (holotype, male) and Hydroporus gyllenhalii (right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987)." httpUri="https://zenodo.org/record/228925/files/figure.png" pageId="4" pageNumber="94">Fig. 1</figureCitation>
, left), distinct from anterior to posterior angle; in anterior half bead becoming gradually narrower in dorsal view (not in lateral view). Whole surface microreticulated with polygonal meshes except before continuous anterior line of small punctures (here meshes transverse); meshes more weakly impressed than on head except medially along posterior margin and laterally. On anterior part of disc meshes of about same size as on head; posteriorly becoming progressively smaller. On disc with moderately dense punctation (
<figureCitation id="130D2A30FFB9FFAEFB30FC56D99CFC50" box="[1152,1221,903,928]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,552,575]" captionTargetBox="[267,1317,196,525]" captionTargetId="figure@5.[264,1322,193,532]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Microsculpture of pronotal disc (2, 3), anterior juxta-sutural region of elytral disc (4, 5) and antero-lateral region of metacoxal plate (6, 7) of (2, 4, 6) Hydroporus rifensis sp. n. (2, 4: paratype, male; 6: paratype, female) and (3, 5, 7) H. gyllenhalii (3, 5: male specimen from Rigny-Ussé, Touraine, France; 7: female specimen from same locality). Figures 2 – 5 were done using a stereomicroscope with a nearly horizontal light source; pictures 6 – 7 are SEM views. All pictures shown at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228926/files/figure.png" pageId="4" pageNumber="94">Fig. 2</figureCitation>
); punctures of two size categories: small punctures of diameter about 1 mesh of microreticulation and larger coarse punctures with diameter equalling 1.5–2.5 meshes of microreticulation, latter more numerous; average distance between punctures about 3–4 meshes. Laterally punctation slightly denser with all punctures resembling larger punctures of disc. Posterior third of pronotum with small punctures and with rather dense and very coarse large punctures of diameter reaching about 3 meshes of microreticulation. Laterally before posterior margin punctation denser in shallow transverse depression. Punctures all bearing a conspicuous seta.
</paragraph>
<paragraph id="8B8936B5FFB9FFAEFF77FBAEDD9DFA0B" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="94">
<emphasis id="B942EAA7FFB9FFAEFF77FBAEDC4DFB68" bold="true" box="[199,276,1151,1176]" pageId="4" pageNumber="94">Elytra</emphasis>
brown, distinctly paler than pronotum, with diffusely delimited paler areas on lateral margins and apex but without any defined drawing (
<figureCitation id="130D2A30FFB9FFAEFDA8FB72DF02FB4C" box="[536,603,1187,1212]" captionStart="FIGURE 1" captionStartId="3.[151,250,1842,1864]" captionTargetBox="[184,1398,783,1798]" captionTargetId="figure@3.[170,1417,767,1820]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Hydroporus rifensis sp. n. (left) (holotype, male) and Hydroporus gyllenhalii (right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987)." httpUri="https://zenodo.org/record/228925/files/figure.png" pageId="4" pageNumber="94">Fig. 1</figureCitation>
, left). In lateral view elytral margin distinctly ascending towards humeral angle; epipleuron not visible until humeral angle. Elytral bead much thinner than pronotal bead in lateral view. Dorsal surface of elytra in anterior half rather regularly convex in profile as well as in frontal view. Microreticulation well-defined but rather finely impressed (meshes contiguous), with meshes distinctly larger than on pronotum (
<figureCitation id="130D2A30FFB9FFAEFE8BFAE0DCDBFABA" box="[315,386,1329,1354]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,552,575]" captionTargetBox="[267,1317,196,525]" captionTargetId="figure@5.[264,1322,193,532]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Microsculpture of pronotal disc (2, 3), anterior juxta-sutural region of elytral disc (4, 5) and antero-lateral region of metacoxal plate (6, 7) of (2, 4, 6) Hydroporus rifensis sp. n. (2, 4: paratype, male; 6: paratype, female) and (3, 5, 7) H. gyllenhalii (3, 5: male specimen from Rigny-Ussé, Touraine, France; 7: female specimen from same locality). Figures 2 – 5 were done using a stereomicroscope with a nearly horizontal light source; pictures 6 – 7 are SEM views. All pictures shown at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228926/files/figure.png" pageId="4" pageNumber="94">Fig. 4</figureCitation>
); towards apex meshes more strongly impressed. Elytral punctation (
<figureCitation id="130D2A30FFB9FFAEFB2DFAE0D9BBFABA" box="[1181,1250,1329,1354]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,552,575]" captionTargetBox="[267,1317,196,525]" captionTargetId="figure@5.[264,1322,193,532]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Microsculpture of pronotal disc (2, 3), anterior juxta-sutural region of elytral disc (4, 5) and antero-lateral region of metacoxal plate (6, 7) of (2, 4, 6) Hydroporus rifensis sp. n. (2, 4: paratype, male; 6: paratype, female) and (3, 5, 7) H. gyllenhalii (3, 5: male specimen from Rigny-Ussé, Touraine, France; 7: female specimen from same locality). Figures 2 – 5 were done using a stereomicroscope with a nearly horizontal light source; pictures 6 – 7 are SEM views. All pictures shown at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228926/files/figure.png" pageId="4" pageNumber="94">Fig. 4</figureCitation>
) rather fine and moderately dense; punctures homogeneous in size, their average diameter about 25 µm (2 meshes), with rare interspersed micropunctures; on elytral disc mean distance between punctures about twice diameter of punctures. Inner puncture line weakly discernible in anterior quarter; puncture lines otherwise indistinct. Pubescence welldeveloped (
<figureCitation id="130D2A30FFB9FFAEFEABFA6EDC06FA27" box="[283,351,1471,1496]" captionStart="FIGURE 1" captionStartId="3.[151,250,1842,1864]" captionTargetBox="[184,1398,783,1798]" captionTargetId="figure@3.[170,1417,767,1820]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Hydroporus rifensis sp. n. (left) (holotype, male) and Hydroporus gyllenhalii (right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987)." httpUri="https://zenodo.org/record/228925/files/figure.png" pageId="4" pageNumber="94">Fig. 1</figureCitation>
, left), each puncture bearing a rather long yellow seta; average length of elytral setae about 70 µm.
