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<document id="62B5F9CF9556AB8970F20E844A7A58F3" ID-CLB-Dataset="42309" ID-DOI="10.11646/zootaxa.3878.5.4" ID-GBIF-Dataset="697c91ee-9d8b-4095-bd15-e55709b2b6bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="229037" ID-ZooBank="377F9037-D039-44A5-B562-051560CBC9E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460949457252" checkinUser="plazi" docAuthor="Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick &amp; Nel, André" docDate="2014" docId="03923C218622FF8BFF45FA98009DFCA6" docLanguage="en" docName="zt03878p470.pdf" docOrigin="Zootaxa 3878 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aviologus duquesnei Coty, Háva, Prokop, Roques &amp; Nel, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="468" masterDocId="FFAB44598627FF8DFFD2FFED015AFFFB" masterDocTitle="New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)" masterLastPageNumber="470" masterPageNumber="462" pageNumber="467" updateTime="1698585716371" updateUser="ExternalLinkService">
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<mods:title id="D369CF304C0EDD6C0EC1CA246A83B38B">New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)</mods:title>
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<mods:name id="3D41169C65E30217381D43D88E4BB71D" type="personal">
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<mods:namePart id="A913DDD4F94AE91B2DF750C430CEE8FB">Coty, David</mods:namePart>
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<mods:namePart id="6A71485060BAF1E502F9AB0A7E0172A8">Háva, Jiří</mods:namePart>
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<mods:namePart id="F2834F5519ED88FF2C46D4FD06FD5B2C">Prokop, Jakub</mods:namePart>
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<mods:namePart id="4FBEB2D7FB288C5CB00B28F05284B1EC">Roques, Patrick</mods:namePart>
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<mods:namePart id="69785C2C15E4429E4E2D49BFC3543430">Nel, André</mods:namePart>
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<mods:title id="A253C135784458079F525613B73FE72F">Zootaxa</mods:title>
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<mods:date id="4790718F49B19B5B5BB11B63E28DEA9A">2014</mods:date>
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<mods:number id="F7A3807733913C6E80EDE4A53B472324">3878</mods:number>
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<treatment id="03923C218622FF8BFF45FA98009DFCA6" ID-DOI="http://doi.org/10.5281/zenodo.6142671" ID-GBIF-Taxon="119591900" ID-Zenodo-Dep="6142671" LSID="urn:lsid:plazi:treatment:03923C218622FF8BFF45FA98009DFCA6" httpUri="http://treatment.plazi.org/id/03923C218622FF8BFF45FA98009DFCA6" lastPageId="6" lastPageNumber="468" pageId="5" pageNumber="467">
<subSubSection id="C321DEBC8622FF88FF45FA9801B1FA4B" pageId="5" pageNumber="467" type="nomenclature">
<paragraph id="8B848D378622FF88FF45FA9800AFFA74" blockId="5.[151,501,1397,1457]" box="[151,501,1397,1423]" pageId="5" pageNumber="467">
<heading id="D0CC3A5B8622FF88FF45FA9800AFFA74" bold="true" box="[151,501,1397,1423]" fontSize="11" level="1" pageId="5" pageNumber="467" reason="1">
<emphasis id="B94F51258622FF88FF45FA9800AFFA74" bold="true" box="[151,501,1397,1423]" pageId="5" pageNumber="467">
<taxonomicName id="4C3BF6B48622FF88FF45FA9800CBFA74" box="[151,401,1397,1423]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="5" pageNumber="467" phylum="Arthropoda" rank="species" species="duquesnei" status="sp. nov.">
<emphasis id="B94F51258622FF88FF45FA9800CBFA74" bold="true" box="[151,401,1397,1423]" italics="true" pageId="5" pageNumber="467">Aviologus duquesnei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27CEC5E8622FF88FE45FA9800AFFA74" box="[407,501,1397,1423]" pageId="5" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B848D378622FF88FF45FA7501B1FA4B" blockId="5.[151,501,1397,1457]" box="[151,235,1432,1457]" pageId="5" pageNumber="467">
(
<figureCitation id="130091B28622FF88FF4DFA7501B9FA4B" box="[159,227,1432,1457]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1924]" captionTargetBox="[151,1436,916,1873]" captionTargetId="figure@6.[151,1436,916,1878]" captionTargetPageId="6" captionText="FIGURE 4. Aviologus duquesnei gen. nov. et sp. nov., holotype Avion 0 0 0 34 (print and counterprint) stored in MNHN coll. finally deposited in the Musée Géologique Pierre Vetter, Decazeville, France. A, photograph of forewing venation; B, line drawing of forewing venation. Scale bar = 5 mm." httpUri="https://zenodo.org/record/229040/files/figure.png" pageId="5" pageNumber="467">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C321DEBC8622FF88FF45FA0D032DF99F" pageId="5" pageNumber="467" type="materials_examined">
<paragraph id="8B848D378622FF88FF45FA0D0262F9E7" blockId="5.[151,1437,1504,2033]" pageId="5" pageNumber="467">
<emphasis id="B94F51258622FF88FF45FA0D0004FA02" bold="true" box="[151,350,1504,1529]" pageId="5" pageNumber="467">Material studied</emphasis>
:
<typeStatus id="548033958622FF88FEBFFA0D008FFA02" box="[365,469,1504,1529]" pageId="5" pageNumber="467" type="holotype">Holotype</typeStatus>
specimen Avion 0 0 0 34 (print and counterprint), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville,
<collectingCountry id="F32CCDA78622FF88FD36F9E8026EF9E7" box="[740,820,1541,1564]" name="France" pageId="5" pageNumber="467">France</collectingCountry>
.
</paragraph>
<paragraph id="8B848D378622FF88FF15F9C5032DF99F" blockId="5.[151,1437,1504,2033]" pageId="5" pageNumber="467">
<emphasis id="B94F51258622FF88FF15F9C500D6F9BA" bold="true" box="[199,396,1576,1601]" pageId="5" pageNumber="467">Age and outcrop</emphasis>
.
<typeStatus id="548033958622FF88FE49F9C40089F9BA" box="[411,467,1577,1601]" pageId="5" pageNumber="467">Type</typeStatus>
locality and horizon. Moscovian (Westphalian C/D equivalent to Bolsovian/Asturian), Terril N°7, Avion, Pas-de-Calais,
<collectingCountry id="F32CCDA78622FF88FDF6F9A0032EF99F" box="[548,628,1613,1636]" name="France" pageId="5" pageNumber="467">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C321DEBC8622FF88FF15F99D034DF957" pageId="5" pageNumber="467" type="etymology">
<paragraph id="8B848D378622FF88FF15F99D034DF957" blockId="5.[151,1437,1504,2033]" pageId="5" pageNumber="467">
<emphasis id="B94F51258622FF88FF15F99D001CF972" bold="true" box="[199,326,1648,1673]" pageId="5" pageNumber="467">Etymology</emphasis>
. Named after Mr. Hervé Duquesne, who is a well known collector of plants and insects in the Carboniferous of North of
<collectingCountry id="F32CCDA78622FF88FE11F9780349F957" box="[451,531,1685,1708]" name="France" pageId="5" pageNumber="467">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C321DEBC8622FF88FF15F9550352F92A" box="[199,520,1720,1745]" pageId="5" pageNumber="467" type="diagnosis">
<paragraph id="8B848D378622FF88FF15F9550352F92A" blockId="5.[151,1437,1504,2033]" box="[199,520,1720,1745]" pageId="5" pageNumber="467">
<emphasis id="B94F51258622FF88FF15F9550060F92A" bold="true" box="[199,314,1720,1745]" pageId="5" pageNumber="467">Diagnosis</emphasis>
.
