322 lines
34 KiB
XML
322 lines
34 KiB
XML
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<mods:title id="544A8FB3D3CCB088F848A4FD70C69922">Snailfishes of the genus Careproctus (Perciformes: Liparidae) with a reduced pelvic disk: three new species and new records from the western North Pacific with comments on their phenotypic diversity</mods:title>
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<mods:namePart id="3D90F2E45B1415EF03B65D81A88BCF10">Kai, Yoshiaki</mods:namePart>
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<mods:affiliation id="BE2ECD3EA477EA1B94349F4782C0DA4B">Maizuru Fisheries Research Station, Field Science Education and Research Center, Kyoto University, Nagahama, Maizuru, Kyoto 625 - 0086, Japan</mods:affiliation>
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<mods:nameIdentifier id="0BD5918F7E01ECDEFBBEABF3E7C00C35" type="email">kai.yoshiaki.4c@kyoto-u.ac.Jp</mods:nameIdentifier>
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<mods:namePart id="E404DF20C8E6BC8763461BE60F05C072">Matsuzaki, Koji</mods:namePart>
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<mods:affiliation id="10FBC5DB7B35F4A9E84C645A181C2D2F">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
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<mods:nameIdentifier id="7D4BF785198A7280156D855839F7449A" type="email">matsu@aquamarine.or.Jp</mods:nameIdentifier>
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<mods:namePart id="B8C5FF7A156B9F79D586AE10ABA24528">Mori, Toshiaki</mods:namePart>
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<mods:affiliation id="C280DDB7B2CA8C0CEB231821772F0938">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
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<mods:nameIdentifier id="0DFBC494E7BA2837DD01A837179FE9CE" type="email">mori.toshiaki@aquamarine.or.Jp</mods:nameIdentifier>
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<mods:namePart id="F8B42E5440DE3379F0E4175E826957CE">Pitruk, Dmitry L.</mods:namePart>
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<mods:nameIdentifier id="E3DDC9092A3446CCB121507699DCF72B" type="ORCID">0009-0003-4449-8453</mods:nameIdentifier>
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<mods:affiliation id="461574579225BC6922A3C2F03A1DBED2">A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch of the Russian Academy of Sciences, ul. Palchevskogo 17, Vladivostok 690041, Russia</mods:affiliation>
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<mods:nameIdentifier id="16DF91C784806B1FE12FD43A1A27FC6C" type="email">pitruk@mail.ru</mods:nameIdentifier>
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<mods:namePart id="162FE389A1BF8A64F5C4B288EFA76FA0">Misawa, Ryo</mods:namePart>
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<mods:affiliation id="4FA2B70A3DD4F11BE24FBEFAB2CAD7AB">Demersal Fish Resources Division, Fisheries Stock Assessment Center, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, National Research and Development Agency, 25 - 259 Shimomekurakubo, Same, Hachinohe, Aomori 031 - 0841, Japan</mods:affiliation>
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<mods:nameIdentifier id="1B5B9CF5780A69EE41D9495F40064D3C" type="email">batoideafish@gmail.com</mods:nameIdentifier>
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<mods:namePart id="ACB181D1C0929122B9E27B25DF922F32">Tashiro, Fumihito</mods:namePart>
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<mods:nameIdentifier id="9EA1872D2AC067E6A040C7F9AE6BE7DD" type="ORCID">0000-0003-0640-8456</mods:nameIdentifier>
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<mods:affiliation id="8E1FBEFB1859CDEADAF9EE5CC53C2784">Fisheries Science Center, The Hokkaido University Museum, 3 - 1 - 1 Minato-cho, Hakodate, Hokkaido 041 - 8611, Japan</mods:affiliation>
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<mods:nameIdentifier id="6A68FE310F7DC9F1EBF779D089D25E45" type="email">fumihito.tashiro@gmail.com</mods:nameIdentifier>
