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<document id="E0DA23C98256697D9FCE5F7DC0DC931E" ID-CLB-Dataset="5940" ID-DOI="10.11646/zootaxa.3761.1.1" ID-GBIF-Dataset="5444e27e-3e2a-429b-931b-823a2086e3c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909057" ID-ZooBank="82E0F1DC-BC98-4E8A-A3D5-21ECB392CC0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623107451891" checkinUser="felipe" docAuthor="Luk, Stephen P. L. &amp; Marshall, Stephen A." docDate="2014" docId="038487F1FF8CFF99FDC7FB34FAF20B6D" docLanguage="en" docName="zt03761p156.pdf" docOrigin="Zootaxa 3761 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aptilotella umbracatus Luk &amp; Marshall 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FFBDFF89FFA5FFB3FD50FE2BFFC70D18" masterDocTitle="A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="42" updateTime="1699002622211" updateUser="ExternalLinkService">
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<mods:title id="D00F6FA8F4865607F047DA05FB70154B">A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)</mods:title>
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<mods:namePart id="3A05E54A2B1F9EC2B7C1B4D24BFF9ADA">Luk, Stephen P. L.</mods:namePart>
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<mods:namePart id="67A9B9FC911E84988E3C50FC930225E9">Marshall, Stephen A.</mods:namePart>
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<mods:title id="1118ED9E16BDC696A310097A07C03E5F">Zootaxa</mods:title>
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<mods:date id="8E71365ECE69CFC62D1AFF1D12446564">2014</mods:date>
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<treatment id="038487F1FF8CFF99FDC7FB34FAF20B6D" ID-DOI="http://doi.org/10.5281/zenodo.5042262" ID-GBIF-Taxon="183203087" ID-Zenodo-Dep="5042262" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D" httpUri="http://treatment.plazi.org/id/038487F1FF8CFF99FDC7FB34FAF20B6D" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<subSubSection id="C337656CFF8CFF9AFDC7FB34FD170821" box="[151,720,1311,1337]" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="8B9236E7FF8CFF9AFDC7FB34FD170821" blockId="41.[151,720,1311,1371]" box="[151,720,1311,1337]" pageId="41" pageNumber="42">
<heading id="D0DA818BFF8CFF9AFDC7FB34FD170821" bold="true" box="[151,720,1311,1337]" fontSize="11" level="2" pageId="41" pageNumber="42" reason="4">
<emphasis id="B959EAF5FF8CFF9AFDC7FB34FD170821" bold="true" box="[151,720,1311,1337]" pageId="41" pageNumber="42">
<taxonomicName id="4C2D4D64FF8CFF9AFDC7FB34FD460821" authority="Luk &amp; Marshall" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,641,1311,1337]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="umbracatus" status="sp. nov.">
<emphasis id="B959EAF5FF8CFF9AFDC7FB34FE6D0821" bold="true" box="[151,426,1311,1337]" italics="true" pageId="41" pageNumber="42">Aptilotella umbracatus</emphasis>
Luk &amp; Marshall
</taxonomicName>
,
<taxonomicNameLabel id="A26A578EFF8CFF9AFFDDFB34FD170821" box="[653,720,1311,1337]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C337656CFF8CFF99FDC7FB69FCCD0EBD" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="8B9236E7FF8CFF9AFDC7FB69FE340843" blockId="41.[151,720,1311,1371]" box="[151,499,1346,1371]" pageId="41" pageNumber="42">
<figureCitation id="13162A62FF8CFF9AFDC7FB69FEC50843" box="[151,258,1346,1371]" captionStart="FIGURE 4" captionStartId="41.[151,249,1214,1236]" captionTargetId="figure-455@41.[390,1196,591,1192]" captionTargetPageId="41" captionText="FIGURE 4. Aptilotella umbracatus Luk &amp; Marshall, sp. n., paratype male." figureDoi="http://doi.org/10.5281/zenodo.5042243" httpUri="https://zenodo.org/record/5042243/files/figure.png" pageId="41" pageNumber="42">Figures 4</figureCitation>
,
<figureCitation id="13162A62FF8CFF9AFC5EFB69FE910843" box="[270,342,1346,1371]" captionStart-0="FIGURES 4950" captionStart-1="FIGURE 51" captionStartId-0="79.[151,264,1834,1857]" captionStartId-1="80.[151,249,1500,1522]" captionTargetBox-0="[151,1436,193,1813]" captionTargetBox-1="[151,1436,193,1478]" captionTargetId-0="figure-35@79.[151,1436,193,1813]" captionTargetId-1="figure-54@80.[151,1436,193,1478]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 4950. Aptilotella umbracatus Luk &amp; Marshall, sp. n., male: 49—habitus, left lateral; 50—habitus, dorsal." captionText-1="FIGURE 51. Aptilotella umbracatus Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SPN = spinules of synsternite 6+7, SUR = surstylus, VFS = ventral flanking sclerites of distiphallus." figureDoi-0="http://doi.org/10.5281/zenodo.4909161" figureDoi-1="http://doi.org/10.5281/zenodo.4909167" httpUri-0="https://zenodo.org/record/4909161/files/figure.png" httpUri-1="https://zenodo.org/record/4909167/files/figure.png" pageId="41" pageNumber="42">4951</figureCitation>
, and 295305
</paragraph>
<paragraph id="8B9236E7FF8CFF9AFDC7FBA1FE160A6F" blockId="41.[151,1437,1418,2019]" pageId="41" pageNumber="42">
<emphasis id="B959EAF5FF8CFF9AFDC7FBA1FEE408BB" bold="true" box="[151,291,1418,1443]" italics="true" pageId="41" pageNumber="42">Description.</emphasis>
Habitus as in
<figureCitation id="13162A62FF8CFF9AFC94FBA1FDE908BB" box="[452,558,1418,1443]" captionStart="FIGURE 4" captionStartId="41.[151,249,1214,1236]" captionTargetId="figure-455@41.[390,1196,591,1192]" captionTargetPageId="41" captionText="FIGURE 4. Aptilotella umbracatus Luk &amp; Marshall, sp. n., paratype male." figureDoi="http://doi.org/10.5281/zenodo.5042243" httpUri="https://zenodo.org/record/5042243/files/figure.png" pageId="41" pageNumber="42">Figures 4</figureCitation>
,
<figureCitation id="13162A62FF8CFF9AFF6BFBA1FD9E08BB" box="[571,601,1418,1443]" captionStart="FIGURES 4950" captionStartId="79.[151,264,1834,1857]" captionTargetBox="[151,1436,193,1813]" captionTargetId="figure-35@79.[151,1436,193,1813]" captionTargetPageId="79" captionText="FIGURES 4950. Aptilotella umbracatus Luk &amp; Marshall, sp. n., male: 49—habitus, left lateral; 50—habitus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4909161" httpUri="https://zenodo.org/record/4909161/files/figure.png" pageId="41" pageNumber="42">49</figureCitation>
, and 50. Body length 1.0
<quantity id="4CD59B02FF8CFF9AFED1FBA1FC1F08BA" box="[897,984,1418,1443]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="41" pageNumber="42" unit="mm" value="1.3">1.3 mm</quantity>
. Head ground color yellow. Frons finely rugose, its pale areas uniformly constricted by brown stripes; medial brown stripes broad, each approximately onequarter the width of frons, diffuse along the anterior margin; orbital stripes darker, ending at ocular emargination. Ocellar tubercle slightly raised and overlapping medial stripes; ocellar bristle two-thirds the length of frons. Orbital bristle absent; orbital setulae inconspicuous, in three pairs. Interfrontal setae minute, in two pairs. Facial tubercle, excavation and gena shining. Antenna yellow, first flagellomere brown in distal two-thirds. Scutum and scutellum dark reddish-brown to black, shining. Scutum uniformly, sparsely setose, with minute patch of tomentosity on anterior margin laterally and medially; posterior margin tomentose. Scutellum microtrichose; 2.5 times wider than long, 0.7 times the width of scutum. Apical scutellar bristles 2.5 times as long as basal. Pleuron black; anepisternum sometimes with a faint pruinose stripe. Legs yellow; fore coxa brown, mid and hind coxae dark brown; mid tibia and tarsus darker; mid tibia with two anterodorsal bristles. Wing rudiment yellowish. Abdomen black, shining; tergites distally with a row of evenly-spaced setae, syntergite with 2 rows; sternites finely microtrichose, with rows of longer setae posteriorly. Terminalia reddish-brown; epandrium densely microtrichose; cercus and surstylus yellow.
