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<document id="E3AFFB136AE1FEC218665BB9D1BA0232" ID-DOI="10.5281/zenodo.14662132" ID-ISSN="1942-1354" ID-Zenodo-Dep="14662132" ID-ZooBank="5671A77B-2ECB-445F-8F61-246A9E442CDE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736984354913" checkinUser="felipe" docAuthor="Eya, Bryan K." docDate="2024" docId="038487F8020FFFE5FF140C18FA49FB4B" docLanguage="en" docName="InsectaMundi.2024.1061.1-94.pdf" docOrigin="Insecta Mundi 2024 (61)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Cyphosterna Chevrolat 1862" docType="key" docVersion="2" lastPageNumber="84" masterDocId="FFBDFF80025AFFB0FFD40F39FFE9FFB1" masterDocTitle="Revision of trachyderines related to Sphaenothecus from North America with description of three new genera, and new species of Lophalia Casey, 1912, Mannophorus LeConte, 1854, and Ischnocnemis Thomson, 1864 (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="84" updateTime="1736985555110" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="AAB51DD35C851087F88FF8361EF4C932">Revision of trachyderines related to Sphaenothecus from North America with description of three new genera, and new species of Lophalia Casey, 1912, Mannophorus LeConte, 1854, and Ischnocnemis Thomson, 1864 (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini)</mods:title>
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<mods:namePart id="DB6A9A46E425BBDF80A78B25EB3378E6">Eya, Bryan K.</mods:namePart>
<mods:affiliation id="B7C7C87E611E673A9420364FF1A4DEE9">California Academy of Sciences 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118 USA</mods:affiliation>
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<mods:title id="B54E7448AFD0ECC12D5DF19DB50C46BA">Insecta Mundi</mods:title>
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<mods:date id="8ADFF8E51D86439DA3BA47C75BAD326F">2024</mods:date>
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<treatment id="038487F8020FFFE5FF140C18FA49FB4B" LSID="urn:lsid:plazi:treatment:038487F8020FFFE5FF140C18FA49FB4B" httpUri="http://treatment.plazi.org/id/038487F8020FFFE5FF140C18FA49FB4B" lastPageNumber="84" pageId="85" pageNumber="84">
<subSubSection id="C3376565020FFFE5FF140C18FC62FC8C" box="[192,907,800,829]" pageId="85" pageNumber="84" type="nomenclature">
<paragraph id="8B9236EE020FFFE5FF140C18FC62FC8C" blockId="85.[192,907,800,829]" box="[192,907,800,829]" pageId="85" pageNumber="84">
<heading id="D0DA8182020FFFE5FF140C18FC62FC8C" box="[192,907,800,829]" fontSize="12" level="2" pageId="85" pageNumber="84" reason="2">
<emphasis id="B959EAFC020FFFE5FF140C18FC62FC8C" bold="true" box="[192,907,800,829]" pageId="85" pageNumber="84">
Key to species of the genus
<taxonomicName id="4C2D4D6D020FFFE5FDCE0C19FC62FC8C" ID-CoL="3YRT" authority="Chevrolat, 1862" authorityName="Chevrolat" authorityYear="1862" box="[538,907,800,829]" class="Insecta" family="Cerambycidae" genus="Cyphosterna" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="B959EAFC020FFFE5FDCE0C19FD50FC8C" bold="true" box="[538,697,800,829]" italics="true" pageId="85" pageNumber="84">Cyphosterna</emphasis>
<bibRefCitation id="EFBC4B1F020FFFE5FD140C19FC62FC8C" author="Chevrolat LAA" box="[704,907,800,829]" pageId="85" pageNumber="84" pagination="747 - 763" refId="ref70371" refString="Chevrolat LAA. 1862. Revision des genres Eriphus et Mallosoma Serville, du groupe des Clytides, et description de trois nouveaux genres dont un doit etre rapporte au groupe des Callidiites. Annales de la Societe Entomologique de France (4) 2: 747 - 763." type="journal article" year="1862">Chevrolat, 1862</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3376565020FFFE5FF140C76FA49FB4B" pageId="85" pageNumber="84" type="key">
<keyStep id="30D92E4B020FFFE5FF140C76FA49FC69" pageId="85" pageNumber="84">
<paragraph id="8B9236EE020FFFE5FF140C76FA49FCD8" blockId="85.[192,1440,847,873]" box="[192,1440,847,873]" pageId="85" pageNumber="84">
<keyLead id="30DC95DB020FFFE5FF140C76FA49FCD8" box="[192,1440,847,873]" pageId="85" pageNumber="84">
