156 lines
20 KiB
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156 lines
20 KiB
XML
<document id="E588A329415BE62458D38A783275995C" ID-CLB-Dataset="21415" ID-DOI="10.5852/ejt.2020.691" ID-GBIF-Dataset="918d86bb-f11d-414e-9c1e-21ec17737792" ID-Zenodo-Dep="3961840" ID-ZooBank="CC8487E0-169D-4B32-A2AB-F9B34DFB3F3B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595862647445" checkinUser="valdenar" docAuthor="Grego, Jozef" docDate="2020" docId="039F79458404FFDEBFEDFA1A622920FB" docLanguage="en" docName="ejt-691_grego.pdf" docOrigin="European Journal of Taxonomy 691" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Travunijana robusta subsp. asculpta" docType="treatment" docVersion="8" lastPageNumber="44" masterDocId="FFA6013D842EFFF5BE26FFE6663C236D" masterDocTitle="Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions" masterLastPageNumber="56" masterPageNumber="1" pageNumber="43" updateTime="1698844003826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="98347F6459E2AB389C0C32D7405523E4">
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<mods:title id="826135AA54D73B2B357BEFD2196DB559">Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions</mods:title>
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<mods:namePart id="BA505F9AD9FC5BDBC34B4363F1BA0CA2">Grego, Jozef</mods:namePart>
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<mods:affiliation id="273161CA4021D3C8AE32BB22AA840890">Horná Mičiná, SK- 97401 Banská Bystrica, Slovakia.</mods:affiliation>
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<mods:nameIdentifier id="BBD75D7BC47BA123C7DBF4BAC05E9664" type="email">jozef.grego@gmail.com</mods:nameIdentifier>
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<mods:title id="0BDE987A705F0C9C3A664FD180E5DB76">European Journal of Taxonomy</mods:title>
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<mods:part id="8807BF864EC2694F22267D49EAADB6F2">
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<mods:date id="EA48DDB8816882D26A99FF94C3BFDBE0">2020</mods:date>
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<mods:number id="3B142A6AB49A74D7851EB77B4C88BC60">2020-07-27</mods:number>
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<mods:number id="9C864F1559A015671681034F91C7953D">691</mods:number>
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<mods:identifier id="D835B90BBCABB3D7CF2BE20ED5FC8031" type="DOI">10.5852/ejt.2020.691</mods:identifier>
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<treatment id="039F79458404FFDEBFEDFA1A622920FB" ID-DOI="http://doi.org/10.5281/zenodo.4336044" ID-GBIF-Taxon="166003137" ID-Zenodo-Dep="4336044" LSID="urn:lsid:plazi:treatment:039F79458404FFDEBFEDFA1A622920FB" httpUri="http://treatment.plazi.org/id/039F79458404FFDEBFEDFA1A622920FB" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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<subSubSection id="C32C9BD88404FFDFBFEDFA1A6255257A" box="[459,1129,1532,1559]" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="8B89C8538404FFDFBFEDFA1A6255257A" blockId="42.[459,1129,1532,1595]" box="[459,1129,1532,1559]" pageId="42" pageNumber="43">
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<heading id="D0C17F3F8404FFDFBFEDFA1A6255257A" box="[459,1129,1532,1559]" centered="true" fontSize="11" level="2" pageId="42" pageNumber="43" reason="2">
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<taxonomicName id="4C36B3D08404FFDFBFEDFA1A65E3257B" ID-CoL="7V3M6" authority="(Schutt, 1972)" baseAuthorityName="Schutt" baseAuthorityYear="1972" box="[459,991,1532,1559]" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="42" pageNumber="43" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
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<emphasis id="B94214418404FFDFBFEDFA1A650D257A" bold="true" box="[459,817,1532,1559]" italics="true" pageId="42" pageNumber="43">Travunijana robusta asculpta</emphasis>
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(Schütt, 1972)
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</taxonomicName>