</paragraph>
<paragraph id="8B9236E7FF8CFF99FD97F9A9FE9F0F9B" blockId="41.[151,1437,1418,2019]" lastBlockId="42.[151,1437,151,1653]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="B959EAF5FF8CFF9AFD97F9A9FE4B0A83" bold="true" box="[199,396,1922,1947]" italics="true" pageId="41" pageNumber="42">Male terminalia.</emphasis>
Sternite 5 (
<figureCitation id="13162A62FF8CFF9AFF4BF9A9FDB90A83" box="[539,638,1922,1947]" captionStart="FIGURES 295298" captionStartId="143.[151,264,1115,1138]" captionTargetBox="[384,1206,197,1090]" captionTargetId="figure-42@143.[376,1212,193,1094]" captionTargetPageId="143" captionText="FIGURES 295298. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 295—male terminalia, posterior; 296—male terminalia, left lateral; 297—male synsternite 6+7; 298—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909302" httpUri="https://zenodo.org/record/4909302/files/figure.png" pageId="41" pageNumber="42">Fig. 298</figureCitation>
) irregularly shallowly excised in posteromedial third, with a simple membranous lining. Synsternite 6+7 (
<figureCitation id="13162A62FF8CFF9AFF1EF98DFD770AA7" box="[590,688,1958,1983]" captionStart="FIGURE 51" captionStartId="80.[151,249,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure-54@80.[151,1436,193,1478]" captionTargetPageId="80" captionText="FIGURE 51. Aptilotella umbracatus Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SPN = spinules of synsternite 6+7, SUR = surstylus, VFS = ventral flanking sclerites of distiphallus." figureDoi="http://doi.org/10.5281/zenodo.4909167" httpUri="https://zenodo.org/record/4909167/files/figure.png" pageId="41" pageNumber="42">Figs. 51</figureCitation>
,
<figureCitation id="13162A62FF8CFF9AFF90F98DFD2B0AA7" box="[704,748,1958,1983]" captionStart="FIGURES 295298" captionStartId="143.[151,264,1115,1138]" captionTargetBox="[384,1206,197,1090]" captionTargetId="figure-42@143.[376,1212,193,1094]" captionTargetPageId="143" captionText="FIGURES 295298. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 295—male terminalia, posterior; 296—male terminalia, left lateral; 297—male synsternite 6+7; 298—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909302" httpUri="https://zenodo.org/record/4909302/files/figure.png" pageId="41" pageNumber="42">297</figureCitation>
) densely microtrichose, with a field of minute spinules on medial bridge. Cercus (
<figureCitation id="13162A62FF8CFF9AFCF0F9E1FE390AFB" box="[416,510,1994,2019]" captionStart="FIGURE 51" captionStartId="80.[151,249,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure-54@80.[151,1436,193,1478]" captionTargetPageId="80" captionText="FIGURE 51. Aptilotella umbracatus Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SPN = spinules of synsternite 6+7, SUR = surstylus, VFS = ventral flanking sclerites of distiphallus." figureDoi="http://doi.org/10.5281/zenodo.4909167" httpUri="https://zenodo.org/record/4909167/files/figure.png" pageId="41" pageNumber="42">Figs. 51</figureCitation>
,
<figureCitation id="13162A62FF8CFF9AFF5BF9E1FDB70AFB" box="[523,624,1994,2019]" captionStart="FIGURES 295298" captionStartId="143.[151,264,1115,1138]" captionTargetBox="[384,1206,197,1090]" captionTargetId="figure-42@143.[376,1212,193,1094]" captionTargetPageId="143" captionText="FIGURES 295298. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 295—male terminalia, posterior; 296—male terminalia, left lateral; 297—male synsternite 6+7; 298—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909302" httpUri="https://zenodo.org/record/4909302/files/figure.png" pageId="41" pageNumber="42">295, 296</figureCitation>
) 1.3 times as long as basal width; base triangular, with two inner setae; distal half narrow and curved, basally with a long seta half the length of cercus, and a shorter seta below. Surstylus (
<figureCitation id="13162A62FF8FFF99FDCFFE97FF390DCD" box="[159,254,188,213]" captionStart="FIGURE 51" captionStartId="80.[151,249,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure-54@80.[151,1436,193,1478]" captionTargetPageId="80" captionText="FIGURE 51. Aptilotella umbracatus Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SPN = spinules of synsternite 6+7, SUR = surstylus, VFS = ventral flanking sclerites of distiphallus." figureDoi="http://doi.org/10.5281/zenodo.4909167" httpUri="https://zenodo.org/record/4909167/files/figure.png" pageId="42" pageNumber="43">Figs. 51</figureCitation>
,
<figureCitation id="13162A62FF8FFF99FC5CFE97FEB50DCD" box="[268,370,188,213]" captionStart="FIGURES 295298" captionStartId="143.[151,264,1115,1138]" captionTargetBox="[384,1206,197,1090]" captionTargetId="figure-42@143.[376,1212,193,1094]" captionTargetPageId="143" captionText="FIGURES 295298. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 295—male terminalia, posterior; 296—male terminalia, left lateral; 297—male synsternite 6+7; 298—male sternite 5." figureDoi="http://doi.org/10.5281/zenodo.4909302" httpUri="https://zenodo.org/record/4909302/files/figure.png" pageId="42" pageNumber="43">295, 296</figureCitation>
) irregularly angular; posteromedially with a large setaceous tubercle, beneath which are four stout setae of which the anterior one is largest; distal margin squared, with a few small sensory setae. Postgonite (
<figureCitation id="13162A62FF8FFF99FDCFFF2FFF3B0C05" box="[159,252,260,285]" captionStart="FIGURE 51" captionStartId="80.[151,249,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure-54@80.[151,1436,193,1478]" captionTargetPageId="80" captionText="FIGURE 51. Aptilotella umbracatus Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SPN = spinules of synsternite 6+7, SUR = surstylus, VFS = ventral flanking sclerites of distiphallus." figureDoi="http://doi.org/10.5281/zenodo.4909167" httpUri="https://zenodo.org/record/4909167/files/figure.png" pageId="42" pageNumber="43">Figs. 51</figureCitation>