</paragraph>
<paragraph id="8B8936B5FFB9FFAEFF77F9D7DEBDF802" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="94">
<emphasis id="B942EAA7FFB9FFAEFF77F9D7DC02F9EF" bold="true" box="[199,347,1542,1567]" pageId="4" pageNumber="94">Ventral side</emphasis>
. Head including genae and gula rufo-testaceous; contact line between gena and gula darker. Elytral epipleura testaceous. Ventral surface of prothorax rufo-testaceous except dark ferrugineous median region of prosternum. Rest of ventral surface black, except ferrugineous posterior part of metacoxal processes, light brown broad lateral markings and narrow posterior band on abdominal ventrites 3–4, and light brown abdominal ventrites 5–6, these darker medially along anterior margin. Genae with weakly impressed microreticulation; gula surface smooth with dense very small micropunctures, anterior half with large coarse punctures. Prosternal process (i.e. blade) ferrugineous, oblong-rhomboidal, gradually tapering backwards; in cross section rather wide, broadly and regularly convex (not roof like and not carinate) (
<figureCitation id="130D2A30FFB9FFAEFD73F8D3DE5EF8EA" box="[707,775,1794,1819]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[267,1320,745,1895]" captionTargetId="figure@5.[264,1322,741,1898]" captionTargetPageId="5" captionText="FIGURES 8 – 13. SEM views of selected structures of the ventral side of (8 – 10) Hydroporus rifensis sp. n. (8, 10: paratype, female; 9: paratype, male) and (11 – 13) H. gyllenhalii (female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm." httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="4" pageNumber="94">Fig. 8</figureCitation>
); its surface anteriorly microgranulose, posteriorly smooth and shining with a few small punctures; margin distinctly beaded; apex rounded (
<figureCitation id="130D2A30FFB9FFAEFBC0F8F7D9E3F8CF" box="[1136,1210,1830,1855]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[267,1320,745,1895]" captionTargetId="figure@5.[264,1322,741,1898]" captionTargetPageId="5" captionText="FIGURES 8 – 13. SEM views of selected structures of the ventral side of (8 – 10) Hydroporus rifensis sp. n. (8, 10: paratype, female; 9: paratype, male) and (11 – 13) H. gyllenhalii (female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm." httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="4" pageNumber="94">Fig. 8</figureCitation>
). Entire surface of prosternal process provided with long golden setae. Between procoxae with well-defined transverse prominence, in ventral view slightly curved (
<figureCitation id="130D2A30FFB9FFAEFE55F8BFDF14F877" box="[485,589,1902,1927]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[267,1320,745,1895]" captionTargetId="figure@5.[264,1322,741,1898]" captionTargetPageId="5" captionText="FIGURES 8 – 13. SEM views of selected structures of the ventral side of (8 – 10) Hydroporus rifensis sp. n. (8, 10: paratype, female; 9: paratype, male) and (11 – 13) H. gyllenhalii (female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm." httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="4" pageNumber="94">Figs 8, 9</figureCitation>
). File between prosternal base and blade narrow, its sides parallel; surface smooth and shining with several weakly impressed (almost indistinct) transverse grooves (
<figureCitation id="130D2A30FFB9FFAEFB2AF843D986F85A" box="[1178,1247,1938,1963]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[267,1320,745,1895]" captionTargetId="figure@5.[264,1322,741,1898]" captionTargetPageId="5" captionText="FIGURES 8 – 13. SEM views of selected structures of the ventral side of (8 – 10) Hydroporus rifensis sp. n. (8, 10: paratype, female; 9: paratype, male) and (11 – 13) H. gyllenhalii (female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm." httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="4" pageNumber="94">Fig. 9</figureCitation>
); file flanked by deeply excavated cavities in front of procoxa. Between anterior extremity of file and anterior prosternal margin surface microgranulose with small coarse punctures and short golden setae.
</paragraph>
<caption id="DF49663DFFB8FFAFFF27FDF9DCC5FD4A" httpUri="https://zenodo.org/record/228926/files/figure.png" pageId="5" pageNumber="95" targetBox="[267,1317,196,525]" targetPageId="5">
<paragraph id="8B8936B5FFB8FFAFFF27FDF9DCC5FD4A" blockId="5.[151,1437,552,698]" pageId="5" pageNumber="95">
<emphasis id="B942EAA7FFB8FFAFFF27FDF9DC64FDCE" bold="true" box="[151,317,552,575]" pageId="5" pageNumber="95">FIGURES 2–7.</emphasis>
Microsculpture of pronotal disc (2, 3), anterior juxta-sutural region of elytral disc (4, 5) and antero-lateral region of metacoxal plate (6, 7) of (2, 4, 6) 
<taxonomicName id="4C364D36FFB8FFAFFDF9FD96DE4AFDAD" box="[585,787,583,605]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFB8FFAFFDF9FD96DE4AFDAD" box="[585,787,583,605]" italics="true" pageId="5" pageNumber="95">Hydroporus rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFB8FFAFFCA9FD99DE0BFDAD" box="[793,850,584,605]" pageId="5" pageNumber="95" rank="species">
<emphasis id="B942EAA7FFB8FFAFFCA9FD99DE0BFDAD" bold="true" box="[793,850,584,605]" pageId="5" pageNumber="95">sp. n.</emphasis>
</taxonomicNameLabel>
(2, 4: paratype, male; 6: paratype, female) and (3, 5, 7) 
<taxonomicName id="4C364D36FFB8FFAFFA33FD98DC5AFD8C" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="gyllenhalii">
<emphasis id="B942EAA7FFB8FFAFFA33FD98DC5AFD8C" italics="true" pageId="5" pageNumber="95">H. gyllenhalii</emphasis>
</taxonomicName>
(3, 5: male specimen from Rigny-Ussé, Touraine, France; 7: female specimen from same locality). Figures 2–5 were done using a stereomicroscope with a nearly horizontal light source; pictures 6–7 are SEM views. All pictures shown at same scale (scale bar: 0.1 mm).
</paragraph>
</caption>
<caption id="DF49663DFFB8FFAFFF27F8AED9F2F802" httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="5" pageNumber="95" targetBox="[267,1320,745,1895]" targetPageId="5">
<paragraph id="8B8936B5FFB8FFAFFF27F8AED9F2F802" blockId="5.[151,1436,1919,2034]" pageId="5" pageNumber="95">
<emphasis id="B942EAA7FFB8FFAFFF27F8AEDC1BF864" bold="true" box="[151,322,1919,1942]" pageId="5" pageNumber="95">FIGURES 8–13</emphasis>
. SEM views of selected structures of the ventral side of (8–10) 
<taxonomicName id="4C364D36FFB8FFAFFC6CF8AED9F1F865" box="[988,1192,1919,1941]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFB8FFAFFC6CF8AED9F1F865" box="[988,1192,1919,1941]" italics="true" pageId="5" pageNumber="95">Hydroporus rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFB8FFAFFB00F851D9B2F865" box="[1200,1259,1920,1941]" pageId="5" pageNumber="95" rank="species">
<emphasis id="B942EAA7FFB8FFAFFB00F851D9B2F865" bold="true" box="[1200,1259,1920,1941]" pageId="5" pageNumber="95">sp. n.</emphasis>
</taxonomicNameLabel>
(8, 10: paratype, female; 9: paratype, male) and (11–13) 
<taxonomicName id="4C364D36FFB8FFAFFD96F84EDFE8F844" box="[550,689,1950,1972]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="95" phylum="Arthropoda" rank="species" species="gyllenhalii">
<emphasis id="B942EAA7FFB8FFAFFD96F84EDFE8F844" box="[550,689,1950,1972]" italics="true" pageId="5" pageNumber="95">H. gyllenhalii</emphasis>
</taxonomicName>
(female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm.
