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<document id="EDBEB6D568FB2CCEA51C8B695B7F11A0" ID-CLB-Dataset="6241" ID-DOI="10.11646/zootaxa.4973.1.1" ID-GBIF-Dataset="d0967b08-98d1-4ccd-b472-a678e0e57eda" ID-ISSN="1175-5326" ID-Zenodo-Dep="4771958" ID-ZooBank="42EB4BF2-A571-4894-9EEF-783649A27E4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1621409551899" checkinUser="plazi" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander &amp; Auffenberg, Kurt" docDate="2021" docId="038487E62F0B2416FF12B589FD70FC11" docLanguage="en" docName="zootaxa.4973.1.1.pdf" docOrigin="Zootaxa 4973 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dicharax verrucosus Pall-Gergely &amp; Hunyadi 2021, n. sp." docType="treatment" docVersion="6" lastPageNumber="45" masterDocId="FFBDFF9E2F20243AFF85B51EFF9BFFD8" masterDocTitle="Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)" masterLastPageNumber="61" masterPageNumber="1" pageNumber="44" updateTime="1698940034724" updateUser="ExternalLinkService">
<mods:mods id="3E22B65CAACE12005B43B3C565F747ED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B7FEE7FFAD3A97CB62662DBD8297767E">
<mods:title id="89930D010F0ACF231E5A35B590ADB6AC">Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)</mods:title>
</mods:titleInfo>
<mods:name id="0B51DD4DB1C8559D013698491646E95A" type="personal">
<mods:role id="95105C8A45AB274BC0AFD2D0D88C9886">
<mods:roleTerm id="3C01AA76AB17AD50D13694E84F7D8878">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E62C1C84AC8D9C0356330F1A8B78A684">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="2F796CFC551BFBC07A131CFFB8480F8D" type="personal">
<mods:role id="53985235D0AF479F0C08039066C23CC8">
<mods:roleTerm id="AF4CDE8D54DEE5BD54B32E732B150158">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72E6A0360A045CDB4E151175EB59E7D0">Hunyadi, András</mods:namePart>
<mods:nameIdentifier id="CEF496164DD19368CE70158B6F0EAABE" type="ORCID">0000-0003-3488-2817</mods:nameIdentifier>
<mods:nameIdentifier id="3B7EB532F1991448C11643005D8AF7B0" type="email">hunand@freemail.hu</mods:nameIdentifier>
</mods:name>
<mods:name id="7941E8764808CDE63B332CB1DE86686F" type="personal">
<mods:role id="93CD2ACBEF12007016187A757D58A257">
<mods:roleTerm id="7A7C99CF44371841BB96531CC479D808">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD9C25A15A7701CBFC6FD09D446C7DD7">Grego, Jozef</mods:namePart>
<mods:nameIdentifier id="9C41D0871A8D1CA21BED9B5A5E27DDBF" type="ORCID">0000-0002-4977-0415</mods:nameIdentifier>
<mods:nameIdentifier id="D6A7477EEF55536AB71BC3A036905BB1" type="email">jozef.grego@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="A80AFADF016306C0D145F2BBADB7A78F" type="personal">
<mods:role id="E263C4AB8E3B69741AA3174A11C8CE24">
<mods:roleTerm id="6572983B58B9A49D7960C2AC8AFB16EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="11F9AD5A897F9A5FF86787B0EAF3E07F">Reischütz, Alexander</mods:namePart>
<mods:nameIdentifier id="E9251703978E667C8299CCE69F3F39F2" type="ORCID">0000-0002-8844-7004</mods:nameIdentifier>
<mods:nameIdentifier id="3F71BC49FA5ECBEC3C61AF34040DCB7B" type="email">alexander.reischuetz@gmx.at</mods:nameIdentifier>
</mods:name>
<mods:name id="C98D383178099729E0C3C38F392EBD29" type="personal">
<mods:role id="D1173B77DB719CEC425EC0FAE8FBAE95">
<mods:roleTerm id="0E2BA97FBF716578F3FB5E3525CC7EBB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7A12D005E572C4C527A7F8B09FF29771">Auffenberg, Kurt</mods:namePart>
<mods:nameIdentifier id="EEA7C7343F059812066BECB3FB86AC35" type="ORCID">0000-0001-9660-6776</mods:nameIdentifier>
<mods:nameIdentifier id="8D2A3E3D6B56A1C01E23E0CDFA6C6036" type="email">kurtauffenberg@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B697B6A0EA9A047D0607A8FB845413D2">text</mods:typeOfResource>
<mods:relatedItem id="D6CC76B8B2C02F1305E7D02153B42DFE" type="host">
<mods:titleInfo id="62F84A352219226F5F3548B933E99241">
<mods:title id="33387DA985BDDBB5D2433FD74A07983C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CE5460418ACA54641D758FEAD705C5F1">
<mods:date id="55EBFD50FAE1FAF77F36DAD63F4AC22F">2021</mods:date>
<mods:detail id="80A9AD551572BE74B6F0E54CE8CD70E8" type="pubDate">
<mods:number id="7736948CB0AFC7DB9C636784AE4BEF53">2021-05-18</mods:number>
</mods:detail>
<mods:detail id="77FC23480E4CC46F280729B43B3F3214" type="volume">
<mods:number id="DD0C3293F9583437B270ECF1A1A45BD9">4973</mods:number>
</mods:detail>
<mods:detail id="7E9F65E153D1884343CA534BDFB77F23" type="issue">
<mods:number id="740910CEB3CFFC9107E3B461D9286224">1</mods:number>
</mods:detail>
<mods:extent id="E42CFE2A115301985E708DAC7C8A07E8" unit="page">
<mods:start id="EDA31BDCCF2DB9CD8A6808426AE53647">1</mods:start>
<mods:end id="AE3D3D599A6F87C0B6B6DFE67CB3EC36">61</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1896DCB0B56FE991E191D17EECBEC08A">journal article</mods:classification>
<mods:identifier id="95C7006F4451E6E0433D9966C6956D46" type="CLB-Dataset">6241</mods:identifier>
<mods:identifier id="688532E01A842679F00D743273985FF0" type="DOI">10.11646/zootaxa.4973.1.1</mods:identifier>
<mods:identifier id="6638C56C59CD5962C8D706C1E6D003A3" type="GBIF-Dataset">d0967b08-98d1-4ccd-b472-a678e0e57eda</mods:identifier>
<mods:identifier id="E1C9710DECAE0906484A8CD0F5159DC5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9342C331874FFEA75EC42F61DC2D0299" type="Zenodo-Dep">4771958</mods:identifier>
<mods:identifier id="DE78AC73633B6A5559966E9DD8EF1B29" type="ZooBank">42EB4BF2-A571-4894-9EEF-783649A27E4F</mods:identifier>
</mods:mods>
<treatment id="038487E62F0B2416FF12B589FD70FC11" ID-DOI="http://doi.org/10.5281/zenodo.4916583" ID-GBIF-Taxon="182640526" ID-Zenodo-Dep="4916583" LSID="urn:lsid:plazi:treatment:038487E62F0B2416FF12B589FD70FC11" httpUri="http://treatment.plazi.org/id/038487E62F0B2416FF12B589FD70FC11" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
<subSubSection id="C337657B2F0B2411FF12B589FC83FF69" box="[151,792,150,178]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8B9236F02F0B2411FF12B589FC83FF69" blockId="43.[151,792,150,213]" box="[151,792,150,178]" pageId="43" pageNumber="44">
<heading id="D0DA819C2F0B2411FF12B589FC83FF69" bold="true" box="[151,792,150,178]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="4">
<emphasis id="B959EAE22F0B2411FF12B589FC83FF69" bold="true" box="[151,792,150,178]" pageId="43" pageNumber="44">
<taxonomicName id="4C2D4D732F0B2411FF12B589FD55FF69" authority="Pall-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2021" box="[151,718,150,178]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="verrucosus" status="sp. nov.">
<emphasis id="B959EAE22F0B2411FF12B589FE0EFF69" bold="true" box="[151,405,151,178]" italics="true" pageId="43" pageNumber="44">Dicharax verrucosus</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel id="A26A57992F0B2411FD53B589FC83FF69" box="[726,792,151,177]" pageId="43" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F0B2411FF12B5A5FF7CFF0D" box="[151,231,187,213]" pageId="43" pageNumber="44" type="description">
<paragraph id="8B9236F02F0B2411FF12B5A5FF7CFF0D" blockId="43.[151,792,150,213]" box="[151,231,187,213]" pageId="43" pageNumber="44">
<figureCitation id="13162A752F0B2411FF12B5A5FF7CFF0D" box="[151,231,187,213]" captionStart="FIGURE 34" captionStartId="43.[151,250,1952,1976]" captionTargetBox="[151,1436,489,1928]" captionTargetId="figure-127@43.[151,1436,489,1928]" captionTargetPageId="43" captionText="FIGURE 34. Dicharax verrucosus Páll-Gergely &amp; Hunyadi n. sp., A: holotype (HNHM 104862), B: specimen from sample 2018/40 (coll. HA)." figureDoi="http://doi.org/10.5281/zenodo.4772035" httpUri="https://zenodo.org/record/4772035/files/figure.png" pageId="43" pageNumber="44">Fig. 34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F0B2411FF12B41DFE73FE09" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8B9236F02F0B2411FF12B41DFE73FE09" blockId="43.[151,1437,259,466]" pageId="43" pageNumber="44">
<emphasis id="B959EAE22F0B2411FF12B41DFE18FEC5" bold="true" box="[151,387,259,285]" pageId="43" pageNumber="44">Material examined.</emphasis>
<materialsCitation id="3B453CAD2F0B2411FE09B41DFCF2FEBE" ID-GBIF-Occurrence="3128117356" collectingDate="2018-10-08" collectionCode="HNHM" collectorName="A. Hunyadi &amp; K. Okubo &amp; J. U. Otani" country="Myanmar" county="Taunggyi" elevation="1380" latitude="20.791483" location="Dragon Cave" longLatPrecision="1" longitude="97.0506" municipality="Aye Say Tee Pagoda" pageId="43" pageNumber="44" specimenCode="HNHM 104862" specimenCount="2" stateProvince="Shan" typeStatus="holotype">
<emphasis id="B959EAE22F0B2411FE09B41DFD99FEC5" bold="true" box="[396,514,259,285]" pageId="43" pageNumber="44">
<typeStatus id="549688522F0B2411FE09B41DFE66FEC5" box="[396,509,259,285]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F33A76602F0B2411FD8EB41AFDE1FEC6" box="[523,634,260,286]" name="Myanmar" pageId="43" pageNumber="44">Myanmar</collectingCountry>
,
<collectingRegion id="49E9F8122F0B2411FD02B41DFC98FEC5" box="[647,771,259,285]" country="Myanmar" name="Shan" pageId="43" pageNumber="44">Shan State</collectingRegion>
,
<collectingCounty id="62F34E7C2F0B2411FC95B41DFCE7FEC5" box="[784,892,259,285]" pageId="43" pageNumber="44">Taunggyi</collectingCounty>
, hill above
<collectingMunicipality id="6BF6AC8A2F0B2411FB86B41DFB6AFEC5" box="[1027,1265,259,286]" pageId="43" pageNumber="44">Aye Say Tee Pagoda</collectingMunicipality>
,
<location id="8EF2602B2F0B2411FB7BB41AFA03FEC5" LSID="urn:lsid:plazi:treatment:038487E62F0B2416FF12B589FD70FC11:8EF2602B2F0B2411FB7BB41AFA03FEC5" box="[1278,1432,259,286]" country="Myanmar" county="Taunggyi" latitude="20.791483" longLatPrecision="1" longitude="97.0506" municipality="Aye Say Tee Pagoda" name="Dragon Cave" pageId="43" pageNumber="44" stateProvince="Shan">Dragon Cave</location>
,
<geoCoordinate id="EE1950372F0B2411FF12B439FEB7FE99" box="[151,300,295,321]" degrees="20" direction="north" minutes="47.489" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="20.791483">20°47.489’N</geoCoordinate>
,
<geoCoordinate id="EE1950372F0B2411FEBCB439FE51FE99" box="[313,458,295,321]" degrees="97" direction="east" minutes="03.036" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="97.0506">97°03.036’E</geoCoordinate>
,
<elevation id="0000D1C32F0B2411FE5CB439FDF5FE99" box="[473,622,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="43" pageNumber="44" unit="m" value="1380.0">
<quantity id="4CD59B152F0B2411FE5CB439FDABFE9A" box="[473,560,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="43" pageNumber="44" unit="m" value="1380.0">1380 m</quantity>
a.s.l.
</elevation>
(locality code: 2018/44),
<date id="FF9310302F0B2411FC1FB439FBA8FE99" box="[922,1075,295,321]" pageId="43" pageNumber="44" value="2018-10-08">
<collectingDate id="EFD7E9D82F0B2411FC1FB439FBA8FE99" box="[922,1075,295,321]" pageId="43" pageNumber="44" value="2018-10-08">08 Oct. 2018</collectingDate>
</date>
,
<collectorName id="26D853262F0B2411FBBAB439FB5CFE99" box="[1087,1223,295,321]" pageId="43" pageNumber="44">A. Hunyadi</collectorName>
,
<collectorName id="26D853262F0B2411FB50B436FADDFE99" box="[1237,1350,295,322]" pageId="43" pageNumber="44">K. Okubo</collectorName>
&amp;
<collectorName id="26D853262F0B2411FAEAB436FF4EFEBD" pageId="43" pageNumber="44">J.U. Otani</collectorName>
leg.,
<specimenCode id="DB8B9E8B2F0B2411FE96B452FE58FEBD" box="[275,451,331,358]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="43" pageNumber="44">HNHM 104862</specimenCode>
(
<typeStatus id="549688522F0B2411FE54B455FDA2FEBD" box="[465,569,331,357]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
: D:
<quantity id="4CD59B152F0B2411FDE0B455FD50FEBE" box="[613,715,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="2.83" pageId="43" pageNumber="44" unit="mm" value="2.83">2.83 mm</quantity>
, H:
<quantity id="4CD59B152F0B2411FD7CB455FCFBFEBE" box="[761,864,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="2.14" pageId="43" pageNumber="44" unit="mm" value="2.14">2.14 mm</quantity>
)
</materialsCitation>
.
<materialsCitation id="3B453CAD2F0B2411FCF6B455FA02FEBD" ID-GBIF-Occurrence="3128117311" box="[883,1433,331,358]" collectingDate="2018-10-08" collectionCode="HA" collectorName="A. Hunyadi &amp; K. Okubo &amp; J. U. Otani" country="Myanmar" county="Taunggyi" elevation="1380" latitude="20.791483" location="Dragon Cave" longLatPrecision="1" longitude="97.0506" municipality="Aye Say Tee Pagoda" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Shan" typeStatus="paratype">
<emphasis id="B959EAE22F0B2411FCF6B455FC6FFEBD" bold="true" box="[883,1012,331,357]" pageId="43" pageNumber="44">
<typeStatus id="549688522F0B2411FCF6B455FC74FEBD" box="[883,1007,331,357]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
</emphasis>
10 shells, same data as holotype,
<collectionCode id="ED3CAE352F0B2411FAE8B455FA02FEBD" box="[1389,1433,331,357]" pageId="43" pageNumber="44">HA</collectionCode>
</materialsCitation>
.
<emphasis id="B959EAE22F0B2411FF12B471FECFFE51" bold="true" box="[151,340,367,393]" pageId="43" pageNumber="44">Other material:</emphasis>
<materialsCitation id="3B453CAD2F0B2411FEDEB471FE7FFE09" ID-GBIF-Occurrence="3128117328" collectingDate="2018-10-07" collectionCode="HA" collectorName="A. Hunyadi &amp; K. Okubo &amp; J. U. Otani" country="Myanmar" elevation="400" latitude="20.13305" location="Hsihseng" longLatPrecision="1" longitude="97.302414" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Shan">
2 shells,
<collectingCountry id="F33A76602F0B2411FE3BB46EFDB6FE52" box="[446,557,368,394]" name="Myanmar" pageId="43" pageNumber="44">Myanmar</collectingCountry>
,
<collectingRegion id="49E9F8122F0B2411FDBDB471FD28FE51" box="[568,691,367,393]" country="Myanmar" name="Shan" pageId="43" pageNumber="44">Shan State</collectingRegion>
,
<location id="8EF2602B2F0B2411FD3BB471FCBDFE51" LSID="urn:lsid:plazi:treatment:038487E62F0B2416FF12B589FD70FC11:8EF2602B2F0B2411FD3BB471FCBDFE51" box="[702,806,367,393]" country="Myanmar" latitude="20.13305" longLatPrecision="1" longitude="97.302414" name="Hsihseng" pageId="43" pageNumber="44" stateProvince="Shan">Hsihseng</location>
centre E ca.
<quantity id="4CD59B152F0B2411FC3CB471FC6EFE51" box="[953,1013,367,393]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="43" pageNumber="44" unit="km" value="6.0">6 km</quantity>
, right side of rd. +
<quantity id="4CD59B152F0B2411FB49B471FA88FE52" box="[1228,1299,367,394]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="43" pageNumber="44" unit="m" value="400.0">
<elevation id="0000D1C32F0B2411FB49B471FA88FE52" box="[1228,1299,367,394]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="43" pageNumber="44" unit="m" value="400.0">400 m</elevation>
</quantity>
on unpaved rd., limestone hill,
<geoCoordinate id="EE1950372F0B2411FEE9B48DFE69FE75" box="[364,498,403,429]" degrees="20" direction="north" minutes="7.983" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="20.13305">20°7.983’N</geoCoordinate>
,
<geoCoordinate id="EE1950372F0B2411FE7BB48DFD0BFE76" box="[510,656,403,430]" degrees="97" direction="east" minutes="18.145" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="97.302414">97°18.145’E</geoCoordinate>
,
<elevation id="0000D1C32F0B2411FD1EB48DFCB6FE75" box="[667,813,403,430]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="43" pageNumber="44" unit="m" value="1010.0">
<quantity id="4CD59B152F0B2411FD1EB48DFD6AFE76" box="[667,753,403,430]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="43" pageNumber="44" unit="m" value="1010.0">1010 m</quantity>
a.s.l.
</elevation>
(locality code: 2018/40),
<date id="FF9310302F0B2411FBD5B48DFB7EFE75" box="[1104,1253,403,429]" pageId="43" pageNumber="44" value="2018-10-07">
<collectingDate id="EFD7E9D82F0B2411FBD5B48DFB7EFE75" box="[1104,1253,403,429]" pageId="43" pageNumber="44" value="2018-10-07">07 Oct. 2018</collectingDate>
</date>
,
<collectorName id="26D853262F0B2411FB6AB48DFAEEFE75" box="[1263,1397,403,429]" pageId="43" pageNumber="44">A. Hunyadi</collectorName>
,
<collectorName id="26D853262F0B2411FA04B48AFF78FE09" pageId="43" pageNumber="44">K. Okubo</collectorName>
&amp;
<collectorName id="26D853262F0B2411FE82B4A6FEE2FE09" box="[263,377,439,466]" pageId="43" pageNumber="44">J.U. Otani</collectorName>
leg.,
<collectionCode id="ED3CAE352F0B2411FE3DB4A9FE7FFE09" box="[440,484,439,465]" pageId="43" pageNumber="44">HA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5266782F0B2411FF12B2BEFEFBF804" ID-DOI="http://doi.org/10.5281/zenodo.4772035" ID-Zenodo-Dep="4772035" httpUri="https://zenodo.org/record/4772035/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1952,1976]" targetBox="[151,1436,489,1928]" targetPageId="43">
<paragraph id="8B9236F02F0B2411FF12B2BEFEFBF804" blockId="43.[151,1436,1952,2012]" pageId="43" pageNumber="44">
<emphasis id="B959EAE22F0B2411FF12B2BEFEB9F860" bold="true" box="[151,290,1952,1976]" pageId="43" pageNumber="44">FIGURE 34.</emphasis>
<taxonomicName id="4C2D4D732F0B2411FEACB2BEFC9FF860" authority="Pall-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2021" box="[297,772,1952,1976]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="verrucosus" status="sp. nov.">
<emphasis id="B959EAE22F0B2411FEACB2BEFD9BF860" box="[297,512,1952,1976]" italics="true" pageId="43" pageNumber="44">Dicharax verrucosus</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel id="A26A57992F0B2411FC89B2BFFCDDF860" box="[780,838,1953,1976]" pageId="43" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
, A: holotype (HNHM 104862), B: specimen from sample 2018/40 (coll. HA).
</paragraph>
</caption>
<subSubSection id="C337657B2F0C2416FF42B589FE17FF0D" pageId="44" pageNumber="45" type="etymology">
<paragraph id="8B9236F02F0C2416FF42B589FE17FF0D" blockId="44.[151,1437,151,969]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B589FED5FF69" bold="true" box="[199,334,151,177]" pageId="44" pageNumber="45">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2D4D732F0C2416FDC5B589FD20FF68" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2021" box="[576,699,151,176]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="verrucosus">
<emphasis id="B959EAE22F0C2416FDC5B589FD20FF68" box="[576,699,151,176]" italics="true" pageId="44" pageNumber="45">verrucosus</emphasis>
</taxonomicName>
(Latin for excrescence/wart) refers to the small swelling on R3, reminiscent of a wart.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F0C2416FF42B5C0FD36FEC5" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8B9236F02F0C2416FF42B5C0FD36FEC5" blockId="44.[151,1437,151,969]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B5C0FED9FF20" bold="true" box="[199,322,222,248]" pageId="44" pageNumber="45">Diagnosis.</emphasis>
A
<taxonomicName id="4C2D4D732F0C2416FEE6B5C1FDBDFF21" box="[355,550,223,249]" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B959EAE22F0C2416FEE6B5C1FE56FF21" box="[355,461,223,249]" italics="true" pageId="44" pageNumber="45">Dicharax</emphasis>
species
</taxonomicName>
with a relatively high spire, glossy, smooth R1, R2+R3 about a quarter whorl (R3 shorter than R2), and blunt swelling on R3.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F0C2416FF42B438FE4FFD15" pageId="44" pageNumber="45" type="description">
<paragraph id="8B9236F02F0C2416FF42B438FB3AFD5D" blockId="44.[151,1437,151,969]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B438FEC2FE98" bold="true" box="[199,345,294,320]" pageId="44" pageNumber="45">Description.</emphasis>
Shell light grey, possibly translucent when fresh; shell outline rounded in dorsal view, spire elevated, body whorl rounded; protoconch low, rather glossy, 1.5 whorls; R1 approximately 2.25 whorls, smooth, glossy, with occasional, weak growth lines; boundary between R1 and R2 conspicuous due to dense R2 ribs; R2 ribs curved toward aperture, ca. 36–38 R2 ribs present; R2 ribs situated close to each other, spaces between ribs much less than rib width; R2+R3 covers ca. quarter whorl; R3 very short; about half as long as R2; sculpture of R3 similar to R1, boundary between R2 and R3 distinct, indicated by deep constriction; middle of R3 with prominent, blunt swelling; aperture slightly oblique to shell axis; rounded, with slight upper incision in parieto-palatal region; boundary between inner and outer peristomes distinct, especially at columellar portion; inner peristome slim, slightly protruding, outer peristome strongly thickened, expanded and reflected, especially toward umbilicus (in sample 2018/40 more conspicuously than in 2018/44); umbilicus relatively narrow, less than a third of shell width.
</paragraph>
<paragraph id="8B9236F02F0C2416FF42B791FA07FD71" blockId="44.[151,1437,151,969]" box="[199,1436,654,681]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B791FEE5FD71" bold="true" box="[199,382,655,681]" pageId="44" pageNumber="45">Measurements.</emphasis>
D:
<quantity id="4CD59B152F0C2416FE22B791FDD7FD71" box="[423,588,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.24" metricValueMin="2.76" pageId="44" pageNumber="45" unit="mm" value="3.0" valueMax="3.24" valueMin="2.76">2.76–3.24 mm</quantity>
, H:
<quantity id="4CD59B152F0C2416FDFEB791FC8BFD71" box="[635,784,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" metricValueMax="2.46" metricValueMin="2.1" pageId="44" pageNumber="45" unit="mm" value="2.28" valueMax="2.46" valueMin="2.1">2.1–2.46 mm</quantity>
(2018/44); D:
<quantity id="4CD59B152F0C2416FC32B790FBC7FD71" box="[951,1116,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="3.38" metricValueMax="3.55" metricValueMin="3.21" pageId="44" pageNumber="45" unit="mm" value="3.38" valueMax="3.55" valueMin="3.21">3.21–3.55 mm</quantity>
, H:
<quantity id="4CD59B152F0C2416FB0FB790FA84FD71" box="[1162,1311,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="2.48" metricValueMax="2.5" metricValueMin="2.46" pageId="44" pageNumber="45" unit="mm" value="2.48" valueMax="2.5" valueMin="2.46">2.46–2.5 mm</quantity>
(2018/40).
</paragraph>
<paragraph id="8B9236F02F0C2416FF42B7ACFE4FFD15" blockId="44.[151,1437,151,969]" box="[199,468,690,717]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B7ACFECCFD14" bold="true" box="[199,343,690,716]" pageId="44" pageNumber="45">Operculum.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F0C2416FF42B7C8FD0AFCCD" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8B9236F02F0C2416FF42B7C8FD0AFCCD" blockId="44.[151,1437,151,969]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B7C8FE55FD28" bold="true" box="[199,462,726,752]" pageId="44" pageNumber="45">Differential diagnosis.</emphasis>
The combination of the elevated spire, smooth R2, short R3 with a prominent swelling distinguishes this species from all congeners.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F0C2416FF42B600FD7AFC85" pageId="44" pageNumber="45" type="distribution">
<paragraph id="8B9236F02F0C2416FF42B600FD7AFC85" blockId="44.[151,1437,151,969]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B600FEFAFCE0" bold="true" box="[199,353,798,824]" pageId="44" pageNumber="45">Distribution.</emphasis>
Both known populations occur in the southeastern part of
<collectingRegion id="49E9F8122F0C2416FC7BB601FBADFCE1" box="[1022,1078,799,825]" country="Myanmar" name="Shan" pageId="44" pageNumber="45">Shan</collectingRegion>
States,
<collectingCountry id="F33A76602F0C2416FB14B601FA9BFCE1" box="[1169,1280,799,825]" name="Myanmar" pageId="44" pageNumber="45">Myanmar</collectingCountry>
. The straightline distance between the two is ca.
<quantity id="4CD59B152F0C2416FDAFB65DFDEAFC85" box="[554,625,835,861]" metricMagnitude="4" metricUnit="m" metricValue="7.7" pageId="44" pageNumber="45" unit="km" value="77.0">77 km</quantity>
(
<figureCitation id="13162A752F0C2416FD05B65DFD48FC85" box="[640,723,835,861]" captionStart="FIGURE 17" captionStartId="23.[151,250,1861,1885]" captionTargetBox="[202,1386,1025,1838]" captionTargetId="figure-32@23.[202,1386,1025,1838]" captionTargetPageId="23" captionText="FIGURE 17. Distribution of alycaeid species in northern Thailand and Myanmar. filled circle: Dicharax admirandus Páll- Gergely &amp; Auffenberg, n. sp., empty square: Dicharax ebrius Páll-Gergely &amp; Auffenberg, n. sp., filled triangle: Dicharax gnomus Páll-Gergely, n. sp., filled square: Dicharax mirounga Páll-Gergely, n. sp., inverted empty triangles: Dicharax verruco- sus Páll-Gergely &amp; Hunyadi n. sp., rhombus: Dicharax stuparum Páll-Gergely &amp; Hunyadi, 2018, empty triangle: Metalycaeus anas Páll-Gergely &amp; A. Reischütz, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4771996" httpUri="https://zenodo.org/record/4771996/files/figure.png" pageId="44" pageNumber="45">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F0C2416FF42B679FD70FC11" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8B9236F02F0C2416FF42B679FD70FC11" blockId="44.[151,1437,151,969]" pageId="44" pageNumber="45">
<emphasis id="B959EAE22F0C2416FF42B679FEA0FC59" bold="true" box="[199,315,871,897]" pageId="44" pageNumber="45">Remarks.</emphasis>
The typical shells from Taunggyi are slightly smaller, and have less reflected peristomes than specimens from Hsihseng. However, since all other shell characters (shape, ratios of the shell regions, and shell sculpture) are identical, we treat them as the same species.
</paragraph>
</subSubSection>
</treatment>
</document>
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