,
<figureCitation id="13162A62FF8FFF99FC57FF2FFEF50C05" box="[263,306,260,285]" captionStart="FIGURES 299301" captionStartId="144.[151,264,1074,1097]" captionTargetBox="[443,1147,174,1050]" captionTargetId="figure-85@144.[437,1151,167,1053]" captionTargetPageId="144" captionText="FIGURES 299301. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 299—aedeagus with postgonites removed; 300—left postgonite; 301—hypandrium. Abbreviations: BP = basiphallus, DS = dorsal sclerite, LFS = lateral flanking sclerite, VFS = ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909304" httpUri="https://zenodo.org/record/4909304/files/figure.png" pageId="42" pageNumber="43">300</figureCitation>
) sinuous posteriorly, descending arm one-third the total length, with a basal sensory seta; articulatory process for pregonite hooked upward and truncate; articulatory process for basiphallus reduced to a small knob. Hypandrium (
<figureCitation id="13162A62FF8FFF99FC64FF67FE530C7D" box="[308,404,332,357]" captionStart="FIGURES 299301" captionStartId="144.[151,264,1074,1097]" captionTargetBox="[443,1147,174,1050]" captionTargetId="figure-85@144.[437,1151,167,1053]" captionTargetPageId="144" captionText="FIGURES 299301. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 299—aedeagus with postgonites removed; 300—left postgonite; 301—hypandrium. Abbreviations: BP = basiphallus, DS = dorsal sclerite, LFS = lateral flanking sclerite, VFS = ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909304" httpUri="https://zenodo.org/record/4909304/files/figure.png" pageId="42" pageNumber="43">Fig. 301</figureCitation>
) with medial rod straight and slightly slanted to the right; hypandrial arms apparently fused to broad base, slender, with the apex rounded and bent back; pregonite minute, triangular, and fused to base. Aedeagus as in
<figureCitation id="13162A62FF8FFF99FC03FFBFFE130CB5" box="[339,468,404,429]" captionStart="FIGURES 299301" captionStartId="144.[151,264,1074,1097]" captionTargetBox="[443,1147,174,1050]" captionTargetId="figure-85@144.[437,1151,167,1053]" captionTargetPageId="144" captionText="FIGURES 299301. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 299—aedeagus with postgonites removed; 300—left postgonite; 301—hypandrium. Abbreviations: BP = basiphallus, DS = dorsal sclerite, LFS = lateral flanking sclerite, VFS = ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909304" httpUri="https://zenodo.org/record/4909304/files/figure.png" pageId="42" pageNumber="43">Figure 299</figureCitation>
. Basiphallus cylindrical; posteroventrally humped, appearing boot-shaped in lateral profile; articulatory process for postgonite rounded and divergent. Ejaculatory apodeme very slender-tipped. Lateral flanking sclerites with dark dorsal and ventral margins, very narrowly fused posterodorsally and more broadly posteroventrally; dorsum densely armed with heavily-sclerotized teeth. Dorsal sclerite very dark; flattened; apex curved upwards and broadly rounded, with a rudder-like ventral keel. Ventral flanking sclerites paddleshaped; the triangular medial article sharply pointed beneath the basal article; the distal article darkest and irregularly ovate.
</paragraph>
<paragraph id="8B9236E7FF8FFF99FD97FCA4FBF20E0D" blockId="42.[151,1437,151,1653]" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FD97FCA4FE630FB0" bold="true" box="[199,420,655,680]" italics="true" pageId="42" pageNumber="43">Female terminalia.</emphasis>
Tergite 7 divided medially; distal half densely microtrichose, with several marginal setae. Epiproct semicircular. Each half of tergite 8 (
<figureCitation id="13162A62FF8FFF99FFF2FC9FFC800FD5" box="[674,839,692,717]" captionStart="FIGURES 302305" captionStartId="144.[151,264,1978,2001]" captionTargetBox="[405,1183,1214,1954]" captionTargetId="figure-87@144.[397,1189,1208,1957]" captionTargetPageId="144" captionText="FIGURES 302305. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 302—female terminalia, dorsal; 303—female terminalia, left lateral; 304—female terminalia, ventral; 305—spermathecae. Abbreviation: S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909306" httpUri="https://zenodo.org/record/4909306/files/figure.png" pageId="42" pageNumber="43">Figs. 302304</figureCitation>
) rhomboid, with a weak medial ridge. Cercus three times as long as wide; truncate. Hypoproct a narrow band. Spermathecae (
<figureCitation id="13162A62FF8FFF99FE83FCFCFBF50FE8" box="[979,1074,727,752]" captionStart="FIGURES 302305" captionStartId="144.[151,264,1978,2001]" captionTargetBox="[405,1183,1214,1954]" captionTargetId="figure-87@144.[397,1189,1208,1957]" captionTargetPageId="144" captionText="FIGURES 302305. Aptilotella umbracatus Luk &amp; Marshall, sp. n.: 302—female terminalia, dorsal; 303—female terminalia, left lateral; 304—female terminalia, ventral; 305—spermathecae. Abbreviation: S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909306" httpUri="https://zenodo.org/record/4909306/files/figure.png" pageId="42" pageNumber="43">Fig. 305</figureCitation>
) ridged; collar with two rings of minute stubs; sclerotized ducts very short, less than the diameter of a spermatheca.
</paragraph>
<paragraph id="8B9236E7FF8FFF99FD97FD34FCCD0EBD" blockId="42.[151,1437,151,1653]" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FD97FD34FEFE0E20" bold="true" box="[199,313,799,824]" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FD97FD34FEF50E20" bold="true" box="[199,306,799,824]" italics="true" pageId="42" pageNumber="43">Variation</emphasis>
.