1. Integument black, each elytron with prominent submarginal and subsutural eburneous costae......
<emphasis id="B959EAFC020FFFE5FA460C76FA49FCD8" bold="true" box="[1426,1440,847,873]" pageId="85" pageNumber="84">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9236EE020FFFE5FF140C45FA49FC69" blockId="85.[192,1441,891,985]" pageId="85" pageNumber="84">
<keyLead id="30DC95DB020FFFE5FF140C45FA49FC69" pageId="85" pageNumber="84">
— Integument black with dull greenish purple overtone, elytra black, without eburneous costae, each elytron with four prominent costae, submarginal costa joined to a shorter lateral one near apex (
<figureCitation id="13162A6B020FFFE5FAA10CA4FDF6FC69" captionStart="Figures 489499" captionStartId="91.[192,271,1212,1236]" captionTargetBox="[194,1440,240,1182]" captionTargetId="figure-104@91.[192,1440,229,1185]" captionTargetPageId="91" captionText="Figures 489499. Cyphosterna atropurpureum (Chemsak and Noguera) comb. nov., dorsal and lateral images: 489490) Male, 20.5 mm, La Paz, HND. Head: 491493) Front vertical, subquadrate with a deep pit on each side above postclypeus (white arrow), genae subquadrate, prominent (red arrow), scape carinate (yellow arrow). Pronotum: 494) Disc moderately inflated, anterior half carinate in middle with callus on each side (yellow arrows), posterior half with an opaque, impunctate callus in middle and obliquely, elongate, coarsely punctate callus on each side (white arrows), sides tuberculate (red arrow) and laterally excavated above (green arrow). Thorax lateral image: 495) Pronotal disc with base behind lateral and median calli narrowly impressed (purple arrow), mesosternal process protuberant (blue arrow).Metatarsi:496) Metatarsomere 1 (T1) subequal (red brackets) to tarsomeres 2+3 (T2+3), T1/ T2+3: 1.0. Elytra dorsal and lateral images: 497) Apices obliquely sinuate truncate with outer angles obtusely dentate. 498499) Each elytron with four prominent costae, a subsutural costa extending from base almost to apex (red arrow), submarginal costa starting behind humerus attaining apical 1/6 (blue arrow) with lateral branching costa starting on inside at apical 1/5 reaching basal ¼ (green arrow), and another costa between subsutural one and submarginal branch starting from basal 1/5 attaining apical 1/6 (yellow arrow)." figureDoi="http://doi.org/10.5281/zenodo.14662252" httpUri="https://zenodo.org/record/14662252/files/figure.png" pageId="85" pageNumber="84">Fig. 489490, 498499</figureCitation>
)..................
<emphasis id="B959EAFC020FFFE5FCCB0C87FA49FC69" bold="true" box="[799,1440,958,985]" pageId="85" pageNumber="84">
<taxonomicName id="4C2D4D6D020FFFE5FCCB0C87FAFEFC69" authority="(Chemsak and Noguera)" authorityName="Eya" authorityYear="2024" baseAuthorityName=", Chemsak and Noguera" baseAuthorityYear="2024" box="[799,1303,958,985]" class="Insecta" family="Cerambycidae" genus="Cyphosterna" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="atropurpureum" status="comb. nov.">
<emphasis id="B959EAFC020FFFE5FCCB0C87FC18FC68" bold="true" box="[799,1009,958,985]" italics="true" pageId="85" pageNumber="84">C. atropurpureum</emphasis>
(Chemsak and Noguera)
</taxonomicName>
,
<taxonomicNameLabel id="A26A5787020FFFE5FAF60C87FA49FC69" box="[1314,1440,958,984]" pageId="85" pageNumber="84" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D92E4B020FFFE5FF140CD2FA49FB4B" pageId="85" pageNumber="84">
<paragraph id="8B9236EE020FFFE5FF140CD2FA49FBD8" blockId="85.[192,1440,1002,1130]" pageId="85" pageNumber="84">
<keyLead id="30DC95DB020FFFE5FF140CD2FA49FBD8" pageId="85" pageNumber="84">
2(1). Pronotum coarsely punctate with lateral margins of disc and ventral surface of body including prosternum clothed with dense, appressed golden pubescence; elytral apices tapered, narrowly rounded with minute sutural spine; pronotum of females black (
<figureCitation id="13162A6B020FFFE5FC670B17FBAEFBF9" box="[947,1095,1070,1096]" captionStart="Figures 478488" captionStartId="89.[192,271,1028,1052]" captionTargetBox="[195,1440,242,986]" captionTargetId="figure-312@89.[192,1440,229,998]" captionTargetPageId="89" captionText="Figures 478488. Cyphosterna sinaloanum (Chemsak and Linsley) comb. nov., dorsal and lateral images: 478479) Male, 22 mm, Chamela, Jalisco, MEX. 480481) Female, 18 mm, Chamela, Jalisco, MEX. Head:482484) Front subvertical, subquadrate, seamlessly fused with postclypeus, genae subquadrate, prominent (red arrow), scape carinate (yellow arrow). Pronotum: 485) Disc moderately inflated, excavated laterally (green arrow) and sides tuberculate (red arrow). Thorax lateral image:486) Pronotal disc with prominent dorsal callus, mesosternal process protuberant (blue arrow). Elytra: 487) Each elytron with prominent paired eburneous ridges, apices tapered, narrowly rounded (red arrows) with minute sutural spine (green arrow) and appressed silver pubescence (blue arrow). Metatarsi:488) Metatarsomeres narrow, elongate, tarsomere 1 (T1) longer (red brackets) than tarsomeres 2+3 (T2+3), T1/ T2+3: 1.2." figureDoi="http://doi.org/10.5281/zenodo.14662250" httpUri="https://zenodo.org/record/14662250/files/figure.png" pageId="85" pageNumber="84">Fig. 478488</figureCitation>
)..................................................................
<emphasis id="B959EAFC020FFFE5FC8E0B76FA49FBD8" bold="true" box="[858,1440,1103,1130]" pageId="85" pageNumber="84">
<taxonomicName id="4C2D4D6D020FFFE5FC8E0B76FAFFFBD8" authority="(Chemsak and Linsley)" authorityName="Eya" authorityYear="2024" baseAuthorityName="Chemsak and Linsley" baseAuthorityYear="2024" box="[858,1302,1103,1130]" class="Insecta" family="Cerambycidae" genus="Cyphosterna" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="sinaloanum" status="comb. nov.">
<emphasis id="B959EAFC020FFFE5FC8E0B76FBEBFBDB" bold="true" box="[858,1026,1103,1130]" italics="true" pageId="85" pageNumber="84">C. sinaloanum</emphasis>
(Chemsak and Linsley)
</taxonomicName>
,
<taxonomicNameLabel id="A26A5787020FFFE5FAF60B76FA49FBD8" box="[1314,1440,1103,1129]" pageId="85" pageNumber="84" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9236EE020FFFE5FF140B45FA49FB4B" blockId="85.[192,1442,1148,1275]" pageId="85" pageNumber="84">
<keyLead id="30DC95DB020FFFE5FF140B45FA49FB4B" pageId="85" pageNumber="84">
— Pronotum less coarsely punctate, disc lacking dense, appressed golden pubescent margin, prosternum thinly pubescent without dense appressed golden pubescence; elytral apices obliquely emarginate; pronotum of females reddish with apical and basal margins narrowly black (
<figureCitation id="13162A6B020FFFE5FB750B86FADDFB6B" box="[1185,1332,1215,1242]" captionStart="Figures 459477" captionStartId="86.[192,271,1591,1615]" captionTargetBox="[196,1440,236,1560]" captionTargetId="figure-13@86.[192,1440,229,1561]" captionTargetPageId="86" captionText="Figures 459477. Cyphosterna quadrilineatum Chevrolat, dorsal and lateral image: 459460) Male, 19 mm, 23 mi S Matias Romero, Oaxaca, MEX. 461462) Female, 22 mm, 17 km N Ariaga, Chiapas, MEX. 463) Female, ventral image. 464) C. quadrilineatum syntype, Yucatan, MEX. (Bezark 2024, id: 6392). Head: 465467) Front subvertical, subquadrate, seamlessly fused with postclypeus, genae subquadrate, prominent (red arrow). 467468) Scape carinate (red arrow). Pronotum: 469, 471) Disc laterally excavated (green arrow) and sides tuberculate (red arrow). Thorax lateral images: 470, 472) Pronotal disc inflated with prominent dorsal callus, mesosternal process protuberant (red arrow). Elytra: 473474) Each elytron with prominent paired eburneous ridges, apices emarginate (red arrows) with subsutural dentation (green arrow) and appressed silver pubescence (yellow arrow).475) Elytral apices of syntype (Bezark 2024, id: 6392). Metatarsi:476477) Metatarsomeres narrow, elongate, tarsomere 1 (T1) longer (red brackets) than tarsomeres 2+3 (T2+3). 476) Male, T1/ T2+3: 1.4. 477) Female, T1/ T2+3: 1.3." figureDoi="http://doi.org/10.5281/zenodo.14662248" httpUri="https://zenodo.org/record/14662248/files/figure.png" pageId="85" pageNumber="84">Fig. 459475</figureCitation>
)..................................................................
<taxonomicName id="4C2D4D6D020FFFE5FB850BD9FA49FB4B" authority="Chevrolat" authorityName="Chevrolat. Lateral" authorityYear="1862" box="[1105,1440,1248,1275]" class="Insecta" family="Cerambycidae" genus="Cyphosterna" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="84" phylum="Arthropoda" rank="species" species="quadrilineatum">
<emphasis id="B959EAFC020FFFE5FB850BD9FA49FB4B" bold="true" box="[1105,1440,1248,1275]" pageId="85" pageNumber="84">
<emphasis id="B959EAFC020FFFE5FB850BD9FACCFB4A" bold="true" box="[1105,1317,1248,1275]" italics="true" pageId="85" pageNumber="84">C. quadrilineatum</emphasis>
Chevrolat
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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