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<mods:title id="232A8D734C36FE2C7644166D893B6A98">Zootaxa</mods:title>
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<mods:part id="06B5CA12FF7C000E34EE3622BE729802">
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<mods:date id="8078C571C96863D28EE95681F3CA1AA2">2024</mods:date>
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<mods:number id="4A962F0842D4E6FC7A5D7EEF43AE61D0">2024-08-05</mods:number>
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<mods:number id="50395B644872DC99DF49C10FA07D2593">5492</mods:number>
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<mods:url id="A8B46C9437BF176FFECF6305335C7516">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
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<mods:classification id="339E685AFFDA0F486E285A4D64ABF30E">journal article</mods:classification>
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<mods:identifier id="F614BEB8E1B245A60A3D8BEBDCD19C01" type="DOI">10.11646/zootaxa.5492.2.3</mods:identifier>
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<treatment id="038487F4FF87386B8BCDAD76D5AAFE6D" LSID="urn:lsid:plazi:treatment:038487F4FF87386B8BCDAD76D5AAFE6D" httpUri="http://treatment.plazi.org/id/038487F4FF87386B8BCDAD76D5AAFE6D" lastPageId="3" lastPageNumber="194" pageId="2" pageNumber="193">
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<subSubSection id="C3376569FF87386A8BCDAD76D247FBE6" box="[151,624,1086,1113]" pageId="2" pageNumber="193" type="nomenclature">
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<paragraph id="8B9236E2FF87386A8BCDAD76D247FBE6" blockId="2.[151,624,1086,1113]" box="[151,624,1086,1113]" pageId="2" pageNumber="193">
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<heading id="D0DA818EFF87386A8BCDAD76D247FBE6" bold="true" box="[151,624,1086,1113]" fontSize="11" level="1" pageId="2" pageNumber="193" reason="1">
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<emphasis id="B959EAF0FF87386A8BCDAD76D247FBE6" bold="true" box="[151,624,1086,1113]" pageId="2" pageNumber="193">
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The
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<taxonomicName id="4C2D4D61FF87386A8B95AD76D1F6FBE6" ID-CoL="69BGL" authority="Burke, 1912" authorityName="Burke" authorityYear="1912" box="[207,449,1086,1113]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="gilberti">
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<emphasis id="B959EAF0FF87386A8B95AD76D1F6FBE6" bold="true" box="[207,449,1086,1113]" italics="true" pageId="2" pageNumber="193">Careproctus gilberti</emphasis>
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species group
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</paragraph>
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<subSubSection id="C3376569FF87386A8BCDADCFD12DFAB2" pageId="2" pageNumber="193" type="diagnosis">
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<paragraph id="8B9236E2FF87386A8BCDADCFD12DFAB2" blockId="2.[151,1437,1158,1725]" pageId="2" pageNumber="193">
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<emphasis id="B959EAF0FF87386A8BCDADCFD125FB1F" bold="true" box="[151,274,1158,1184]" pageId="2" pageNumber="193">Diagnosis.</emphasis>
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A species group of
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<taxonomicName id="4C2D4D61FF87386A895EADCED2B9FB1E" authorityName="Kroyer" authorityYear="1862" box="[516,654,1159,1185]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="genus">
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<emphasis id="B959EAF0FF87386A895EADCED2B9FB1E" box="[516,654,1159,1185]" italics="true" pageId="2" pageNumber="193">Careproctus</emphasis>
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</taxonomicName>