</emphasis>
Some series consist of darker and barer individuals. These specimens, while possessing the unmistakable dorsal sclerite which characterises
<taxonomicName id="4C2D4D64FF8FFF99FF9BFD6FFC170E44" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[715,976,836,860]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="umbracatus">
<emphasis id="B959EAF5FF8FFF99FF9BFD6FFC170E44" box="[715,976,836,860]" italics="true" pageId="42" pageNumber="43">Aptilotella umbracatus</emphasis>
</taxonomicName>
, differ by most to all of the following: darker overall colour; legs dull yellow to orange, often paler distally; all coxae dark; scutum lacking anteromedial patch of tomentosity; scutellum lacking microtrichosity.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8FFF99FD97FD84FB5B0EF5" pageId="42" pageNumber="43" type="etymology">
<paragraph id="8B9236E7FF8FFF99FD97FD84FB5B0EF5" blockId="42.[151,1437,151,1653]" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FD97FD84FE800ED0" bold="true" box="[199,327,943,968]" italics="true" pageId="42" pageNumber="43">Etymology.</emphasis>
The species epithet is derived from the Latin
<emphasis id="B959EAF5FF8FFF99FE27FD9BFC070ED0" box="[887,960,944,968]" italics="true" pageId="42" pageNumber="43">umbra</emphasis>
, “shadow,” and
<emphasis id="B959EAF5FF8FFF99F9D8FD9AFB160ED0" box="[1160,1233,945,968]" italics="true" pageId="42" pageNumber="43">acatus</emphasis>
, “a boat or light vessel,” in reference to the starkly contrasting, rudder-like dorsal sclerite of the distiphallus.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8FFF99FD97FDD3FBFD0B10" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8B9236E7FF8FFF99FD97FDD3FE97092D" blockId="42.[151,1437,151,1653]" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FD97FDD3FEB60908" bold="true" box="[199,369,1015,1040]" italics="true" pageId="42" pageNumber="43">Type material.</emphasis>
<materialsCitation id="3B453CBAFF8FFF99FC2DFDDCFE97092D" ID-GBIF-Occurrence="3311767356" collectingDate="1977-05-23" collectingDateMax="1977-05-28" collectingDateMin="1977-05-23" collectionCode="DEBU" collectorName="S. Peck." country="Panama" elevation="2500" location="Cerro Punta" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="holotype">
<typeStatus id="54968845FF8FFF99FC2DFDDCFE220908" box="[381,485,1015,1040]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2BFD6EFF8FFF99FCA2FDD3FDCE0917" box="[498,521,1016,1039]" pageId="42" pageNumber="43" type="male"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FF4AFDD3FDAD0917" box="[538,618,1016,1039]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
.
<collectingCountry id="F33A7677FF8FFF99FF29FDDCFD330908" box="[633,756,1015,1040]" name="Panama" pageId="42" pageNumber="43">PANAMA</collectingCountry>
:
<collectingRegion id="49E9F805FF8FFF99FE54FDDCFCA30908" box="[772,868,1015,1040]" country="Panama" name="Chiriqui" pageId="42" pageNumber="43">Chiriquí</collectingRegion>
,
<quantity id="4CD59B02FF8FFF99FE24FDDCFC0F0908" box="[884,968,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="42" pageNumber="43" unit="km" value="4.5">4.5 km</quantity>
E
<location id="8EF2603CFF8FFF99FEA1FDDCFB470917" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FEA1FDDCFB470917" box="[1009,1152,1015,1040]" country="Panama" name="Cerro Punta" pageId="42" pageNumber="43" stateProvince="Chiriqui">Cerro Punta</location>
,
<quantity id="4CD59B02FF8FFF99F9C0FDDCFB2B0917" box="[1168,1260,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="42" pageNumber="43" unit="m" value="2500.0">
<elevation id="0000D1D4FF8FFF99F9C0FDDCFB2B0917" box="[1168,1260,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="42" pageNumber="43" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99F9ADFDDCFA5F0908" box="[1277,1432,1015,1040]" pageId="42" pageNumber="43" value="1977-05-23" valueMax="1977-05-28" valueMin="1977-05-23">
<collectingDate id="EFD7E9CFFF8FFF99F9ADFDDCFA5F0908" box="[1277,1432,1015,1040]" pageId="42" pageNumber="43" value="1977-05-23" valueMax="1977-05-28" valueMin="1977-05-23">2328.v.1977</collectingDate>
</date>
, carrion,
<collectorName id="26D85331FF8FFF99FDA4FA37FE97092D" box="[244,336,1052,1077]" pageId="42" pageNumber="43">S. Peck.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9236E7FF8FFF99FD97FA6BFBFD0B10" blockId="42.[151,1437,151,1653]" pageId="42" pageNumber="43">
<materialsCitation id="3B453CBAFF8FFF99FD97FA6BFC3B0940" ID-GBIF-Occurrence="3311767433" box="[199,1020,1087,1112]" collectionCode="DEBU" country="Panama" location="Chiriqui" pageId="42" pageNumber="43" specimenCount="13" specimenCount-female="4" specimenCount-male="9" stateProvince="Chiriqui" typeStatus="paratype">
<emphasis id="B959EAF5FF8FFF99FD97FA6BFEF8094F" bold="true" box="[199,319,1088,1111]" italics="true" pageId="42" pageNumber="43">
<typeStatus id="54968845FF8FFF99FD97FA6BFEFC094F" box="[199,315,1088,1111]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F33A7677FF8FFF99FC16FA14FE060940" box="[326,449,1087,1112]" name="Panama" pageId="42" pageNumber="43">PANAMA</collectingCountry>
:
<collectingRegion id="49E9F805FF8FFF99FC9BFA14FDEC0940" box="[459,555,1087,1112]" country="Panama" name="Chiriqui" pageId="42" pageNumber="43">Chiriquí</collectingRegion>
, same label as
<typeStatus id="54968845FF8FFF99FF80FA14FCF50940" box="[720,818,1087,1112]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D2BFD6EFF8FFF99FE11FA14FCA00940" box="[833,871,1087,1112]" pageId="42" pageNumber="43" type="male">9♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FE22FA14FC500940" box="[882,919,1087,1112]" pageId="42" pageNumber="43" type="female">4♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FEF3FA14FC330940" box="[931,1012,1087,1112]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99F956FA14FD610965" ID-GBIF-Occurrence="3311767406" collectingDate="1977-05-28" collectingDateMax="1977-06-08" collectingDateMin="1977-05-28" collectionCode="DEBU" collectorName="Peck" country="Panama" elevation="2200" location="2 km E Cerro Punta" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiriqui" typeStatus="paratype">
<location id="8EF2603CFF8FFF99F956FA14FB21094F" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99F956FA14FB21094F" box="[1030,1254,1087,1112]" country="Panama" name="2 km E Cerro Punta" pageId="42" pageNumber="43" stateProvince="Chiriqui">
<quantity id="4CD59B02FF8FFF99F956FA14FBF80940" box="[1030,1087,1087,1112]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="km" value="2.0">2 km</quantity>
E Cerro Punta
</location>
,
<quantity id="4CD59B02FF8FFF99F9A0FA14FA81094F" box="[1264,1350,1087,1112]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="42" pageNumber="43" unit="m" value="2200.0">
<elevation id="0000D1D4FF8FFF99F9A0FA14FA81094F" box="[1264,1350,1087,1112]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="42" pageNumber="43" unit="m" value="2200.0">2200 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99F801FA14FEC40965" pageId="42" pageNumber="43" value="1977-05-28" valueMax="1977-06-08" valueMin="1977-05-28">
<collectingDate id="EFD7E9CFFF8FFF99F801FA14FEC40965" pageId="42" pageNumber="43" value="1977-05-28" valueMax="1977-06-08" valueMin="1977-05-28">28.v. 8.vi.1977</collectingDate>
</date>
,
<collectingMethod id="526C4EF0FF8FFF99FC5FFA4FFE400963" box="[271,391,1124,1149]" pageId="42" pageNumber="43">dung traps</collectingMethod>
, S.
<collectorName id="26D85331FF8FFF99FCE1FA4FFE200965" box="[433,487,1124,1149]" pageId="42" pageNumber="43">Peck</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FCA7FA4FFDDA0965" box="[503,541,1124,1149]" pageId="42" pageNumber="43" type="male">2♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FF79FA4FFDF90963" box="[553,574,1124,1147]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FF1DFA4FFD590965" box="[589,670,1124,1149]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FFE1FA4FFEDB09B8" ID-GBIF-Occurrence="3311767312" collectingDate="1977-06-01" collectingDateMax="1977-06-08" collectingDateMin="1977-06-01" collectionCode="DEBU" collectorName="Peck" country="Panama" elevation="2200" location="2 km E Cerro Punta" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
same locality as previous label,
<date id="FF931027FF8FFF99F948FA4FFB670965" box="[1048,1184,1124,1149]" pageId="42" pageNumber="43" value="1977-06-01" valueMax="1977-06-08" valueMin="1977-06-01">
<collectingDate id="EFD7E9CFFF8FFF99F948FA4FFB670965" box="[1048,1184,1124,1149]" pageId="42" pageNumber="43" value="1977-06-01" valueMax="1977-06-08" valueMin="1977-06-01">18.vi.1977</collectingDate>
</date>
,
<collectingMethod id="526C4EF0FF8FFF99F9FCFA4FFAFB0963" box="[1196,1340,1124,1149]" pageId="42" pageNumber="43">carrion traps</collectingMethod>
, S.
<collectorName id="26D85331FF8FFF99F836FA4FFA5B0965" box="[1382,1436,1124,1149]" pageId="42" pageNumber="43">Peck</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FDF0FAACFF7009B8" box="[160,183,1159,1184]" pageId="42" pageNumber="43" type="male"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FD93FAACFED309B8" box="[195,276,1159,1184]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FC76FAA3FB8209B8" ID-GBIF-Occurrence="3311767346" box="[294,1093,1159,1184]" collectingDate="1977-06-08" collectingDateMax="1977-06-08" collectingDateMin="1977-06-01" collectionCode="DEBU" collectorName="Peck" country="Panama" elevation="2200" location="2 km E Cerro Punta" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
same locality as previous label,
<date id="FF931027FF8FFF99FFD8FAACFD3209B8" box="[648,757,1159,1184]" pageId="42" pageNumber="43" value="1977-06-08">
<collectingDate id="EFD7E9CFFF8FFF99FFD8FAACFD3209B8" box="[648,757,1159,1184]" pageId="42" pageNumber="43" value="1977-06-08">8.vi.1977</collectingDate>
</date>
, dung, S.
<collectorName id="26D85331FF8FFF99FE32FAACFC5F09B8" box="[866,920,1159,1184]" pageId="42" pageNumber="43">Peck</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FEF8FAACFC7809B8" box="[936,959,1159,1184]" pageId="42" pageNumber="43" type="male"></specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FE9AFAA3FC180987" box="[970,991,1160,1183]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FEBCFAACFBFA09B8" box="[1004,1085,1159,1184]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99F900FAA3FC6109DD" ID-GBIF-Occurrence="3311767310" collectingDate="1995-06-19" collectionCode="DEBU" collectorName="R. S. Anderson" country="Panama" elevation="1500" location="Boquete" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
<location id="8EF2603CFF8FFF99F900FAA3FB770987" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99F900FAA3FB770987" box="[1104,1200,1160,1183]" country="Panama" name="Boquete" pageId="42" pageNumber="43" stateProvince="Chiriqui">Boquete</location>
,
<quantity id="4CD59B02FF8FFF99F9EBFAACFACF09B8" box="[1211,1288,1159,1184]" metricMagnitude="3" metricUnit="m" metricValue="5.7" pageId="42" pageNumber="43" unit="km" value="5.7">5.7 km</quantity>
NE,
<quantity id="4CD59B02FF8FFF99F811FAACFA500987" box="[1345,1431,1159,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="m" value="1500.0">
<elevation id="0000D1D4FF8FFF99F811FAACFA500987" box="[1345,1431,1159,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99FDC7FA87FED609DD" box="[151,273,1196,1221]" pageId="42" pageNumber="43" value="1995-06-19">
<collectingDate id="EFD7E9CFFF8FFF99FDC7FA87FED609DD" box="[151,273,1196,1221]" pageId="42" pageNumber="43" value="1995-06-19">19.vi.1995</collectingDate>
</date>
, mixed oak forest, leaf litter,
<collectorName id="26D85331FF8FFF99FF05FA87FD3E09DD" box="[597,761,1196,1221]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FE58FA87FCD809DD" box="[776,799,1196,1221]" pageId="42" pageNumber="43" type="male"></specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FE7BFA87FC8709DB" box="[811,832,1196,1219]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FE1DFA87FC5909DD" box="[845,926,1196,1221]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FEE1FA87FDFA09F0" ID-GBIF-Occurrence="3311767355" collectingDate="1996-06-14" collectionCode="DEBU" collectorName="R. S. Anderson" country="Panama" location="Boquete" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui" typeStatus="paratype">
<location id="8EF2603CFF8FFF99FEE1FA87FBD609DB" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FEE1FA87FBD609DB" box="[945,1041,1196,1219]" country="Panama" name="Boquete" pageId="42" pageNumber="43" stateProvince="Chiriqui">Boquete</location>
,
<quantity id="4CD59B02FF8FFF99F94BFA87FBAF09DD" box="[1051,1128,1196,1221]" metricMagnitude="3" metricUnit="m" metricValue="5.8" pageId="42" pageNumber="43" unit="km" value="5.8">5.8 km</quantity>
NE,
<date id="FF931027FF8FFF99F9F1FA87FADC09DD" box="[1185,1307,1196,1221]" pageId="42" pageNumber="43" value="1996-06-14">
<collectingDate id="EFD7E9CFFF8FFF99F9F1FA87FADC09DD" box="[1185,1307,1196,1221]" pageId="42" pageNumber="43" value="1996-06-14">14.vi.1996</collectingDate>
</date>
, oak forest, leaf litter,
<collectorName id="26D85331FF8FFF99FC5BFAE4FE7709F0" box="[267,432,1231,1256]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FC92FAE4FE1109F0" box="[450,470,1231,1256]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FCB3FAE4FDF309F0" box="[483,564,1231,1256]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FF19FAE4FCFD0815" ID-GBIF-Occurrence="3311767329" collectingDate="1995-06-14" collectionCode="DEBU" collectorName="R. S. Anderson" country="Panama" elevation="2400" location="Volcan Baru National Park" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiriqui" typeStatus="paratype">
