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<document id="613AC1E6D977C5D6F2A8DF8CA750E2E9" ID-CLB-Dataset="4095" ID-DOI="10.5852/ejt.2021.764.1477" ID-GBIF-Dataset="e9baa250-ec84-440f-96bf-06b55f6738e3" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514464" ID-ZooBank="8B5799F9-487D-4A9F-BF9F-54ADA06056FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1631926753962" checkinUser="felipe" docAuthor="Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan &amp; Yang, Ding" docDate="2021" docId="038487C8FF8CFFA7FD8FFB65FD59FEE8" docLanguage="en" docName="ejt-764_152-192.pdf" docOrigin="European Journal of Taxonomy 764 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.764.1477" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tipula (Vestiplex) motuoensis Yang &amp; Pan &amp; Liu &amp; Yang 2021, sp. nov." docType="treatment" docUuid="6536F0F9-0D40-490C-BEF5-06A3A1153033" docUuidSource="ZooBank" docVersion="8" lastPageNumber="188" masterDocId="FFBDFFB0FFACFF81FFBAFFA1FFE6FF82" masterDocTitle="Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet" masterLastPageNumber="192" masterPageNumber="152" pageNumber="184" updateTime="1699449403584" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A09B65C67FEEBB0C8D3EFE93689332C3">Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet</mods:title>
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<mods:namePart id="03DF2C808176C47A1CF11C8EFF43560C">Yang, Qi-cheng</mods:namePart>
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<mods:affiliation id="2A5B9FB2578144314DE4C51CB817CBE4">Hubei Insect Resources Utilization and Sustainable Pest Management Key Laboratory, College of Plant Science &amp; Technology of Huazhong Agriculture University, Wuhan, Hubei, 430070, China. Plant Protection College, Shenyang Agricultural University, Shenyang, Liaoning, 110866, China. Institute of Plateau Ecology, Tibet Agriculture &amp; Animal Husbandry University, Linzhi, Xizang, P. R. 860000, China. China Agricultural University, Beijing, 100193, China.</mods:affiliation>
<mods:nameIdentifier id="7EC9D7796DBA84FCC970BB625D1D7A27" type="email">qichengyang@foxmail.com&amp;liyan_syau@163.com</mods:nameIdentifier>
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<mods:namePart id="D4A705A8563E2477FE34A34D009A1F4A">Pan, Zhao-hui</mods:namePart>
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<mods:nameIdentifier id="979C36B8DAB0FD5BE7B36296D9A4F118" type="email">panzhaohui2005@163.com</mods:nameIdentifier>
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<mods:namePart id="5E3512EAEE7C86630144CE1A419D734C">Liu, Xiao-yan</mods:namePart>
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<mods:affiliation id="4A0CFE857C36381D2BCB621FEDC91118">Hubei Insect Resources Utilization and Sustainable Pest Management Key Laboratory, College of Plant Science &amp; Technology of Huazhong Agriculture University, Wuhan, Hubei, 430070, China. Plant Protection College, Shenyang Agricultural University, Shenyang, Liaoning, 110866, China. Institute of Plateau Ecology, Tibet Agriculture &amp; Animal Husbandry University, Linzhi, Xizang, P. R. 860000, China. China Agricultural University, Beijing, 100193, China.</mods:affiliation>
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<mods:namePart id="DBFC208E835ED0D206180C3820A4F352">Yang, Ding</mods:namePart>
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<mods:title id="D0D608C1BA5F03FF42EBD116EC6F877A">European Journal of Taxonomy</mods:title>
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<treatment id="038487C8FF8CFFA7FD8FFB65FD59FEE8" ID-DOI="http://doi.org/10.5281/zenodo.5514513" ID-GBIF-Taxon="188606732" ID-Zenodo-Dep="5514513" LSID="urn:lsid:zoobank.org:act:6536F0F9-0D40-490C-BEF5-06A3A1153033" httpUri="http://treatment.plazi.org/id/038487C8FF8CFFA7FD8FFB65FD59FEE8" lastPageId="38" lastPageNumber="188" pageId="32" pageNumber="184">
<subSubSection id="C3376555FF8CFFA1FD8FFB65FC18FB5C" box="[565,1022,1220,1247]" pageId="32" pageNumber="184" type="nomenclature">
<paragraph id="8B9236DEFF8CFFA1FD8FFB65FC18FB5C" blockId="32.[374,1213,1220,1317]" box="[565,1022,1220,1247]" pageId="32" pageNumber="184">
<heading id="D0DA81B2FF8CFFA1FD8FFB65FC18FB5C" box="[565,1022,1220,1247]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="184" reason="2">
<taxonomicName id="4C2D4D5DFF8CFFA1FD8FFB65FC7DFB5C" authority="Yang &amp; Pan &amp; Liu &amp; Yang, 2021" authorityName="Yang &amp; Pan &amp; Liu &amp; Yang" authorityYear="2021" box="[565,923,1220,1247]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="184" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="B959EACCFF8CFFA1FD8FFB65FC7DFB5C" bold="true" box="[565,923,1220,1247]" italics="true" pageId="32" pageNumber="184">Tipula (Vestiplex) motuoensis</emphasis>