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with the following characters: reduced pelvic disk, less than 2.7% SL in length, usually round, sometimes incomplete (see each description of species); gill slit moderately large, extending ventrally to pectoral-fin rays 4–19, not reaching to base of lower pectoral-fin lobe; teeth simple or weakly shouldered.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376569FF87386A8B9DAC5FD36AFAEA" pageId="2" pageNumber="193" type="taxon_list">
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<paragraph id="8B9236E2FF87386A8B9DAC5FD36AFAEA" blockId="2.[151,1437,1158,1725]" pageId="2" pageNumber="193">
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<emphasis id="B959EAF0FF87386A8B9DAC5FD1A1FA8F" bold="true" box="[199,406,1302,1328]" pageId="2" pageNumber="193">Species included.</emphasis>
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In addition to
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<taxonomicName id="4C2D4D61FF87386A891EAC5ED28EFA8E" authorityName="Burke" authorityYear="1912" box="[580,697,1303,1329]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="gilberti">
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<emphasis id="B959EAF0FF87386A891EAC5ED28EFA8E" box="[580,697,1303,1329]" italics="true" pageId="2" pageNumber="193">C. gilberti</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2D4D61FF87386A8990AC5ED3E1FA8E" authorityName="Kai & Matsuzaki & Mori & Pitruk & Misawa & Tashiro" authorityYear="2024" box="[714,982,1303,1329]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
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<emphasis id="B959EAF0FF87386A8990AC5ED3E1FA8E" box="[714,982,1303,1329]" italics="true" pageId="2" pageNumber="193">Careproctus barbatulus</emphasis>
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</taxonomicName>
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<emphasis id="B959EAF0FF87386A88BAAC5ED40BFA8E" bold="true" box="[992,1084,1303,1329]" pageId="2" pageNumber="193">
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<taxonomicNameLabel id="A26A578BFF87386A88BAAC5ED40BFA8E" box="[992,1084,1303,1329]" pageId="2" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C2D4D61FF87386A8F17AC5ED53AFA8E" authorityName="Kai & Matsuzaki & Mori & Pitruk & Misawa & Tashiro" authorityYear="2024" box="[1101,1293,1303,1329]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="longibarbatus" status="sp. nov.">
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<emphasis id="B959EAF0FF87386A8F17AC5ED53AFA8E" box="[1101,1293,1303,1329]" italics="true" pageId="2" pageNumber="193">C. longibarbatus</emphasis>
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</taxonomicName>
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<emphasis id="B959EAF0FF87386A8E4CAC5ED545FA8E" bold="true" box="[1302,1394,1303,1329]" pageId="2" pageNumber="193">
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<taxonomicNameLabel id="A26A578BFF87386A8E4CAC5ED545FA8E" box="[1302,1394,1303,1329]" pageId="2" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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,
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<taxonomicName id="4C2D4D61FF87386A8ED9AC5ED0D2FAEA" authorityName="Schmidt" authorityYear="1916" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="mederi">
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<emphasis id="B959EAF0FF87386A8ED9AC5ED0D2FAEA" italics="true" pageId="2" pageNumber="193">C. mederi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2D4D61FF87386A8BA9AC72D14EFAEB" authorityName="Gilbert" authorityYear="1896" box="[243,377,1339,1365]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="ostentum">
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<emphasis id="B959EAF0FF87386A8BA9AC72D14EFAEB" box="[243,377,1339,1365]" italics="true" pageId="2" pageNumber="193">C. ostentum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C2D4D61FF87386A8ADDAC72D21FFAEA" authorityName="Imamura & Nobetsu" authorityYear="2002" box="[391,552,1339,1365]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="parvidiscus">