<location id="8EF2603CFF8FFF99FF19FAE4FCBE09F0" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FF19FAE4FCBE09F0" box="[585,889,1231,1256]" country="Panama" name="Volcan Baru National Park" pageId="42" pageNumber="43" stateProvince="Chiriqui">Volcán Barú National Park</location>
,
<locationDeviation id="9A202FC5FF8FFF99FED5FAE4FBB909FF" box="[901,1150,1231,1256]" location="Cerro Punta" pageId="42" pageNumber="43">
<quantity id="4CD59B02FF8FFF99FED5FAE4FC1309F0" box="[901,980,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="42" pageNumber="43" unit="km" value="5.9">5.9 km</quantity>
E Cerro Punta
</locationDeviation>
,
<quantity id="4CD59B02FF8FFF99F9D9FAE4FB2609FF" box="[1161,1249,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="42" pageNumber="43" unit="m" value="2400.0">
<elevation id="0000D1D4FF8FFF99F9D9FAE4FB2609FF" box="[1161,1249,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="42" pageNumber="43" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99F9BDFAE4FAA009F0" box="[1261,1383,1231,1256]" pageId="42" pageNumber="43" value="1995-06-14">
<collectingDate id="EFD7E9CFFF8FFF99F9BDFAE4FAA009F0" box="[1261,1383,1231,1256]" pageId="42" pageNumber="43" value="1995-06-14">14.vi.1995</collectingDate>
</date>
, oak ridge-bamboo forest, litter,
<collectorName id="26D85331FF8FFF99FC80FADFFDB00815" box="[464,631,1268,1293]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FFD9FADFFD690815" box="[649,686,1268,1293]" pageId="42" pageNumber="43" type="male">2♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FFECFADFFD160813" box="[700,721,1268,1291]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FFB1FADFFCF50815" box="[737,818,1268,1293]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
; same as previous label but at
<quantity id="4CD59B02FF8FFF99F9CDFADFFB310813" box="[1181,1270,1268,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="42" pageNumber="43" unit="m" value="2150.0">2150 m</quantity>
, riparian alder forest (
<specimenCount id="9D2BFD6EFF8FFF99FDB9FB3CFF3A0828" box="[233,253,1303,1328]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FC59FB3CFE9D0828" box="[265,346,1303,1328]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
);
<materialsCitation id="3B453CBAFF8FFF99FC3DFB3CFDCD084D" ID-GBIF-Occurrence="3311767357" collectingDate="1995-06-18" collectionCode="DEBU" collectorName="R. S. Anderson" country="Panama" elevation="2150" location="Volcan Baru National Park" pageId="42" pageNumber="43" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Chiriqui" typeStatus="paratype">
<location id="8EF2603CFF8FFF99FC3DFB3CFD5B0828" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FC3DFB3CFD5B0828" box="[365,668,1303,1328]" country="Panama" name="Volcan Baru National Park" pageId="42" pageNumber="43" stateProvince="Chiriqui">Volcán Barú National Park</location>
,
<locationDeviation id="9A202FC5FF8FFF99FFF7FB3CFCB30837" box="[679,884,1303,1328]" location="Boquete" pageId="42" pageNumber="43">
<quantity id="4CD59B02FF8FFF99FFF7FB3CFD2A0828" box="[679,749,1303,1328]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="42" pageNumber="43" unit="km" value="11.0">11 km</quantity>
W Boquete
</locationDeviation>
,
<quantity id="4CD59B02FF8FFF99FE2FFB3CFC110837" box="[895,982,1303,1328]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="42" pageNumber="43" unit="m" value="2150.0">
<elevation id="0000D1D4FF8FFF99FE2FFB3CFC110837" box="[895,982,1303,1328]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="42" pageNumber="43" unit="m" value="2150.0">2150 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99FEB2FB3CFB9B0828" box="[994,1116,1303,1328]" pageId="42" pageNumber="43" value="1995-06-18">
<collectingDate id="EFD7E9CFFF8FFF99FEB2FB3CFB9B0828" box="[994,1116,1303,1328]" pageId="42" pageNumber="43" value="1995-06-18">18.vi.1995</collectingDate>
</date>
, mixed oak forest, leaf litter,
<collectorName id="26D85331FF8FFF99FDC7FB17FEFB084D" box="[151,316,1340,1365]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FC1DFB17FEB5084D" box="[333,370,1340,1365]" pageId="42" pageNumber="43" type="male">3♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FC2FFB17FE63084D" box="[383,420,1340,1365]" pageId="42" pageNumber="43" type="female">4♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FCE1FB17FDC5084D" box="[433,514,1340,1365]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FF46FB17FDC00860" ID-GBIF-Occurrence="3311767444" collectingDate="1996-06-17" collectionCode="DEBU" collectorName="R. S. Anderson" country="Panama" elevation="2120" location="Cerro" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui" typeStatus="paratype">
<location id="8EF2603CFF8FFF99FF46FB17FD91084D" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FF46FB17FD91084D" box="[534,598,1340,1365]" country="Panama" name="Cerro" pageId="42" pageNumber="43" stateProvince="Chiriqui">Cerro</location>
Pando,
<locationDeviation id="9A202FC5FF8FFF99FFE3FB17FC71084D" box="[691,950,1340,1365]" location="Santa Clara" pageId="42" pageNumber="43">
<quantity id="4CD59B02FF8FFF99FFE3FB17FD3C084D" box="[691,763,1340,1365]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="42" pageNumber="43" unit="km" value="12.0">12 km</quantity>
NE Santa Clara
</locationDeviation>
,
<quantity id="4CD59B02FF8FFF99FE91FB17FBDE084B" box="[961,1049,1340,1365]" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="42" pageNumber="43" unit="m" value="2120.0">
<elevation id="0000D1D4FF8FFF99FE91FB17FBDE084B" box="[961,1049,1340,1365]" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="42" pageNumber="43" unit="m" value="2120.0">2120 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99F976FB17FB67084D" box="[1062,1184,1340,1365]" pageId="42" pageNumber="43" value="1996-06-17">