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<taxonomicNameLabel id="A26A57B7FF8CFFA1FC18FB64FC18FB5C" box="[930,1022,1221,1247]" pageId="32" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3376555FF8CFFA1FECCFB46FCBDFAA7" pageId="32" pageNumber="184" type="description">
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urn:lsid:zoobank.org:act:
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<paragraph id="8B9236DEFF8CFFA1FD62FAABFCBDFAA7" blockId="32.[374,1213,1220,1317]" box="[728,859,1290,1317]" pageId="32" pageNumber="184">
<figureCitation id="13162A5BFF8CFFA1FD62FAABFCBDFAA7" box="[728,859,1290,1317]" captionStart-0="Figs 74–76" captionStart-1="Figs 77–78" captionStart-2="Figs 79–83" captionStartId-0="33.[189,242,1922,1948]" captionStartId-1="34.[189,242,1954,1980]" captionStartId-2="35.[189,242,1888,1914]" captionTargetBox-0="[260,1330,289,1812]" captionTargetBox-1="[290,1329,290,1893]" captionTargetBox-2="[255,1300,294,1823]" captionTargetId-0="figure-16@33.[243,1344,265,1884]" captionTargetId-1="figure-12@34.[232,1355,265,1917]" captionTargetId-2="figure-15@35.[255,1332,265,1850]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionText-0="Figs 74–76. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 74. Habitus, lateral view. 75. Male habitus, dorsal view. 76. Wing. Abbreviations: see Material and methods. Scale bars = 1.0 mm." captionText-1="Figs 77–78. Tipula (Vestiplex) motuoensis sp. nov., allotype, ♀ (CAU). 77. Habitus, lateral view. 78. Ovipositor, lateral view. Scale bars = 1.0 mm." captionText-2="Figs 79–83. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 79. Hypopygium, lateral view. 80–81. Outer and inner gonostylus, lateral external view and lateral dorsal view. 82. Tergite nine, dorsal view. 83. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 79, 82–83 = 0.5 mm; 80–81 = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5514515" figureDoi-1="http://doi.org/10.5281/zenodo.5514517" figureDoi-2="http://doi.org/10.5281/zenodo.5514521" httpUri-0="https://zenodo.org/record/5514515/files/figure.png" httpUri-1="https://zenodo.org/record/5514517/files/figure.png" httpUri-2="https://zenodo.org/record/5514521/files/figure.png" pageId="32" pageNumber="184">Figs 75–83</figureCitation>
</paragraph>
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<subSubSection id="C3376555FF8CFFA1FF07FAEAFB55F9C2" pageId="32" pageNumber="184" type="diagnosis">
<paragraph id="8B9236DEFF8CFFA1FF07FAEAFED0FAE7" blockId="32.[189,310,1355,1381]" box="[189,310,1355,1381]" pageId="32" pageNumber="184">
<heading id="D0DA81B2FF8CFFA1FF07FAEAFED0FAE7" bold="true" box="[189,310,1355,1381]" fontSize="11" level="3" pageId="32" pageNumber="184" reason="3">
<emphasis id="B959EACCFF8CFFA1FF07FAEAFED0FAE7" bold="true" box="[189,310,1355,1381]" pageId="32" pageNumber="184">Diagnosis</emphasis>
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<paragraph id="8B9236DEFF8CFFA1FF07FAD8FB55F9C2" blockId="32.[187,1399,1401,1601]" pageId="32" pageNumber="184">Proboscis relatively short. Pronotum brownish yellow with dark yellow setae. Prescutum mainly reddish yellow with four olive-brown striped pruinescence, velvety, two middle stripes virtually fused. Gonocoxite ends hook-like, not separated from sternite 9; tergite 9 with two pairs of lobes, longitudinally divided into two parts, slightly connected, pair of emarginate arms arising from ventral surface of tergite 9. Inner gonostylus with posterior crest produced backward, base of inner gonostylus and outer basal lobe of inner gonostylus rolled inward, produced depression with row of short setulae.</paragraph>
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<subSubSection id="C3376555FF8CFFA1FF07F9C5FCF0F92E" box="[189,790,1636,1708]" pageId="32" pageNumber="184" type="etymology">
<paragraph id="8B9236DEFF8CFFA1FF07F9C5FEA2F9FC" blockId="32.[189,324,1636,1662]" box="[189,324,1636,1662]" pageId="32" pageNumber="184">
<heading id="D0DA81B2FF8CFFA1FF07F9C5FEA2F9FC" bold="true" box="[189,324,1636,1662]" fontSize="11" level="3" pageId="32" pageNumber="184" reason="3">
<emphasis id="B959EACCFF8CFFA1FF07F9C5FEA2F9FC" bold="true" box="[189,324,1636,1662]" pageId="32" pageNumber="184">Etymology</emphasis>
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The species is named after the
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locality Motuo.