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<emphasis id="B959EAF0FF87386A8ADDAC72D21FFAEA" box="[391,552,1339,1365]" italics="true" pageId="2" pageNumber="193">C. parvidiscus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C2D4D61FF87386A893CAC72D2CAFAEA" authorityName="Kai & Matsuzaki & Mori & Pitruk & Misawa & Tashiro" authorityYear="2024" box="[614,765,1339,1365]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
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<emphasis id="B959EAF0FF87386A893CAC72D2CAFAEA" box="[614,765,1339,1365]" italics="true" pageId="2" pageNumber="193">C. spinulosus</emphasis>
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</taxonomicName>
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<emphasis id="B959EAF0FF87386A885EAC72D36AFAEA" bold="true" box="[772,861,1339,1365]" pageId="2" pageNumber="193">
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<taxonomicNameLabel id="A26A578BFF87386A885EAC72D36AFAEA" box="[772,861,1339,1365]" pageId="2" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3376569FF87386A8B9DAC16D240F902" pageId="2" pageNumber="193" type="discussion">
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<paragraph id="8B9236E2FF87386A8B9DAC16D240F902" blockId="2.[151,1437,1158,1725]" pageId="2" pageNumber="193">
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<emphasis id="B959EAF0FF87386A8B9DAC16D10CFAC6" bold="true" box="[199,315,1375,1401]" pageId="2" pageNumber="193">Remarks.</emphasis>
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Among other liparids, species of
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<taxonomicName id="4C2D4D61FF87386A89F4AC16D308FAC6" authorityName="Gilbert & Burke" authorityYear="1912" box="[686,831,1375,1401]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="genus">
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<emphasis id="B959EAF0FF87386A89F4AC16D308FAC6" box="[686,831,1375,1401]" italics="true" pageId="2" pageNumber="193">Elassodiscus</emphasis>
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</taxonomicName>
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, which are also characterized by a reduced pelvic disk, are most similar to the above species group but can be distinguished on the basis of having a black orobranchial cavity (vs. pale orobranchial cavity), a gill slit above the pectoral fin or extending ventrally to pectoral-fin rays 1–7, 4 proximal pectoral radials with no interradial fenestrae (vs.proximal pectoral radials 3–4 with interradial fenestrae), and no pleural ribs (vs. usually 1–3 pleural ribs) (
|
||
<bibRefCitation id="EFBC4B13FF87386A8819ACA6D3F7F9B6" author="Kido, K." box="[835,960,1519,1545]" pageId="2" pageNumber="193" pagination="125 - 256" refId="ref18070" refString="Kido, K. (1988) Phylogeny of the family Liparididae, with the taxonomy of the species found around Japan. Memories of the Faculty of Fisheries, Hokkaido University, 35, 125 - 256." type="journal article" year="1988">Kido 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBC4B13FF87386A8897ACA6D4E1F9B6" author="Pitruk, D. L. & Fedorov, V. V." box="[973,1238,1519,1545]" pageId="2" pageNumber="193" refId="ref18902" refString="Pitruk, D. L. & Fedorov, V. V. (1993) The validity of the genus Elassodiscus Gilbert and Burke, 1912 (Scorpaeniformes, Liparidae)" type="book" year="1993">Pitruk & Fedorov 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBC4B13FF87386A8FB9ACA6D5A6F9B6" author="Kai, Y. & Matsuzaki, K. & Orr, J. W. & Mori, T. & Kamiunten, M." box="[1251,1425,1519,1545]" pageId="2" pageNumber="193" pagination="55 - 66" refId="ref17899" refString="Kai, Y., Matsuzaki, K., Orr, J. W., Mori, T. & Kamiunten, M. (2020) A new species of Elassodiscus (Cottoidei: Liparidae) from the North Pacific with an emended diagnosis of the genus. Ichthyological Research, 68, 55 - 66. https: // doi. org / 10.1007 / s 10228 - 020 - 00764 - 4" type="journal article" year="2020">
|
||
Kai
|
||
<emphasis id="B959EAF0FF87386A8E49ACA6D572F9B6" box="[1299,1349,1519,1545]" italics="true" pageId="2" pageNumber="193">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