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comb. nov.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C9BD88404FFDFBCEDF9C765552556" box="[715,873,1568,1595]" pageId="42" pageNumber="43" type="description">
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<paragraph id="8B89C8538404FFDFBCEDF9C765552556" blockId="42.[459,1129,1532,1595]" box="[715,873,1568,1595]" pageId="42" pageNumber="43">
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<figureCitation id="130DD4D68404FFDFBCEDF9C765102556" box="[715,812,1569,1595]" captionStart="Fig" captionStartId="15.[189,232,1572,1598]" captionTargetBox="[189,1398,261,1936]" captionTargetId="figure@16.[189,1398,261,1936]" captionTargetPageId="16" captionText="Fig. 7 (next page). A. Plagigeyeria mostarensis, Kuščer 1933, holotype, from Kuščer 1933, assigned to Izvor Buna in Blagaj but rather represents the form from near spring Bunica in Hodbina. B. P. mostarensis, hypotype of Schütt, 1970, from spring Bunica (SMF 221236). C. P. mostarensis, likely topotype from spring Bunica (JG F1146). D–E. P. mostarensis var., from spring of Buna assigned as type locality (JG F0536). Fa–Fb. Paladilhiopsis (Costellina) turrita (Kuščer, 1933) paratype and holotype from Kuščer 1933, spring Izvor in valley of river Jadro near Split. G. P. pseudocostellina sp. nov., left bank side spring of river Buna 400 m from main spring (holotype HNHM-MOLL-104164). H. P. pseudocostellina sp. nov., left bank side spring of river Buna 200 m from main spring (paratype JG F1140). I. Travunijana ovalis (Kuščer, 1933), holotype from Kuščer, assigned to Izvor Buna in Blagaj but not found in assigned locality, only in spring Bunica in Hodbina. J–K. T. ovalis (Kuščer, 1933), spring Bunica near Hodbina (JG F1145). L. T. robusta (Schütt, 1959), Trebinje district, Tučevac spring at right bank of Trebišnjica River (JG F1163). M. T. robusta, Trebinje district, Gorica, cave Vruljak 1 (JG F0831). N. T. robusta asculpta (Schütt, 1972), spring Ombla (Dubrovačka Rijeka) at Komolac near Dubrovnik (JG F1080)." figureDoi="http://doi.org/10.5281/zenodo.3962588" httpUri="https://zenodo.org/record/3962588/files/figure.png" pageId="42" pageNumber="43">Figs 7N</figureCitation>
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,
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<figureCitation id="130DD4D68404FFDFBD11F9C665552556" box="[823,873,1568,1595]" captionStart="Fig" captionStartId="44.[189,232,1860,1886]" captionTargetBox="[227,1361,265,1822]" captionTargetId="figure@44.[227,1361,265,1825]" captionTargetPageId="44" captionText="Fig. 15. A. Travunijana robusta (Schütt, 1959), Trebinje, IzvorTučevac, SBMNH 626406. B. Travunijana robusta (Schütt, 1959), Trebinje, Vrelo Vruljak 1, SBMNH 625919. C. Travunijana robusta asculpta (Schütt, 1972), Croatia, Dubrovnik, Komolac, Izvor Ombla, SBMNH 625896. D. Travunijana gloeri sp. nov., Bjeljani, Dabarsko Polje, Vrelo Vrijeka, paratype SBMNH 632722. Scale bars = 1mm. (SEM SBMNH Vanessa Delnavaz)." figureDoi="http://doi.org/10.5281/zenodo.3961864" httpUri="https://zenodo.org/record/3961864/files/figure.png" pageId="42" pageNumber="43">15C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C9BD88404FFDFBE9BF982641E258B" pageId="42" pageNumber="43" type="reference_group">
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<paragraph id="8B89C8538404FFDFBE9BF98262382513" blockId="42.[189,1028,1635,1662]" box="[189,1028,1635,1662]" pageId="42" pageNumber="43">
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<treatmentCitationGroup id="AB26EF7D8404FFDFBE9BF98262382513" box="[189,1028,1635,1662]" pageId="42" pageNumber="43">
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<treatmentCitation id="0A97EE428404FFDFBE9BF982650F2513" box="[189,819,1635,1662]" pageId="42" pageNumber="43">
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<taxonomicName id="4C36B3D08404FFDFBE9BF98264CB2513" ID-CoL="7TZQN" authority="Schutt, 1972: 115" authorityName="Schutt" authorityPageNumber="115" authorityYear="1972" box="[189,759,1635,1662]" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="42" pageNumber="43" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