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<heading id="D0DA81B2FF8CFFA1FF07F96EFE4CF96B" bold="true" box="[189,426,1743,1769]" fontSize="11" level="3" pageId="32" pageNumber="184" reason="3">
<emphasis id="B959EACCFF8CFFA1FF07F96EFE4CF96B" bold="true" box="[189,426,1743,1769]" pageId="32" pageNumber="184">Material examined</emphasis>
</heading>
</paragraph>
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<materialsCitation id="3B453C83FF8CFFA1FF50F95CFE5EF8DE" ID-GBIF-Occurrence="3382544301" collectingDate="2017-06-13" collectingMethod="light trap" collectionCode="CAU" collectorName="Qicheng Yang" country="China" elevation="2020" latitude="29.658054" location="Motuo" longLatPrecision="20" longitude="95.49" pageId="32" pageNumber="184" specimenCount="1" specimenCount-male="1" stateProvince="Tibet" typeStatus="holotype">
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<typeStatus id="5496887CFF8CFFA1FF50F95CFEBAF895" box="[234,348,1789,1815]" pageId="32" pageNumber="184" type="holotype">Holotype</typeStatus>
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<collectingCountry id="F33A764EFF8CFFA1FF07F881FEFEF8B8" box="[189,280,1824,1850]" name="China" pageId="32" pageNumber="184">CHINA</collectingCountry>
•
<specimenCount id="9D2BFD57FF8CFFA1FE8BF881FEACF8B8" box="[305,330,1824,1850]" pageId="32" pageNumber="184" type="male">♂</specimenCount>
;
<collectingRegion id="49E9F83CFF8CFFA1FEEDF8BEFE71F8BB" box="[343,407,1823,1849]" country="China" name="Xizang" pageId="32" pageNumber="184">Tibet</collectingRegion>
,
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,
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,
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;
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a.s.l.
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;
<collectorName id="26D85308FF8CFFA1FB6AF8BEFA91F8B8" box="[1232,1399,1823,1850]" pageId="32" pageNumber="184">Qicheng Yang</collectorName>
leg.;
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;
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.
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<materialsCitation id="3B453C83FF8CFFA1FF50F8DEFCB5F83E" ID-GBIF-Occurrence="3382544311" collectingDate="2017-06-13" collectingMethod="light trap" collectionCode="CAU" collectorName="Qicheng Yang" country="China" elevation="2020" latitude="29.658054" location="CHINA" longLatPrecision="20" longitude="95.49" pageId="32" pageNumber="184" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="allotype">
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<typeStatus id="5496887CFF8CFFA1FF50F8DEFEB2F81B" box="[234,340,1919,1945]" pageId="32" pageNumber="184" type="allotype">Allotype</typeStatus>
</emphasis>
<collectingCountry id="F33A764EFF8CFFA1FF07F803FEFEF83E" box="[189,280,1954,1980]" name="China" pageId="32" pageNumber="184">CHINA</collectingCountry>
•
<specimenCount id="9D2BFD57FF8CFFA1FE8BF802FEAEF83F" box="[305,328,1955,1981]" pageId="32" pageNumber="184" type="female">♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED3CAE1BFF8CFFA1FCB7F803FCA9F83E" box="[781,847,1954,1980]" collectionName="China, Beijing, China Agricultural University" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="32" pageNumber="184">CAU</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="DF526656FF8DFFA0FF07F823FA91F842" ID-DOI="http://doi.org/10.5281/zenodo.5514515" ID-Zenodo-Dep="5514515" httpUri="https://zenodo.org/record/5514515/files/figure.png" pageId="33" pageNumber="185" startId="33.[189,242,1922,1948]" targetBox="[260,1330,289,1812]" targetPageId="33">
<paragraph id="8B9236DEFF8DFFA0FF07F823FA91F842" blockId="33.[189,1399,1922,1984]" pageId="33" pageNumber="185">
<emphasis id="B959EACCFF8DFFA0FF07F823FEA9F81E" bold="true" box="[189,335,1922,1948]" pageId="33" pageNumber="185">Figs 74–76.</emphasis>
<taxonomicName id="4C2D4D5DFF8DFFA0FEE6F823FD25F81E" authority="Yang &amp; Pan &amp; Liu &amp; Yang, 2021" authorityName="Yang &amp; Pan &amp; Liu &amp; Yang" authorityYear="2021" box="[348,707,1922,1948]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="185" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="B959EACCFF8DFFA0FEE6F823FE4EF81E" box="[348,424,1922,1948]" italics="true" pageId="33" pageNumber="185">Tipula</emphasis>
(
<emphasis id="B959EACCFF8DFFA0FE04F823FDC1F81E" box="[446,551,1922,1948]" italics="true" pageId="33" pageNumber="185">Vestiplex</emphasis>
)
<emphasis id="B959EACCFF8DFFA0FD84F822FD25F81E" box="[574,707,1923,1948]" italics="true" pageId="33" pageNumber="185">motuoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57B7FF8DFFA0FD75F823FCD4F81E" box="[719,818,1922,1948]" pageId="33" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (CAU).