<collectingDate id="EFD7E9CFFF8FFF99F976FB17FB67084D" box="[1062,1184,1340,1365]" pageId="42" pageNumber="43" value="1996-06-17">17.vi.1996</collectingDate>
</date>
, wet cloud forest, leaf litter,
<collectorName id="26D85331FF8FFF99FD88FB74FEBB0860" box="[216,380,1375,1400]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FCDDFB74FE670860" box="[397,416,1375,1400]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FCFDFB74FE390860" box="[429,510,1375,1400]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FF41FB74FDD50885" ID-GBIF-Occurrence="3311767447" collectingDate="1995-06-16" collectionCode="DEBU" collectorName="R. S. Anderson" country="Panama" elevation="1800" location="Hartmann's Finca" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
<location id="8EF2603CFF8FFF99FF41FB74FD1C0860" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FF41FB74FD1C0860" box="[529,731,1375,1400]" country="Panama" name="Hartmann's Finca" pageId="42" pageNumber="43" stateProvince="Chiriqui">Hartmanns Finca</location>
,
<locationDeviation id="9A202FC5FF8FFF99FFB6FB74FC7F0860" box="[742,952,1375,1400]" location="Volcan" pageId="42" pageNumber="43">
<quantity id="4CD59B02FF8FFF99FFB6FB74FC860860" box="[742,833,1375,1400]" metricMagnitude="4" metricUnit="m" metricValue="3.07" pageId="42" pageNumber="43" unit="km" value="30.7">30.7 km</quantity>
W Volcán
</locationDeviation>
,
<quantity id="4CD59B02FF8FFF99FE93FB74FBDE086F" box="[963,1049,1375,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="42" pageNumber="43" unit="m" value="1800.0">
<elevation id="0000D1D4FF8FFF99FE93FB74FBDE086F" box="[963,1049,1375,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="42" pageNumber="43" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99F974FB74FB580860" box="[1060,1183,1375,1400]" pageId="42" pageNumber="43" value="1995-06-16">
<collectingDate id="EFD7E9CFFF8FFF99F974FB74FB580860" box="[1060,1183,1375,1400]" pageId="42" pageNumber="43" value="1995-06-16">16.vi.1995</collectingDate>
</date>
, mixed oak forest, leaf litter,
<collectorName id="26D85331FF8FFF99FD8BFBAFFE460885" box="[219,385,1412,1437]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FCC4FBAFFE6C0885" box="[404,427,1412,1437]" pageId="42" pageNumber="43" type="male"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FCE9FBAFFDCD0885" box="[441,522,1412,1437]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FF70FBAFFDF808D8" ID-GBIF-Occurrence="3311767399" collectingDate="1995-06-11" collectionCode="DEBU" collectorName="R. S. Anderson" country="Switzerland" elevation="1525" location="Finca La" municipality="La Fortuna area" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiriqui" typeStatus="paratype">
<collectingMunicipality id="6BF6AC9DFF8FFF99FF70FBAFFD1F0883" box="[544,728,1412,1435]" pageId="42" pageNumber="43">La Fortuna area</collectingMunicipality>
,
<location id="8EF2603CFF8FFF99FFB6FBAFFC8D0883" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FFB6FBAFFC8D0883" box="[742,842,1412,1437]" country="Switzerland" municipality="La Fortuna area" name="Finca La" pageId="42" pageNumber="43" stateProvince="Chiriqui">Finca La</location>
<collectingCountry id="F33A7677FF8FFF99FE04FBAFFC590885" box="[852,926,1412,1437]" name="Switzerland" pageId="42" pageNumber="43">Suisse</collectingCountry>
,
<quantity id="4CD59B02FF8FFF99FEFCFBAFFB8C0883" box="[940,1099,1412,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="42" pageNumber="43" unit="m" value="1525.0" valueMax="1600.0" valueMin="1450.0">
<elevation id="0000D1D4FF8FFF99FEFCFBAFFB8C0883" box="[940,1099,1412,1437]" metricMagnitude="3" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="42" pageNumber="43" unit="m" value="1525.0" valueMax="1600.0" valueMin="1450.0">14501600 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99F90AFBAFFB150885" box="[1114,1234,1412,1437]" pageId="42" pageNumber="43" value="1995-06-11">
<collectingDate id="EFD7E9CFFF8FFF99F90AFBAFFB150885" box="[1114,1234,1412,1437]" pageId="42" pageNumber="43" value="1995-06-11">11.vi.1995</collectingDate>
</date>
, oak ridge forest, litter,
<collectorName id="26D85331FF8FFF99FD8AFB8CFEB808D8" box="[218,383,1447,1472]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FCC0FB8CFE7208D8" box="[400,437,1447,1472]" pageId="42" pageNumber="43" type="male">2♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FC92FB83FE1008A7" box="[450,471,1448,1471]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FCB6FB8CFDF008D8" box="[486,567,1447,1472]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8FFF99FF1CFB83FDA408FD" ID-GBIF-Occurrence="3311767397" collectingDate="1995-06-12" collectionCode="DEBU" collectorName="R. S. Anderson" country="Switzerland" elevation="1450" location="Finca La" municipality="La Fortuna area" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui" typeStatus="paratype">
<collectingMunicipality id="6BF6AC9DFF8FFF99FF1CFB83FCC508A7" box="[588,770,1448,1471]" pageId="42" pageNumber="43">La Fortuna area</collectingMunicipality>
,
<location id="8EF2603CFF8FFF99FE5EFB8CFCB508A7" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99FE5EFB8CFCB508A7" box="[782,882,1447,1472]" country="Switzerland" municipality="La Fortuna area" name="Finca La" pageId="42" pageNumber="43" stateProvince="Chiriqui">Finca La</location>
<collectingCountry id="F33A7677FF8FFF99FE2AFB8CFC0308D8" box="[890,964,1447,1472]" name="Switzerland" pageId="42" pageNumber="43">Suisse</collectingCountry>
,
<quantity id="4CD59B02FF8FFF99FE80FB8CFBEE08A7" box="[976,1065,1447,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="42" pageNumber="43" unit="m" value="1450.0">