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<emphasis id="B94214418404FFDFBE9BF98264232513" box="[189,543,1636,1662]" italics="true" pageId="42" pageNumber="43">Plagigeyeria robusta asculpta</emphasis>
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Schütt, 1972: 115
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</taxonomicName>
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, 122
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</treatmentCitation>
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, pl. 8, figs 25–26.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B89C8538404FFDFBE9BF940641E258B" blockId="42.[189,1398,1702,1766]" pageId="42" pageNumber="43">
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<treatmentCitationGroup id="AB26EF7D8404FFDFBE9BF940641E258B" pageId="42" pageNumber="43">
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<taxonomicName id="4C36B3D08404FFDFBE9BF940642325AD" ID-CoL="7TZQN" authorityName="Schutt" authorityYear="1972" box="[189,543,1702,1728]" class="Gastropoda" family="Hydrobiidae" genus="Plagigeyeria" kingdom="Animalia" order="Littorinimorpha" pageId="42" pageNumber="43" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
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<emphasis id="B94214418404FFDFBE9BF940642325AD" box="[189,543,1702,1728]" italics="true" pageId="42" pageNumber="43">Plagigeyeria robusta asculpta</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A97EE428404FFDFBC1BF94065BA25AC" author="Bole J. & Velkovrh F." box="[573,902,1702,1729]" page="202" pageId="42" pageNumber="43" year="1986">
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<bibRefCitation id="EFA7B5A28404FFDFBC1BF94065BA25AC" author="Bole J. & Velkovrh F." box="[573,902,1702,1729]" pageId="42" pageNumber="43" pagination="177 - 206" refId="ref24755" refString="Bole J. & Velkovrh F. 1986. Mollusca from continental subterranean aquatic habitats. In: Botosaneanu P. (ed.) Stygofauna mundi: 177 - 206. E. J. Brill, Leiden." type="book chapter" year="1986">Bole & Velkovrh 1986: 202</bibRefCitation>
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</treatmentCitation>
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. —
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||
<bibRefCitation id="EFA7B5A28404FFDFBD90F940620A25AC" author="Bank R. A." box="[950,1078,1702,1729]" pageId="42" pageNumber="43" refId="ref24339" refString="Bank R. A. 2013. Fauna Europaea: Mollusca. Fauna Europaea version 2.6. 2. https: // fauna-eu. org / [accessed 11 Nov. 2019]" type="url" year="2013">Bank 2013</bibRefCitation>
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: Fauna Europaea v. 2.6. —
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<treatmentCitation id="0A97EE428404FFDFBECCF92D6423258B" author="Bank R. A. & Neubert E." box="[234,543,1739,1766]" page="25" pageId="42" pageNumber="43" year="2017">
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<bibRefCitation id="EFA7B5A28404FFDFBECCF92D6423258B" author="Bank R. A. & Neubert E." box="[234,543,1739,1766]" pageId="42" pageNumber="43" refId="ref24372" refString="Bank R. A. & Neubert E. 2017. Checklist of the land and freshwater Gastropoda of Europe. Available from http: // www. molluscabase. org / aphia. php? p = sourcedetails & id = 279050 [last update: 16 Jul. 2017; accessed 11 Nov. 2019]." type="url" year="2017">Bank & Neubert 2017: 25</bibRefCitation>
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32C9BD88404FFDFBE9BF8E867342486" pageId="42" pageNumber="43" type="diagnosis">
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<paragraph id="8B89C8538404FFDFBE9BF8E8670A2445" blockId="42.[189,310,1806,1832]" box="[189,310,1806,1832]" pageId="42" pageNumber="43">
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<heading id="D0C17F3F8404FFDFBE9BF8E8670A2445" bold="true" box="[189,310,1806,1832]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
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<emphasis id="B94214418404FFDFBE9BF8E8670A2445" bold="true" box="[189,310,1806,1832]" pageId="42" pageNumber="43">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B89C8538404FFDFBE9BF8D867342486" blockId="42.[189,1399,1854,2027]" pageId="42" pageNumber="43">Shell very similar to nominotypical subspecies but having in general a more elevated, more conical spire with blunter apex. Surface can be smooth or finely ribbed as well as in the nominotypical morphotype. Aperture ear-shaped and missing the slightly protruded columellar fold of the nominotypical subspecies. Lateral labral profile straight and columellar labrum possesses a weak cut sinuation. Protoconch surface pitted.</paragraph>
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</subSubSection>
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<subSubSection id="C32C9BD88405FFDEBE9BFEED652522F0" pageId="43" pageNumber="44" type="distribution">
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<paragraph id="8B89C8538405FFDEBE9BFEED676A2248" blockId="43.[189,342,267,293]" box="[189,342,267,293]" pageId="43" pageNumber="44">
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<heading id="D0C17F3F8405FFDEBE9BFEED676A2248" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
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<emphasis id="B94214418405FFDEBE9BFEED676A2248" bold="true" box="[189,342,267,293]" pageId="43" pageNumber="44">Distribution</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B89C8538405FFDEBE9BFEDD652522F0" blockId="43.[189,1399,314,413]" pageId="43" pageNumber="44">
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<taxonomicName id="4C36B3D08405FFDEBE9BFEDD642B2239" baseAuthorityName="Schutt" baseAuthorityYear="1972" box="[189,535,314,340]" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
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<emphasis id="B94214418405FFDEBE9BFEDD642B2239" box="[189,535,314,340]" italics="true" pageId="43" pageNumber="44">Travunijana robusta asculpta</emphasis>
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</taxonomicName>
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can be found in the thanatocoenoses of the Vrelo “Ombla” near Komolac (
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<quantity id="4CCE65B68405FFDEBEE0FEB866CE2214" box="[198,242,350,377]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="43" pageNumber="44" unit="m" value="7.0">7 m</quantity>
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a.s.l.) and Vrelo “Mlin” (
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||
<quantity id="4CCE65B68405FFDEBC3EFEB864782214" box="[536,580,350,377]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="m" value="2.0">2 m</quantity>
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||
a.s.l.) at
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<collectingCountry id="F32188C38405FFDEBC8AFEB864DF2215" box="[684,739,350,376]" name="Serbia" pageId="43" pageNumber="44">Mali</collectingCountry>
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||
Zaton north of Dubrovnik. Springs are draining the karst area of upper-to-central Popovo Polje (
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||
<quantity id="4CCE65B68405FFDEBCA3FE6464ED22F0" box="[645,721,386,413]" metricMagnitude="2" metricUnit="m" metricValue="2.27" pageId="43" pageNumber="44" unit="m" value="227.0">227 m</quantity>
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a.s.l).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32C9BD88405FFDEBE9BFE22622920FB" pageId="43" pageNumber="44" type="discussion">
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<paragraph id="8B89C8538405FFDEBE9BFE22671322B3" blockId="43.[189,303,452,478]" box="[189,303,452,478]" pageId="43" pageNumber="44">
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||
<heading id="D0C17F3F8405FFDEBE9BFE22671322B3" bold="true" box="[189,303,452,478]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
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||
<emphasis id="B94214418405FFDEBE9BFE22671322B3" bold="true" box="[189,303,452,478]" pageId="43" pageNumber="44">Remarks</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B89C8538405FFDEBE9BFE15622920FB" blockId="43.[189,1399,499,919]" pageId="43" pageNumber="44">
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||
The morphological variability of nominotypical subspecies and
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||
<taxonomicName id="4C36B3D08405FFDEBD82FE1262BE2160" baseAuthorityName="Schutt" baseAuthorityYear="1972" box="[932,1154,499,525]" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
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||
<emphasis id="B94214418405FFDEBD82FE1262BE2160" box="[932,1154,499,525]" italics="true" pageId="43" pageNumber="44">T. robusta asculpta</emphasis>
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||
</taxonomicName>
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||
overlap significantly and even
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||
<bibRefCitation id="EFA7B5A28405FFDEBF08FDF167F7215C" author="Schutt H." box="[302,459,535,561]" pageId="43" pageNumber="44" pagination="203 - 215" refId="ref26197" refString="Schutt H. 2000. Die Hohlenmollusken der Ombla-Quelle. Natura Croatica 9 (3): 203 - 215." type="journal article" year="2000">Schütt (2000)</bibRefCitation>
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||
assigned some of specimens of
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<taxonomicName id="4C36B3D08405FFDEBD63FDFE621D215C" baseAuthorityName="Schutt" baseAuthorityYear="1972" box="[837,1057,535,561]" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
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||
<emphasis id="B94214418405FFDEBD63FDFE621D215C" box="[837,1057,535,561]" italics="true" pageId="43" pageNumber="44">P. robusta asculpta</emphasis>
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||
</taxonomicName>
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from the
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<typeStatus id="548D76F18405FFDEBAB0FDF162FB215C" box="[1174,1223,535,561]" pageId="43" pageNumber="44">type</typeStatus>
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locality (Vrelo “Ombla”) to the nominotypical subspecies based upon the presence of ribs, and conversely, the few specimens collected from Trebišnjica Springs in Bileća were assigned to
|
||
<taxonomicName id="4C36B3D08405FFDEBD5BFDB962692115" baseAuthorityName="Schutt" baseAuthorityYear="1972" box="[893,1109,606,632]" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
|
||
<emphasis id="B94214418405FFDEBD5BFDB962692115" box="[893,1109,606,632]" italics="true" pageId="43" pageNumber="44">T. robusta asculpta</emphasis>
|
||
</taxonomicName>
|
||
just for their lack of ribs. However the presence or absence of the fine shell ribs on the eroded shells found in the thanatocoenoses has to be carefully evaluated by their remains around the suture, as upon first sight both ribbed and smooth forms can be found in the thanatocoenoses within the springs draining the whole Eastern Trebišnjica aquifer. The main distinguishing feature of
|
||
<taxonomicName id="4C36B3D08405FFDEBC32FD0864D5206A" baseAuthorityName="Schutt" baseAuthorityYear="1972" box="[532,745,749,775]" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="subSpecies" species="robusta" subSpecies="asculpta">
|
||
<emphasis id="B94214418405FFDEBC32FD0864D5206A" box="[532,745,749,775]" italics="true" pageId="43" pageNumber="44">T. robusta asculpta</emphasis>
|
||
</taxonomicName>
|
||
is in general the more elevated and more conical spire and slightly more elongated shell shape, but even this feature possesses a high variability within each locality. Until live material becomes available and larger shells material can be examined, I prefer to separate both in a provisional way on a subspecies level, based only on their zoogeographical data. It is possible that further investigation could reveal several related species in the
|
||
<taxonomicName id="4C36B3D08405FFDEBD15FC9B659820FA" authorityName="Grego" authorityYear="2020" baseAuthorityName="Schutt" baseAuthorityYear="1959" box="[819,932,893,919]" class="Gastropoda" family="Hydrobiidae" genus="Travunijana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="robusta">
|
||
<emphasis id="B94214418405FFDEBD15FC9B659820FA" box="[819,932,893,919]" italics="true" pageId="43" pageNumber="44">T. robusta</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |