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<document id="4AB22E71A5EA67775F0E58892E0D85D9" ID-CLB-Dataset="5940" ID-DOI="10.11646/zootaxa.3761.1.1" ID-GBIF-Dataset="5444e27e-3e2a-429b-931b-823a2086e3c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909057" ID-ZooBank="82E0F1DC-BC98-4E8A-A3D5-21ECB392CC0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623107451891" checkinUser="felipe" docAuthor="Luk, Stephen P. L. &amp; Marshall, Stephen A." docDate="2014" docId="038487F1FF88FF9DFDC7F8ECFDFA0A55" docLanguage="en" docName="zt03761p156.pdf" docOrigin="Zootaxa 3761 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Aptilotella macula Luk &amp; Marshall 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="47" masterDocId="FFBDFF89FFA5FFB3FD50FE2BFFC70D18" masterDocTitle="A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="46" updateTime="1699002622211" updateUser="ExternalLinkService">
<mods:mods id="9097B023043C558511F9E93ACB362E94" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ECE8CE7A001E3B51D1B4A2B3EC0057D5">
<mods:title id="E24F95A3B823CDC51470AEA52619EE27">A revision of the New World genus Aptilotella Duda (Sphaeroceridae: Limosininae)</mods:title>
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<mods:name id="3ECD62C0E3B4575F61C160D2AB774A2E" type="personal">
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<mods:namePart id="C51AD0280CF8718AC2EAEACF15EE357E">Luk, Stephen P. L.</mods:namePart>
<mods:nameIdentifier id="EAAA99182811375D34E3E01B448D3AF8" type="email">luk@uoguelph.ca</mods:nameIdentifier>
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<mods:namePart id="862479492868C9E82007AE38487219EA">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="898E258EAEB526DF1F61079DB4D1B2D2" type="email">luk@uoguelph.ca</mods:nameIdentifier>
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<mods:title id="56F4D4437EC3DC1DD20721FC682B283F">Zootaxa</mods:title>
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<mods:part id="3ACF2E253975C27F8E02202480B459D3">
<mods:date id="3721306C5F682D4E63A7F416032ACDED">2014</mods:date>
<mods:detail id="12690569C0F05F3D404C5B7D286EE08B" type="pubDate">
<mods:number id="0598E94A06B05295686C0D28F0673D66">2014-02-05</mods:number>
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<mods:number id="A8A0AFAC3D175213BA817B8003129C24">3761</mods:number>
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<mods:number id="572877CF25B3B597D125281385C84051">1</mods:number>
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<mods:classification id="B56EA7504B375D64A3B70B7887E77E52">journal article</mods:classification>
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<mods:identifier id="35584D34C134FEBC66301A2C10F4F50B" type="DOI">10.11646/zootaxa.3761.1.1</mods:identifier>
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<treatment id="038487F1FF88FF9DFDC7F8ECFDFA0A55" ID-DOI="http://doi.org/10.5281/zenodo.4909163" ID-GBIF-Taxon="183203090" ID-Zenodo-Dep="4909163" LSID="urn:lsid:plazi:treatment:038487F1FF88FF9DFDC7F8ECFDFA0A55" httpUri="http://treatment.plazi.org/id/038487F1FF88FF9DFDC7F8ECFDFA0A55" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<subSubSection id="C337656CFF88FF9EFDC7F8ECFD5D0BFA" box="[151,666,1735,1762]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8B9236E7FF88FF9EFDC7F8ECFD5D0BFA" blockId="45.[151,666,1735,1795]" box="[151,666,1735,1762]" pageId="45" pageNumber="46">
<heading id="D0DA818BFF88FF9EFDC7F8ECFD5D0BFA" bold="true" box="[151,666,1735,1762]" fontSize="11" level="2" pageId="45" pageNumber="46" reason="4">
<emphasis id="B959EAF5FF88FF9EFDC7F8ECFD5D0BFA" bold="true" box="[151,666,1735,1762]" pageId="45" pageNumber="46">
<taxonomicName id="4C2D4D64FF88FF9EFDC7F8ECFD8C0BFA" authority="Luk &amp; Marshall" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[151,587,1735,1762]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="B959EAF5FF88FF9EFDC7F8ECFEB20BF9" bold="true" box="[151,373,1735,1761]" italics="true" pageId="45" pageNumber="46">Aptilotella macula</emphasis>
Luk &amp; Marshall
</taxonomicName>
,
<taxonomicNameLabel id="A26A578EFF88FF9EFF08F8E3FD5D0BFA" box="[600,666,1736,1762]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF88FF9DFDC7F8C1FB7B084D" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="8B9236E7FF88FF9EFDC7F8C1FE330A1B" blockId="45.[151,666,1735,1795]" box="[151,500,1770,1795]" pageId="45" pageNumber="46">
<figureCitation id="13162A62FF88FF9EFDC7F8C1FEC70A1B" box="[151,256,1770,1795]" captionStart="FIGURE 1" captionStartId="3.[151,249,871,893]" captionTargetId="figure-234@3.[403,1184,193,850]" captionTargetPageId="3" captionText="FIGURE 1. Aptilotella macula Luk &amp; Marshall, sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.5042233" httpUri="https://zenodo.org/record/5042233/files/figure.png" pageId="45" pageNumber="46">Figures 1</figureCitation>
,
<figureCitation id="13162A62FF88FF9EFC5EF8C1FE910A1B" box="[270,342,1770,1795]" captionStart-0="FIGURES 58–59" captionStart-1="FIGURE 60" captionStartId-0="85.[151,264,1998,2021]" captionStartId-1="86.[151,249,1500,1522]" captionTargetBox-0="[201,1386,193,1978]" captionTargetBox-1="[151,1436,193,1478]" captionTargetId-0="figure-34@85.[201,1386,193,1978]" captionTargetId-1="figure-41@86.[151,1436,193,1478]" captionTargetPageId-0="85" captionTargetPageId-1="86" captionText-0="FIGURES 58–59. Aptilotella macula Luk &amp; Marshall, sp. n., male: 58—habitus, left lateral; 59—habitus, dorsal." captionText-1="FIGURE 60. Aptilotella macula Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SUR = surstylus." figureDoi-0="http://doi.org/10.5281/zenodo.4909177" figureDoi-1="http://doi.org/10.5281/zenodo.4909179" httpUri-0="https://zenodo.org/record/4909177/files/figure.png" httpUri-1="https://zenodo.org/record/4909179/files/figure.png" pageId="45" pageNumber="46">58–60</figureCitation>
, and 337–347
</paragraph>
<paragraph id="8B9236E7FF88FF9DFDC7F919FC150C90" blockId="45.[151,1437,1842,2011]" lastBlockId="46.[151,1437,151,1869]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<emphasis id="B959EAF5FF88FF9EFDC7F919FEE40A53" bold="true" box="[151,291,1842,1867]" italics="true" pageId="45" pageNumber="46">Description.</emphasis>
Habitus as in
<figureCitation id="13162A62FF88FF9EFC93F919FDEB0A53" box="[451,556,1842,1867]" captionStart="FIGURE 1" captionStartId="3.[151,249,871,893]" captionTargetId="figure-234@3.[403,1184,193,850]" captionTargetPageId="3" captionText="FIGURE 1. Aptilotella macula Luk &amp; Marshall, sp. n., holotype male." figureDoi="http://doi.org/10.5281/zenodo.5042233" httpUri="https://zenodo.org/record/5042233/files/figure.png" pageId="45" pageNumber="46">Figures 1</figureCitation>
,
<figureCitation id="13162A62FF88FF9EFF6AF919FD9F0A53" box="[570,600,1842,1867]" captionStart="FIGURES 58–59" captionStartId="85.[151,264,1998,2021]" captionTargetBox="[201,1386,193,1978]" captionTargetId="figure-34@85.[201,1386,193,1978]" captionTargetPageId="85" captionText="FIGURES 58–59. Aptilotella macula Luk &amp; Marshall, sp. n., male: 58—habitus, left lateral; 59—habitus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4909177" httpUri="https://zenodo.org/record/4909177/files/figure.png" pageId="45" pageNumber="46">58</figureCitation>
, and 59. Body length
<quantity id="4CD59B02FF88FF9EFE1DF919FC100A52" box="[845,983,1842,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="45" pageNumber="46" unit="mm" value="1.25" valueMax="1.4" valueMin="1.1">1.1–1.4 mm</quantity>
. Head ground color orange. Frons finely rugose, with five silvery spots: one before each ocular emargination, one diffusely at each hind corner and slightly on occiput, and one on interfrontal plate before tubercle. Ocellar tubercle scarcely raised, with minute medial seta; ocellar bristle greater than the length of frons. Orbital bristle absent; orbital setulae inconspicuous, in four pairs. Interfrontal setae in three pairs. Face shining; lunule with silver spot; gena finely rugose, posterior margin black. Antenna yellow, distal half of first flagellomere brown, scape black. Scutum and scutellum black, with metallic sheen. Scutum uniformly, sparsely setose; posterior dorsocentral bristles as long as scutum. Scutellum bare; three times wider than long, 0.8 times the width of scutum; margins carinate. Scutellar bristles subequal in length. Pleuron black, weakly shining; upper anepisternum and anepimeron with pale pruinose stripe. Legs brown; mid and hind coxae black; front and mid femora and distal half of hind femur orange; hind tarsus light brown; mid tibia with two anterodorsal bristles. Wing rudiment black. Abdomen black, with metallic sheen; tergites setose, densely microtrichose; sternites finely microtrichose. Cercus and surstylus brown.
</paragraph>
<paragraph id="8B9236E7FF8BFF9DFD97FFBFFEB70940" blockId="46.[151,1437,151,1869]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97FFBFFE4A0CB5" bold="true" box="[199,397,404,429]" italics="true" pageId="46" pageNumber="47">Male terminalia.</emphasis>
Sternite 5 (
<figureCitation id="13162A62FF8BFF9DFF4EFFBFFD430CB5" box="[542,644,404,429]" captionStart="FIGURES 337–340" captionStartId="152.[151,264,1266,1289]" captionTargetBox="[370,1212,202,1237]" captionTargetId="figure-44@152.[365,1222,193,1245]" captionTargetPageId="152" captionText="FIGURES 337–340. Aptilotella macula Luk &amp; Marshall, sp. n.: 337—male terminalia, posterior; 338—male terminalia, left lateral; 339—male sternite 5; 340—male synsternite 6+7." figureDoi="http://doi.org/10.5281/zenodo.4909326" httpUri="https://zenodo.org/record/4909326/files/figure.png" pageId="46" pageNumber="47">Fig. 339</figureCitation>
) lunate; posteromedial third weakly dilated, concave and polished, flanked by a patch of setae and four strong marginal setae, medially with four marginal setae. Synsternite 6+7 as in
<figureCitation id="13162A62FF8BFF9DFDC7FFF7FED20CED" box="[151,277,476,501]" captionStart="FIGURES 337–340" captionStartId="152.[151,264,1266,1289]" captionTargetBox="[370,1212,202,1237]" captionTargetId="figure-44@152.[365,1222,193,1245]" captionTargetPageId="152" captionText="FIGURES 337–340. Aptilotella macula Luk &amp; Marshall, sp. n.: 337—male terminalia, posterior; 338—male terminalia, left lateral; 339—male sternite 5; 340—male synsternite 6+7." figureDoi="http://doi.org/10.5281/zenodo.4909326" httpUri="https://zenodo.org/record/4909326/files/figure.png" pageId="46" pageNumber="47">Figure 340</figureCitation>
. Cercus (
<figureCitation id="13162A62FF8BFF9DFC2FFFF7FE1A0CED" box="[383,477,476,501]" captionStart="FIGURE 60" captionStartId="86.[151,249,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure-41@86.[151,1436,193,1478]" captionTargetPageId="86" captionText="FIGURE 60. Aptilotella macula Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SUR = surstylus." figureDoi="http://doi.org/10.5281/zenodo.4909179" httpUri="https://zenodo.org/record/4909179/files/figure.png" pageId="46" pageNumber="47">Figs. 60</figureCitation>
,
<figureCitation id="13162A62FF8BFF9DFCBAFFF7FD880CED" box="[490,591,476,501]" captionStart="FIGURES 337–340" captionStartId="152.[151,264,1266,1289]" captionTargetBox="[370,1212,202,1237]" captionTargetId="figure-44@152.[365,1222,193,1245]" captionTargetPageId="152" captionText="FIGURES 337–340. Aptilotella macula Luk &amp; Marshall, sp. n.: 337—male terminalia, posterior; 338—male terminalia, left lateral; 339—male sternite 5; 340—male synsternite 6+7." figureDoi="http://doi.org/10.5281/zenodo.4909326" httpUri="https://zenodo.org/record/4909326/files/figure.png" pageId="46" pageNumber="47">337, 338</figureCitation>
) stout, triangular, apically truncate; inner margin setulose. Surstylus (
<figureCitation id="13162A62FF8BFF9DF834FFF7FF710F00" captionStart="FIGURE 60" captionStartId="86.[151,249,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure-41@86.[151,1436,193,1478]" captionTargetPageId="86" captionText="FIGURE 60. Aptilotella macula Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SUR = surstylus." figureDoi="http://doi.org/10.5281/zenodo.4909179" httpUri="https://zenodo.org/record/4909179/files/figure.png" pageId="46" pageNumber="47">Figs. 60</figureCitation>
,
<figureCitation id="13162A62FF8BFF9DFD96FFD4FEE80F00" box="[198,303,511,536]" captionStart="FIGURES 337–340" captionStartId="152.[151,264,1266,1289]" captionTargetBox="[370,1212,202,1237]" captionTargetId="figure-44@152.[365,1222,193,1245]" captionTargetPageId="152" captionText="FIGURES 337–340. Aptilotella macula Luk &amp; Marshall, sp. n.: 337—male terminalia, posterior; 338—male terminalia, left lateral; 339—male sternite 5; 340—male synsternite 6+7." figureDoi="http://doi.org/10.5281/zenodo.4909326" httpUri="https://zenodo.org/record/4909326/files/figure.png" pageId="46" pageNumber="47">337, 338</figureCitation>
) basally swollen and divergent, bending at a nearly right angle at midpoint, then rounded and upwardly curved apically; base setulose; inner margin with scattered rows of sensory setae before the bend. Postgonite (
<figureCitation id="13162A62FF8BFF9DFC4FFC6CFEB90F78" box="[287,382,583,608]" captionStart="FIGURE 60" captionStartId="86.[151,249,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure-41@86.[151,1436,193,1478]" captionTargetPageId="86" captionText="FIGURE 60. Aptilotella macula Luk &amp; Marshall, sp. n., male terminalia. Abbreviations: CER = cercus, PG = postgonite, SUR = surstylus." figureDoi="http://doi.org/10.5281/zenodo.4909179" httpUri="https://zenodo.org/record/4909179/files/figure.png" pageId="46" pageNumber="47">Figs. 60</figureCitation>
,
<figureCitation id="13162A62FF8BFF9DFCDDFC6CFE7F0F78" box="[397,440,583,608]" captionStart="FIGURES 341–343" captionStartId="153.[151,264,1134,1157]" captionTargetBox="[436,1149,197,1113]" captionTargetId="figure-82@153.[431,1155,193,1114]" captionTargetPageId="153" captionText="FIGURES 341–343. Aptilotella macula Luk &amp; Marshall, sp. n.: 341—aedeagus with postgonites removed; 342—left postgonite; 343—hypandrium. Abbreviations: BP = basiphallus, CDS = curved dorsal sclerite, LFS = lateral flanking sclerite, MS = membranous sheet, VBA = basal article of ventral flanking sclerite, VBS = ventrobasal sclerite, VMA = medial article of ventral flanking sclerite, VDA = distal article of ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909328" httpUri="https://zenodo.org/record/4909328/files/figure.png" pageId="46" pageNumber="47">342</figureCitation>
) 1.8 times as long as wide; posterior margin medially with large tooth-like projection; preapically with three sensory setae; articulatory processes rounded, each with an internal tooth; margin shallowly concave between the two processes. Hypandrium (
<figureCitation id="13162A62FF8BFF9DFFB6FCA4FC8E0FB0" box="[742,841,655,680]" captionStart="FIGURES 341–343" captionStartId="153.[151,264,1134,1157]" captionTargetBox="[436,1149,197,1113]" captionTargetId="figure-82@153.[431,1155,193,1114]" captionTargetPageId="153" captionText="FIGURES 341–343. Aptilotella macula Luk &amp; Marshall, sp. n.: 341—aedeagus with postgonites removed; 342—left postgonite; 343—hypandrium. Abbreviations: BP = basiphallus, CDS = curved dorsal sclerite, LFS = lateral flanking sclerite, MS = membranous sheet, VBA = basal article of ventral flanking sclerite, VBS = ventrobasal sclerite, VMA = medial article of ventral flanking sclerite, VDA = distal article of ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909328" httpUri="https://zenodo.org/record/4909328/files/figure.png" pageId="46" pageNumber="47">Fig. 343</figureCitation>
) with medial rod pinched in distal three-fifths and apically dilated to the right into a rounded fan; posteromedial fork indistinct but swollen; hypandrial arms irregularly triangular, partially fused to base; pregonite inconspicuous. Aedeagus as in
<figureCitation id="13162A62FF8BFF9DF9DAFCFCFACC0FE8" box="[1162,1291,727,752]" captionStart="FIGURES 341–343" captionStartId="153.[151,264,1134,1157]" captionTargetBox="[436,1149,197,1113]" captionTargetId="figure-82@153.[431,1155,193,1114]" captionTargetPageId="153" captionText="FIGURES 341–343. Aptilotella macula Luk &amp; Marshall, sp. n.: 341—aedeagus with postgonites removed; 342—left postgonite; 343—hypandrium. Abbreviations: BP = basiphallus, CDS = curved dorsal sclerite, LFS = lateral flanking sclerite, MS = membranous sheet, VBA = basal article of ventral flanking sclerite, VBS = ventrobasal sclerite, VMA = medial article of ventral flanking sclerite, VDA = distal article of ventral flanking sclerite." figureDoi="http://doi.org/10.5281/zenodo.4909328" httpUri="https://zenodo.org/record/4909328/files/figure.png" pageId="46" pageNumber="47">Figure 341</figureCitation>
. Basiphallus stout, cylindrical; articulatory process for postgonite short, divergent; articulation with distiphallus dorsally projected. Ventrobasal sclerite weakly divided. Lateral flanking sclerites very narrowly fused ventrobasally; dorsal margin slightly curved, diverging beyond basal two-fifths and descending, then hooked upward and cradling a membranous sheet in between; the sheet laterally lobed and studded with numerous round or elongate sclerotized scales. Ventral flanking sclerites darker; the basal article fused along ventral margin of lateral flanking sclerite, medially bearing a dorsal arm; the triangular medial article articulating with basal article, its inner corner sharply pointed, its top corner rising to just beneath the hooked portion of lateral flanking sclerite; the membranous distal article with a darker, sickle-shaped ascending arm. Curved dorsal sclerites dark, originating from within hooked portion of lateral flanking sclerite, curving inward and converging, their slightly swollen apices protruding between membranous sheet.
</paragraph>
<paragraph id="8B9236E7FF8BFF9DFD97FA4FFE290815" blockId="46.[151,1437,151,1869]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97FA4FFE610965" bold="true" box="[199,422,1124,1149]" italics="true" pageId="46" pageNumber="47">Female terminalia.</emphasis>
Epiproct (
<figureCitation id="13162A62FF8BFF9DFF71FA4FFD000965" box="[545,711,1124,1149]" captionStart="FIGURES 344–347" captionStartId="154.[151,264,1260,1283]" captionTargetBox="[397,1197,199,1232]" captionTargetId="figure-50@154.[384,1202,193,1239]" captionTargetPageId="154" captionText="FIGURES 344–347. Aptilotella macula Luk &amp; Marshall, sp. n.: 344—female terminalia, dorsal; 345—female terminalia, left lateral; 346—female terminalia, ventral; 347—spermathecae. Abbreviations: EP = epiproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909330" httpUri="https://zenodo.org/record/4909330/files/figure.png" pageId="46" pageNumber="47">Figs. 344, 345</figureCitation>
) semicircular. Each half of tergite 8 (
<figureCitation id="13162A62FF8BFF9DF927FA4FFAD90965" box="[1143,1310,1124,1149]" captionStart="FIGURES 344–347" captionStartId="154.[151,264,1260,1283]" captionTargetBox="[397,1197,199,1232]" captionTargetId="figure-50@154.[384,1202,193,1239]" captionTargetPageId="154" captionText="FIGURES 344–347. Aptilotella macula Luk &amp; Marshall, sp. n.: 344—female terminalia, dorsal; 345—female terminalia, left lateral; 346—female terminalia, ventral; 347—spermathecae. Abbreviations: EP = epiproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909330" httpUri="https://zenodo.org/record/4909330/files/figure.png" pageId="46" pageNumber="47">Figs. 344, 345</figureCitation>
) triangular with broadly rounded apex, convex; dorsal half with several long setae. Cercus 2.5 times as long as wide; with two long preapical setae. Sternite 8 (
<figureCitation id="13162A62FF8BFF9DFF54FA87FD6E09DD" box="[516,681,1196,1221]" captionStart="FIGURES 344–347" captionStartId="154.[151,264,1260,1283]" captionTargetBox="[397,1197,199,1232]" captionTargetId="figure-50@154.[384,1202,193,1239]" captionTargetPageId="154" captionText="FIGURES 344–347. Aptilotella macula Luk &amp; Marshall, sp. n.: 344—female terminalia, dorsal; 345—female terminalia, left lateral; 346—female terminalia, ventral; 347—spermathecae. Abbreviations: EP = epiproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909330" httpUri="https://zenodo.org/record/4909330/files/figure.png" pageId="46" pageNumber="47">Figs. 345, 346</figureCitation>
) divided; each half boomerang-shaped, apically microtrichose and with one preapical seta. Spermathecae (
<figureCitation id="13162A62FF8BFF9DFF01FAE4FD7709F0" box="[593,688,1231,1256]" captionStart="FIGURES 344–347" captionStartId="154.[151,264,1260,1283]" captionTargetBox="[397,1197,199,1232]" captionTargetId="figure-50@154.[384,1202,193,1239]" captionTargetPageId="154" captionText="FIGURES 344–347. Aptilotella macula Luk &amp; Marshall, sp. n.: 344—female terminalia, dorsal; 345—female terminalia, left lateral; 346—female terminalia, ventral; 347—spermathecae. Abbreviations: EP = epiproct, S8 = sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4909330" httpUri="https://zenodo.org/record/4909330/files/figure.png" pageId="46" pageNumber="47">Fig. 347</figureCitation>
) finely ridged; sclerotized ducts extremely long, six to seven times the diameter of a spermatheca.
</paragraph>
<paragraph id="8B9236E7FF8BFF9DFD97FB3CFB7B084D" blockId="46.[151,1437,151,1869]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97FB3CFEFE0828" bold="true" box="[199,313,1303,1328]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97FB3CFEF50828" bold="true" box="[199,306,1303,1328]" italics="true" pageId="46" pageNumber="47">Variation</emphasis>
.
</emphasis>
The reflectiveness of the silvery spots varies from faint to very strong. Infrequently, the hind femur is more or fully orange. Teneral specimens have a brownish body but retain the metallic sheen.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8BFF9DFD97FB74FBF20860" box="[199,1077,1375,1400]" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8B9236E7FF8BFF9DFD97FB74FBF20860" blockId="46.[151,1437,151,1869]" box="[199,1077,1375,1400]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97FB74FDD80860" box="[199,543,1375,1400]" italics="true" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97FB74FE800860" bold="true" box="[199,327,1375,1400]" italics="true" pageId="46" pageNumber="47">Etymology.</emphasis>
<taxonomicName id="4C2D4D64FF8BFF9DFC1FFB4BFDD80860" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[335,543,1376,1400]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="macula">Aptilotella macula</taxonomicName>
</emphasis>
is named for the unique patterning of the frons.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8BFF9DFD97FBAFFE980B35" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8B9236E7FF8BFF9DFD97FBAFFC8108D8" blockId="46.[151,1437,151,1869]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97FBAFFEAB0885" bold="true" box="[199,364,1412,1437]" italics="true" pageId="46" pageNumber="47">Type material.</emphasis>
<materialsCitation id="3B453CBAFF8BFF9DFC23FBAFFC8108D8" ID-GBIF-Occurrence="3311767361" collectingDate="2001-04-05" collectingDateMax="2001-04-16" collectingDateMin="2001-04-05" collectionCode="CBFC" collectorName="La Paz &amp; S. A. Marshall." country="Bolivia" county="Cerro Uchumachi" elevation="2550" latitude="-16.211945" location="La Paz" longLatPrecision="21" longitude="-67.71361" municipality="Coroico" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54968845FF8BFF9DFC23FBAFFE1C0885" box="[371,475,1412,1437]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2BFD6EFF8BFF9DFCB2FBAFFE3E0883" box="[482,505,1412,1435]" pageId="46" pageNumber="47" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE22FF8BFF9DFF55FBAFFD970885" box="[517,592,1412,1437]" pageId="46" pageNumber="47">CBFC</collectionCode>
.
<collectingCountry id="F33A7677FF8BFF9DFF0BFBAFFD140885" box="[603,723,1412,1437]" name="Bolivia" pageId="46" pageNumber="47">BOLIVIA</collectingCountry>
:
<collectorName id="26D85331FF8BFF9DFF8DFBAFFCEB0883" box="[733,812,1412,1435]" pageId="46" pageNumber="47">
<location id="8EF2603CFF8BFF9DFF8DFBAFFCEB0883" LSID="urn:lsid:plazi:treatment:038487F1FF88FF9DFDC7F8ECFDFA0A55:8EF2603CFF8BFF9DFF8DFBAFFCEB0883" box="[733,812,1412,1435]" country="Bolivia" county="Cerro Uchumachi" latitude="-16.211945" longLatPrecision="21" longitude="-67.71361" municipality="Coroico" name="La Paz" pageId="46" pageNumber="47">La Paz</location>
</collectorName>
,
<collectingMunicipality id="6BF6AC9DFF8BFF9DFE68FBAFFC520885" box="[824,917,1412,1437]" pageId="46" pageNumber="47">Coroico</collectingMunicipality>
,
<collectingCounty id="62F34E6BFF8BFF9DFEF0FBAFFBAC0885" box="[928,1131,1412,1437]" pageId="46" pageNumber="47">Cerro Uchumachi</collectingCounty>
,
<geoCoordinate id="EE195020FF8BFF9DF926FBAFFB3A0885" box="[1142,1277,1412,1437]" degrees="16" direction="south" minutes="12" orientation="latitude" pageId="46" pageNumber="47" precision="15" seconds="43" value="-16.211945">16°12’43”S</geoCoordinate>
,
<geoCoordinate id="EE195020FF8BFF9DF858FBAFFA500885" box="[1288,1431,1412,1437]" degrees="67" direction="west" minutes="42" orientation="longitude" pageId="46" pageNumber="47" precision="15" seconds="49" value="-67.71361">67°42’49”W</geoCoordinate>
,
<quantity id="4CD59B02FF8BFF9DFDC7FB8CFF2908A7" box="[151,238,1447,1472]" metricMagnitude="3" metricUnit="m" metricValue="2.55" pageId="46" pageNumber="47" unit="m" value="2550.0">
<elevation id="0000D1D4FF8BFF9DFDC7FB8CFF2908A7" box="[151,238,1447,1472]" metricMagnitude="3" metricUnit="m" metricValue="2.55" pageId="46" pageNumber="47" unit="m" value="2550.0">2550 m</elevation>
</quantity>
,
<date id="FF931027FF8BFF9DFDAAFB8CFE4908D8" box="[250,398,1447,1472]" pageId="46" pageNumber="47" value="2001-04-05" valueMax="2001-04-16" valueMin="2001-04-05">
<collectingDate id="EFD7E9CFFF8BFF9DFDAAFB8CFE4908D8" box="[250,398,1447,1472]" pageId="46" pageNumber="47" value="2001-04-05" valueMax="2001-04-16" valueMin="2001-04-05">5–16.iv.2001</collectingDate>
</date>
, elfin forest, dung pans,
<collectorName id="26D85331FF8BFF9DFFF2FB8CFC8108D8" box="[674,838,1447,1472]" pageId="46" pageNumber="47">S.A. Marshall.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9236E7FF8BFF9DFD97FBE7FE980B35" blockId="46.[151,1437,151,1869]" pageId="46" pageNumber="47">
<materialsCitation id="3B453CBAFF8BFF9DFD97FBE7FB3608FD" ID-GBIF-Occurrence="3311767453" box="[199,1265,1484,1509]" collectionCode="DEBU, CBFC" collectorName="La Paz" country="Bolivia" location="BOLIVIA" pageId="46" pageNumber="47" specimenCount="43" specimenCount-female="18" specimenCount-male="25" typeStatus="paratype">
<emphasis id="B959EAF5FF8BFF9DFD97FBE7FEF808FB" bold="true" box="[199,319,1484,1507]" italics="true" pageId="46" pageNumber="47">
<typeStatus id="54968845FF8BFF9DFD97FBE7FEFC08FB" box="[199,315,1484,1507]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F33A7677FF8BFF9DFC1AFBE7FE0508FD" box="[330,450,1484,1509]" name="Bolivia" pageId="46" pageNumber="47">BOLIVIA</collectingCountry>
:
<collectorName id="26D85331FF8BFF9DFC9FFBE7FDE408FB" box="[463,547,1484,1507]" pageId="46" pageNumber="47">La Paz</collectorName>
, same label as
<typeStatus id="54968845FF8BFF9DFF89FBE7FCFC08FD" box="[729,827,1484,1509]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D2BFD6EFF8BFF9DFE1EFBE7FC4508FD" box="[846,898,1484,1509]" pageId="46" pageNumber="47" type="male">21♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8BFF9DFEC2FBE7FC0308FD" box="[914,964,1484,1509]" pageId="46" pageNumber="47" type="female">13♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8BFF9DFE84FBE7FBE308FB" box="[980,1060,1484,1507]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="46" pageNumber="47">DEBU</collectionCode>
;
<specimenCount id="9D2BFD6EFF8BFF9DF963FBE7FB9E08FD" box="[1075,1113,1484,1509]" pageId="46" pageNumber="47" type="male">4♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8BFF9DF937FBE7FB4B08FD" box="[1127,1164,1484,1509]" pageId="46" pageNumber="47" type="female">5♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8BFF9DF9CCFBE7FB2E08FD" box="[1180,1257,1484,1509]" pageId="46" pageNumber="47">CBFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8BFF9DF850FBE7FC1B0B10" ID-GBIF-Occurrence="3311767409" collectingDate="2001-01-28" collectionCode="DEBU" collectorName="R. S. Anderson" country="Bolivia" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
same label as
<typeStatus id="54968845FF8BFF9DFDC7FBC4FF3E0B10" box="[151,249,1519,1544]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
but
<date id="FF931027FF8BFF9DFC7BFBC4FE500B10" box="[299,407,1519,1544]" pageId="46" pageNumber="47" value="2001-01-28">
<collectingDate id="EFD7E9CFFF8BFF9DFC7BFBC4FE500B10" box="[299,407,1519,1544]" pageId="46" pageNumber="47" value="2001-01-28">28.i.2001</collectingDate>
</date>
, cloud forest, leaf litter,
<collectorName id="26D85331FF8BFF9DFFFAFBC4FC880B10" box="[682,847,1519,1544]" pageId="46" pageNumber="47">R.S. Anderson</collectorName>
(
<specimenCount id="9D2BFD6EFF8BFF9DFE0FFBC4FCB10B10" box="[863,886,1519,1544]" pageId="46" pageNumber="47" type="male">♂</specimenCount>
,
<collectionCode id="ED3CAE22FF8BFF9DFED3FBC4FC130B10" box="[899,980,1519,1544]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="46" pageNumber="47">DEBU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B453CBAFF8BFF9DFEB7FBDBFE9C0B35" ID-GBIF-Occurrence="3311767436" collectingDate="2011-04-05" collectingDateMax="2011-04-06" collectingDateMin="2011-04-05" collectionCode="DEBU" country="Bolivia" pageId="46" pageNumber="47" specimenCount="10" specimenCount-female="4" specimenCount-male="6" typeStatus="holotype">
same label as
<typeStatus id="54968845FF8BFF9DF9D4FBC4FB210B10" box="[1156,1254,1519,1544]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
but
<date id="FF931027FF8BFF9DF849FBC4FA5B0B10" box="[1305,1436,1519,1544]" pageId="46" pageNumber="47" value="2011-04-05" valueMax="2011-04-06" valueMin="2011-04-05">
<collectingDate id="EFD7E9CFFF8BFF9DF849FBC4FA5B0B10" box="[1305,1436,1519,1544]" pageId="46" pageNumber="47" value="2011-04-05" valueMax="2011-04-06" valueMin="2011-04-05">5–6.iv.2011</collectingDate>
</date>
(
<specimenCount id="9D2BFD6EFF8BFF9DFDF0F83FFF020B35" box="[160,197,1556,1581]" pageId="46" pageNumber="47" type="male">6♂</specimenCount>
,
<specimenCount id="9D2BFD6EFF8BFF9DFD81F83FFF310B35" box="[209,246,1556,1581]" pageId="46" pageNumber="47" type="female">4♀</specimenCount>
,
<collectionCode id="ED3CAE22FF8BFF9DFC52F83FFE940B35" box="[258,339,1556,1581]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="46" pageNumber="47">DEBU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337656CFF8BFF9DFD97F81CFDFA0A55" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8B9236E7FF8BFF9DFD97F81CFDFA0A55" blockId="46.[151,1437,151,1869]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97F81CFE8F0B48" bold="true" box="[199,328,1591,1616]" pageId="46" pageNumber="47">
<emphasis id="B959EAF5FF8BFF9DFD97F81CFE850B48" bold="true" box="[199,322,1591,1616]" italics="true" pageId="46" pageNumber="47">Comments</emphasis>
.
</emphasis>
<taxonomicName id="4C2D4D64FF8BFF9DFC00F813FDE50B48" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[336,546,1592,1616]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="macula">
<emphasis id="B959EAF5FF8BFF9DFC00F813FDE50B48" box="[336,546,1592,1616]" italics="true" pageId="46" pageNumber="47">Aptilotella macula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D64FF8BFF9DFF0AF813FD780B48" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[602,703,1592,1616]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="macta">
<emphasis id="B959EAF5FF8BFF9DFF0AF813FD780B48" box="[602,703,1592,1616]" italics="true" pageId="46" pageNumber="47">A. macta</emphasis>
</taxonomicName>
are the southernmost members of the genus; their
<typeStatus id="54968845FF8BFF9DF9AEF813FAE90B48" box="[1278,1326,1592,1616]" pageId="46" pageNumber="47">type</typeStatus>
localities in the Bolivian Altiplano are over
<quantity id="4CD59B02FF8BFF9DFF76F877FD540B6D" box="[550,659,1628,1653]" metricMagnitude="6" metricUnit="m" metricValue="2.1" pageId="46" pageNumber="47" unit="km" value="2100.0">2,100 km</quantity>
southeast of the described Ecuadorian
<taxonomicName id="4C2D4D64FF8BFF9DF909F877FAEB0B6D" box="[1113,1324,1628,1653]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B959EAF5FF8BFF9DF909F877FB080B6C" box="[1113,1231,1628,1652]" italics="true" pageId="46" pageNumber="47">Aptilotella</emphasis>
species
</taxonomicName>
, although undescribed species are known from
<collectingCountry id="F33A7677FF8BFF9DFF68F8ABFDA80B8F" box="[568,623,1664,1687]" name="Peru" pageId="46" pageNumber="47">Peru</collectingCountry>
. These two species are probably very recently diverged from each other, and are found just
<quantity id="4CD59B02FF8BFF9DFC39F88FFE770BA5" box="[361,432,1700,1725]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="km" value="50.0">50 km</quantity>
apart. Both are readily recognized by the spotted frons and very shiny metallic and squat body. The cercal chaetotaxy is simple, and the surstyli are boomerang-shaped. The male distiphallus is distinctive for the scale-studded membranous sheet between the lateral flanking sclerites, and the claw-like fused medial and distal articles of the ventral flanking sclerites.
<taxonomicName id="4C2D4D64FF8BFF9DFFF7F93BFCBE0A30" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[679,889,1808,1832]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="macula">
<emphasis id="B959EAF5FF8BFF9DFFF7F93BFCBE0A30" box="[679,889,1808,1832]" italics="true" pageId="46" pageNumber="47">Aptilotella macula</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C2D4D64FF8BFF9DF940F93BFBB20A30" authorityName="Luk &amp; Marshall" authorityYear="2014" box="[1040,1141,1808,1832]" class="Insecta" family="Sphaeroceridae" genus="Aptilotella" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="macta">
<emphasis id="B959EAF5FF8BFF9DF940F93BFBB20A30" box="[1040,1141,1808,1832]" italics="true" pageId="46" pageNumber="47">A. macta</emphasis>
</taxonomicName>
by the five-spotted frons, larger size, and robust male surstylus.
</paragraph>
</subSubSection>
</treatment>
</document>
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