<elevation id="0000D1D4FF8FFF99FE80FB8CFBEE08A7" box="[976,1065,1447,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="42" pageNumber="43" unit="m" value="1450.0">1450 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99F965FB8CFB6808D8" box="[1077,1199,1447,1472]" pageId="42" pageNumber="43" value="1995-06-12">
<collectingDate id="EFD7E9CFFF8FFF99F965FB8CFB6808D8" box="[1077,1199,1447,1472]" pageId="42" pageNumber="43" value="1995-06-12">12.vi.1995</collectingDate>
</date>
, wet montane forest, leaf litter,
<collectorName id="26D85331FF8FFF99FC4AFBE7FE0108FD" box="[282,454,1484,1509]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FCB1FBE7FE3208FD" box="[481,501,1484,1509]" pageId="42" pageNumber="43" type="female"></specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FF5AFBE7FD9C08FD" box="[522,603,1484,1509]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B453CBAFF8FFF99FF25FBE7FBF10B10" ID-GBIF-Occurrence="3311767377" collectingDate="2005-02-26" collectingDateMax="2005-02-27" collectingDateMin="2005-02-26" collectionCode="DEBU" collectorName="R. S. Anderson" country="Costa Rica" county="Valle del Silencio" elevation="2473" latitude="9.110278" location="La Estacion" longLatPrecision="21" longitude="-82.961945" municipality="La Estacion" pageId="42" pageNumber="43" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Limon" typeStatus="paratype">
<collectingCountry id="F33A7677FF8FFF99FF25FBE7FCE208FD" box="[629,805,1484,1509]" name="Costa Rica" pageId="42" pageNumber="43">COSTA RICA</collectingCountry>
:
<collectingRegion id="49E9F805FF8FFF99FE68FBE7FC4108FD" box="[824,902,1484,1509]" country="Costa Rica" name="Limon" pageId="42" pageNumber="43">Limón</collectingRegion>
,
<collectingCounty id="62F34E6BFF8FFF99FEC9FBE7FBB508FD" box="[921,1138,1484,1509]" pageId="42" pageNumber="43">Valle del Silencio</collectingCounty>
,
<location id="8EF2603CFF8FFF99F9D5FBE7FAD308FD" LSID="urn:lsid:plazi:treatment:038487F1FF8CFF99FDC7FB34FAF20B6D:8EF2603CFF8FFF99F9D5FBE7FAD308FD" box="[1157,1300,1484,1509]" country="Costa Rica" county="Valle del Silencio" latitude="9.110278" longLatPrecision="21" longitude="-82.961945" municipality="La Estacion" name="La Estacion" pageId="42" pageNumber="43" stateProvince="Limon">
<collectingMunicipality id="6BF6AC9DFF8FFF99F9D5FBE7FAD308FD" box="[1157,1300,1484,1509]" pageId="42" pageNumber="43">La Estación</collectingMunicipality>
</location>
,
<geoCoordinate id="EE195020FF8FFF99F878FBE7FA5008FD" box="[1320,1431,1484,1509]" degrees="9" direction="north" minutes="6" orientation="latitude" pageId="42" pageNumber="43" precision="15" seconds="37" value="9.110278">9°637”N</geoCoordinate>
,
<geoCoordinate id="EE195020FF8FFF99FDC7FBC4FEE10B10" box="[151,294,1519,1544]" degrees="82" direction="west" minutes="57" orientation="longitude" pageId="42" pageNumber="43" precision="15" seconds="43" value="-82.961945">82°5743”W</geoCoordinate>
,
<quantity id="4CD59B02FF8FFF99FC62FBC4FE4E0B1F" box="[306,393,1519,1544]" metricMagnitude="3" metricUnit="m" metricValue="2.473" pageId="42" pageNumber="43" unit="m" value="2473.0">
<elevation id="0000D1D4FF8FFF99FC62FBC4FE4E0B1F" box="[306,393,1519,1544]" metricMagnitude="3" metricUnit="m" metricValue="2.473" pageId="42" pageNumber="43" unit="m" value="2473.0">2473 m</elevation>
</quantity>
,
<date id="FF931027FF8FFF99FCC5FBC4FDF50B10" box="[405,562,1519,1544]" pageId="42" pageNumber="43" value="2005-02-26" valueMax="2005-02-27" valueMin="2005-02-26">
<collectingDate id="EFD7E9CFFF8FFF99FCC5FBC4FDF50B10" box="[405,562,1519,1544]" pageId="42" pageNumber="43" value="2005-02-26" valueMax="2005-02-27" valueMin="2005-02-26">2627.ii.2005</collectingDate>
</date>
, forest litter,
<collectorName id="26D85331FF8FFF99FF96FBC4FCAC0B10" box="[710,875,1519,1544]" pageId="42" pageNumber="43">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8FFF99FE2BFBC4FC670B10" box="[891,928,1519,1544]" pageId="42" pageNumber="43" type="male">4♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8FFF99FEFCFBC4FC170B10" box="[940,976,1519,1544]" pageId="42" pageNumber="43" type="female">2♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8FFF99FE8DFBC4FBE90B10" box="[989,1070,1519,1544]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="42" pageNumber="43">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8FFF99FD97F83FFAF20B6D" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8B9236E7FF8FFF99FD97F83FFAF20B6D" blockId="42.[151,1437,151,1653]" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FD97F83FFE8F0B35" bold="true" box="[199,328,1556,1581]" pageId="42" pageNumber="43">
<emphasis id="B959EAF5FF8FFF99FD97F83FFE850B35" bold="true" box="[199,322,1556,1581]" italics="true" pageId="42" pageNumber="43">Comments</emphasis>
.
</emphasis>
<taxonomicName id="4C2D4D64FF8FFF99FC03F83FFD900B34" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[339,599,1556,1580]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="umbracatus">
<emphasis id="B959EAF5FF8FFF99FC03F83FFD900B34" box="[339,599,1556,1580]" italics="true" pageId="42" pageNumber="43">Aptilotella umbracatus</emphasis>
</taxonomicName>
has a known range occupying the southerly slopes of the Cordillera de Talamanca in
<collectingCountry id="F33A7677FF8FFF99FC64F813FE4A0B57" box="[308,397,1592,1615]" name="Panama" pageId="42" pageNumber="43">Panama</collectingCountry>
and neighbouring
<collectingCountry id="F33A7677FF8FFF99FF31F81CFD180B48" box="[609,735,1591,1616]" name="Costa Rica" pageId="42" pageNumber="43">Costa Rica</collectingCountry>
. This species is significantly smaller than
<taxonomicName id="4C2D4D64FF8FFF99F9EBF813FA860B48" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[1211,1345,1592,1616]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="B959EAF5FF8FFF99F9EBF813FA860B48" box="[1211,1345,1592,1616]" italics="true" pageId="42" pageNumber="43">A. quadrata</emphasis>
</taxonomicName>
and not as boldly marked. The scutal tomentosity and rudder-like dorsal sclerite of the distiphallus are diagnostic.
</paragraph>
</subSubSection>
</treatment>
</document>