<collectingCountry id="F32CCDA78622FF88FE9AF954003DF92A" box="[328,359,1721,1745]" name="American Samoa" pageId="5" pageNumber="467">As</collectingCountry>
for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C321DEBC8622FF8BFF15F93100C1FEBB" lastPageId="6" lastPageNumber="468" pageId="5" pageNumber="467" type="description">
<paragraph id="8B848D378622FF8BFF15F93100C1FEBB" blockId="5.[151,1437,1504,2033]" lastBlockId="6.[151,1437,151,861]" lastPageId="6" lastPageNumber="468" pageId="5" pageNumber="467">
<emphasis id="B94F51258622FF88FF15F931000BF90E" bold="true" box="[199,337,1756,1781]" pageId="5" pageNumber="467">Description</emphasis>
. A nearly complete forewing perfectly preserved with only fragments of mid part missing. Preserved length
<quantity id="4CC320D28622FF88FE88F8ED009BF8E3" box="[346,449,1792,1817]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="5" pageNumber="467" unit="mm" value="22.6">22.6 mm</quantity>
, width
<quantity id="4CC320D28622FF88FDC6F8ED0337F8E3" box="[532,621,1792,1817]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="467" unit="mm" value="6.9">6.9 mm</quantity>
; no defined ScA; ScP anteriorly pectinate, ending on RA, with branches simple or forked and more or less sigmoidal; area between costal margin and ScP
<quantity id="4CC320D28622FF88FBFAF8C90527F8C7" box="[1064,1149,1828,1852]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="467" unit="mm" value="0.8">0.8 mm</quantity>
wide; base of
<collectingCountry id="F32CCDA78622FF88FACCF8C8041AF8C7" box="[1310,1344,1829,1852]" name="Poland" pageId="5" pageNumber="467">RP</collectingCountry>
<quantity id="4CC320D28622FF88FA95F8C904C7F8C7" box="[1351,1437,1828,1853]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="467" unit="mm" value="5.3">5.3 mm</quantity>
distal of wing base; ScP and R (+ M + CuA) basally separated; presence of a common stem R + (M + CuA); RA convex, simple; crossveins in area between RA and
<collectingCountry id="F32CCDA78622FF88FD38F8800256F87F" box="[746,780,1901,1924]" name="Poland" pageId="5" pageNumber="467">RP</collectingCountry>
not preserved, if present;
<collectingCountry id="F32CCDA78622FF88FBEAF8800500F87F" box="[1080,1114,1901,1924]" name="Poland" pageId="5" pageNumber="467">RP</collectingCountry>
rather concave; first fork of
<collectingCountry id="F32CCDA78622FF88FF45F87C01E3F853" box="[151,185,1937,1960]" name="Poland" pageId="5" pageNumber="467">RP</collectingCountry>
<quantity id="4CC320D28622FF88FF10F87D0043F853" box="[194,281,1936,1961]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="467" unit="mm" value="6.5">6.5 mm</quantity>
from its base,
<collectingCountry id="F32CCDA78622FF88FE16F87C00BCF853" box="[452,486,1937,1960]" name="Poland" pageId="5" pageNumber="467">RP</collectingCountry>
with seven branches on wing margin; first branch short before its fusion with MA; CuA separated from M in basal third of wing; M dichotomously ramified into neutral MA and concave MP, far,
<quantity id="4CC320D28622FF88FAABF859019FF80B" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="467" unit="mm" value="5.3">5.3 mm</quantity>
, from its base; MA distally fused with first branch of
<collectingCountry id="F32CCDA78622FF88FCFCF834020AF80B" box="[814,848,2009,2032]" name="Poland" pageId="5" pageNumber="467">RP</collectingCountry>
, simple; MP with two branches near posterior wing margin; CuA fused with CuPaα at base of M; CuA + CuPaα posteriorly pectinate, with its first posterior branch concave; CuPaα short basal of its fusion with CuA; CuPaβ present, simple; a series of short crossveins in area between CuA + CuPaα and CuPaβ; division of CuP into CuPa and CuPb
<quantity id="4CC320D28621FF8BFC08FF0D0568FF0C" box="[986,1074,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="468" unit="mm" value="2.8">2.8 mm</quantity>
from wing base; concave CuPb simple; area between CuPa and CuPb narrow; three anal veins ending on posterior wing margin, all simple and more convex than CuP.
</paragraph>
</subSubSection>
<subSubSection id="C321DEBC8621FF8BFF15FEA1009DFCA6" pageId="6" pageNumber="468" type="discussion">
<paragraph id="8B848D378621FF8BFF15FEA1009DFCA6" blockId="6.[151,1437,151,861]" pageId="6" pageNumber="468">
<emphasis id="B94F51258621FF8BFF15FEA1001FFE9E" bold="true" box="[199,325,332,357]" pageId="6" pageNumber="468">Discussion</emphasis>
. This forewing corresponds to that of a clade Panorthoptera for the same reasons as above (except for the ScP ending on RA). The veins CuPaβ and CuA + CuPaα not strongly approximate exclude affinities with the Caloneurodea.
<taxonomicName id="4C3BF6B48621FF8BFEB9FE790082FE56" box="[363,472,404,429]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFEB9FE790082FE56" box="[363,472,404,429]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27CEC5E8621FF8BFE30FE780313FE57" box="[482,585,405,429]" pageId="6" pageNumber="468" rank="genus">gen. nov.</taxonomicNameLabel>
shares with the
<taxonomicName id="4C3BF6B48621FF8BFCD3FE790220FE56" box="[769,890,404,429]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
the veins CuPaβ, CuPb and 1A simple, and MP divided into two small distal branches. Nevertheless
<taxonomicName id="4C3BF6B48621FF8BFD37FE5A0209FE2B" box="[741,851,439,464]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFD37FE5A0209FE2B" box="[741,851,439,464]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
differs from the
<taxonomicName id="4C3BF6B48621FF8BFBC2FE5A05F8FE2B" box="[1040,1186,439,464]" class="Insecta" family="Oedischiidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="family">Oedischiidae</taxonomicName>
in the basal fork of M far from point of separation between M and CuA. Also its fusion of MA with first posterior branch of
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is quite different from the stem of
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and MA strongly approximate or fused for a short distance of the
<taxonomicName id="4C3BF6B48621FF8BFAD1FE1204C3FDE3" box="[1283,1433,511,536]" class="Insecta" family="Oedischiidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="family">Oedischiidae</taxonomicName>
.
<taxonomicName id="4C3BF6B48621FF8BFF45FDC9005FFDC6" box="[151,261,548,573]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFF45FDC9005FFDC6" box="[151,261,548,573]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
differs from the
<taxonomicName id="4C3BF6B48621FF8BFE1EFDC90360FDC6" box="[460,570,548,573]" class="Insecta" family="Geraridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="family">Geraridae</taxonomicName>
in the three characters: RA without well defined branches, MA fused with first posterior branch of
<collectingCountry id="F32CCDA78621FF8BFE67FDA5008DFDA4" box="[437,471,584,607]" name="Poland" pageId="6" pageNumber="468">RP</collectingCountry>
instead of being fused or approximating the stem of
<collectingCountry id="F32CCDA78621FF8BFB9FFDA50535FDA4" box="[1101,1135,584,607]" name="Poland" pageId="6" pageNumber="468">RP</collectingCountry>
, and CuPb simple. Lastly
<emphasis id="B94F51258621FF8BFF45FD8100CAFD7E" box="[151,400,620,645]" italics="true" pageId="6" pageNumber="468">Heterologus duyiwuer</emphasis>
and
<emphasis id="B94F51258621FF8BFE15FD8103A9FD7E" box="[455,755,620,645]" italics="true" pageId="6" pageNumber="468">Heterologus langfordorum</emphasis>
share with
<taxonomicName id="4C3BF6B48621FF8BFCA4FD8102BEFD7E" box="[886,996,620,645]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFCA4FD8102BEFD7E" box="[886,996,620,645]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
a simple CuPaβ, and a long stem of M before its first fork; but both strongly differ from
<taxonomicName id="4C3BF6B48621FF8BFD15FD62026FFD53" box="[711,821,655,680]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFD15FD62026FFD53" box="[711,821,655,680]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
in the median and radial veins well separated and MP with numerous branches.
<taxonomicName id="4C3BF6B48621FF8BFE10FD590319FD36" box="[450,579,692,717]" class="Insecta" family="Tococladidae" genus="Bruaylogus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="6" pageNumber="465" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFE10FD590319FD36" box="[450,579,692,717]" italics="true" pageId="6" pageNumber="468">Bruaylogus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C3BF6B48621FF8BFD31FD59020BFD36" box="[739,849,692,717]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFD31FD59020BFD36" box="[739,849,692,717]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
in median and
<collectingCountry id="F32CCDA78621FF8BFBDFFD580575FD37" box="[1037,1071,693,716]" name="Poland" pageId="6" pageNumber="468">RP</collectingCountry>
veins well separated, MP with numerous branches, M with a long stem, plus the presence of only two anal veins ending on posterior wing margin instead of three in
<taxonomicName id="4C3BF6B48621FF8BFEBBFD11008DFCEE" box="[361,471,764,789]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFEBBFD11008DFCEE" box="[361,471,764,789]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3BF6B48621FF8BFE37FD110309FCEE" box="[485,595,764,789]" class="Insecta" family="Geraridae" genus="Aviologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geraroptera" pageId="6" pageNumber="468" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFE37FD110309FCEE" box="[485,595,764,789]" italics="true" pageId="6" pageNumber="468">Aviologus</emphasis>
</taxonomicName>
corresponds to a new genus but its family affinities remains undetermined since a new phylogenetic analysis is not performed, as for
<taxonomicName id="4C3BF6B48621FF8BFCF0FCF202F9FCC3" box="[802,931,799,824]" class="Insecta" family="Tococladidae" genus="Bruaylogus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="6" pageNumber="465" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F51258621FF8BFCF0FCF202F9FCC3" box="[802,931,799,824]" italics="true" pageId="6" pageNumber="468">Bruaylogus</emphasis>
</taxonomicName>
. It is even not sure if they belong to the same or to two separate families.
</paragraph>
</subSubSection>
</treatment>
</document>