<emphasis id="B959EACCFF8DFFA0FBF9F823FB86F81E" bold="true" box="[1091,1120,1922,1948]" pageId="33" pageNumber="185">74</emphasis>
. Habitus, lateral view.
<emphasis id="B959EACCFF8DFFA0FF07F804FF3CF83D" bold="true" box="[189,218,1957,1983]" pageId="33" pageNumber="185">75</emphasis>
. Male habitus, dorsal view.
<emphasis id="B959EACCFF8DFFA0FDA5F804FDDAF83D" bold="true" box="[543,572,1957,1983]" pageId="33" pageNumber="185">76</emphasis>
. Wing. Abbreviations: see Material and methods. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF526656FF8EFFA3FF07F803FCE5F862" ID-DOI="http://doi.org/10.5281/zenodo.5514517" ID-Zenodo-Dep="5514517" httpUri="https://zenodo.org/record/5514517/files/figure.png" pageId="34" pageNumber="186" startId="34.[189,242,1954,1980]" targetBox="[290,1329,290,1893]" targetPageId="34">
<paragraph id="8B9236DEFF8EFFA3FF07F803FCE5F862" blockId="34.[189,1399,1954,2016]" pageId="34" pageNumber="186">
<emphasis id="B959EACCFF8EFFA3FF07F803FEB6F83E" bold="true" box="[189,336,1954,1980]" pageId="34" pageNumber="186">Figs 77–78.</emphasis>
<taxonomicName id="4C2D4D5DFF8EFFA3FEE7F803FD23F83E" authority="Yang &amp; Pan &amp; Liu &amp; Yang, 2021" authorityName="Yang &amp; Pan &amp; Liu &amp; Yang" authorityYear="2021" box="[349,709,1954,1981]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="186" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="B959EACCFF8EFFA3FEE7F803FE4FF83E" box="[349,425,1954,1980]" italics="true" pageId="34" pageNumber="186">Tipula</emphasis>
(
<emphasis id="B959EACCFF8EFFA3FE7AF803FDCFF83E" box="[448,553,1954,1980]" italics="true" pageId="34" pageNumber="186">Vestiplex</emphasis>
)
<emphasis id="B959EACCFF8EFFA3FDFAF802FD23F83E" box="[576,709,1955,1980]" italics="true" pageId="34" pageNumber="186">motuoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57B7FF8EFFA3FD68F802FCD3F83F" box="[722,821,1955,1981]" pageId="34" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
, allotype, ♀ (CAU).
<emphasis id="B959EACCFF8EFFA3FBFBF802FBB8F83F" bold="true" box="[1089,1118,1955,1981]" pageId="34" pageNumber="186">77</emphasis>
. Habitus, lateral view.
<emphasis id="B959EACCFF8EFFA3FF07F864FF3CF85D" bold="true" box="[189,218,1989,2015]" pageId="34" pageNumber="186">78</emphasis>
. Ovipositor, lateral view. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF526656FF8FFFA2FF07F8C1FE16F866" ID-DOI="http://doi.org/10.5281/zenodo.5514521" ID-Zenodo-Dep="5514521" httpUri="https://zenodo.org/record/5514521/files/figure.png" pageId="35" pageNumber="187" startId="35.[189,242,1888,1914]" targetBox="[255,1300,294,1823]" targetPageId="35">
<paragraph id="8B9236DEFF8FFFA2FF07F8C1FE16F866" blockId="35.[188,1399,1888,2020]" pageId="35" pageNumber="187">
<emphasis id="B959EACCFF8FFFA2FF07F8C1FEACF8F8" bold="true" box="[189,330,1888,1914]" pageId="35" pageNumber="187">Figs 79–83.</emphasis>
<taxonomicName id="4C2D4D5DFF8FFFA2FEE8F8C1FD49F8F8" authority="Yang &amp; Pan &amp; Liu &amp; Yang, 2021" authorityName="Yang &amp; Pan &amp; Liu &amp; Yang" authorityYear="2021" box="[338,687,1888,1914]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="187" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="B959EACCFF8FFFA2FEE8F8C1FE78F8F8" box="[338,414,1888,1914]" italics="true" pageId="35" pageNumber="187">Tipula</emphasis>
(
<emphasis id="B959EACCFF8FFFA2FE15F8C1FDFEF8F8" box="[431,536,1888,1914]" italics="true" pageId="35" pageNumber="187">Vestiplex</emphasis>
)
<emphasis id="B959EACCFF8FFFA2FD90F8C0FD49F8F8" box="[554,687,1889,1914]" italics="true" pageId="35" pageNumber="187">motuoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57B7FF8FFFA2FD0DF8C1FCF2F8F8" box="[695,788,1888,1914]" pageId="35" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (CAU).
<emphasis id="B959EACCFF8FFFA2FBA8F8C1FBC9F8F8" bold="true" box="[1042,1071,1888,1914]" pageId="35" pageNumber="187">79</emphasis>
. Hypopygium, lateral view.
<emphasis id="B959EACCFF8FFFA2FF07F822FEE0F81F" bold="true" box="[189,262,1923,1950]" pageId="35" pageNumber="187">80–81</emphasis>
. Outer and inner gonostylus, lateral external view and lateral dorsal view.
<emphasis id="B959EACCFF8FFFA2FBD0F822FB61F81F" bold="true" box="[1130,1159,1923,1949]" pageId="35" pageNumber="187">82</emphasis>
. Tergite nine, dorsal view.
<emphasis id="B959EACCFF8FFFA2FEB8F807FEF9F842" bold="true" box="[258,287,1958,1984]" pageId="35" pageNumber="187">83</emphasis>
. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 79, 82–83 = 0.5 mm; 80–81 = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B9236DEFF88FFA5FF50FEA8FC8FFEC7" blockId="36.[189,873,265,326]" pageId="36" pageNumber="188">
<materialsCitation id="3B453C83FF88FFA5FF50FEA8FC83FEC7" ID-GBIF-Occurrence="3382544306" collectingDate="2017-06-13" collectingMethod="light trap" collectionCode="CAU" collectorName="Qicheng Yang" country="China" elevation="2020" latitude="29.658054" location="CHINA" longLatPrecision="20" longitude="95.49" pageId="36" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="B959EACCFF88FFA5FF50FEA8FEBAFEA1" bold="true" box="[234,348,265,291]" pageId="36" pageNumber="188">
<typeStatus id="5496887CFF88FFA5FF50FEA8FEBAFEA1" box="[234,348,265,291]" pageId="36" pageNumber="188" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F33A764EFF88FFA5FF07FE8AFEFEFEC7" box="[189,280,299,325]" name="China" pageId="36" pageNumber="188">CHINA</collectingCountry>
•
<specimenCount id="9D2BFD57FF88FFA5FE8BFE8AFE86FEC4" box="[305,352,299,326]" pageId="36" pageNumber="188" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED3CAE1BFF88FFA5FC99FE8AFC83FEC7" box="[803,869,299,325]" collectionName="China, Beijing, China Agricultural University" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="36" pageNumber="188">CAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376555FF88FFA5FF07FEC9FD46F926" pageId="36" pageNumber="188" type="description">
<paragraph id="8B9236DEFF88FFA5FF07FEC9FEA8FE00" blockId="36.[189,334,360,386]" box="[189,334,360,386]" pageId="36" pageNumber="188">
<heading id="D0DA81B2FF88FFA5FF07FEC9FEA8FE00" bold="true" box="[189,334,360,386]" fontSize="11" level="3" pageId="36" pageNumber="188" reason="3">
<emphasis id="B959EACCFF88FFA5FF07FEC9FEA8FE00" bold="true" box="[189,334,360,386]" pageId="36" pageNumber="188">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF50FE36FECCFE33" blockId="36.[189,1280,407,468]" box="[234,298,407,433]" pageId="36" pageNumber="188">
<emphasis id="B959EACCFF88FFA5FF50FE36FECCFE33" bold="true" box="[234,298,407,433]" pageId="36" pageNumber="188">Male</emphasis>
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07FE1BFAE6FE51" blockId="36.[189,1280,407,468]" box="[189,1280,441,468]" pageId="36" pageNumber="188">MEASUREMENTS. Body length 12.0 mm, wing length 15.0 mm, antenna length 4.0 mm (n = 2).</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07FE5AFB64FDD9" blockId="36.[188,1399,506,603]" pageId="36" pageNumber="188">
HEAD (
<figureCitation id="13162A5BFF88FFA5FEB4FE5BFE71FD97" box="[270,407,506,533]" captionStart="Figs 74–76" captionStartId="33.[189,242,1922,1948]" captionTargetBox="[260,1330,289,1812]" captionTargetId="figure-16@33.[243,1344,265,1884]" captionTargetPageId="33" captionText="Figs 74–76. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 74. Habitus, lateral view. 75. Male habitus, dorsal view. 76. Wing. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514515" httpUri="https://zenodo.org/record/5514515/files/figure.png" pageId="36" pageNumber="188">Figs 74–75</figureCitation>
). Mainly dark yellow. Vertex with dark brown stripe. Frontal tubercle relatively high. Proboscis relatively short, rostrum and nasus dark yellow. Setae on head dark brown. Scape yellow, pedicel pale yellow, flagellum dark brown. Proboscis and palpus brownish yellow.
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07FD22FC45FC34" blockId="36.[189,1400,642,951]" pageId="36" pageNumber="188">
THORAX (
<figureCitation id="13162A5BFF88FFA5FE90FD23FE52FD1F" box="[298,436,642,669]" captionStart="Figs 74–76" captionStartId="33.[189,242,1922,1948]" captionTargetBox="[260,1330,289,1812]" captionTargetId="figure-16@33.[243,1344,265,1884]" captionTargetPageId="33" captionText="Figs 74–76. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 74. Habitus, lateral view. 75. Male habitus, dorsal view. 76. Wing. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514515" httpUri="https://zenodo.org/record/5514515/files/figure.png" pageId="36" pageNumber="188">Figs 74–75</figureCitation>
). Mainly dark yellow, heavily pruinescence. Pronotum brownish yellow with dark yellow setae. Prescutum mainly reddish yellow with four olive-brown striped pruinescence, velvety, two middle stripes virtually fused. Scutum with four reddish brown spots, with olive-brown striped pruinescence. Scutellum and mediotergite yellow with pale brown mid-longitudinal stripes. Thorax yellow in lateral view. Pleura below anterior thoracic spiracle to middle with an irregular brownish black spot; anepisternum and katepisternum pale brownish yellow. Setae on thorax yellow. Legs with coxae and bases of femora pale yellow, apices of femora brown; tibiae and tarsi dark brown. Setae on legs brownish black. Wing light brown; pterostigma brown; petiole of cell m 1 shorter than discal cell (
<figureCitation id="13162A5BFF88FFA5FF72FC3DFEC6FC35" box="[200,288,924,951]" captionStart="Figs 74–76" captionStartId="33.[189,242,1922,1948]" captionTargetBox="[260,1330,289,1812]" captionTargetId="figure-16@33.[243,1344,265,1884]" captionTargetPageId="33" captionText="Figs 74–76. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 74. Habitus, lateral view. 75. Male habitus, dorsal view. 76. Wing. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514515" httpUri="https://zenodo.org/record/5514515/files/figure.png" pageId="36" pageNumber="188">Fig. 76</figureCitation>
). Halter with stem dark yellow, knob brownish yellow.
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07FC7FFA91FB99" blockId="36.[189,1399,989,1051]" pageId="36" pageNumber="188">
ABDOMEN (
<figureCitation id="13162A5BFF88FFA5FEF9FC7CFE35FC7A" box="[323,467,989,1016]" captionStart="Figs 74–76" captionStartId="33.[189,242,1922,1948]" captionTargetBox="[260,1330,289,1812]" captionTargetId="figure-16@33.[243,1344,265,1884]" captionTargetPageId="33" captionText="Figs 74–76. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 74. Habitus, lateral view. 75. Male habitus, dorsal view. 76. Wing. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514515" httpUri="https://zenodo.org/record/5514515/files/figure.png" pageId="36" pageNumber="188">Figs 74–75</figureCitation>
). Mainly yellow. Abdominal dorsum with three brown longitudinal stripes, abdominal segment 6 brownish yellow, segments 7–9 brown. Setae on abdomen dark brown and yellow.
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07FBE2FA8BFAAD" blockId="36.[189,1401,1090,1328]" pageId="36" pageNumber="188">
HYPOPYGIUM (
<figureCitation id="13162A5BFF88FFA5FEE0FBE3FE39FBDE" box="[346,479,1090,1116]" captionStart="Figs 79–83" captionStartId="35.[189,242,1888,1914]" captionTargetBox="[255,1300,294,1823]" captionTargetId="figure-15@35.[255,1332,265,1850]" captionTargetPageId="35" captionText="Figs 79–83. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 79. Hypopygium, lateral view. 80–81. Outer and inner gonostylus, lateral external view and lateral dorsal view. 82. Tergite nine, dorsal view. 83. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 79, 82–83 = 0.5 mm; 80–81 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514521" httpUri="https://zenodo.org/record/5514521/files/figure.png" pageId="36" pageNumber="188">Figs 79–83</figureCitation>
). Gonocoxite ends hook-like, not separated from sternite 9, tergite 9 bearing two pairs of lobes, longitudinally divided into two parts, slightly connected, inner margin with dense setae; a pair of emarginate arms arising from ventral surface of tergite 9, inner margin black, other areas yellow. Sternite 9 with a median prominence, middle of sternite 9 slightly raised. Sternite 8 with abundant setae. Outer gonostylus slender and small, apex curly (
<figureCitation id="13162A5BFF88FFA5FCB9FB6EFC6BFB6B" box="[771,909,1231,1257]" captionStart="Figs 79–83" captionStartId="35.[189,242,1888,1914]" captionTargetBox="[255,1300,294,1823]" captionTargetId="figure-15@35.[255,1332,265,1850]" captionTargetPageId="35" captionText="Figs 79–83. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 79. Hypopygium, lateral view. 80–81. Outer and inner gonostylus, lateral external view and lateral dorsal view. 82. Tergite nine, dorsal view. 83. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 79, 82–83 = 0.5 mm; 80–81 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514521" httpUri="https://zenodo.org/record/5514521/files/figure.png" pageId="36" pageNumber="188">Figs 79–81</figureCitation>
). Inner gonostylus, dorsal crest produced into fishtail, base of inner gonostylus and outer basal lobe of inner gonostylus inward-rolled, produced depression with a row of short setulae, outer basal lobe of inner gonostylus large triangular (
<figureCitation id="13162A5BFF88FFA5FB60FAB4FA86FAAD" box="[1242,1376,1301,1327]" captionStart="Figs 79–83" captionStartId="35.[189,242,1888,1914]" captionTargetBox="[255,1300,294,1823]" captionTargetId="figure-15@35.[255,1332,265,1850]" captionTargetPageId="35" captionText="Figs 79–83. Tipula (Vestiplex) motuoensis sp. nov., holotype, ♂ (CAU). 79. Hypopygium, lateral view. 80–81. Outer and inner gonostylus, lateral external view and lateral dorsal view. 82. Tergite nine, dorsal view. 83. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 79, 82–83 = 0.5 mm; 80–81 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514521" httpUri="https://zenodo.org/record/5514521/files/figure.png" pageId="36" pageNumber="188">Figs 80–81</figureCitation>
).
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF50FAF6FEA3FAF3" blockId="36.[189,1236,1367,1429]" box="[234,325,1367,1393]" pageId="36" pageNumber="188">
<emphasis id="B959EACCFF88FFA5FF50FAF6FEA3FAF3" bold="true" box="[234,325,1367,1393]" pageId="36" pageNumber="188">Female</emphasis>
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07FADBFB32FA16" blockId="36.[189,1236,1367,1429]" box="[189,1236,1402,1429]" pageId="36" pageNumber="188">
MEASUREMENTS. Body length
<quantity id="4CD59B3BFF88FFA5FDAFFADBFD8CFA16" box="[533,618,1402,1428]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="36" pageNumber="188" unit="mm" value="13.0">13 mm</quantity>
, wing length
<quantity id="4CD59B3BFF88FFA5FCB2FADBFCB8FA16" box="[776,862,1402,1429]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="188" unit="mm" value="15.0">15 mm</quantity>
, antenna length 2.4 mm (n = 1).
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07FA1AFCB0F9C2" blockId="36.[189,1400,1467,1600]" pageId="36" pageNumber="188">
COLORATION. Body color slightly paler than male. Antenna shorter than male, dark yellow. Legs paler than male, tibiae and tarsi dark yellow. Abdominal tergites each with three stripes, middle stripe brown, lateral stripes brownish black, abdominal 7–8 segments yellowish brown, 8th segment dark brown. Junction of each 3–9 segments brownish black (
<figureCitation id="13162A5BFF88FFA5FD4BF984FCAEF9C2" box="[753,840,1573,1600]" captionStart="Figs 77–78" captionStartId="34.[189,242,1954,1980]" captionTargetBox="[290,1329,290,1893]" captionTargetId="figure-12@34.[232,1355,265,1917]" captionTargetPageId="34" captionText="Figs 77–78. Tipula (Vestiplex) motuoensis sp. nov., allotype, ♀ (CAU). 77. Habitus, lateral view. 78. Ovipositor, lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514517" httpUri="https://zenodo.org/record/5514517/files/figure.png" pageId="36" pageNumber="188">Fig. 77</figureCitation>
).
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07F9C6FD46F926" blockId="36.[189,1398,1638,1700]" pageId="36" pageNumber="188">
OVIPOSITOR. Tergite 9 yellowish brown, tergite 10 yellow. Cercus dark yellow, slightly up-turned, outer margin without rough serration (
<figureCitation id="13162A5BFF88FFA5FD81F92BFD75F926" box="[571,659,1674,1700]" captionStart="Figs 77–78" captionStartId="34.[189,242,1954,1980]" captionTargetBox="[290,1329,290,1893]" captionTargetId="figure-12@34.[232,1355,265,1917]" captionTargetPageId="34" captionText="Figs 77–78. Tipula (Vestiplex) motuoensis sp. nov., allotype, ♀ (CAU). 77. Habitus, lateral view. 78. Ovipositor, lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514517" httpUri="https://zenodo.org/record/5514517/files/figure.png" pageId="36" pageNumber="188">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3376555FF88FFA5FF07F96AFE85F895" box="[189,355,1739,1815]" pageId="36" pageNumber="188" type="distribution">
<paragraph id="8B9236DEFF88FFA5FF07F96AFEB0F967" blockId="36.[189,342,1739,1765]" box="[189,342,1739,1765]" pageId="36" pageNumber="188">
<heading id="D0DA81B2FF88FFA5FF07F96AFEB0F967" bold="true" box="[189,342,1739,1765]" fontSize="11" level="3" pageId="36" pageNumber="188" reason="3">
<emphasis id="B959EACCFF88FFA5FF07F96AFEB0F967" bold="true" box="[189,342,1739,1765]" pageId="36" pageNumber="188">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9236DEFF88FFA5FF07F95DFE85F895" blockId="36.[189,355,1788,1815]" box="[189,355,1788,1815]" pageId="36" pageNumber="188">
<collectingCountry id="F33A764EFF88FFA5FF07F95DFEE5F894" box="[189,259,1788,1814]" name="China" pageId="36" pageNumber="188">China</collectingCountry>
(
<collectingRegion id="49E9F83CFF88FFA5FEA9F95DFEB1F895" box="[275,343,1788,1815]" country="China" name="Xizang" pageId="36" pageNumber="188">Tibet</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3376555FF88FFA7FF07F89BFD59FEE8" lastPageId="38" lastPageNumber="190" pageId="36" pageNumber="188" type="discussion">
<paragraph id="8B9236DEFF88FFA5FF07F89BFEC9F8D6" blockId="36.[189,303,1850,1876]" box="[189,303,1850,1876]" pageId="36" pageNumber="188">
<heading id="D0DA81B2FF88FFA5FF07F89BFEC9F8D6" bold="true" box="[189,303,1850,1876]" fontSize="11" level="3" pageId="36" pageNumber="188" reason="3">
<emphasis id="B959EACCFF88FFA5FF07F89BFEC9F8D6" bold="true" box="[189,303,1850,1876]" pageId="36" pageNumber="188">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9236DEFF88FFA7FF07F8C9FD59FEE8" blockId="36.[189,1401,1896,1993]" lastBlockId="38.[189,1398,265,362]" lastPageId="38" lastPageNumber="190" pageId="36" pageNumber="188">
The new species is similar to
<emphasis id="B959EACCFF88FFA5FD93F8C8FDD1F800" box="[553,567,1897,1922]" italics="true" pageId="36" pageNumber="188">T</emphasis>
. (
<emphasis id="B959EACCFF88FFA5FDE8F8C8FD82F800" box="[594,612,1897,1922]" italics="true" pageId="36" pageNumber="188">V</emphasis>
.)
<emphasis id="B959EACCFF88FFA5FDC4F8C9FCF0F800" box="[638,790,1896,1922]" italics="true" pageId="36" pageNumber="188">theowaldana</emphasis>
Alexander, 1964 from
<collectingRegion id="49E9F83CFF88FFA5FB88F8C9FB63F800" box="[1074,1157,1896,1922]" country="India" name="Assam" pageId="36" pageNumber="188">Assam</collectingRegion>
,
<collectingCountry id="F33A764EFF88FFA5FB28F8C9FB37F800" box="[1170,1233,1896,1922]" name="India" pageId="36" pageNumber="188">India</collectingCountry>
, but the latter differs in the following characters: basal abdominal tergites yellow; gonocoxite ends sharp and not hooked; tergite 9 with a pair of larger and elongate lobes. The species may form a new species group, perhaps together with
<emphasis id="B959EACCFF8AFFA7FE78FEABFE34FEA1" box="[450,466,266,291]" italics="true" pageId="38" pageNumber="190">T</emphasis>
. (
<emphasis id="B959EACCFF8AFFA7FE51FEABFE1BFEA1" box="[491,509,266,291]" italics="true" pageId="38" pageNumber="190">V</emphasis>
.)
<emphasis id="B959EACCFF8AFFA7FDAFFEA8FD4BFEA1" box="[533,685,265,291]" italics="true" pageId="38" pageNumber="190">theowaldana</emphasis>
. However, we have not examined the specimens of the latter species and can therefore not say with certainty if these two species share characteristics that would separate them from existing species groups.
</paragraph>
</subSubSection>
</treatment>
</document>
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