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<document id="C85A35F280E7BCD5298653DC94F0F8D1" ID-DOI="10.5281/zenodo.14662132" ID-ISSN="1942-1354" ID-Zenodo-Dep="14662132" ID-ZooBank="5671A77B-2ECB-445F-8F61-246A9E442CDE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736984354913" checkinUser="felipe" docAuthor="Eya, Bryan K." docDate="2024" docId="038487F80244FFAFFF140CB4FB6CFBBD" docLanguage="en" docName="InsectaMundi.2024.1061.1-94.pdf" docOrigin="Insecta Mundi 2024 (61)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Lophaliamorpha Eya 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="30" masterDocId="FFBDFF80025AFFB0FFD40F39FFE9FFB1" masterDocTitle="Revision of trachyderines related to Sphaenothecus from North America with description of three new genera, and new species of Lophalia Casey, 1912, Mannophorus LeConte, 1854, and Ischnocnemis Thomson, 1864 (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="29" updateTime="1736985555110" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="901CCA37A8F61CEA2DDB127ED827BF26">Revision of trachyderines related to Sphaenothecus from North America with description of three new genera, and new species of Lophalia Casey, 1912, Mannophorus LeConte, 1854, and Ischnocnemis Thomson, 1864 (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)</mods:title>
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<mods:namePart id="3065BB0CE61717E9D5928F90E3F3AD87">Eya, Bryan K.</mods:namePart>
<mods:affiliation id="6509835DE891EA541237BB00DB41DCB7">California Academy of Sciences 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118 USA</mods:affiliation>
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<mods:title id="558D9594E8FD36920A1F221B1C670D4C">Insecta Mundi</mods:title>
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<mods:part id="BF71590F2F1E90F7E66F7B824E07FE2D">
<mods:date id="833B2986F1503BF3AEB188EC7F13E5FA">2024</mods:date>
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<mods:number id="2796DF4DC7E510D9F5F2328BB3E1C72B">2024-07-26</mods:number>
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<mods:number id="88C472F8EA90C4537347B03CC016EEF5">2024</mods:number>
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<mods:identifier id="9521BB9AA31A766462228CDBBA9825D8" type="ISSN">1942-1354</mods:identifier>
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<treatment id="038487F80244FFAFFF140CB4FB6CFBBD" ID-DOI="http://doi.org/10.5281/zenodo.14662157" ID-Zenodo-Dep="14662157" LSID="urn:lsid:plazi:treatment:038487F80244FFAFFF140CB4FB6CFBBD" httpUri="http://treatment.plazi.org/id/038487F80244FFAFFF140CB4FB6CFBBD" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29">
<subSubSection id="C33765650244FFAEFF140CB4FDB5FC1B" box="[192,604,909,938]" pageId="30" pageNumber="29" type="nomenclature">
<paragraph id="8B9236EE0244FFAEFF140CB4FDB5FC1B" blockId="30.[192,604,909,938]" box="[192,604,909,938]" pageId="30" pageNumber="29">
<heading id="D0DA81820244FFAEFF140CB4FDB5FC1B" box="[192,604,909,938]" fontSize="12" level="2" pageId="30" pageNumber="29" reason="2">
<emphasis id="B959EAFC0244FFAEFF140CB4FDB5FC1B" bold="true" box="[192,604,909,938]" pageId="30" pageNumber="29">
<taxonomicName id="4C2D4D6D0244FFAEFF140CB4FE22FC1B" authority="Eya" authorityName="Eya" authorityYear="2024" box="[192,459,909,938]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B959EAFC0244FFAEFF140CB4FE7DFC1B" bold="true" box="[192,404,909,938]" italics="true" pageId="30" pageNumber="29">Lophaliamorpha</emphasis>
Eya
</taxonomicName>
,
<taxonomicNameLabel id="A26A57870244FFAEFE030CB7FDB5FC1B" box="[471,604,910,938]" pageId="30" pageNumber="29" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33765650244FFAEFF140C80FCCAFB89" pageId="30" pageNumber="29" type="type_taxon">
<paragraph id="8B9236EE0244FFAEFF140C80FD0AFC65" blockId="30.[192,1442,953,1080]" box="[192,739,953,980]" pageId="30" pageNumber="29">
<emphasis id="B959EAFC0244FFAEFF140C80FEB0FC62" bold="true" box="[192,345,953,979]" pageId="30" pageNumber="29">
<typeStatus id="5496884C0244FFAEFF140C80FF10FC62" box="[192,249,953,979]" pageId="30" pageNumber="29">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C2D4D6D0244FFAEFE8B0C80FD0AFC65" ID-CoL="4YY4Q" authority="Bates, 1885" authorityName="Bates" authorityYear="1885" box="[351,739,953,980]" class="Insecta" family="Cerambycidae" genus="Sphenothecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="luteicollis">
<emphasis id="B959EAFC0244FFAEFE8B0C80FDB7FC62" box="[351,606,953,979]" italics="true" pageId="30" pageNumber="29">Sphenothecus luteicollis</emphasis>
<bibRefCitation id="EFBC4B1F0244FFAEFDB10C80FD0AFC65" author="Bates HW" box="[613,739,953,980]" pageId="30" pageNumber="29" refId="ref69673" refString="Bates HW. 1885. Supplement to Longicornia. p. 225 - 436 pls. XVII - XXIV. In: Bates HW, Sharp D. Biologia Centrali-Americana. Insecta, Coleoptera, Vol. 5 Porter RH.; London; xii + 526 p. + 26 pl." type="book" year="1885">Bates, 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B9236EE0244FFAEFED40CE2FCCAFB89" blockId="30.[192,1442,953,1080]" pageId="30" pageNumber="29">
A new genus
<taxonomicName id="4C2D4D6D0244FFAEFE490CE3FDBDFC45" box="[413,596,986,1012]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0244FFAEFE490CE3FDBDFC45" box="[413,596,986,1012]" italics="true" pageId="30" pageNumber="29">Lophaliamorpha</emphasis>
</taxonomicName>
is created to include
<taxonomicName id="4C2D4D6D0244FFAEFC9D0CE3FB62FC44" ID-CoL="3Q4TD" authority="Bates" authorityName="Bates" box="[841,1163,986,1013]" class="Insecta" family="Cerambycidae" genus="Ischnocnemis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="luteicollis">
<emphasis id="B959EAFC0244FFAEFC9D0CE3FBA0FC45" box="[841,1097,986,1012]" italics="true" pageId="30" pageNumber="29">Ischnocnemis luteicollis</emphasis>
Bates
</taxonomicName>
and
<taxonomicName id="4C2D4D6D0244FFAEFB130CE3FF14FBA6" ID-CoL="3VZWB" authority="Bates" authorityName="Bates" authorityYear="1892" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="cribricollis">
<emphasis id="B959EAFC0244FFAEFB130CE3FA4BFC45" box="[1223,1442,986,1012]" italics="true" pageId="30" pageNumber="29">Lophalia cribricollis</emphasis>
Bates
</taxonomicName>
, and the following new combinations are proposed:
<taxonomicName id="4C2D4D6D0244FFAEFCE90CC5FB1FFBA6" authority="(Bates, 1885)" authorityName="Eya" authorityYear="2024" baseAuthorityName="Chemsak and Noguera" baseAuthorityYear="1885" box="[829,1270,1020,1047]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="luteicollis" status="comb. nov.">
<emphasis id="B959EAFC0244FFAEFCE90CC5FBB6FBA7" box="[829,1119,1020,1046]" italics="true" pageId="30" pageNumber="29">Lophaliamorpha luteicollis</emphasis>
(
<bibRefCitation id="EFBC4B1F0244FFAEFBBA0CC5FB06FBA6" author="Bates HW" box="[1134,1263,1020,1047]" pageId="30" pageNumber="29" refId="ref69673" refString="Bates HW. 1885. Supplement to Longicornia. p. 225 - 436 pls. XVII - XXIV. In: Bates HW, Sharp D. Biologia Centrali-Americana. Insecta, Coleoptera, Vol. 5 Porter RH.; London; xii + 526 p. + 26 pl." type="book" year="1885">Bates, 1885</bibRefCitation>
)
</taxonomicName>
<emphasis id="B959EAFC0244FFAEFB2F0CC5FF19FB86" bold="true" pageId="30" pageNumber="29">
<taxonomicNameLabel id="A26A57870244FFAEFB2F0CC5FF19FB86" pageId="30" pageNumber="29" rank="species">new combination</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C2D4D6D0244FFAEFEF90B26FDA5FB89" ID-CoL="3VZWB" authority="(Bates 1892)" authorityName="Eya" authorityYear="2024" baseAuthorityName="Bates" baseAuthorityYear="1892" box="[301,588,1054,1080]" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="cribricollis" status="new combination">
<emphasis id="B959EAFC0244FFAEFEF90B26FE52FB89" box="[301,443,1054,1080]" italics="true" pageId="30" pageNumber="29">L. cribricollis</emphasis>
(
<bibRefCitation id="EFBC4B1F0244FFAEFE1D0B27FDADFB89" author="Bates HW" box="[457,580,1054,1080]" pageId="30" pageNumber="29" pagination="143 - 183" refId="ref69726" refString="Bates HW. 1892. Additions to the Longicornia of Mexico and Central America, with remarks on some of the previously recorded species. Transactions of the Entomological Society of London 2: 143 - 183, 3 pl." type="journal article" year="1892">Bates 1892</bibRefCitation>
)
</taxonomicName>
<emphasis id="B959EAFC0244FFAEFD860B27FCF4FB86" bold="true" box="[594,797,1053,1080]" pageId="30" pageNumber="29">
<taxonomicNameLabel id="A26A57870244FFAEFD860B27FCF4FB86" box="[594,797,1053,1080]" pageId="30" pageNumber="29" rank="species">new combination</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C33765650244FFAEFF140B7EFACDF90F" pageId="30" pageNumber="29" type="description">
<paragraph id="8B9236EE0244FFAEFF140B7EFACDF90F" blockId="30.[190,1442,1095,1727]" pageId="30" pageNumber="29">
<emphasis id="B959EAFC0244FFAEFF140B7EFE7EFBD0" bold="true" box="[192,407,1095,1121]" pageId="30" pageNumber="29">Description. Form</emphasis>
moderate sized, sides subparallel.
<emphasis id="B959EAFC0244FFAEFCD90B7EFCA2FBD0" bold="true" box="[781,843,1095,1121]" pageId="30" pageNumber="29">Head</emphasis>
with front short, declivous in middle, mid-cranial sulcus (or median line) deeply canaliculate or well defined, extending from postclypeus towards vertex past antennal tubercles, each side above postclypeus below antennal insertion with a deep pit, frontoclypeal sulcus transversely excavated between pits; genae very short, anterior margins of lower eye lobes almost contiguous with base of mandibles; palpi short, subequal, last segments not expanded, outer edges of maxillary palpi impressed, apices truncate; mandibles with sides strongly angulated near base then arcuate towards apices, outer edges vaguely impressed and striate-punctate; apices simple, narrowly rounded or truncate; eyes moderately large, finely faceted, upper lobes small, well separated, lower lobes large; antennal tubercles divergent, moderately elevated above vertex, apices angulate; integument between tubercles deeply impressed; antennae elongate, slender, 11-segmented, 11
<superScript id="7C589BA60244FFAEFF0F0A4CFF00FA32" attach="left" box="[219,233,1397,1411]" fontSize="6" pageId="30" pageNumber="29">th</superScript>
antennomere vaguely to non-appendiculate.
<emphasis id="B959EAFC0244FFAEFD080A4FFCBDFA21" bold="true" box="[732,852,1398,1424]" pageId="30" pageNumber="29">Pronotum</emphasis>
hexagonal, slightly broader than long, narrower than base of elytra at humeri, sides obtusely tuberculate to angulate; disc arcuately impressed above lateral tubercles and basal two-thirds distinctly flattened or impressed in middle; prosternum with intercoxal process level with coxae, narrower than coxal cavity, occasionally impressed medially, apex vertically concave, coxal cavities wide open behind; mesosternum with intercoxal process prominent, protuberant above coxae, vertical or abruptly declivous and excavated anteriorly; posterior-lateral margins vaguely overlapping coxae.
<emphasis id="B959EAFC0244FFAEFB7B0924FACCF986" bold="true" box="[1199,1317,1565,1591]" pageId="30" pageNumber="29">Scutellum</emphasis>
triangular, glabrous, apex attenuating to a narrow point.
<emphasis id="B959EAFC0244FFAEFD650906FD10F9E8" bold="true" box="[689,761,1599,1625]" pageId="30" pageNumber="29">Elytra</emphasis>
each with three, distinct, longitudinal, glabrous costae; apices obliquely truncate, serrate with sutural angles minutely dentate, exterior angles obtusely angulate to rounded or minutely dentate.
<emphasis id="B959EAFC0244FFAEFE5C09BAFE54F92C" bold="true" box="[392,445,1667,1693]" pageId="30" pageNumber="29">Legs</emphasis>
moderately short; metafemora falling short of elytral apices in both sexes; metatarsomere I subequal to or slightly longer than tarsomeres II and III combined.
<emphasis id="B959EAFC0244FFAEFC6A099DFBDBF90F" bold="true" box="[958,1074,1700,1726]" pageId="30" pageNumber="29">Abdomen</emphasis>
normally segmented.
</paragraph>
</subSubSection>
<subSubSection id="C33765650244FFAFFF1409F4FB7CFC11" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29" type="discussion">
<paragraph id="8B9236EE0244FFAFFF1409F4FBF2FE78" blockId="30.[192,1442,1741,1903]" lastBlockId="31.[190,1442,230,929]" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29">
<emphasis id="B959EAFC0244FFAEFF1409F4FEAEF956" bold="true" box="[192,327,1741,1767]" pageId="30" pageNumber="29">Discussion.</emphasis>
<taxonomicName id="4C2D4D6D0244FFAEFE8509F7FDD0F959" authority="Eya" authorityName="Eya" authorityYear="2024" box="[337,569,1742,1768]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B959EAFC0244FFAEFE8509F7FDE1F959" box="[337,520,1742,1768]" italics="true" pageId="30" pageNumber="29">Lophaliamorpha</emphasis>
Eya
</taxonomicName>
<emphasis id="B959EAFC0244FFAEFD9009F7FD28F959" bold="true" box="[580,705,1742,1768]" pageId="30" pageNumber="29">
<taxonomicNameLabel id="A26A57870244FFAEFD9009F7FD28F959" box="[580,705,1742,1768]" pageId="30" pageNumber="29" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
is characterized as follows: (1) head with front short, declivous in middle (
<figureCitation id="13162A6B0244FFAEFE9409D6FE76F8B8" box="[320,415,1775,1801]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="30" pageNumber="29">Fig. 121</figureCitation>
,
<figureCitation id="13162A6B0244FFAEFE7909D6FE31F8BB" box="[429,472,1775,1802]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="30" pageNumber="29">135</figureCitation>
); (2) mid-cranial sulcus (median line) deeply canaliculate or well defined, extending from postclypeus to vertex past the antennal tubercles (
<figureCitation id="13162A6B0244FFAEFCEA0828FC74F89A" box="[830,925,1809,1835]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="30" pageNumber="29">Fig. 119</figureCitation>
); (5) mandibles with sides strongly angulated near base (
<figureCitation id="13162A6B0244FFAEFEE8080BFE73F8FC" box="[316,410,1842,1869]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="30" pageNumber="29">Fig. 121</figureCitation>
,
<figureCitation id="13162A6B0244FFAEFE73080BFE3BF8FC" box="[423,466,1842,1869]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="30" pageNumber="29">135</figureCitation>
); (3) genae narrow, lower eye lobes almost contiguous with mandibles (
<figureCitation id="13162A6B0244FFAEFB2B080BFAB4F8FC" box="[1279,1373,1842,1869]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="30" pageNumber="29">Fig. 120</figureCitation>
,
<figureCitation id="13162A6B0244FFAEFABD080AFA7CF8FC" box="[1385,1429,1843,1869]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="30" pageNumber="29">134</figureCitation>
); (5) antennal tubercles moderately elevated, divergent, and the integument between tubercles deeply impressed forming a V-shaped valley sloping down to mid-cranial sulcus (
<figureCitation id="13162A6B0245FFAFFCAF0FDFFC3FFEB1" box="[891,982,230,256]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 121</figureCitation>
,
<figureCitation id="13162A6B0245FFAFFC340FDFFBE2FEB0" box="[992,1035,230,257]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="31" pageNumber="30">135</figureCitation>
); (6) pronotum hexagonal with sides impressed above lateral tubercles (
<figureCitation id="13162A6B0245FFAFFDE80E3EFD71FE90" box="[572,664,263,289]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 122</figureCitation>
,
<figureCitation id="13162A6B0245FFAFFD770E3EFD24FE90" box="[675,717,263,289]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="31" pageNumber="30">136</figureCitation>
) and basal two-thirds distinctly flattened or impressed in middle (
<figureCitation id="13162A6B0245FFAFFF1C0E10FECDFEF2" box="[200,292,297,323]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 123</figureCitation>
,
<figureCitation id="13162A6B0245FFAFFEFB0E10FEB2FEF2" box="[303,347,297,323]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="31" pageNumber="30">137</figureCitation>
); (7) mesosternal intercoxal process that is protuberant above coxae, vertical or abruptly declivous anteriorly (
<figureCitation id="13162A6B0245FFAFFEE80E73FE70FED4" box="[316,409,330,357]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 123</figureCitation>
,
<figureCitation id="13162A6B0245FFAFFE700E72FE39FED4" box="[420,464,331,357]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="31" pageNumber="30">137</figureCitation>
); (8) each elytron with three, distinct, longitudinal, glabrous costae (
<figureCitation id="13162A6B0245FFAFFB190E73FAC0FED4" box="[1229,1321,330,357]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 124</figureCitation>
,
<figureCitation id="13162A6B0245FFAFFAE10E72FAB6FED4" box="[1333,1375,331,357]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="31" pageNumber="30">138</figureCitation>
) with apex truncate to obliquely truncate with exterior angle obtusely angulate (
<figureCitation id="13162A6B0245FFAFFBD40E55FBB5FE37" box="[1024,1116,364,390]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 126</figureCitation>
,
<figureCitation id="13162A6B0245FFAFFBBD0E55FB7CFE37" box="[1129,1173,364,390]" captionStart="Figures 129138" captionStartId="34.[195,274,1689,1713]" captionTargetBox="[198,1442,979,1659]" captionTargetId="figure-266@34.[195,1443,964,1659]" captionTargetPageId="34" captionText="Figures 129138. Lophaliamorpha cribricollis (Bates) comb. nov., dorsal and lateral images: 129130) Male, 17 mm, Guerrero, MEX. 131132) Female, 17 mm, Guerrero, MEX. Head: 133) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 134) Lateral profile with gena narrow (green arrow). 135) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), sides of mandibles angulate (blue arrow). Pronotum: 136) Disc hexagonal, sides arcuately impressed above lateral tubercles (red arrow). Thorax lateral image: 137) Basal 2/ rd of pronotal disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), 3 submarginal costa of each elytron starting below humerus (green arrow). Elytra:138) Each elytron with three distinct raised costae (green arrows), elytral apices obliquely truncate, exterior angle obtusely angulate (red arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662183" httpUri="https://zenodo.org/record/14662183/files/figure.png" pageId="31" pageNumber="30">138</figureCitation>
), minutely dentate (
<figureCitation id="13162A6B0245FFAFFAA10E55FF02FE18" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 125</figureCitation>
) or rounded; (8) scutellum triangular with apex attenuating to a narrow point; and (10) metatarsomere I subequal to or slightly longer than tarsomeres II and III combined (
<figureCitation id="13162A6B0245FFAFFC640E96FBE7FE78" box="[944,1038,431,457]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 127</figureCitation>
).
</paragraph>
<paragraph id="8B9236EE0245FFAFFED40EE8FB7CFC11" blockId="31.[190,1442,230,929]" pageId="31" pageNumber="30">
<taxonomicName id="4C2D4D6D0245FFAFFED40EE8FE5EFE5A" box="[256,439,465,491]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFED40EE8FE5EFE5A" box="[256,439,465,491]" italics="true" pageId="31" pageNumber="30">Lophaliamorpha</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C2D4D6D0245FFAFFD680EE8FC8FFE5A" authority="Casey" authorityName="Casey. Similarity" authorityYear="1912" box="[700,870,465,491]" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFD680EE8FCF4FE5A" box="[700,797,465,491]" italics="true" pageId="31" pageNumber="30">Lophalia</emphasis>
Casey
</taxonomicName>
by the three, distinct, longitudinal, glabrous costae on each elytron, hexagonal pronotum that is cribriform or alveolate punctate with basal two-thirds distinctly flattened or impressed in middle, and obliquely truncate elytral apices with exterior angles usually angulate and unarmed. The front is declivous in middle and the mid-cranial sulcus is deeply canaliculate from the frontoclypeal sulcus on to vertex. The mandibles are strongly angulated at sides, and genae narrow with anterior margins of lower eyes almost contiguous with the base of mandibles. The antennal tubercles are moderately elevated and divergent with the integument between tubercles deeply impressed.
<taxonomicName id="4C2D4D6D0245FFAFFC6A0DA3FBF6FD05" authorityName="Casey. Similarity" authorityYear="1912" box="[958,1055,666,692]" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFC6A0DA3FBF6FD05" box="[958,1055,666,692]" italics="true" pageId="31" pageNumber="30">Lophalia</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C2D4D6D0245FFAFFA870DA3FEDBFD67" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFA870DA3FEDBFD67" italics="true" pageId="31" pageNumber="30">Lophaliamorpha</emphasis>
</taxonomicName>
by the subparallel form, dark polished integument, broadly rounded sides of pronotum, and strongly dentate exterior angle of elytral apices. The front of
<taxonomicName id="4C2D4D6D0245FFAFFD2A0DE7FCB6FD49" authorityName="Casey. Similarity" authorityYear="1912" box="[766,863,734,760]" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFD2A0DE7FCB6FD49" box="[766,863,734,760]" italics="true" pageId="31" pageNumber="30">Lophalia</emphasis>
</taxonomicName>
is convex and subvertical with integument between antennal tubercles is not or barely impressed.
<taxonomicName id="4C2D4D6D0245FFAFFD150DC6FCCBFCA8" authorityName="Casey. Similarity" authorityYear="1912" box="[705,802,767,793]" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFD150DC6FCCBFCA8" box="[705,802,767,793]" italics="true" pageId="31" pageNumber="30">Lophalia</emphasis>
</taxonomicName>
has mandibles that are arcuate at sides, and the mid-cranial sulcus is shallow or vague. The metatarsomeres are more elongate in
<taxonomicName id="4C2D4D6D0245FFAFFC310C18FB75FC8A" box="[997,1180,801,827]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFC310C18FB75FC8A" box="[997,1180,801,827]" italics="true" pageId="31" pageNumber="30">Lophaliamorpha</emphasis>
</taxonomicName>
(
<figureCitation id="13162A6B0245FFAFFB7F0C18FAEFFC8A" box="[1195,1286,801,827]" captionStart="Figures 109127" captionStartId="32.[191,270,1287,1311]" captionTargetBox="[195,1432,244,1255]" captionTargetId="figure-122@32.[192,1440,229,1257]" captionTargetPageId="32" captionText="Figures 109127. Lophaliamorpha luteicollis (Bates) comb. nov., dorsal and lateral images: 109110) Male, 19 mm, Jalisco, MEX.111112) Female, 20 mm, Jalisco, MEX.113114) Male,19 mm, Jalisco, MEX.115116) Female, 20 mm, Jalisco, MEX. 117118) Male, 19 mm, Jalisco, MEX. Head: 119) Front short, mid-cranial sulcus (red arrow) deeply canaliculate extending from frontoclypeal sulcus (green arrow) to vertex. 120) Lateral profile with gena narrow (green arrow). 121) Frontolateral profile with antennal tubercles elevated, divergent (red arrows), side of mandibles angulate (blue arrow). Pronotum: 122) Disc hexagonal, sides arcuately impressed (red arrow) above lateral tubercles. Thorax lateral image:123) Basal 2/ rd of pronotal 3 disc impressed in middle (red arrow), mesosternal intercoxal process protuberant (blue arrow), submarginal costa of each elytron starting below humerus (green arrow). Elytra:124) Each elytron with three distinct costae (blue arrows). 125) Apices obliquely truncate, exterior angle minutely dentate (red arrow). 126) Apices with exterior angle obtusely angulate (red arrow). Metatarsi: 127) Metatarsomere 1 (T1) subequal (green brackets) to or longer than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.10. Comparison of metatarsi to Lophalia: 128) Metatarsi of Lophalia cyanicollis, male, metatarsomere 1 (T1) subequal (green brackets) to or shorter than tarsomeres 2+3 (T2+3), T1/ T2+3: 0.92." figureDoi="http://doi.org/10.5281/zenodo.14662181" httpUri="https://zenodo.org/record/14662181/files/figure.png" pageId="31" pageNumber="30">Fig. 127</figureCitation>
) compared to
<taxonomicName id="4C2D4D6D0245FFAFFF6B0C7BFEC9FCED" authorityName="Casey. Similarity" authorityYear="1912" box="[191,288,834,860]" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFF6B0C7BFEC9FCED" box="[191,288,834,860]" italics="true" pageId="31" pageNumber="30">Lophalia</emphasis>
</taxonomicName>
(Fig. 128).
<taxonomicName id="4C2D4D6D0245FFAFFE490C7BFDBDFCED" box="[413,596,834,860]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFE490C7BFDBDFCED" box="[413,596,834,860]" italics="true" pageId="31" pageNumber="30">Lophaliamorpha</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C2D4D6D0245FFAFFCAA0C7BFC09FCED" authorityName="Casey" authorityYear="1912" box="[894,992,834,860]" class="Insecta" family="Cerambycidae" genus="Zalophia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFCAA0C7BFC09FCED" box="[894,992,834,860]" italics="true" pageId="31" pageNumber="30">Zalophia</emphasis>
</taxonomicName>
by the elongate and slender basal antennomeres (
<figureCitation id="13162A6B0245FFAFFEFA0C5DFE94FCCF" box="[302,381,868,894]" captionStart="Figures 6568" captionStartId="21.[192,271,1089,1113]" captionTargetBox="[193,1440,229,1059]" captionTargetId="figure-315@21.[192,1440,229,1059]" captionTargetPageId="21" captionText="Figures 6568. Antennae of Sphaenothecus-like trachyderines with protuberant mesosternal process. 65) Zalophia funebris (Bates) male, GTM. 66) Zalophia auricomis (Chemsak and Linsley, 1979) comb. nov., male, El Taladro, HND. 67) Lophalia cyanicollis (Dupont), male, Tamaulipas, MEX 68) Lophaliamorpha luteicollis (Bates, 1885) comb. nov., male, Jalisco, MEX. 6566) Antennomeres IV short, stout, densely clothed with suberect setae. 6768) Antennomeres IV slender, elongate, clothed with minute, dark, setae." figureDoi="http://doi.org/10.5281/zenodo.14662169" httpUri="https://zenodo.org/record/14662169/files/figure.png" pageId="31" pageNumber="30">Fig. 68</figureCitation>
) and cribriform or alveolate punctate pronotum that are not obscured by pubescence.
<taxonomicName id="4C2D4D6D0245FFAFFAEA0C5DFA49FCCF" authorityName="Casey" authorityYear="1912" box="[1342,1440,868,894]" class="Insecta" family="Cerambycidae" genus="Zalophia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFAEA0C5DFA49FCCF" box="[1342,1440,868,894]" italics="true" pageId="31" pageNumber="30">Zalophia</emphasis>
</taxonomicName>
has short and stout basal antennomeres (
<figureCitation id="13162A6B0245FFAFFD520CBFFD3CFC10" box="[646,725,902,929]" captionStart="Figures 6568" captionStartId="21.[192,271,1089,1113]" captionTargetBox="[193,1440,229,1059]" captionTargetId="figure-315@21.[192,1440,229,1059]" captionTargetPageId="21" captionText="Figures 6568. Antennae of Sphaenothecus-like trachyderines with protuberant mesosternal process. 65) Zalophia funebris (Bates) male, GTM. 66) Zalophia auricomis (Chemsak and Linsley, 1979) comb. nov., male, El Taladro, HND. 67) Lophalia cyanicollis (Dupont), male, Tamaulipas, MEX 68) Lophaliamorpha luteicollis (Bates, 1885) comb. nov., male, Jalisco, MEX. 6566) Antennomeres IV short, stout, densely clothed with suberect setae. 6768) Antennomeres IV slender, elongate, clothed with minute, dark, setae." figureDoi="http://doi.org/10.5281/zenodo.14662169" httpUri="https://zenodo.org/record/14662169/files/figure.png" pageId="31" pageNumber="30">Fig. 65</figureCitation>
) and pronotum that are densely hirsute.
</paragraph>
</subSubSection>
<subSubSection id="C33765650245FFAFFF140C96FB6CFBBD" pageId="31" pageNumber="30" type="etymology">
<paragraph id="8B9236EE0245FFAFFF140C96FB6CFBBD" blockId="31.[192,1441,943,1037]" pageId="31" pageNumber="30">
<emphasis id="B959EAFC0245FFAFFF140C96FEADFC78" bold="true" box="[192,324,943,969]" pageId="31" pageNumber="30">Etymology.</emphasis>
The etymology of the name
<taxonomicName id="4C2D4D6D0245FFAFFDA90C96FCDDFC78" box="[637,820,943,969]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFDA90C96FCDDFC78" box="[637,820,943,969]" italics="true" pageId="31" pageNumber="30">Lophaliamorpha</emphasis>
</taxonomicName>
refers to the shape or appearance that resembles
<taxonomicName id="4C2D4D6D0245FFAFFA850C96FF35FC5A" authorityName="Casey. Similarity" authorityYear="1912" class="Insecta" family="Cerambycidae" genus="Lophalia" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFA850C96FF35FC5A" italics="true" pageId="31" pageNumber="30">Lophalia</emphasis>
</taxonomicName>
. The suffix “
<emphasis id="B959EAFC0245FFAFFEA40CE8FE39FC5A" box="[368,464,977,1003]" italics="true" pageId="31" pageNumber="30">-morpha</emphasis>
” is Latin from “-
<emphasis id="B959EAFC0245FFAFFD4E0CE8FD12FC5A" box="[666,763,977,1003]" italics="true" pageId="31" pageNumber="30">morphus</emphasis>
” or “shaped” from Ancient Greek μορφή (
<emphasis id="B959EAFC0245FFAFFB3F0CE8FAA9FC5C" box="[1259,1344,977,1005]" italics="true" pageId="31" pageNumber="30">morphḗ</emphasis>
, “shape, appearance”).
<taxonomicName id="4C2D4D6D0245FFAFFE8B0CCBFDFFFBBD" box="[351,534,1010,1036]" class="Insecta" family="Cerambycidae" genus="Lophaliamorpha" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC0245FFAFFE8B0CCBFDFFFBBD" box="[351,534,1010,1036]" italics="true" pageId="31" pageNumber="30">Lophaliamorpha</emphasis>
</taxonomicName>
is a genus-group name with Latinized feminine ending.
</paragraph>
</subSubSection>
</treatment>
</document>