</paragraph>
</caption>
<caption id="DF49663DFFBBFFACFF27FB9CD9F8FB51" httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="6" pageNumber="96" targetBox="[229,1359,194,1079]" targetPageId="6">
<paragraph id="8B8936B5FFBBFFACFF27FB9CD9F8FB51" blockId="6.[151,1436,1101,1185]" pageId="6" pageNumber="96">
<emphasis id="B942EAA7FFBBFFACFF27FB9CDC16FB93" bold="true" box="[151,335,1101,1124]" pageId="6" pageNumber="96">FIGURES 14–17</emphasis>
. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14–15) 
<taxonomicName id="4C364D36FFBBFFACFA93FB9FDDBBFB73" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="96" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFBBFFACFA93FB9FDDBBFB73" italics="true" pageId="6" pageNumber="96">Hydroporus rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFBBFFACFF57FBBCDC46FB72" box="[231,287,1133,1154]" pageId="6" pageNumber="96" rank="species">
<emphasis id="B942EAA7FFBBFFACFF57FBBCDC46FB72" bold="true" box="[231,287,1133,1154]" pageId="6" pageNumber="96">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes) and (16–17) 
<taxonomicName id="4C364D36FFBBFFACFDADFBBFDFFEFB73" box="[541,679,1133,1155]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="96" phylum="Arthropoda" rank="species" species="gyllenhalii">
<emphasis id="B942EAA7FFBBFFACFDADFBBFDFFEFB73" box="[541,679,1133,1155]" italics="true" pageId="6" pageNumber="96">H. gyllenhalii</emphasis>
</taxonomicName>
(16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="DF49663DFFBBFFACFF27F8A6DD97F83B" httpUri="https://zenodo.org/record/228929/files/figure.png" pageId="6" pageNumber="96" targetBox="[274,1284,1260,1890]" targetPageId="6">
<paragraph id="8B8936B5FFBBFFACFF27F8A6DD97F83B" blockId="6.[151,1436,1911,1995]" pageId="6" pageNumber="96">
<emphasis id="B942EAA7FFBBFFACFF27F8A6DC16F87D" bold="true" box="[151,335,1911,1934]" pageId="6" pageNumber="96">FIGURES 18–21</emphasis>
. Drawings of fifth protarsomere and anterior claw (top) and posterior claw (bottom) in male (18, 20) and female (19, 21) of (18–19) 
<taxonomicName id="4C364D36FFBBFFACFE03F847DFD9F85C" box="[435,640,1942,1964]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="96" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFBBFFACFE03F847DFD9F85C" box="[435,640,1942,1964]" italics="true" pageId="6" pageNumber="96">Hydroporus rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFBBFFACFD37F846DF9BF85C" box="[647,706,1943,1964]" pageId="6" pageNumber="96" rank="species">
<emphasis id="B942EAA7FFBBFFACFD37F846DF9BF85C" bold="true" box="[647,706,1943,1964]" pageId="6" pageNumber="96">sp. n.</emphasis>
</taxonomicNameLabel>
and (20–21) 
<emphasis id="B942EAA7FFBBFFACFCFFF849DEBDF85C" box="[847,996,1942,1965]" italics="true" pageId="6" pageNumber="96">
<taxonomicName id="4C364D36FFBBFFACFCFFF849DEB9F85C" box="[847,992,1942,1965]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="96" phylum="Arthropoda" rank="species" species="gyllenhalii">H. gyllenhalii</taxonomicName>
.
</emphasis>
All drawings at same scale (scale bar: 0.1 mm).
</paragraph>
</caption>
<caption id="DF49663DFFBAFFADFF27FBDCDC46FB6F" httpUri="https://zenodo.org/record/228930/files/figure.png" pageId="7" pageNumber="97" targetBox="[180,1414,199,1006]" targetPageId="7">
<paragraph id="8B8936B5FFBAFFADFF27FBDCDC46FB6F" blockId="7.[151,1436,1037,1183]" pageId="7" pageNumber="97">
<emphasis id="B942EAA7FFBAFFADFF27FBDCDC16FBD3" bold="true" box="[151,335,1037,1060]" pageId="7" pageNumber="97">FIGURES 22–31</emphasis>
. Male and female genitalia of (22–26) 
<taxonomicName id="4C364D36FFBAFFADFD40FBDFDEE7FBD4" box="[752,958,1038,1060]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="97" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFBAFFADFD40FBDFDEE7FBD4" box="[752,958,1038,1060]" italics="true" pageId="7" pageNumber="97">Hydroporus rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFBAFFADFC77FBDED95BFBD4" box="[967,1026,1039,1060]" pageId="7" pageNumber="97" rank="species">
<emphasis id="B942EAA7FFBAFFADFC77FBDED95BFBD4" bold="true" box="[967,1026,1039,1060]" pageId="7" pageNumber="97">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes) and (27–31) 
<taxonomicName id="4C364D36FFBAFFADFABEFBDED8C5FBD4" box="[1294,1436,1038,1060]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="97" phylum="Arthropoda" rank="species" species="gyllenhalii">
<emphasis id="B942EAA7FFBAFFADFABEFBDED8C5FBD4" box="[1294,1436,1038,1060]" italics="true" pageId="7" pageNumber="97">H. gyllenhalii</emphasis>
</taxonomicName>
(specimens from Plévenon, Bretagne, France). (22, 27) Median lobe of aedeagus in profile view; (23, 28) median lobe of aedeagus in ventral view; (24, 29) left paramere (external view); (25, 30) left gonocoxosternum (external view); (26, 31) gonocoxa (ventral view). Homologous structures shown at same scale for both species. Scale bars: 22–25, 27–30: 0.1 mm; 26, 31: 0.05 mm.
</paragraph>
</caption>
<caption id="DF49663DFFBAFFADFF27F85FDFC2F854" box="[151,667,1934,1956]" pageId="7" pageNumber="97">
<paragraph id="8B8936B5FFBAFFADFF27F85FDFC2F854" blockId="7.[151,667,1934,1956]" box="[151,667,1934,1956]" pageId="7" pageNumber="97">
<emphasis id="B942EAA7FFBAFFADFF27F85FDC40F854" bold="true" box="[151,281,1934,1956]" pageId="7" pageNumber="97">FIGURE 32</emphasis>
. Habitat of 
<taxonomicName id="4C364D36FFBAFFADFE22F85FDF05F854" box="[402,604,1934,1956]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="97" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFBAFFADFE22F85FDF05F854" box="[402,604,1934,1956]" italics="true" pageId="7" pageNumber="97">Hydroporus rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFBAFFADFDD2F85EDFC2F854" box="[610,667,1935,1956]" pageId="7" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8936B5FFB5FFA2FF77FF46DE5AFD50" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="98">
Whole surface of metacoxal plates microreticulated with finely impressed meshes; punctures coarse, moderately large and dense (
<figureCitation id="130D2A30FFB5FFA2FE4FFF6ADF14FF24" box="[511,589,187,212]" captionStart="FIGURES 2 – 7" captionStartId="5.[151,264,552,575]" captionTargetBox="[267,1317,196,525]" captionTargetId="figure@5.[264,1322,193,532]" captionTargetPageId="5" captionText="FIGURES 2 – 7. Microsculpture of pronotal disc (2, 3), anterior juxta-sutural region of elytral disc (4, 5) and antero-lateral region of metacoxal plate (6, 7) of (2, 4, 6) Hydroporus rifensis sp. n. (2, 4: paratype, male; 6: paratype, female) and (3, 5, 7) H. gyllenhalii (3, 5: male specimen from Rigny-Ussé, Touraine, France; 7: female specimen from same locality). Figures 2 – 5 were done using a stereomicroscope with a nearly horizontal light source; pictures 6 – 7 are SEM views. All pictures shown at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228926/files/figure.png" pageId="8" pageNumber="98">Fig. 6</figureCitation>
). Elytral epipleura with obsolete microreticulation; median region of metaventrite and most of interlinear space of metacoxal processes smooth, latter with a few small punctures; rest of ventral metathorax surface with finely impressed microreticulation; on sides of metaventrite punctures coarser and denser than on metacoxal plates. Midline of metacoxal processes deeply impressed over its entire length. Lateral lines of metacoxal processes distinct, weakly divergent anteriad (
<figureCitation id="130D2A30FFB5FFA2FCC9FE98DE94FE91" box="[889,973,329,354]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[267,1320,745,1895]" captionTargetId="figure@5.[264,1322,741,1898]" captionTargetPageId="5" captionText="FIGURES 8 – 13. SEM views of selected structures of the ventral side of (8 – 10) Hydroporus rifensis sp. n. (8, 10: paratype, female; 9: paratype, male) and (11 – 13) H. gyllenhalii (female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm." httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="8" pageNumber="98">Fig. 10</figureCitation>
), ending shortly behind posterior margin of metaventrite; metacoxal lines internally bordered by narrow grooves. Posterior margin of metacoxal processes medially slightly prolonged backwards, very slightly concavely sinuate along sides (
<figureCitation id="130D2A30FFB5FFA2FBD7FE41D9E7FE58" box="[1127,1214,400,425]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[267,1320,745,1895]" captionTargetId="figure@5.[264,1322,741,1898]" captionTargetPageId="5" captionText="FIGURES 8 – 13. SEM views of selected structures of the ventral side of (8 – 10) Hydroporus rifensis sp. n. (8, 10: paratype, female; 9: paratype, male) and (11 – 13) H. gyllenhalii (female specimen from Rigny-Ussé, Touraine, France). (8, 11) Prosternal process; (9, 12) anterior extremity of prosternal process and file in front of prosternal process; (10, 13) metacoxal lines and metacoxal apophysis. Anterior on top in all pictures. Scale bars: 8, 10, 11, 13: 0.1 mm; 9, 12: 0.05 mm." httpUri="https://zenodo.org/record/228927/files/figure.png" pageId="8" pageNumber="98">Fig. 10</figureCitation>
). Whole surface of abdominal ventrites 1–5 finely microreticulated; on abdominal ventrite 6 microreticulation much more strongly impressed. On abdominal ventrites 3–5 and anterior part of abdominal ventrite 6, meshes with transverse orientation. Punctures coarse and dense on abdominal ventrites 1–2; punctures much smaller and sparser on abdominal ventrites 3–4; on abdominal ventrite 5 punctation denser; on abdominal ventrite 6 punctures distinctly larger and much denser (average distance between punctures about one diameter of a puncture). Distinct and rather long setae present everywhere except on metepisterna and sides of metaventrite; midline of abdominal ventrites 4 and 5 bearing a small dense brush of long golden setae.
</paragraph>
<paragraph id="8B8936B5FFB5FFA2FF77FD7DD836FBF3" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="98">
<emphasis id="B942EAA7FFB5FFA2FF77FD7DDC00FD35" bold="true" box="[199,345,684,709]" pageId="8" pageNumber="98">Appendages</emphasis>
. Antennomeres 1–4 testaceous; antennomeres 5–10 dark brown with basal third testaceous; antennomere 11 dark brown with basal quarter testaceous (
<figureCitation id="130D2A30FFB5FFA2FC82FD1EDE2CFD18" box="[818,885,719,744]" captionStart="FIGURE 1" captionStartId="3.[151,250,1842,1864]" captionTargetBox="[184,1398,783,1798]" captionTargetId="figure@3.[170,1417,767,1820]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Hydroporus rifensis sp. n. (left) (holotype, male) and Hydroporus gyllenhalii (right) (male specimen from Puente de Sanabria, Zamora province, Spain, Fery leg. 22 VII 1987)." httpUri="https://zenodo.org/record/228925/files/figure.png" pageId="8" pageNumber="98">Fig. 1</figureCitation>
, left). Antennomeres 5–10 elongate, their length equalling 2.5–3 times their width. Palpi testaceous; last article of maxillary palpi infuscate near apex. Legs uniformly rufo-testaceous. Ventral surface of pro- and mesotarsomeres 1–3 bearing numerous densely packed small trumpet-like (distally enlarged) setae (
<figureCitation id="130D2A30FFB5FFA2FD2EFCE8DFAFFCA2" box="[670,758,825,850]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Fig. 14</figureCitation>
); central ventral surface of first pro- and mesotarsomeres provided with 6 sucker cups arranged in two longitudinal rows in an area devoid of small trumpet-like setae; 2 additional sucker cups inserted on ventral surface of second pro- and mesotarsomeres towards their proximal extremity (
<figureCitation id="130D2A30FFB5FFA2FEA5FC75DC32FC4D" box="[277,363,932,957]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Fig. 14</figureCitation>
). Ventral surface of fifth protarsomere provided with 4–6 thick setae (
<figureCitation id="130D2A30FFB5FFA2FB2DFC75D9AEFC4D" box="[1181,1271,932,957]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Figs 14</figureCitation>
, 
<figureCitation id="130D2A30FFB5FFA2FAB7FC74D87FFC4D" box="[1287,1318,933,957]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[274,1284,1260,1890]" captionTargetId="figure@6.[264,1323,1235,1890]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Drawings of fifth protarsomere and anterior claw (top) and posterior claw (bottom) in male (18, 20) and female (19, 21) of (18 – 19) Hydroporus rifensis sp. n. and (20 – 21) H. gyllenhalii. All drawings at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228929/files/figure.png" pageId="8" pageNumber="98">18</figureCitation>
). Anterior protarsal claw in dorsal or ventral view distinctly thicker and shorter than posterior claw (this character visible on 
<figureCitation id="130D2A30FFB5FFA2FF27FC3ADDB0FBF3" box="[151,233,1003,1028]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Fig. 14</figureCitation>
); in lateral view broader in proximal half and much more strongly curved than posterior claw (
<figureCitation id="130D2A30FFB5FFA2FABFFC3AD83BFBF3" box="[1295,1378,1003,1028]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[274,1284,1260,1890]" captionTargetId="figure@6.[264,1323,1235,1890]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Drawings of fifth protarsomere and anterior claw (top) and posterior claw (bottom) in male (18, 20) and female (19, 21) of (18 – 19) Hydroporus rifensis sp. n. and (20 – 21) H. gyllenhalii. All drawings at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228929/files/figure.png" pageId="8" pageNumber="98">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8B8936B5FFB5FFA2FF77FBDFD9BCFBD7" blockId="8.[151,1437,151,2035]" box="[199,1253,1038,1063]" pageId="8" pageNumber="98">
<emphasis id="B942EAA7FFB5FFA2FF77FBDFDC63FBD7" bold="true" box="[199,314,1038,1063]" pageId="8" pageNumber="98">Aedeagus</emphasis>
. Median lobe in lateral and ventral views as in 
<figureCitation id="130D2A30FFB5FFA2FCFDFBDFDE97FBD7" box="[845,974,1038,1063]" captionStart="FIGURES 22 – 31" captionStartId="7.[151,264,1037,1060]" captionTargetBox="[180,1414,199,1006]" captionTargetId="figure@7.[170,1417,193,1017]" captionTargetPageId="7" captionText="FIGURES 22 – 31. Male and female genitalia of (22 – 26) Hydroporus rifensis sp. n. (paratypes) and (27 – 31) H. gyllenhalii (specimens from Plévenon, Bretagne, France). (22, 27) Median lobe of aedeagus in profile view; (23, 28) median lobe of aedeagus in ventral view; (24, 29) left paramere (external view); (25, 30) left gonocoxosternum (external view); (26, 31) gonocoxa (ventral view). Homologous structures shown at same scale for both species. Scale bars: 22 – 25, 27 – 30: 0.1 mm; 26, 31: 0.05 mm." httpUri="https://zenodo.org/record/228930/files/figure.png" pageId="8" pageNumber="98">Figs 22–23</figureCitation>
; parameres as in 
<figureCitation id="130D2A30FFB5FFA2FB3EFBDFD9B8FBD7" box="[1166,1249,1038,1063]" captionStart="FIGURES 22 – 31" captionStartId="7.[151,264,1037,1060]" captionTargetBox="[180,1414,199,1006]" captionTargetId="figure@7.[170,1417,193,1017]" captionTargetPageId="7" captionText="FIGURES 22 – 31. Male and female genitalia of (22 – 26) Hydroporus rifensis sp. n. (paratypes) and (27 – 31) H. gyllenhalii (specimens from Plévenon, Bretagne, France). (22, 27) Median lobe of aedeagus in profile view; (23, 28) median lobe of aedeagus in ventral view; (24, 29) left paramere (external view); (25, 30) left gonocoxosternum (external view); (26, 31) gonocoxa (ventral view). Homologous structures shown at same scale for both species. Scale bars: 22 – 25, 27 – 30: 0.1 mm; 26, 31: 0.05 mm." httpUri="https://zenodo.org/record/228930/files/figure.png" pageId="8" pageNumber="98">Fig. 24</figureCitation>
.
</paragraph>
<paragraph id="8B8936B5FFB5FFA2FF77FBE3DEF9FB62" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="98">
<emphasis id="B942EAA7FFB5FFA2FF77FBE3DC2FFBBB" bold="true" box="[199,374,1074,1099]" pageId="8" pageNumber="98">Measurements</emphasis>
. 
<typeStatus id="548D8817FFB5FFA2FE39FBE3DCAFFBBB" box="[393,502,1074,1099]" pageId="8" pageNumber="98" type="holotype">Holotype</typeStatus>
: total length (TL) 
<quantity id="4CCE9B50FFB5FFA2FD64FBE3DE68FBBA" box="[724,817,1074,1098]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="98" unit="mm" value="4.3">4.3 mm</quantity>
, total length without head 
<quantity id="4CCE9B50FFB5FFA2FBC2FBE3D996FBBA" box="[1138,1231,1074,1098]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="98" unit="mm" value="3.8">3.8 mm</quantity>
, maximum width (MW) 
<quantity id="4CCE9B50FFB5FFA2FF54FB87DC13FB9E" box="[228,330,1110,1135]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="8" pageNumber="98" unit="mm" value="2.05">2.05 mm</quantity>
, ratio TL/MW 2.08. 
<typeStatus id="548D8817FFB5FFA2FD83FB86DFFCFB9F" box="[563,677,1111,1135]" pageId="8" pageNumber="98" type="paratype">Paratypes</typeStatus>
: TL 4.0– 
<quantity id="4CCE9B50FFB5FFA2FCB9FB87DE35FB9E" box="[777,876,1110,1135]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" pageId="8" pageNumber="98" unit="mm" value="4.35">4.35 mm</quantity>
(4.15 ± 0.09), TL without head 
<quantity id="4CCE9B50FFB5FFA2FB64FB87D802FB9E" box="[1236,1371,1110,1135]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="8" pageNumber="98" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6–3.9 mm</quantity>
(3.75 ± 0.07), MW 1.9–2.1 (2.02 ± 0.05), TL/MW 2.01–2.15 (2.05 ± 0.03).
</paragraph>
<paragraph id="8B8936B5FFB5FFA2FF77FB4FDE55FAB7" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="98">
<emphasis id="B942EAA7FFB5FFA2FF77FB4FDC70FB47" bold="true" box="[199,297,1182,1207]" pageId="8" pageNumber="98">Females</emphasis>
. Second protarsomere longer than in male (compare 
<figureCitation id="130D2A30FFB5FFA2FC33FB4FDE8DFB47" box="[899,980,1182,1207]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Fig. 15</figureCitation>
and 
<figureCitation id="130D2A30FFB5FFA2FBBCFB4FD906FB47" box="[1036,1119,1182,1207]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Fig. 14</figureCitation>
). Fifth protarsomere shorter than in male (compare 
<figureCitation id="130D2A30FFB5FFA2FE29FB13DCB1FB2B" box="[409,488,1218,1243]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Fig. 15</figureCitation>
vs. 14, and 
<figureCitation id="130D2A30FFB5FFA2FDDFFB13DFE6FB2A" box="[623,703,1218,1243]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[274,1284,1260,1890]" captionTargetId="figure@6.[264,1323,1235,1890]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Drawings of fifth protarsomere and anterior claw (top) and posterior claw (bottom) in male (18, 20) and female (19, 21) of (18 – 19) Hydroporus rifensis sp. n. and (20 – 21) H. gyllenhalii. All drawings at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228929/files/figure.png" pageId="8" pageNumber="98">Fig. 19</figureCitation>
vs. 18); its ventral surface bearing only two (exceptionally three) thin setae inserted towards distal extremity (
<figureCitation id="130D2A30FFB5FFA2FD3AFB37DFB8FB0F" box="[650,737,1254,1279]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Figs 15</figureCitation>
, 
<figureCitation id="130D2A30FFB5FFA2FD5EFB36DE54FB0F" box="[750,781,1255,1279]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[274,1284,1260,1890]" captionTargetId="figure@6.[264,1323,1235,1890]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Drawings of fifth protarsomere and anterior claw (top) and posterior claw (bottom) in male (18, 20) and female (19, 21) of (18 – 19) Hydroporus rifensis sp. n. and (20 – 21) H. gyllenhalii. All drawings at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228929/files/figure.png" pageId="8" pageNumber="98">19</figureCitation>
). Anterior and posterior protarsal claws of identical shape and size (
<figureCitation id="130D2A30FFB5FFA2FEBFFADBDC33FAD2" box="[271,362,1290,1315]" captionStart="FIGURES 18 – 21" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[274,1284,1260,1890]" captionTargetId="figure@6.[264,1323,1235,1890]" captionTargetPageId="6" captionText="FIGURES 18 – 21. Drawings of fifth protarsomere and anterior claw (top) and posterior claw (bottom) in male (18, 20) and female (19, 21) of (18 – 19) Hydroporus rifensis sp. n. and (20 – 21) H. gyllenhalii. All drawings at same scale (scale bar: 0.1 mm)." httpUri="https://zenodo.org/record/228929/files/figure.png" pageId="8" pageNumber="98">Fig. 19</figureCitation>
). Ventral central surface of pro- and mesotarsomeres 1–2 without sucker cups (
<figureCitation id="130D2A30FFB5FFA2FA84FADBD8D6FAD3" box="[1332,1423,1290,1315]" captionStart="FIGURES 14 – 17" captionStartId="6.[151,264,1101,1124]" captionTargetBox="[229,1359,194,1079]" captionTargetId="figure@6.[227,1360,193,1081]" captionTargetPageId="6" captionText="FIGURES 14 – 17. SEM views of ventral side of left protarsus in male (14, 16) and female (15, 17) of (14 – 15) Hydroporus rifensis sp. n. (paratypes) and (16 – 17) H. gyllenhalii (16: male specimen from Fontainebleau, Ile-de-France, France; 17: female specimen from Rigny-Ussé, Touraine, France). Anterior on the left in all pictures. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/228928/files/figure.png" pageId="8" pageNumber="98">Fig. 15</figureCitation>
). Gonocoxosternum as in 
<figureCitation id="130D2A30FFB5FFA2FE18FAFFDCA3FAB7" box="[424,506,1326,1351]" captionStart="FIGURES 22 – 31" captionStartId="7.[151,264,1037,1060]" captionTargetBox="[180,1414,199,1006]" captionTargetId="figure@7.[170,1417,193,1017]" captionTargetPageId="7" captionText="FIGURES 22 – 31. Male and female genitalia of (22 – 26) Hydroporus rifensis sp. n. (paratypes) and (27 – 31) H. gyllenhalii (specimens from Plévenon, Bretagne, France). (22, 27) Median lobe of aedeagus in profile view; (23, 28) median lobe of aedeagus in ventral view; (24, 29) left paramere (external view); (25, 30) left gonocoxosternum (external view); (26, 31) gonocoxa (ventral view). Homologous structures shown at same scale for both species. Scale bars: 22 – 25, 27 – 30: 0.1 mm; 26, 31: 0.05 mm." httpUri="https://zenodo.org/record/228930/files/figure.png" pageId="8" pageNumber="98">Fig. 25</figureCitation>
; gonocoxa as in 
<figureCitation id="130D2A30FFB5FFA2FD05FAFFDE5EFAB7" box="[693,775,1326,1351]" captionStart="FIGURES 22 – 31" captionStartId="7.[151,264,1037,1060]" captionTargetBox="[180,1414,199,1006]" captionTargetId="figure@7.[170,1417,193,1017]" captionTargetPageId="7" captionText="FIGURES 22 – 31. Male and female genitalia of (22 – 26) Hydroporus rifensis sp. n. (paratypes) and (27 – 31) H. gyllenhalii (specimens from Plévenon, Bretagne, France). (22, 27) Median lobe of aedeagus in profile view; (23, 28) median lobe of aedeagus in ventral view; (24, 29) left paramere (external view); (25, 30) left gonocoxosternum (external view); (26, 31) gonocoxa (ventral view). Homologous structures shown at same scale for both species. Scale bars: 22 – 25, 27 – 30: 0.1 mm; 26, 31: 0.05 mm." httpUri="https://zenodo.org/record/228930/files/figure.png" pageId="8" pageNumber="98">Fig. 26</figureCitation>
.
</paragraph>
<paragraph id="8B8936B5FFB5FFA2FF77FA83D8C4F803" blockId="8.[151,1437,151,2035]" pageId="8" pageNumber="98">
<emphasis id="B942EAA7FFB5FFA2FF77FA83DC6CFA9B" bold="true" box="[199,309,1362,1387]" pageId="8" pageNumber="98">Variation</emphasis>
globally weak among 
<typeStatus id="548D8817FFB5FFA2FDF2FA82DF2BFA9B" box="[578,626,1363,1387]" pageId="8" pageNumber="98">type</typeStatus>
series. Lateral outline in some 
<typeStatus id="548D8817FFB5FFA2FC52FA82D917FA9B" box="[994,1102,1363,1387]" pageId="8" pageNumber="98" type="paratype">paratypes</typeStatus>
almost continuous; in others slightly more discontinuous than in 
<typeStatus id="548D8817FFB5FFA2FD89FAA7DFC6FA7F" box="[569,671,1398,1423]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
. In a few specimens colour contrast between pronotum and elytra stronger than in 
<typeStatus id="548D8817FFB5FFA2FEFCFA4BDCF7FA43" box="[332,430,1434,1459]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
(elytra testaceous-brown). On head symmetric brown markings between eyes confluent in most 
<typeStatus id="548D8817FFB5FFA2FF66FA6EDC1CFA27" box="[214,325,1471,1495]" pageId="8" pageNumber="98" type="paratype">paratypes</typeStatus>
; in a few specimens on contrary these markings separated medially. Posterior part of vertex pale brown to dark brown depending on specimens. Extension of paler colour on lateral and posterior part of elytron somewhat variable. In one female 
<typeStatus id="548D8817FFB5FFA2FD9FF9D6DFCBF9EF" box="[559,658,1543,1567]" pageId="8" pageNumber="98" type="paratype">paratype</typeStatus>
, microreticulation on head markedly more strongly impressed and punctation finer than in other specimens of 
<typeStatus id="548D8817FFB5FFA2FD22F9FADF9BF9B3" box="[658,706,1579,1603]" pageId="8" pageNumber="98">type</typeStatus>
series. Antennomeres 5–10 with testaceous colour restricted to basal quarter instead of basal third in some specimens; their shape slightly variable. Thickness of lateral bead of pronotum slightly variable. In some specimens lateral margins of pronotum less convergent in anterior half than in 
<typeStatus id="548D8817FFB5FFA2FF27F947DDA4F95F" box="[151,253,1686,1711]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
. Punctation of pronotal disc quite variable; in some specimens denser than in 
<typeStatus id="548D8817FFB5FFA2FB1EF947D84DF95F" box="[1198,1300,1686,1711]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
; sometimes decisively finer medially than laterally. In some specimens small punctures (1 mesh) not restricted to disc but also present laterally on pronotum. Mean size of elytral punctures slightly variable; in a few 
<typeStatus id="548D8817FFB5FFA2FB31F90ED9B5F907" box="[1153,1260,1759,1783]" pageId="8" pageNumber="98" type="paratype">paratypes</typeStatus>
punctures a bit larger than in 
<typeStatus id="548D8817FFB5FFA2FE84F8D3DCC0F8EB" box="[308,409,1794,1819]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
, in others distinctly finer. In some specimens inner puncture line a bit more distinct than in 
<typeStatus id="548D8817FFB5FFA2FF27F8F7DDA4F8CF" box="[151,253,1830,1855]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
. In some specimens genae slightly darker than gula. In most mature 
<typeStatus id="548D8817FFB5FFA2FBFDF8F6D9E2F8CF" box="[1101,1211,1831,1855]" pageId="8" pageNumber="98" type="paratype">paratypes</typeStatus>
, pale markings on abdominal ventrites 3–6 darker than in 
<typeStatus id="548D8817FFB5FFA2FDDEF89BDF89F893" box="[622,720,1866,1891]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
(ferrugineous instead of pale brown). Some specimens with surface of gula lacking micropunctures or these restricted to posterior gula area. Shape of prosternal process in most specimens slightly narrower than in 
<typeStatus id="548D8817FFB5FFA2FDDFF843DF88F85B" box="[623,721,1938,1963]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
(oblong-lanceolate); in some slightly more narrowly convex in cross section; sometimes apex a bit more acute than in 
<typeStatus id="548D8817FFB5FFA2FC99F867DED6F83F" box="[809,911,1974,1999]" pageId="8" pageNumber="98" type="holotype">holotype</typeStatus>
; ventral surface of prosternal process mostly smooth in many 
<typeStatus id="548D8817FFB5FFA2FEEDF80ADC95F803" box="[349,460,2011,2035]" pageId="8" pageNumber="98" type="paratype">paratypes</typeStatus>
. In some specimens antero-medial region of prosternum (before file) partly smooth.
</paragraph>
<paragraph id="8B8936B5FFB4FFA3FF27FF46DEC5FEED" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="99">
Anterior half of metacoxal lines in most specimens subparallel; sometimes a bit more divergent. Degree of impression of ventral microreticulation somewhat variable, notably on metacoxal plates. Brushes of long golden setae on middle of abdominal ventrites 4–5 lacking in most 
<typeStatus id="548D8817FFB4FFA3FC28FF31D95EFF08" box="[920,1031,224,248]" pageId="9" pageNumber="99" type="paratype">paratypes</typeStatus>
. Some specimens with pro- and mesotarsomeres and apical part of metatarsomeres slightly darkened.
</paragraph>
<paragraph id="8B8936B5FFB4FFA3FF77FEF6D9BDFD3C" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="99">
<emphasis id="B942EAA7FFB4FFA3FF77FEF6DC7AFEB0" bold="true" box="[199,291,295,320]" pageId="9" pageNumber="99">Habitat</emphasis>
. All specimens of the 
<typeStatus id="548D8817FFB4FFA3FD97FEF9DF0EFEB0" box="[551,599,296,320]" pageId="9" pageNumber="99">type</typeStatus>
series were found in 
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May 
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</date>
in a single large pond (dimensions about 
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<date id="FF881075FFB4FFA3FF55FE9CDC4FFE94" box="[229,278,332,356]" pageId="9" pageNumber="99" value="1980-10">
x 
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</date>
m) close to the N2 road at Anasar, Central Rif, ca. 
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WNW Bab Berred, at an altitude of 
<quantity id="4CCE9B50FFB4FFA3FA89FE9DD8C1FE94" box="[1337,1432,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.395" pageId="9" pageNumber="99" unit="m" value="1395.0">1,395 m</quantity>
. Around this pond the vegetation was dominated by shrubs and sparse trees on the upper (north-western) side, while the lower (south-western) side was more impacted by human activities with a prevalence of cultivated fields. At the period of visit, the pond was densely vegetated (
<figureCitation id="130D2A30FFB4FFA3FD75FE66DE41FE20" box="[709,792,439,464]" captionStart="FIGURE 32" captionStartId="7.[151,250,1934,1956]" captionText="FIGURE 32. Habitat of Hydroporus rifensis sp. n." pageId="9" pageNumber="99">Fig. 32</figureCitation>
) with 
<taxonomicName id="4C364D36FFB4FFA3FCD4FE69DEB0FE20" box="[868,1001,440,464]" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" kingdom="Plantae" order="Ranunculales" pageId="9" pageNumber="99" phylum="Tracheophyta" rank="genus">
<emphasis id="B942EAA7FFB4FFA3FCD4FE69DEB0FE20" box="[868,1001,440,464]" italics="true" pageId="9" pageNumber="99">Ranunculus</emphasis>
</taxonomicName>
sp. and 
<taxonomicName id="4C364D36FFB4FFA3FBFCFE69D99CFE20" box="[1100,1221,440,464]" class="Magnoliopsida" family="Plantaginaceae" genus="Callitriche" kingdom="Animalia" order="Lamiales" pageId="9" pageNumber="99" phylum="Tracheophyta" rank="genus">
<emphasis id="B942EAA7FFB4FFA3FBFCFE69D99CFE20" box="[1100,1221,440,464]" italics="true" pageId="9" pageNumber="99">Callitriche</emphasis>
</taxonomicName>
sp. aquatic plants, moderately abundant filamentous green algae, and scattered tufts of 
<taxonomicName id="4C364D36FFB4FFA3FC23FE0CDEB8FE04" box="[915,993,477,500]" class="Liliopsida" family="Juncaceae" genus="Juncus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="99" phylum="Tracheophyta" rank="genus">
<emphasis id="B942EAA7FFB4FFA3FC23FE0CDEB8FE04" box="[915,993,477,500]" italics="true" pageId="9" pageNumber="99">Juncus</emphasis>
</taxonomicName>
sp. The water was very transparent and there was no sign of pollution or other anthropogenic perturbation apart from a few scattered plastic objects. The shallow depth of this lentic water body and the aspect of the vegetation are indicative of a probable temporary nature. During wet periods the water is slowly renewed as deduced from the presence of a little brook leaving the pond towards the South (visible on Google Earth). Within the pond, specimens of 
<taxonomicName id="4C364D36FFB4FFA3FB9DFDBCD9FBFD74" box="[1069,1186,620,644]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="99" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFB4FFA3FB9DFDBCD9FBFD74" box="[1069,1186,620,644]" italics="true" pageId="9" pageNumber="99">H. rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFB4FFA3FB19FDBDD9B1FD75" box="[1193,1256,620,645]" pageId="9" pageNumber="99" rank="species">
<emphasis id="B942EAA7FFB4FFA3FB19FDBDD9B1FD75" bold="true" box="[1193,1256,620,645]" pageId="9" pageNumber="99">sp. n.</emphasis>
</taxonomicNameLabel>
were extremely localised. They were found exclusively in a zone along the northern pond edge characterised by an accumulation of decaying vegetal material under trees (this zone located on the extreme right of the photo 
<figureCitation id="130D2A30FFB4FFA3FB34FD65D981FD3C" box="[1156,1240,692,717]" captionStart="FIGURE 32" captionStartId="7.[151,250,1934,1956]" captionText="FIGURE 32. Habitat of Hydroporus rifensis sp. n." pageId="9" pageNumber="99">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C653EFFB4FFA3FF77FD06D9A7FAC0" pageId="9" pageNumber="99" type="biology_ecology">
<paragraph id="8B8936B5FFB4FFA3FF77FD06DC4EFCE5" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="99">
<emphasis id="B942EAA7FFB4FFA3FF77FD06DC46FD00" bold="true" box="[199,287,727,752]" pageId="9" pageNumber="99">Biology</emphasis>
. About half of the collected specimens were teneral, indicating that May is a period of intense imago emergence.
</paragraph>
<paragraph id="8B8936B5FFB4FFA3FF77FCCED9A7FAC0" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="99">
<emphasis id="B942EAA7FFB4FFA3FF77FCCEDC1EFCC8" bold="true" box="[199,327,799,824]" pageId="9" pageNumber="99">Study area</emphasis>
. Within the Betic-Rif massif, the collecting site belongs to the external domain, separated from the more northern internal domain by the Jebha fault line which runs in a NW-SE orientation a few kilometres north from the sampling site (
<bibRefCitation id="EFA74B44FFB4FFA3FE12FCB6DFCAFC70" author="Chalouan" box="[418,659,871,896]" pageId="9" pageNumber="99" refString="Chalouan, A., Michard, A., El Kadiri, K. &amp; Saddiqi, O. (2011) Nouveaux guides geologiques et miniers du Maroc. Volume 5. Rif central et nord-occidental. Editions du Service Geologique du Maroc, Rabat, 121 pp." type="book" year="2011">
Chalouan 
<emphasis id="B942EAA7FFB4FFA3FDA7FCB8DF09FC70" box="[535,592,872,896]" italics="true" pageId="9" pageNumber="99">et al.</emphasis>
2011
</bibRefCitation>
). The pond where 
<taxonomicName id="4C364D36FFB4FFA3FCD4FCB8DE80FC70" box="[868,985,872,896]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="99" phylum="Arthropoda" rank="species" species="rifensis" status="sp. nov.">
<emphasis id="B942EAA7FFB4FFA3FCD4FCB8DE80FC70" box="[868,985,872,896]" italics="true" pageId="9" pageNumber="99">H. rifensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157DCFFB4FFA3FC51FCB6D979FC70" box="[993,1056,871,896]" pageId="9" pageNumber="99" rank="species">
<emphasis id="B942EAA7FFB4FFA3FC51FCB6D979FC70" bold="true" box="[993,1056,871,896]" pageId="9" pageNumber="99">sp. n.</emphasis>
</taxonomicNameLabel>
was discovered is more precisely located in the intra-Rif structural zone close to its northern margin, at about 
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from the Mediterranean coast. The local geological substrate is a complex assemblage of radiolaritic argilites, calcareous marls and gresocalcareous turbidites of Upper Cretaceous (Cennomanian) age (
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Chalouan 
<emphasis id="B942EAA7FFB4FFA3FC5BFC04D97FFC1C" box="[1003,1062,980,1004]" italics="true" pageId="9" pageNumber="99">et al.</emphasis>
2011
</bibRefCitation>
). The pond is housed in a shallow depression along the southern versant of a mountain which culminates at an altitude of about 
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, just 
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north from the collecting site. The upper part of this mountain (up from the northern side of the road) corresponds to Lower Cretaceous flysh of the Tisirene nappe (
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Chalouan 
<emphasis id="B942EAA7FFB4FFA3FC63FB90D954FBA8" box="[979,1037,1088,1112]" italics="true" pageId="9" pageNumber="99">et al.</emphasis>
2011
</bibRefCitation>
), resting in abnormal contact onto the intra-Rif Cennomanian rocks. Various running water bodies originate from the northern versant of this mountain including an important coastal river, Oued Bouchia. Abundance of water in the area results from Central Rif being the most rainy place of all 
<collectingCountry id="F3217625FFB4FFA3FDFCFB7CDFEDFB34" box="[588,692,1197,1220]" name="Morocco" pageId="9" pageNumber="99">Morocco</collectingCountry>
, with exceptionally high average annual precipitations (between 1,000 and 
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in Ketama, 
<quantity id="4CCE9B50FFB4FFA3FDA9FB01DF3AFB18" box="[537,611,1231,1256]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="99" unit="km" value="40.0">40 km</quantity>
ESE from the 
<typeStatus id="548D8817FFB4FFA3FCA5FB00DE1CFB19" box="[789,837,1233,1257]" pageId="9" pageNumber="99">type</typeStatus>
locality, at a similar distance from the sea and at a comparable altitude). Precipitations are highest from November to April and minimal in July–August (data from fr.meteovista.be). The area is also characterised by cold temperatures in winter owing to the altitude.
</paragraph>
</subSubSection>
<subSubSection id="C32C653EFFB4FFA3FF77FAEDDECAFA88" pageId="9" pageNumber="99" type="etymology">
<paragraph id="8B8936B5FFB4FFA3FF77FAEDDECAFA88" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="99">
<emphasis id="B942EAA7FFB4FFA3FF77FAEDDCF8FAA5" bold="true" box="[199,417,1340,1365]" pageId="9" pageNumber="99">Derivatio nominis</emphasis>
. The specific epithet 
<taxonomicName id="4C364D36FFB4FFA3FD15FAEDDFAEFAA4" box="[677,759,1340,1364]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="99" phylum="Arthropoda" rank="species" species="rifensis">
<emphasis id="B942EAA7FFB4FFA3FD15FAEDDFAEFAA4" box="[677,759,1340,1364]" italics="true" pageId="9" pageNumber="99">rifensis</emphasis>
</taxonomicName>
(“from the Rif”) refers to the geographic area where the species was discovered. It is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C32C653EFFB4FFA3FF77FA55DC54FA30" pageId="9" pageNumber="99" type="distribution">
<paragraph id="8B8936B5FFB4FFA3FF77FA55DC54FA30" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="99">
<emphasis id="B942EAA7FFB4FFA3FF77FA55DC00FA6D" bold="true" box="[199,345,1412,1437]" pageId="9" pageNumber="99">Distribution</emphasis>
. So far only known from the 
<typeStatus id="548D8817FFB4FFA3FD7EFA54DFA7FA6D" box="[718,766,1413,1437]" pageId="9" pageNumber="99">type</typeStatus>
locality, a single pond at Anasar, Central Rif (northern 
<collectingCountry id="F3217625FFB4FFA3FF27FA79DC58FA30" box="[151,257,1448,1472]" name="Morocco" pageId="9" pageNumber="99">Morocco</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>