). The species group is genetically closely related to the
|
||
<taxonomicName id="4C2D4D61FF87386A8851AF5AD434F993" authorityName="Gilbert & Burke" authorityYear="1912" box="[779,1027,1555,1581]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="rastrinus">
|
||
<emphasis id="B959EAF0FF87386A8851AF5AD434F993" box="[779,1027,1555,1581]" italics="true" pageId="2" pageNumber="193">Careproctus rastrinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2D4D61FF87386A8F65AF5AD576F992" authorityName="Kroyer" authorityYear="1862" box="[1087,1345,1555,1581]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="reinhardti">
|
||
<emphasis id="B959EAF0FF87386A8F65AF5AD576F992" box="[1087,1345,1555,1581]" italics="true" pageId="2" pageNumber="193">Careproctus reinhardti</emphasis>
|
||
</taxonomicName>
|
||
species groups (
|
||
<bibRefCitation id="EFBC4B13FF87386A8BA9AF7ED197F9EE" author="Orr, J. W. & Spies, I. & Stevenson, D. E. & Longo, G. C. & Kai, Y. & Ghods, S. & Hollowed, M." box="[243,416,1591,1617]" pageId="2" pageNumber="193" pagination="1 - 79" refId="ref18772" refString="Orr, J. W., Spies, I., Stevenson, D. E., Longo, G. C., Kai, Y., Ghods, S. & Hollowed, M. (2019) Molecular phylogenetics of snailfishes (Liparidae: Cottoidei) based on mtDNA and RADseq genomic analyses, with comments on selected morphological characters. Zootaxa, 4642 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 4642.1.1" type="journal article" year="2019">
|
||
Orr
|
||
<emphasis id="B959EAF0FF87386A8A79AF7ED161F9EE" box="[291,342,1591,1617]" italics="true" pageId="2" pageNumber="193">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
), but is differentiated from both of the latter in having a smaller pelvic disk (vs. 2.2–9.6% SL) and moderately large gill slit (vs. small gill slit, extending ventrally to pectoral-fin ray 10 at most, except for
|
||
<taxonomicName id="4C2D4D61FF87386A8BCDAF36D205F926" authority="Kroyer, 1862" authorityName="Kroyer" authorityYear="1862" box="[151,562,1663,1689]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="reinhardti">
|
||
<emphasis id="B959EAF0FF87386A8BCDAF36D1A1F926" box="[151,406,1663,1689]" italics="true" pageId="2" pageNumber="193">Careproctus reinhardti</emphasis>
|
||
<bibRefCitation id="EFBC4B13FF87386A8AC6AF36D205F926" author="Kroyer H. N." box="[412,562,1663,1689]" pageId="2" pageNumber="193" pagination="233 - 310" refId="ref18109" refString="Kroyer H. N. (1862) Nogle Bidrag til Nordisk ichthyologi. Naturhist Tidsskr (KJObenhavn), Series 3, 1, 233 - 310." type="journal article" year="1862">Krøyer, 1862</bibRefCitation>
|
||
</taxonomicName>
|
||
, which has a large gill slit extending ventrally to the base of the lower pectoral-fin lobe) (
|
||
<bibRefCitation id="EFBC4B13FF87386A8A5FAFEAD181F902" author="Chernova, N. V." box="[261,438,1698,1725]" pageId="2" pageNumber="193" refId="ref17187" refString="Chernova, N. V. (2005) Review of Careproctus (Liparidae) of the North Atlantic and adjacent Arctic, including the generic type C. reinhardti, with rehabilitation of C. gelatinosus (Pallas) from Kamchatka. Journal of Ichthyology, 45 (Supplement 1), S 1 - S 22." type="journal volume" year="2005">Chernova 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBC4B13FF87386A8A98AFEAD25BF902" author="Orr, J. W. & Kai, Y. & Nakabo, T." box="[450,620,1698,1725]" pageId="2" pageNumber="193" pagination="301 - 348" refId="ref18697" refString="Orr, J. W., Kai, Y. & Nakabo, T. (2015) Snailfishes of the Careproctus rastrinus complex (Liparidae): redescriptions of seven species in the North Pacific Ocean region, with the description of a new species from the Beaufort Sea. Zootaxa, 4018 (3), 301 - 348. https: // doi. org / 10.11646 / zootaxa. 4018.3.1" type="journal article" year="2015">
|
||
Orr
|
||
<emphasis id="B959EAF0FF87386A8AAAAFEAD216F902" box="[496,545,1699,1725]" italics="true" pageId="2" pageNumber="193">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3376569FF87386B8BCDAE46D5AAFE6D" lastPageId="3" lastPageNumber="194" pageId="2" pageNumber="193" type="key">
|
||
<paragraph id="8B9236E2FF87386A8BCDAE46D35BF896" blockId="2.[151,876,1806,1833]" box="[151,876,1806,1833]" pageId="2" pageNumber="193">
|
||
<emphasis id="B959EAF0FF87386A8BCDAE46D35BF896" bold="true" box="[151,876,1806,1833]" pageId="2" pageNumber="193">
|
||
Key to the species of the
|
||
<taxonomicName id="4C2D4D61FF87386A8A91AE46D28AF896" ID-CoL="69BGL" authorityName="Burke" authorityYear="1912" box="[459,701,1806,1833]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="gilberti">
|
||
<emphasis id="B959EAF0FF87386A8A91AE46D28AF896" bold="true" box="[459,701,1806,1833]" italics="true" pageId="2" pageNumber="193">Careproctus gilberti</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</paragraph>
|
||
<keyStep id="30D92E47FF87386A8BCDAE1DD5ABF839" pageId="2" pageNumber="193">
|
||
<paragraph id="8B9236E2FF87386A8BCDAE1DD5ABF8D6" blockId="2.[151,1437,1876,2015]" box="[151,1436,1876,1898]" pageId="2" pageNumber="193">
|
||
<keyLead id="30DC95D7FF87386A8BCDAE1DD5ABF8D6" box="[151,1436,1876,1898]" pageId="2" pageNumber="193">1a. Lower part of pectoral girdle protruding anteroventrally below mandible......................................... 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9236E2FF87386A8BCDAE38D5ABF839" blockId="2.[151,1437,1876,2015]" box="[151,1436,1905,1927]" pageId="2" pageNumber="193">
|
||
<keyLead id="30DC95D7FF87386A8BCDAE38D5ABF839" box="[151,1436,1905,1927]" pageId="2" pageNumber="193">1b. Lower part of pectoral girdle not protruding anteroventrally below mandible...................................... 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8B9236E2FF87386A8BCDAEC7D5AAF87E" blockId="2.[151,1437,1876,2015]" pageId="2" pageNumber="193">
|
||
<keyLead id="30DC95D7FF87386A8BCDAEC7D5AAF87E" pageId="2" pageNumber="193">
|
||
<keyStep id="30D92E47FF87386A8BCDAEC7D5AAF87E" pageId="2" pageNumber="193">
|
||
2a. Skin thin, almost naked (covered with extremely deciduous cactus-like prickles); dorsal-fin rays 50; anal-fin rays 44................................................................................................
|
||
<taxonomicName id="4C2D4D61FF87386A8E48AEE5D5AAF87E" ID-CoL="R7TD" authorityName="Imamura & Nobetsu" authorityYear="2002" box="[1298,1437,1964,1985]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="parvidiscus">
|
||
<emphasis id="B959EAF0FF87386A8E48AEE5D5AAF87E" box="[1298,1437,1964,1985]" italics="true" pageId="2" pageNumber="193">C. parvidiscus</emphasis>
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9236E2FF87386B8BCDAE80D5ABFF12" blockId="2.[151,1437,1876,2015]" lastBlockId="3.[151,1437,151,467]" lastPageId="3" lastPageNumber="194" pageId="2" pageNumber="193">
|
||
<keyLead id="30DC95D7FF87386B8BCDAE80D5ABFF12" lastPageId="3" lastPageNumber="194" pageId="2" pageNumber="193">
|
||
2b. Skin relatively thick, covered by cactus-like prickles; dorsal-fin rays> 51; anal-fin rays> 47......................... 4
|
||
<keyStep id="30D92E47FF86386B8BCDA9D1D5ABFF12" box="[151,1436,151,173]" pageId="3" pageNumber="194">3a. Skin thin, prickles absent or deciduous if present............................................................ 5</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9236E2FF86386B8BCDA9FCD5ABFF74" blockId="3.[151,1437,151,467]" box="[151,1436,181,203]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA9FCD5ABFF74" box="[151,1436,181,203]" pageId="3" pageNumber="194">3b. Skin relatively thick, covered by cactus-like prickles......................................................... 6</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30D92E47FF86386B8BCDA99BD5AAFEBA" pageId="3" pageNumber="194">
|
||
<paragraph id="8B9236E2FF86386B8BCDA99BD5AAFF58" blockId="3.[151,1437,151,467]" box="[151,1437,210,232]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA99BD5AAFF58" box="[151,1437,210,232]" pageId="3" pageNumber="194">
|
||
4a. Body white in preservative.......................................................................
|
||
<taxonomicName id="4C2D4D61FF86386B8E64A99BD5AAFF58" ID-CoL="R7SQ" authorityName="Schmidt" authorityYear="1916" box="[1342,1437,210,231]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="mederi">
|
||
<emphasis id="B959EAF0FF86386B8E64A99BD5AAFF58" box="[1342,1437,210,231]" italics="true" pageId="3" pageNumber="194">C. mederi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9236E2FF86386B8BCDA9A6D5AAFEBA" blockId="3.[151,1437,151,467]" box="[151,1437,239,261]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA9A6D5AAFEBA" box="[151,1437,239,261]" pageId="3" pageNumber="194">
|
||
4b. Body dark in preservative.......................................................................
|
||
<taxonomicName id="4C2D4D61FF86386B8E70A9B9D5AAFEBA" ID-CoL="R7T8" authorityName="Gilbert" authorityYear="1896" box="[1322,1437,240,261]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="ostentum">
|
||
<emphasis id="B959EAF0FF86386B8E70A9B9D5AAFEBA" box="[1322,1437,240,261]" italics="true" pageId="3" pageNumber="194">C. ostentum</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D92E47FF86386B8BCDA844D5AAFE80" pageId="3" pageNumber="194">
|
||
<paragraph id="8B9236E2FF86386B8BCDA844D5ABFE9D" blockId="3.[151,1437,151,467]" box="[151,1436,269,291]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA844D5ABFE9D" box="[151,1436,269,291]" pageId="3" pageNumber="194">5a. Gill slit large, extending ventrally to pectoral-fin rays 12–19................................................... 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9236E2FF86386B8BCDA863D5AAFE80" blockId="3.[151,1437,151,467]" box="[151,1437,298,320]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA863D5AAFE80" box="[151,1437,298,320]" pageId="3" pageNumber="194">
|
||
5b. Gill slit moderately large, extending ventrally to pectoral-fin ray 8..................................
|
||
<emphasis id="B959EAF0FF86386B8FA0A863D53DFE80" box="[1274,1290,298,319]" italics="true" pageId="3" pageNumber="194">C</emphasis>
|
||
. sp.
|
||
<taxonomicName id="4C2D4D61FF86386B8E6DA863D5AAFE80" ID-CoL="69BGL" authorityName="Burke" authorityYear="1912" box="[1335,1437,298,320]" family="Liparidae" genus="Careproctus" isUncertain="true" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="gilberti">
|
||
cf.
|
||
<emphasis id="B959EAF0FF86386B8E0CA863D5AAFE80" box="[1366,1437,298,319]" italics="true" pageId="3" pageNumber="194">gilberti</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D92E47FF86386B8BCDA80ED5ABFE28" pageId="3" pageNumber="194">
|
||
<paragraph id="8B9236E2FF86386B8BCDA80ED5ABFEE2" blockId="3.[151,1437,151,467]" box="[151,1436,327,349]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA80ED5ABFEE2" box="[151,1436,327,349]" pageId="3" pageNumber="194">
|
||
6a. Pectoral fin with shallow notch, lower lobe short, extending slightly posterior to anus..............
|
||
<taxonomicName id="4C2D4D61FF86386B8F9FA801D57EFEE3" authorityName="Kai & Matsuzaki & Mori & Pitruk & Misawa & Tashiro" authorityYear="2024" box="[1221,1353,327,349]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
|
||
<emphasis id="B959EAF0FF86386B8F9FA801D57EFEE3" box="[1221,1353,327,349]" italics="true" pageId="3" pageNumber="194">C. barbatulus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B959EAF0FF86386B8E0AA801D5ABFEE2" bold="true" box="[1360,1436,328,349]" pageId="3" pageNumber="194">
|
||
<taxonomicNameLabel id="A26A578BFF86386B8E0AA801D5ABFEE2" box="[1360,1436,328,349]" pageId="3" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9236E2FF86386B8BCDA82CD5ABFE28" blockId="3.[151,1437,151,467]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA82CD5ABFE28" pageId="3" pageNumber="194">
|
||
6b. Pectoral fin deeply notched, lower lobe long, same length as or slightly longer than upper lobe, extending to two-thirds between anus and anal fin origin...............................................................
|
||
<taxonomicName id="4C2D4D61FF86386B8F92A8CBD57EFE28" authorityName="Kai & Matsuzaki & Mori & Pitruk & Misawa & Tashiro" authorityYear="2024" box="[1224,1353,386,407]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
|
||
<emphasis id="B959EAF0FF86386B8F92A8CBD57EFE28" box="[1224,1353,386,407]" italics="true" pageId="3" pageNumber="194">C. spinulosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B959EAF0FF86386B8E0AA8CBD5ABFE28" bold="true" box="[1360,1436,386,407]" pageId="3" pageNumber="194">
|
||
<taxonomicNameLabel id="A26A578BFF86386B8E0AA8CBD5ABFE28" box="[1360,1436,386,407]" pageId="3" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D92E47FF86386B8BCDA8E9D5AAFE6D" pageId="3" pageNumber="194">
|
||
<paragraph id="8B9236E2FF86386B8BCDA8E9D5ABFE0A" blockId="3.[151,1437,151,467]" box="[151,1436,415,437]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA8E9D5ABFE0A" box="[151,1436,415,437]" pageId="3" pageNumber="194">
|
||
7a. Dorsal-fin rays 56–60; anal-fin rays 50–54; peritoneum dark brown..........................
|
||
<taxonomicName id="4C2D4D61FF86386B8FFDA8E9D57DFE0B" authorityName="Kai & Matsuzaki & Mori & Pitruk & Misawa & Tashiro" authorityYear="2024" box="[1191,1354,415,437]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="longibarbatus" status="sp. nov.">
|
||
<emphasis id="B959EAF0FF86386B8FFDA8E9D57DFE0B" box="[1191,1354,415,437]" italics="true" pageId="3" pageNumber="194">C. longibarbatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B959EAF0FF86386B8E0AA8E9D5ABFE0A" bold="true" box="[1360,1436,416,437]" pageId="3" pageNumber="194">
|
||
<taxonomicNameLabel id="A26A578BFF86386B8E0AA8E9D5ABFE0A" box="[1360,1436,416,437]" pageId="3" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B9236E2FF86386B8BCDA8F4D5AAFE6D" blockId="3.[151,1437,151,467]" box="[151,1437,445,467]" pageId="3" pageNumber="194">
|
||
<keyLead id="30DC95D7FF86386B8BCDA8F4D5AAFE6D" box="[151,1437,445,467]" pageId="3" pageNumber="194">
|
||
7b. Dorsal-fin rays 49–55; anal-fin rays 44–48; peritoneum pale or silvery with black dots.......................
|
||
<taxonomicName id="4C2D4D61FF86386B8E60A8F4D5AAFE6D" ID-CoL="69BGL" authorityName="Burke" authorityYear="1912" box="[1338,1437,445,466]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="gilberti">
|
||
<emphasis id="B959EAF0FF86386B8E60A8F4D5AAFE6D" box="[1338,1437,445,466]" italics="true" pageId="3" pageNumber="194">C. gilberti</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |