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<mods:title id="3DC83EE59AF8068D257B954167D2298D">Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea)</mods:title>
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<mods:namePart id="607E89F16417E3F0DC8819A3CF6D9836">Park, Kyu-Tek</mods:namePart>
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<mods:namePart id="68A7F7F37B509769DBD8B0EEB3F6A33F">Koo, Jun-Mo</mods:namePart>
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<mods:namePart id="579AF9BA49A44FE7417C1C68D3235A36">Minet, Joël</mods:namePart>
<mods:affiliation id="8E12A6D233787881CF003CC1DA1436E4">Muséum National dHistoire Naturelle (ISYEB, UMR 7205), Entomologie (C. P. 50), 45, rue Buffon, F- 75005 Paris, France.</mods:affiliation>
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<taxonomicName id="4C3BB3FFFFBEFFD55CE19E04FE51FE7E" ID-CoL="92PKV" authority="Meyrick, 1894" authorityName="Meyrick" authorityYear="1894" box="[151,497,366,392]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBEFFD55CE19E04FE51FE7E" bold="true" box="[151,497,366,392]" pageId="9" pageNumber="160">
<emphasis id="B94F146EFFBEFFD55CE19E04FE96FE7E" bold="true" box="[151,310,366,392]" italics="true" pageId="9" pageNumber="160">TORODORA</emphasis>
<bibRefCitation id="EFAAB58DFFBEFFD55D489E04FE51FE7E" author="Meyrick, E." box="[318,497,366,392]" pageId="9" pageNumber="160" pagination="1 - 29" refId="ref23540" refString="Meyrick, E. (1894) On a collection of Lepidoptera from Upper Burma. Transactions of the entomological Society of London, 1894 (1), 1 - 29." type="journal article" year="1894">Meyrick, 1894</bibRefCitation>
</emphasis>
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<typeStatus id="548076DEFFBEFFD55CE19ED9FF69FE3D" box="[151,201,435,459]" pageId="9" pageNumber="160">Type</typeStatus>
species:
<taxonomicName id="4C3BB3FFFFBEFFD55D509ED9FD3DFE3D" authority="Meyrick, 1894" authorityName="Meyrick" authorityYear="1894" box="[294,669,435,459]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="characteris">
<emphasis id="B94F146EFFBEFFD55D509ED9FE5EFE3D" box="[294,510,435,459]" italics="true" pageId="9" pageNumber="160">Torodora characteris</emphasis>
<bibRefCitation id="EFAAB58DFFBEFFD55E729ED9FD3DFE3D" author="Meyrick, E." box="[516,669,435,459]" pageId="9" pageNumber="160" pagination="1 - 29" refId="ref23540" refString="Meyrick, E. (1894) On a collection of Lepidoptera from Upper Burma. Transactions of the entomological Society of London, 1894 (1), 1 - 29." type="journal article" year="1894">Meyrick, 1894</bibRefCitation>
</taxonomicName>
.
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<paragraph id="8B84C87CFFBEFFD55CE19E93FC84FC35" blockId="9.[151,1437,505,1036]" pageId="9" pageNumber="160">
The
<taxonomicName id="4C3BB3FFFFBEFFD55CBC9E93FEF0FDE5" ID-CoL="92CZT" authorityName="Herrich-Schaffer" authorityYear="1853" box="[202,336,505,531]" class="Insecta" family="Lecithoceridae" genus="Lecithocera" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBEFFD55CBC9E93FEF0FDE5" box="[202,336,505,531]" italics="true" pageId="9" pageNumber="160">Lecithocera</emphasis>
</taxonomicName>
species described by
<bibRefCitation id="EFAAB58DFFBEFFD55E359E93FD6EFDE2" author="Viette, P. E. L." box="[579,718,505,532]" pageId="9" pageNumber="160" pagination="75 - 114" refId="ref24327" refString="Viette, P. E. L. (1955) Nouveaux Tineoidea (s. l.) de Madagascar [Lep.]. Annales de la Societe entomologique de France, 123 (1954), 75 - 114." type="journal article" year="1955">Viette (1955</bibRefCitation>
<bibRefCitation id="EFAAB58DFFBEFFD55EAB9E93FCB6FDE2" author="Viette, P. E. L." box="[733,790,505,532]" pageId="9" pageNumber="160" pagination="99 - 109" refId="ref24718" refString="Viette, P. E. L. (1988) Nouveaux Microlepidopteres de Madagascar (Xyloryctidae, Stenomidae, Lecithoceridae, Gelechiidae) [Lep.]. Bulletin de la Societe entomologique de France, 93 (3 - 4), 99 - 109. https: // doi. org / 10.3406 / linly. 1986.10753" type="journal article" year="1988">1988</bibRefCitation>
) definitely do not belong to the subfamily
<taxonomicName id="4C3BB3FFFFBEFFD558809E93FA3CFDE5" box="[1270,1436,505,531]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="subFamily" subFamily="Lecithocerinae">Lecithocerinae</taxonomicName>
(hence are not members of the genus
<taxonomicName id="4C3BB3FFFFBEFFD55E379D77FC32FDCE" ID-CoL="92CZT" authority="Herrich-Schaffer" authorityName="Herrich-Schaffer" authorityYear="1853" box="[577,914,541,568]" class="Insecta" family="Lecithoceridae" genus="Lecithocera" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBEFFD55E379D77FD67FDC1" box="[577,711,541,567]" italics="true" pageId="9" pageNumber="160">Lecithocera</emphasis>
Herrich-Schäffer
</taxonomicName>
); actually they all belong to the subfamily
<taxonomicName id="4C3BB3FFFFBEFFD559009D77FEA2FDAD" ID-CoL="9DMY8" authorityName="Gozmany" authorityYear="1978" class="Insecta" family="Lecithoceridae" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="subFamily" subFamily="Torodorinae">Torodorinae</taxonomicName>
since they have spinose zones on the abdominal segments (dorsally), and male genitalia possessing an elongate uncus but lacking the well developed costal bar occurring in
<taxonomicName id="4C3BB3FFFFBEFFD55FE99D0FFBE5FD89" box="[927,1093,613,639]" class="Insecta" family="Gelechiidae" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="subFamily" subFamily="Lecithocerinae">Lecithocerinae</taxonomicName>
(i.e. a bar connecting tegumen and valva). The 12
<emphasis id="B94F146EFFBEFFD55D1D9DE3FDB0FD55" box="[363,528,649,675]" italics="true" pageId="9" pageNumber="160">
<taxonomicName id="4C3BB3FFFFBEFFD55D039DE3FDA5FD55" ID-CoL="92CZT" authorityName="Herrich-Schaffer" authorityYear="1853" box="[373,517,649,675]" class="Insecta" family="Lecithoceridae" genus="Lecithocera" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">Lecithocera</taxonomicName>
</emphasis>
species assigned to
<taxonomicName id="4C3BB3FFFFBEFFD55E999DE3FCF7FD55" ID-CoL="92PKV" box="[751,855,649,675]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBEFFD55E999DE3FCF7FD55" box="[751,855,649,675]" italics="true" pageId="9" pageNumber="160">Torodora</emphasis>
</taxonomicName>
may show considerable variations in the venation of both wings: CuA
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and CuA
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stalked or coincident in the forewing; M
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stalked with CuA
<subScript id="17BFCA39FFBEFFD558129DD5FBCCFD3B" attach="left" box="[1124,1132,703,717]" fontSize="6" pageId="9" pageNumber="160">1</subScript>
or M
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entirely merged with CuA
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in
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the hind wing. Although such differences in the wing venation can be easily observed, there are no marked differences in the male genitalia and in the abdominal segments VIIVIII (not specifically modified). Accordingly, all the species are tentatively placed together in a single genus,
<taxonomicName id="4C3BB3FFFFBEFFD55F1D9C73FB9DFCC5" ID-CoL="92PKV" authority="Meyrick" authorityName="Meyrick" box="[875,1085,793,819]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBEFFD55F1D9C73FC73FCC5" box="[875,979,793,819]" italics="true" pageId="9" pageNumber="160">Torodora</emphasis>
Meyrick
</taxonomicName>
, also on account of other similarities, notably in antennae, labial palpi, and wing pattern. The interrelationships of these species will probably be solved by the further analysis of DNA sequences when fresh material can be obtained. All the species assigned to
<taxonomicName id="4C3BB3FFFFBEFFD55CE19CEFFF5FFC69" ID-CoL="92PKV" box="[151,255,901,927]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBEFFD55CE19CEFFF5FFC69" box="[151,255,901,927]" italics="true" pageId="9" pageNumber="160">Torodora</emphasis>
</taxonomicName>
in
<collectingCountry id="F32C88ECFFBEFFD55D549CEFFE09FC69" box="[290,425,901,927]" name="Madagascar" pageId="9" pageNumber="160">Madagascar</collectingCountry>
are here tentatively divided into two species-groups, based on differences in the hind wing venation (M
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stalked with CuA
<subScript id="17BFCA39FFBEFFD55D879CD1FE59FC3F" attach="left" box="[497,505,955,969]" fontSize="6" pageId="9" pageNumber="160">1</subScript>
or M
<subScript id="17BFCA39FFBEFFD55E419CD1FD9FFC3F" attach="left" box="[567,575,955,969]" fontSize="6" pageId="9" pageNumber="160">3</subScript>
merged with CuA
<subScript id="17BFCA39FFBEFFD55F7A9CD1FCB4FC3F" attach="left" box="[780,788,955,969]" fontSize="6" pageId="9" pageNumber="160">1</subScript>
).
</paragraph>
<paragraph id="8B84C87CFFBEFFD55CB19CA7FB3AFBFA" blockId="9.[151,1437,505,1036]" pageId="9" pageNumber="160">
On the other hand, it should be noted that two of these species had already been transferred to the genus
<taxonomicName id="4C3BB3FFFFBEFFD559169CA4FF6CFBFD" ID-CoL="92PKV" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFBEFFD559169CA4FF6CFBFD" italics="true" pageId="9" pageNumber="160">Torodora</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAAB58DFFBEFFD55C809C9BFEDDFBFD" author="Park, K. - T." box="[246,381,1009,1035]" pageId="9" pageNumber="160" pagination="1085 - 1093" refId="ref23829" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park (2018)</bibRefCitation>
, namely
<taxonomicName id="4C3BB3FFFFBEFFD55D949C98FC83FBFA" ID-CoL="9FPWQ" authority="(Viette, 1986)" baseAuthorityName="Viette" baseAuthorityYear="1986" box="[482,803,1009,1036]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="kambanella">
<emphasis id="B94F146EFFBEFFD55D949C98FD21FBFD" box="[482,641,1009,1035]" italics="true" pageId="9" pageNumber="160">T. kambanella</emphasis>
(
<bibRefCitation id="EFAAB58DFFBEFFD55EF99C9BFCBBFBFA" author="Viette, P. E. L." box="[655,795,1009,1036]" pageId="9" pageNumber="160" pagination="109 - 118" refId="ref24658" refString="Viette, P. E. L. (1986) Nouveaux Microlepidopteres de Madagascar (Oecophoridae et Gelechiidae) (New species of Malagasy Microlepidoptera-Oecophoridae and Gelechiidae). Bulletin mensuel de la Societe linneenne de Lyon, 55 (4), 109 - 118. https: // doi. org / 10.3406 / linly. 1986.10753" type="journal article" year="1986">Viette, 1986</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C3BB3FFFFBEFFD55F2F9C98FB35FBFA" ID-CoL="9FPWS" authority="(Viette, 1955)" baseAuthorityName="Viette" baseAuthorityYear="1955" box="[857,1173,1009,1036]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="masoalella">
<emphasis id="B94F146EFFBEFFD55F2F9C98FC50FBFD" box="[857,1008,1009,1035]" italics="true" pageId="9" pageNumber="160">T. masoalella</emphasis>
(
<bibRefCitation id="EFAAB58DFFBEFFD55F889C9BFB2DFBFA" author="Viette, P. E. L." box="[1022,1165,1009,1036]" pageId="9" pageNumber="160" pagination="75 - 114" refId="ref24327" refString="Viette, P. E. L. (1955) Nouveaux Tineoidea (s. l.) de Madagascar [Lep.]. Annales de la Societe entomologique de France, 123 (1954), 75 - 114." type="journal article" year="1955">Viette, 1955</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
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